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<document id="1A8FACCAF0ED609E36141DF2764CCE41" ID-DOI="10.11646/zootaxa.5389.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10410051" ID-ZooBank="3E7237B4-85C5-4581-B469-ED1C00154413" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1703073801695" checkinUser="plazi" docAuthor="Zajonz, Uwe, Bogorodsky, Sergey V., Saeed, Fouad K. N., Aideed, Moteah S. &amp; Lavergne, Edouard" docDate="2023" docId="039D87C9FFA0FF85A0A879A4FA7C2BC8" docLanguage="en" docName="zootaxa.5389.3.1.pdf" docOrigin="Zootaxa 5389 (3)" docSource="https://mapress.com/zt/article/download/zootaxa.5389.3.1/52519" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scarus frenatus Lacepede 1802" docType="treatment" docVersion="2" lastPageNumber="311" masterDocId="FFA4FFB1FFAAFF8FA03F7A0BFFF82D7F" masterDocTitle="Parrotfishes (Teleostei: Labridae: Scarini) of the Socotra Archipelago: Diversity and distributional biogeography, including a range extension of Scarus zufar Randall &amp; Hoover, 1995" masterLastPageNumber="330" masterPageNumber="301" pageNumber="311" updateTime="1703074317466" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7D9752A4E4FCF8BB0B7C95ED58C43D7E">Parrotfishes (Teleostei: Labridae: Scarini) of the Socotra Archipelago: Diversity and distributional biogeography, including a range extension of Scarus zufar Randall &amp; Hoover, 1995</mods:title>
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<mods:namePart id="564A459F1CFF1226E9AC4F63537EC9DB">Zajonz, Uwe</mods:namePart>
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<mods:namePart id="EF6B1AFB4BDF38FAD2E3079B949A681D">Saeed, Fouad K. N.</mods:namePart>
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<treatment id="039D87C9FFA0FF85A0A879A4FA7C2BC8" ID-DOI="http://doi.org/10.5281/zenodo.10410088" ID-Zenodo-Dep="10410088" LSID="urn:lsid:plazi:treatment:039D87C9FFA0FF85A0A879A4FA7C2BC8" httpUri="http://treatment.plazi.org/id/039D87C9FFA0FF85A0A879A4FA7C2BC8" lastPageNumber="311" pageId="10" pageNumber="311">
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<paragraph id="8B8B36DFFFA0FF85A0A879A4FDE72EB7" blockId="10.[151,1031,942,1072]" box="[151,543,942,969]" pageId="10" pageNumber="311">
<heading id="D0C381B3FFA0FF85A0A879A4FDE72EB7" bold="true" box="[151,543,942,969]" fontSize="11" level="1" pageId="10" pageNumber="311" reason="1">
<taxonomicName id="4C344D5CFFA0FF85A0A879A4FDE72EB7" ID-CoL="4V3JK" authority="Lacepede, 1802" authorityName="Lacepede" authorityYear="1802" box="[151,543,942,969]" family="Scaridae" genus="Scarus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="311" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="B940EACDFFA0FF85A0A879A4FDE72EB7" bold="true" box="[151,543,942,969]" pageId="10" pageNumber="311">
<emphasis id="B940EACDFFA0FF85A0A879A4FEAF2EB7" bold="true" box="[151,343,942,969]" italics="true" pageId="10" pageNumber="311">Scarus frenatus</emphasis>
<bibRefCitation id="EFA54B2EFFA0FF85A16179A5FDE72EB7" author="Lacepede, B. G. E." box="[350,543,942,969]" pageId="10" pageNumber="311" refId="ref14616" refString="Lacepede, B. G. E. (1802) Histoire naturelle des poissons. Volume 4. F. - G. Levrault, Paris, 728 pp." type="book" year="1802">Lacépède, 1802</bibRefCitation>
</emphasis>
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<treatmentCitation id="0A9510CEFFA0FF85A0A879C4FE002E98" author="Lacepede, B. G. E." box="[151,504,975,999]" page="3" pageId="10" pageNumber="311" year="1802">
<taxonomicName id="4C344D5CFFA0FF85A0A879C4FE002E98" ID-CoL="4V3JK" authority="Lacepede, 1802: 3" authorityName="Lacepede" authorityPageNumber="3" authorityYear="1802" box="[151,504,975,999]" family="Scaridae" genus="Scarus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="311" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="B940EACDFFA0FF85A0A879C4FECC2E98" box="[151,308,975,999]" italics="true" pageId="10" pageNumber="311">Scarus frenatus</emphasis>
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<bibRefCitation id="EFA54B2EFFA0FF85A10479C4FE002E98" author="Lacepede, B. G. E." box="[315,504,975,999]" pageId="10" pageNumber="311" refId="ref14616" refString="Lacepede, B. G. E. (1802) Histoire naturelle des poissons. Volume 4. F. - G. Levrault, Paris, 728 pp." type="book" year="1802">Lacépède, 1802: 3</bibRefCitation>
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; no
<typeStatus id="548F887DFFA0FF85A21D79DBFDAF2E97" box="[546,599,976,1000]" pageId="10" pageNumber="311">types</typeStatus>
known;
<typeStatus id="548F887DFFA0FF85A28E79DBFD252E97" box="[689,733,976,1000]" pageId="10" pageNumber="311">type</typeStatus>
locality:
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.
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</paragraph>
<paragraph id="8B8B36DFFFA0FF85A0A879FAFE982974" blockId="10.[151,1031,942,1072]" box="[151,352,1009,1035]" pageId="10" pageNumber="311">Bridled Parrotfish</paragraph>
</subSubSection>
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<paragraph id="8B8B36DFFFA0FF85A0A87E1EFF0D294F" blockId="10.[151,1031,942,1072]" box="[151,245,1045,1072]" pageId="10" pageNumber="311">
<figureCitation id="130F2A5AFFA0FF85A0A87E1EFF0D294F" box="[151,245,1045,1072]" captionStart="FIGURE 7" captionStartId="11.[151,250,1689,1713]" captionTargetBox="[151,1436,237,1666]" captionTargetId="figure-52@11.[151,1436,237,1666]" captionTargetPageId="11" captionText="FIGURE 7. Scarus frenatus. A: terminal male, Di Hamri, Socotra Island, 3 m depth; B: terminal male, Qariah, Socotra Island, 5 m depth." figureDoi="http://doi.org/10.5281/zenodo.10410066" httpUri="https://zenodo.org/record/10410066/files/figure.png" pageId="10" pageNumber="311">Figure 7</figureCitation>
</paragraph>
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<emphasis id="B940EACDFFA0FF85A0A87E56FE7B2908" bold="true" box="[151,387,1117,1143]" pageId="10" pageNumber="311">Material examined:</emphasis>
Underwater photographs.
</paragraph>
<paragraph id="8B8B36DFFFA0FF85A0F87E8AFB612854" blockId="10.[151,1437,1117,1720]" pageId="10" pageNumber="311">
<emphasis id="B940EACDFFA0FF85A0F87E8AFE2F29E4" bold="true" box="[199,471,1153,1179]" pageId="10" pageNumber="311">Distinctive characters:</emphasis>
Body depth
<quantity id="4CCC9B3AFFA0FF85A2547E8AFD2629E4" box="[619,734,1153,1179]" metricMagnitude="-2" metricUnit="m" metricValue="6.603999999999999" metricValueMax="7.366" metricValueMin="5.842" pageId="10" pageNumber="311" unit="in" value="2.5999999999999996" valueMax="2.9" valueMin="2.3">2.32.9 in</quantity>
SL; dorsal profile of head of terminal males slightly concave on forehead; posterior nostril subequal to anterior nostril; teeth fully fused to form dental plates, about three-fourths covered by lips; cutting edge of dental plates nearly smooth; upper dental plate with 02 conical teeth posteriorly in both phases; caudal fin double emarginate in terminal males. Meristic values: Pectoral-fin rays 14; median predorsal scales 67, fourth or fifth scale largest; cheek with 3 scale rows, lower row with 24 scales.
</paragraph>
<paragraph id="8B8B36DFFFA0FF85A0F87F3EFAF028C3" blockId="10.[151,1437,1117,1720]" pageId="10" pageNumber="311">
<emphasis id="B940EACDFFA0FF85A0F87F3EFEA82830" box="[199,336,1333,1359]" italics="true" pageId="10" pageNumber="311">Colouration</emphasis>
: Initial phase yellowish gray, paler on abdomen, with six dark brown stripes; caudal fin dark gray, remaining fins reddish. Terminal males green, the scales with small lavender-pink spots, posterior part of body abruptly bright green; head with a broad green stripe below eye bifurcating at corner of mouth to snout and chin, head below stripe with green vermiculations; caudal fin green, with orange-pink or magenta crescent.
</paragraph>
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<emphasis id="B940EACDFFA0FF85A0F87FCEFE9B28A0" bold="true" box="[199,355,1477,1503]" pageId="10" pageNumber="311">Distribution:</emphasis>
Red Sea south to
<collectingCountry id="F323764FFFA0FF85A2197FCEFD4528A0" box="[550,701,1477,1503]" name="Mozambique" pageId="10" pageNumber="311">Mozambique</collectingCountry>
,
<collectingCountry id="F323764FFFA0FF85A2F97FCEFCB728A0" box="[710,847,1477,1503]" name="Madagascar" pageId="10" pageNumber="311">Madagascar</collectingCountry>
, and the Mascarene Islands, east to
<collectingCountry id="F323764FFFA0FF85A4EA7FCEFA6128A0" box="[1237,1433,1477,1503]" name="French Polynesia" pageId="10" pageNumber="311">French Polynesia</collectingCountry>
. Known from the entire
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region, including the eastern Gulf of
<collectingRegion id="49F0F83DFFA0FF85A3827FE2FC022B7C" box="[957,1018,1513,1539]" country="Yemen" name="'Adan" pageId="10" pageNumber="311">Aden</collectingRegion>
and
<collectingCountry id="F323764FFFA0FF85A4097FE2FB762B7C" box="[1078,1166,1513,1539]" name="Yemen" pageId="10" pageNumber="311">Socotra</collectingCountry>
, to the
<taxonomicName id="4C344D5CFFA0FF85A4D77FE2FABA2B7C" box="[1256,1346,1513,1539]" order="Arabian" pageId="10" pageNumber="325" rank="order">Arabian</taxonomicName>
/Persian Gulf (
<bibRefCitation id="EFA54B2EFFA0FF85A0E47C06FE252B58" author="Randall, J. E. &amp; Bruce, R. W." box="[219,477,1549,1576]" pageId="10" pageNumber="311" pagination="1 - 39" refId="ref15077" refString="Randall, J. E. &amp; Bruce, R. W. (1983) The parrotfishes of the subfamily Scarinae of the western Indian Ocean with descriptions of three new species. Ichthyological Bulletin of the J. L. B. Smith Institute of Ichthyology, 47, 1 - 39." type="journal article" year="1983">Randall &amp; Bruce 1983</bibRefCitation>
; Randall 1995;
<bibRefCitation id="EFA54B2EFFA0FF85A2B07C06FC352B58" author="Golani, D. &amp; Bogorodsky, S. V." box="[655,973,1549,1575]" pageId="10" pageNumber="311" pagination="1 - 135" refId="ref14096" refString="Golani, D. &amp; Bogorodsky, S. V. (2010) The fishes of the Red Sea - reappraisal and updated checklist. Zootaxa, 2463 (1), 1 - 135. https: // doi. org / 10.11646 / zootaxa. 2463.1.1" type="journal article" year="2010">Golani &amp; Bogorodsky 2010</bibRefCitation>
); but records from the Gulf of
<collectingCountry id="F323764FFFA0FF85A5177C06FA942B58" box="[1320,1388,1549,1575]" name="Oman" pageId="10" pageNumber="311">Oman</collectingCountry>
and
<taxonomicName id="4C344D5CFFA0FF85A0A87C3AFF092B34" box="[151,241,1585,1611]" order="Arabian" pageId="10" pageNumber="325" rank="order">Arabian</taxonomicName>
/Persian Gulf require verification.
</paragraph>
</subSubSection>
<subSubSection id="C32E6554FFA0FF85A0F87C5EFA7C2BC8" pageId="10" pageNumber="311" type="discussion">
<paragraph id="8B8B36DFFFA0FF85A0F87C5EFA7C2BC8" blockId="10.[151,1437,1117,1720]" pageId="10" pageNumber="311">
<emphasis id="B940EACDFFA0FF85A0F87C5EFEC52B10" bold="true" box="[199,317,1621,1647]" pageId="10" pageNumber="311">Remarks:</emphasis>
Observed individuals match the description of
<taxonomicName id="4C344D5CFFA0FF85A3017C5DFC122B10" authorityName="Lacepede" authorityYear="1802" box="[830,1002,1621,1648]" family="Scaridae" genus="Scarus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="311" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="B940EACDFFA0FF85A3017C5DFC122B10" box="[830,1002,1621,1648]" italics="true" pageId="10" pageNumber="311">Scarus frenatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFA54B2EFFA0FF85A3C87C5EFB172B10" author="Randall, J. E. &amp; Bruce, R. W." box="[1015,1263,1621,1648]" pageId="10" pageNumber="311" pagination="1 - 39" refId="ref15077" refString="Randall, J. E. &amp; Bruce, R. W. (1983) The parrotfishes of the subfamily Scarinae of the western Indian Ocean with descriptions of three new species. Ichthyological Bulletin of the J. L. B. Smith Institute of Ichthyology, 47, 1 - 39." type="journal article" year="1983">Randall &amp; Bruce 1983</bibRefCitation>
; Randall 1995), supporting the record of
<bibRefCitation id="EFA54B2EFFA0FF85A18E7C72FD6E2BEC" author="Zajonz, U. &amp; Lavergne, E. &amp; Bogorodsky, S. V. &amp; Saeed, F. N. &amp; Aideed, M. S. &amp; Krupp, F." box="[433,662,1657,1684]" pageId="10" pageNumber="311" pagination="1 - 108" refId="ref16035" refString="Zajonz, U., Lavergne, E., Bogorodsky, S. V., Saeed, F. N., Aideed, M. S. &amp; Krupp, F. (2019) Coastal fish diversity of the Socotra Archipelago, Yemen. Zootaxa, 4636 (1), 1 - 108. https: // doi. org / 10.11646 / zootaxa. 4636.1.1" type="journal article" year="2019">
Zajonz
<emphasis id="B940EACDFFA0FF85A2387C71FDC12BEC" box="[519,569,1657,1683]" italics="true" pageId="10" pageNumber="311">et al</emphasis>
. (2019)
</bibRefCitation>
. Observed only on
<collectingCountry id="F323764FFFA0FF85A34C7C72FBED2BEC" box="[883,1045,1657,1683]" name="Socotra Island" pageId="10" pageNumber="311">Socotra Island</collectingCountry>
in coral-dominated habitats of the north-eastern coast at depths of
<quantity id="4CCC9B3AFFA0FF85A1C37C96FDBE2BC7" box="[508,582,1693,1720]" metricMagnitude="0" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="10" pageNumber="311" unit="m" value="5.0" valueMax="8.0" valueMin="2.0">
2
<specimenCount id="9D32FD56FFA0FF85A2257C96FDBE2BC7" box="[538,582,1693,1720]" count="8" pageId="10" pageNumber="311" type="generic">8 m</specimenCount>
</quantity>
. Based on monitoring data it is a rare species (Zajonz
<emphasis id="B940EACDFFA0FF85A49B7C95FB2E2BC8" box="[1188,1238,1693,1719]" italics="true" pageId="10" pageNumber="311">et al</emphasis>
., unpublished).
</paragraph>
</subSubSection>
</treatment>
</document>