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<document id="6361C46796A44F6D088368E285EB4849" ID-DOI="10.11646/zootaxa.5389.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10410051" ID-ZooBank="3E7237B4-85C5-4581-B469-ED1C00154413" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1703073801695" checkinUser="plazi" docAuthor="Zajonz, Uwe, Bogorodsky, Sergey V., Saeed, Fouad K. N., Aideed, Moteah S. &amp; Lavergne, Edouard" docDate="2023" docId="039D87C9FFB8FF9DA0A87860FC7828C4" docLanguage="en" docName="zootaxa.5389.3.1.pdf" docOrigin="Zootaxa 5389 (3)" docSource="https://mapress.com/zt/article/download/zootaxa.5389.3.1/52519" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scarus tricolor Bleeker 1847" docType="treatment" docVersion="2" lastPageNumber="319" masterDocId="FFA4FFB1FFAAFF8FA03F7A0BFFF82D7F" masterDocTitle="Parrotfishes (Teleostei: Labridae: Scarini) of the Socotra Archipelago: Diversity and distributional biogeography, including a range extension of Scarus zufar Randall &amp; Hoover, 1995" masterLastPageNumber="330" masterPageNumber="301" pageNumber="319" updateTime="1703074317466" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4A5B573C1E9AA0AA89EC9E40025E7ADC">Parrotfishes (Teleostei: Labridae: Scarini) of the Socotra Archipelago: Diversity and distributional biogeography, including a range extension of Scarus zufar Randall &amp; Hoover, 1995</mods:title>
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<mods:namePart id="8F2790D75FF7CB0323F7DD83BC68A48B">Saeed, Fouad K. N.</mods:namePart>
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<heading id="D0C381B3FFB8FF9DA0A87860FE022FFB" bold="true" box="[151,506,618,645]" fontSize="11" level="1" pageId="18" pageNumber="319" reason="1">
<taxonomicName id="4C344D5CFFB8FF9DA0A87860FE022FFB" ID-CoL="4V3P2" authority="Bleeker, 1847" authorityName="Bleeker" authorityYear="1847" box="[151,506,618,645]" family="Scaridae" genus="Scarus" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="319" phylum="Chordata" rank="species" species="tricolor">
<emphasis id="B940EACDFFB8FF9DA0A87860FE022FFB" bold="true" box="[151,506,618,645]" pageId="18" pageNumber="319">
<emphasis id="B940EACDFFB8FF9DA0A87860FEB42FFA" bold="true" box="[151,332,618,645]" italics="true" pageId="18" pageNumber="319">Scarus tricolor</emphasis>
<bibRefCitation id="EFA54B2EFFB8FF9DA16C7860FE022FFB" author="Bleeker, P." box="[339,506,618,645]" pageId="18" pageNumber="319" pagination="155 - 169" refId="ref13284" refString="Bleeker, P. (1847) Pharyngognathorum Siluroideorumque species novae Javanenses. Natuur- en Geneeskundig Archief voor Neerlandsch Indie, 4 (2), 155 - 169." type="journal article" year="1847">Bleeker, 1847</bibRefCitation>
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<paragraph id="8B8B36DFFFB8FF9DA0A87880FB842FDC" blockId="18.[151,1148,618,747]" box="[151,1148,651,676]" pageId="18" pageNumber="319">
<treatmentCitationGroup id="AB2411F1FFB8FF9DA0A87880FB842FDC" box="[151,1148,651,676]" pageId="18" pageNumber="319">
<treatmentCitation id="0A9510CEFFB8FF9DA0A87880FE022FDC" author="Bleeker, P." box="[151,506,651,675]" page="164" pageId="18" pageNumber="319" year="1847">
<taxonomicName id="4C344D5CFFB8FF9DA0A87880FE022FDC" ID-CoL="4V3P2" authority="Bleeker, 1847: 164" authorityName="Bleeker" authorityPageNumber="164" authorityYear="1847" box="[151,506,651,675]" family="Scaridae" genus="Scarus" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="319" phylum="Chordata" rank="species" species="tricolor">
<emphasis id="B940EACDFFB8FF9DA0A87880FED72FDC" box="[151,303,651,675]" italics="true" pageId="18" pageNumber="319">Scarus tricolor</emphasis>
<materialsCitation id="3B5C3C82FFB8FF9DA1097880FE022FDC" box="[310,506,651,675]" collectionCode="BMNH" country="Indonesia" location="Java" pageId="18" pageNumber="319" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
<bibRefCitation id="EFA54B2EFFB8FF9DA1097880FE022FDC" author="Bleeker, P." box="[310,506,651,675]" pageId="18" pageNumber="319" pagination="155 - 169" refId="ref13284" refString="Bleeker, P. (1847) Pharyngognathorum Siluroideorumque species novae Javanenses. Natuur- en Geneeskundig Archief voor Neerlandsch Indie, 4 (2), 155 - 169." type="journal article" year="1847">Bleeker, 1847: 164</bibRefCitation>
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;
<typeStatus id="548F887DFFB8FF9DA23B7880FD992FDC" box="[516,609,651,675]" pageId="18" pageNumber="319" type="holotype">holotype</typeStatus>
:?BMNH 1862.2.28.9; type locality:
<collectingRegion id="49F0F83DFFB8FF9DA3E67887FBF02FDC" box="[985,1032,652,675]" country="Indonesia" name="Jawa Timur" pageId="18" pageNumber="319">Java</collectingRegion>
,
<collectingCountry id="F323764FFFB8FF9DA42D7880FB802FDC" box="[1042,1144,651,675]" name="Indonesia" pageId="18" pageNumber="319">Indonesia</collectingCountry>
.
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</paragraph>
<paragraph id="8B8B36DFFFB8FF9DA0A878A6FE9E2FB8" blockId="18.[151,1148,618,747]" box="[151,358,685,711]" pageId="18" pageNumber="319">Tricolor parrotfish</paragraph>
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<subSubSection id="C32E6554FFB8FF9DA0A878DAFEFB2F94" box="[151,259,721,747]" pageId="18" pageNumber="319" type="description">
<paragraph id="8B8B36DFFFB8FF9DA0A878DAFEFB2F94" blockId="18.[151,1148,618,747]" box="[151,259,721,747]" pageId="18" pageNumber="319">
<figureCitation id="130F2A5AFFB8FF9DA0A878DAFEFB2F94" box="[151,259,721,747]" captionStart="FIGURE 13" captionStartId="17.[151,250,1563,1587]" captionTargetBox="[164,1423,925,1539]" captionTargetId="figure-39@17.[164,1423,925,1539]" captionTargetPageId="17" captionText="FIGURE 13. Scarus tricolor, terminal male, Roosh, Socotra Island, 14 m depth." figureDoi="http://doi.org/10.5281/zenodo.10410096" httpUri="https://zenodo.org/record/10410096/files/figure.png" pageId="18" pageNumber="319">Figure 13</figureCitation>
</paragraph>
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<paragraph id="8B8B36DFFFB8FF9DA0A87912FD642E4C" blockId="18.[151,1437,793,1468]" box="[151,668,793,819]" pageId="18" pageNumber="319">
<emphasis id="B940EACDFFB8FF9DA0A87912FE7B2E4C" bold="true" box="[151,387,793,819]" pageId="18" pageNumber="319">Material examined:</emphasis>
Underwater photograph.
</paragraph>
<paragraph id="8B8B36DFFFB8FF9DA0F87936FBDA2E98" blockId="18.[151,1437,793,1468]" pageId="18" pageNumber="319">
<emphasis id="B940EACDFFB8FF9DA0F87936FE2C2E28" bold="true" box="[199,468,829,855]" pageId="18" pageNumber="319">Distinctive characters:</emphasis>
Body elongate, its depth
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SL; dorsal profile of head straight; posterior nostril subequal to anterior nostril; teeth fully fused to form dental plates, about two-thirds or more covered by lips; cutting edge of dental plates usually smooth; upper dental plate with 12 conical teeth posteriorly in terminal males and large initial phase; caudal fin lunate in terminal males. Meristic values: Pectoral-fin rays 14; median predorsal scales 57, scales subequal in size; cheek with 3 scale rows, lower row with 36 scales.
</paragraph>
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<emphasis id="B940EACDFFB8FF9DA0F879FAFEA82974" box="[199,336,1009,1035]" italics="true" pageId="18" pageNumber="319">Colouration</emphasis>
: Initial phase with zones of colour from dark brown on back to yellow, green, blue and red on sides and ventrally, the scales rimmed with black; head blackish with a large greenish grey patch dorsally above eye; dental plates pale rose; anal and caudal fins orange-red. Body of terminal males green, the basal half of scales salmon pink, with green back and a green stripe ventrally; head greenish, upper lip salmon pink with a submarginal blue-green stripe that continues across cheek, dorsal part of head with a green band from anterior interorbital area passing through dorsal edge of eye to below origin of dorsal fin; lower lip blue-green followed by pale pink area bordered by blue-green short band on chin; dental plates deep blue-green; caudal fin blue-green with a large middle lavender crescent and narrow pink submarginal band on each lobe.
</paragraph>
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<subSubSection id="C32E6554FFB8FF9DA0F87F1AFD8C2830" pageId="18" pageNumber="319" type="distribution">
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<emphasis id="B940EACDFFB8FF9DA0F87F1AFE9B2854" bold="true" box="[199,355,1297,1323]" pageId="18" pageNumber="319">Distribution:</emphasis>
Widespread from eastern Africa east to the Line and
<collectingCountry id="F323764FFFB8FF9DA3FC7F1AFBF72854" box="[963,1039,1297,1323]" name="Samoa" pageId="18" pageNumber="319">Samoa</collectingCountry>
Islands. No records in the
<taxonomicName id="4C344D5CFFB8FF9DA57D7F1AFA642854" box="[1346,1436,1297,1323]" order="Arabian" pageId="18" pageNumber="325" rank="order">Arabian</taxonomicName>
region except for the
<collectingCountry id="F323764FFFB8FF9DA1B87F3EFE252830" box="[391,477,1333,1359]" name="Yemen" pageId="18" pageNumber="319">Socotra</collectingCountry>
Archipelago.
</paragraph>
</subSubSection>
<subSubSection id="C32E6554FFB8FF9DA0F87F52FC7828C4" pageId="18" pageNumber="319" type="discussion">
<paragraph id="8B8B36DFFFB8FF9DA0F87F52FC7828C4" blockId="18.[151,1437,793,1468]" pageId="18" pageNumber="319">
<emphasis id="B940EACDFFB8FF9DA0F87F52FEC5280C" bold="true" box="[199,317,1369,1395]" pageId="18" pageNumber="319">Remarks:</emphasis>
Photographed individual matches the description of
<taxonomicName id="4C344D5CFFB8FF9DA3877F51FA6F280C" authority="(Randall &amp; Bruce 1983)" baseAuthorityName="Randall &amp; Bruce" baseAuthorityYear="1983" box="[952,1431,1369,1396]" family="Scaridae" genus="Scarus" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="319" phylum="Chordata" rank="species" species="tricolor">
<emphasis id="B940EACDFFB8FF9DA3877F51FB9F280C" box="[952,1127,1369,1396]" italics="true" pageId="18" pageNumber="319">Scarus tricolor</emphasis>
(
<bibRefCitation id="EFA54B2EFFB8FF9DA4427F52FA77280C" author="Randall, J. E. &amp; Bruce, R. W." box="[1149,1423,1369,1396]" pageId="18" pageNumber="319" pagination="1 - 39" refId="ref15077" refString="Randall, J. E. &amp; Bruce, R. W. (1983) The parrotfishes of the subfamily Scarinae of the western Indian Ocean with descriptions of three new species. Ichthyological Bulletin of the J. L. B. Smith Institute of Ichthyology, 47, 1 - 39." type="journal article" year="1983">Randall &amp; Bruce 1983</bibRefCitation>
)
</taxonomicName>
, supporting the tentative record of
<bibRefCitation id="EFA54B2EFFB8FF9DA2327F76FD1028E8" author="Zajonz, U. &amp; Lavergne, E. &amp; Bogorodsky, S. V. &amp; Saeed, F. N. &amp; Aideed, M. S. &amp; Krupp, F." box="[525,744,1405,1432]" pageId="18" pageNumber="319" pagination="1 - 108" refId="ref16035" refString="Zajonz, U., Lavergne, E., Bogorodsky, S. V., Saeed, F. N., Aideed, M. S. &amp; Krupp, F. (2019) Coastal fish diversity of the Socotra Archipelago, Yemen. Zootaxa, 4636 (1), 1 - 108. https: // doi. org / 10.11646 / zootaxa. 4636.1.1" type="journal article" year="2019">
Zajonz
<emphasis id="B940EACDFFB8FF9DA25E7F75FD6928E8" box="[609,657,1405,1431]" italics="true" pageId="18" pageNumber="319">et al</emphasis>
. (2019)
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from Abd al-Kuri
<collectingCountry id="F323764FFFB8FF9DA38A7F76FC0528E8" box="[949,1021,1405,1431]" name="Iceland" pageId="18" pageNumber="319">Island</collectingCountry>
. It was photographed at north-eastern
<collectingCountry id="F323764FFFB8FF9DA0A87FAAFEC328C4" box="[151,315,1441,1467]" name="Socotra Island" pageId="18" pageNumber="319">Socotra Island</collectingCountry>
, Roosh-Haleh, in a coral-rich habitat at
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depth.
</paragraph>
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</treatment>
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