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<document id="F0AC8308282D7667A9425C9F1C3EFA6A" ID-CLB-Dataset="30136" ID-DOI="10.11646/zootaxa.4416.1.1" ID-GBIF-Dataset="aa238a39-adf8-4b94-8808-c788670d0455" ID-ISSN="1175-5326" ID-Zenodo-Dep="1242263" ID-ZooBank="3341A80B-7E54-4C6B-8B46-1336094952E2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1525601053009" checkinUser="plazi" docAuthor="Barbosa, Diego N. &amp; Azevedo, Celso O." docDate="2018" docId="039D87C45C38FF9C48F6FD6BFAAAF98E" docLanguage="en" docName="zootaxa.4416.1.1.pdf" docOrigin="Zootaxa 4416 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Anisepyris aghani Barbosa &amp; Azevedo 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="40" masterDocId="FFA4FFBC5C1FFFBB4861FFDFFFCEFFFD" masterDocTitle="Revision of Anisepyris Kieffer (Hymenoptera, Bethylidae), with description of 135 new species" masterLastPageNumber="258" masterPageNumber="1" pageNumber="40" updateTime="1698542699957" updateUser="ExternalLinkService">
<mods:mods id="EE26A417393EFDBC9DAE554085106A66" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F6E12BB491B7BD7154B1B734675D11AF">
<mods:title id="84147B64101F921559FFA772C8886890">Revision of Anisepyris Kieffer (Hymenoptera, Bethylidae), with description of 135 new species</mods:title>
</mods:titleInfo>
<mods:name id="ED83009F3DF49B887B20DFB4AB00E06A" type="personal">
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<mods:namePart id="8EE7ED054B053E79158ED1580EA14305">Barbosa, Diego N.</mods:namePart>
</mods:name>
<mods:name id="1B31328A0858A831A140462EA98AC5A4" type="personal">
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<mods:roleTerm id="14D432C23C83576A7F200560BFE0BA21">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="016C6A99B665CCCBB88530C8BEADBBFC">Azevedo, Celso O.</mods:namePart>
</mods:name>
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<mods:title id="E7DA7C6EC080E3A7696F0CEC5B6C7F94">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6CCB79E9F632F06B28CD92B8F6A86FE4">
<mods:date id="DC217DD2664CE497F80960998E05677E">2018</mods:date>
<mods:detail id="26655D7BE1B4B00641D858183FE1DE94" type="volume">
<mods:number id="579BDFFC2D3D0DBB6A23797D114C3572">4416</mods:number>
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<mods:classification id="88C8A6193FA60E01607978F39EF0DAF9">journal article</mods:classification>
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<mods:identifier id="FE4653BF744D95570E013A219630D98E" type="DOI">10.11646/zootaxa.4416.1.1</mods:identifier>
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</mods:mods>
<treatment id="039D87C45C38FF9C48F6FD6BFAAAF98E" ID-DOI="http://doi.org/10.5281/zenodo.5961191" ID-GBIF-Taxon="143811747" ID-Zenodo-Dep="5961191" LSID="urn:lsid:plazi:treatment:039D87C45C38FF9C48F6FD6BFAAAF98E" httpUri="http://treatment.plazi.org/id/039D87C45C38FF9C48F6FD6BFAAAF98E" lastPageNumber="40" pageId="39" pageNumber="40">
<subSubSection id="C32E65595C38FF9C48F6FD6BFE45FD12" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="8B8B36D25C38FF9C48F6FD6BFD11FD33" blockId="39.[151,735,692,751]" box="[151,735,692,718]" pageId="39" pageNumber="40">
<heading id="D0C381BE5C38FF9C48F6FD6BFD11FD33" bold="true" box="[151,735,692,718]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
<emphasis id="B940EAC05C38FF9C48F6FD6BFD11FD33" bold="true" box="[151,735,692,718]" pageId="39" pageNumber="40">
<taxonomicName id="4C344D515C38FF9C48F6FD6BFDBAFD33" authority="Barbosa &amp; Azevedo" authorityName="Barbosa &amp; Azevedo" authorityYear="2018" box="[151,628,692,718]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="aghani" status="sp. nov.">
<emphasis id="B940EAC05C38FF9C48F6FD6BFEBCFD33" bold="true" box="[151,370,692,718]" italics="true" pageId="39" pageNumber="40">Anisepyris aghani</emphasis>
Barbosa &amp; Azevedo
</taxonomicName>
,
<taxonomicNameLabel id="A27357BB5C38FF9C4AE0FD6BFD11FD33" box="[641,735,692,718]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8B36D25C38FF9C48F6FD09FE45FD12" blockId="39.[151,735,692,751]" box="[151,395,726,751]" pageId="39" pageNumber="40">
(
<figureCitation id="130F2A575C38FF9C48FEFD09FF36FD12" box="[159,248,726,751]" captionStart="FIGURE6" captionStartId="38.[151,250,1942,1964]" captionTargetBox="[224,1359,232,1878]" captionTargetId="figure@38.[182,1404,193,1921]" captionTargetPageId="38" captionText="FIGURE6. Anisepyris: aeneus species-group.AC.Mesopleuron in frontalprofile; amazonicus species-group.Head in dorsal profile.D. A. aegnori sp. nov.; E. A. aerini sp. nov.; F.A.aghani sp. nov.; G. A.amandili sp. nov.; H. A.amariei sp. nov.; I. A. amrodi sp. nov.; J. A.bregolasi sp. nov.; K.A.broddai sp. nov.; L.A. beori sp. nov." httpUri="https://zenodo.org/record/1242275/files/figure.png" pageId="39" pageNumber="40">Figs 6F</figureCitation>
; 8H; 8P; 9C)
</paragraph>
</subSubSection>
<subSubSection id="C32E65595C38FF9C48F6FCC1FF25FAAE" pageId="39" pageNumber="40" type="description">
<paragraph id="8B8B36D25C38FF9C48F6FCC1FF25FAAE" blockId="39.[151,1437,798,1651]" pageId="39" pageNumber="40">
<emphasis id="B940EAC05C38FF9C48F6FCC1FE44FCCA" bold="true" box="[151,394,798,823]" pageId="39" pageNumber="40">Description, female.</emphasis>
Body length
<quantity id="4CCC9B375C38FF9C4A4EFCC1FD54FCCB" box="[559,666,798,823]" metricMagnitude="-3" metricUnit="m" metricValue="6.85" pageId="39" pageNumber="40" unit="mm" value="6.85">6.85 mm</quantity>
. Head with green reflection; mesosoma with green reflection and metapectal-propodeal disc black; wings subhyaline; metasoma castaneous. Head wide. Mandible with four distal teeth, wide, strongly angled, with teeth different sized. Median clypeal lobe rounded, short, truncate and with blunt median tooth; median clypeal carina hardly distinct from clypeus. Frons
<taxonomicName id="4C344D515C38FF9C4BA9FC55FB8AFC5E" authorityName="Kieffer" authorityYear="1908" box="[968,1092,906,931]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="coriaceous">coriaceous</taxonomicName>
; with frontal process; antennal scrobe carinate, but weak. Eye small. Frontal angle of ocellar triangle acute; ocelli small. Vertex convex. Pronotal disc as long as wide, sparsely punctate; transverse pronotal carina complete; posterior margin with posterior pronotal sulcus. Notaulus very narrow. Mesoscutellar disc with mesoscutellar sulcus narrow. Metapectal-propodeal disc long; metapostnotal median carina complete; longitudinal ridges present; metapostnotal-propodeal suture fused with first metapostnotal carina; paraspiracular carina present and inconspicuous; metapleural carina present and conspicuous; metapectal-propodeal disc with short striae; propodeal declivity strigate. Mesopleuron with anterior fovea closed; mesopleural fovea closed; lower fovea closed; posterior fovea absent; mesopleural suture closed. Forewing with stigma developed; Rs&amp;M vein distinguished. Mesotibia spinose; claws trifid. Metasomal segments sparsely setose.
<emphasis id="B940EAC05C38FF9C49A7FB11FDF5FB1A" bold="true" box="[454,571,1230,1255]" pageId="39" pageNumber="40">Genitalia.</emphasis>
T9 with anterior arm long and overlapping 2vf; 1vv longer than genitalia body; 2vv shorter than 1vv length, smoothly narrowing apicad; 3vv short; 1vf curved downward; 2vf slender and with dorsal lap swollen posteriorly; 1rv not curved, ventral process large, and posterior process short; 2rv not curved.
</paragraph>
</subSubSection>
<subSubSection id="C32E65595C38FF9C48A6FA81FDADFA1E" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="8B8B36D25C38FF9C48A6FA81FDADFA1E" blockId="39.[151,1437,798,1651]" pageId="39" pageNumber="40">
<materialsCitation id="3B5C3C8F5C38FF9C48A6FA81FC8BFA66" ID-GBIF-Occurrence="1841141849" collectingDate="2004-05" collectionCode="AEI, AMNH" collectorName="D. B. Wahl" country="United States of America" county="Material" latitude="29.598223" location="Co" longLatPrecision="1" longitude="-82.36522" municipality="Alachua" pageId="39" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Florida" typeStatus="holotype">
<emphasis id="B940EAC05C38FF9C48A6FA81FE77FA8A" bold="true" box="[199,441,1374,1399]" pageId="39" pageNumber="40">
<collectingCounty id="62EA4E5E5C38FF9C48A6FA81FEE1FA8A" box="[199,303,1374,1399]" pageId="39" pageNumber="40">Material</collectingCounty>
examined.
</emphasis>
<typeStatus id="548F88705C38FF9C49A9FA81FDFDFA8A" box="[456,563,1374,1399]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D32FD5B5C38FF9C4A27FA80FD95FA8B" box="[582,603,1375,1398]" pageId="39" pageNumber="40" type="female"></specimenCount>
,
<collectingCountry id="F32376425C38FF9C4A13FA80FD0FFA8A" box="[626,705,1375,1399]" name="United States of America" pageId="39" pageNumber="40">U.S.A.</collectingCountry>
,
<emphasis id="B940EAC05C38FF9C4AB4FA81FCFCFA8A" box="[725,818,1374,1399]" italics="true" pageId="39" pageNumber="40">
<collectingRegion id="49F0F8305C38FF9C4AB4FA81FCE0FA8A" box="[725,814,1374,1399]" country="United States of America" name="Florida" pageId="39" pageNumber="40">Florida</collectingRegion>
,
</emphasis>
<collectingMunicipality id="6BEFACA85C38FF9C4B20FA81FC6FFA8A" box="[833,929,1374,1399]" pageId="39" pageNumber="40">Alachua</collectingMunicipality>
,
<location id="8EEB60095C38FF9C4BD4FA80FC16FA8A" LSID="urn:lsid:plazi:treatment:039D87C45C38FF9C48F6FD6BFAAAF98E:8EEB60095C38FF9C4BD4FA80FC16FA8A" box="[949,984,1375,1399]" country="United States of America" county="Material" latitude="29.598223" longLatPrecision="1" longitude="-82.36522" municipality="Alachua" name="Co" pageId="39" pageNumber="40" stateProvince="Florida">Co</location>
[unty],
<location id="8EEB60095C38FF9C4C51FA81FB7EFA8A" LSID="urn:lsid:plazi:treatment:039D87C45C38FF9C48F6FD6BFAAAF98E:8EEB60095C38FF9C4C51FA81FB7EFA8A" box="[1072,1200,1374,1399]" country="United States of America" county="Material" latitude="29.598223" longLatPrecision="1" longitude="-82.36522" municipality="Alachua" name="Gainesville" pageId="39" pageNumber="40" stateProvince="Florida">Gainesville</location>
<collectionCode id="ED25AE175C38FF9C4CDEFA80FB3EFA8A" box="[1215,1264,1375,1399]" country="USA" httpUri="http://grbio.org/cool/rn7d-z1r2" name="American Entomological Institute" pageId="39" pageNumber="40">AEI</collectionCode>
,
<geoCoordinate id="EE0050155C38FF9C4D64FA81FA52FA8A" box="[1285,1436,1374,1399]" direction="north" orientation="latitude" pageId="39" pageNumber="40" precision="1" value="29.598223">29°35'53.6''N</geoCoordinate>
<geoCoordinate id="EE0050155C38FF9C48F6FA5DFEFAFA66" box="[151,308,1410,1435]" direction="west" orientation="longitude" pageId="39" pageNumber="40" precision="1" value="-82.36522">82°21'54.8''W</geoCoordinate>
,
<collectingDate id="EFCEE9FA5C38FF9C4921FA5CFE78FA66" box="[320,438,1411,1435]" pageId="39" pageNumber="40" value="2004-05">
<date id="FF8A10125C38FF9C4921FA5CFE78FA66" box="[320,438,1411,1435]" pageId="39" pageNumber="40" value="2004-05">May 2004</date>
</collectingDate>
,
<collectingMethod id="52754EC55C38FF9C49A0FA5DFD81FA67" box="[449,591,1410,1435]" pageId="39" pageNumber="40">Malaise trap</collectingMethod>
,
<collectorName id="26C153045C38FF9C4A3AFA5CFD1CFA66" box="[603,722,1410,1435]" pageId="39" pageNumber="40">D.B. Wahl</collectorName>
(
<collectionCode id="ED25AE175C38FF9C4A80FA5CFCF3FA66" box="[737,829,1411,1435]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="39" pageNumber="40">AMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B5C3C8F5C38FF9C4B2EFA5CFE1FFA42" ID-GBIF-Occurrence="1841141834" collectionCode="AMNH" collectorName="D. R. Smith" country="United States of America" location="Essex Co" pageId="39" pageNumber="40" specimenCount="1" specimenCount-female="1" stateProvince="Virginia" typeStatus="paratype">
<typeStatus id="548F88705C38FF9C4B2EFA5CFC72FA66" box="[847,956,1411,1435]" pageId="39" pageNumber="40" type="paratype">Paratypes</typeStatus>
(3).
<collectingCountry id="F32376425C38FF9C4B93FA5CFB8FFA66" box="[1010,1089,1411,1435]" name="United States of America" pageId="39" pageNumber="40">U.S.A.</collectingCountry>
,
<collectingRegion id="49F0F8305C38FF9C4C2CFA5BFB69FA66" box="[1101,1191,1412,1435]" country="United States of America" name="Virginia" pageId="39" pageNumber="40">
<emphasis id="B940EAC05C38FF9C4C2CFA5BFB69FA66" box="[1101,1191,1412,1435]" italics="true" pageId="39" pageNumber="40">Virginia</emphasis>
</collectingRegion>
,
<specimenCount id="9D32FD5B5C38FF9C4CD7FA5CFB2EFA67" box="[1206,1248,1411,1435]" pageId="39" pageNumber="40" type="female">1 ♀</specimenCount>
,
<location id="8EEB60095C38FF9C4C8FFA5CFA97FA66" LSID="urn:lsid:plazi:treatment:039D87C45C38FF9C48F6FD6BFAAAF98E:8EEB60095C38FF9C4C8FFA5CFA97FA66" box="[1262,1369,1411,1435]" country="United States of America" name="Essex Co" pageId="39" pageNumber="40" stateProvince="Virginia">Essex Co</location>
[unty] 1999,
<collectorName id="26C153045C38FF9C48BCFA78FE93FA42" box="[221,349,1446,1471]" pageId="39" pageNumber="40">D.R. Smith</collectorName>
(
<collectionCode id="ED25AE175C38FF9C490CFA78FE07FA42" box="[365,457,1447,1471]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="39" pageNumber="40">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5C3C8F5C38FF9C49BDFA79FD91FA1E" ID-GBIF-Occurrence="1841141861" collectingDate="2004-05" collectionCode="AEI, AMNH" collectorName="D. B. Wahl" country="United States of America" county="Alachua" latitude="29.598223" location="Gainesville" longLatPrecision="1" longitude="-82.36522" municipality="Co" pageId="39" pageNumber="40" specimenCount="2" specimenCount-female="2" stateProvince="Florida" typeStatus="paratype">
<collectingRegion id="49F0F8305C38FF9C49BDFA79FDFFFA42" box="[476,561,1446,1471]" country="United States of America" name="Florida" pageId="39" pageNumber="40">
<emphasis id="B940EAC05C38FF9C49BDFA79FDFFFA42" box="[476,561,1446,1471]" italics="true" pageId="39" pageNumber="40">Florida</emphasis>
</collectingRegion>
,
<specimenCount id="9D32FD5B5C38FF9C4A20FA78FDA2FA43" box="[577,620,1447,1471]" pageId="39" pageNumber="40" type="female">2 ♀</specimenCount>
,
<collectingCounty id="62EA4E5E5C38FF9C4A1AFA79FD15FA42" box="[635,731,1446,1471]" pageId="39" pageNumber="40">Alachua</collectingCounty>
,
<collectingMunicipality id="6BEFACA85C38FF9C4A86FA78FCC4FA42" box="[743,778,1447,1471]" pageId="39" pageNumber="40">Co</collectingMunicipality>
[unty],
<location id="8EEB60095C38FF9C4B3AFA79FC15FA42" LSID="urn:lsid:plazi:treatment:039D87C45C38FF9C48F6FD6BFAAAF98E:8EEB60095C38FF9C4B3AFA79FC15FA42" box="[859,987,1446,1471]" country="United States of America" county="Alachua" latitude="29.598223" longLatPrecision="1" longitude="-82.36522" municipality="Co" name="Gainesville" pageId="39" pageNumber="40" stateProvince="Florida">Gainesville</location>
<collectionCode id="ED25AE175C38FF9C4B83FA78FBDAFA42" box="[994,1044,1447,1471]" country="USA" httpUri="http://grbio.org/cool/rn7d-z1r2" name="American Entomological Institute" pageId="39" pageNumber="40">AEI</collectionCode>
,
<geoCoordinate id="EE0050155C38FF9C4C41FA79FB79FA42" box="[1056,1207,1446,1471]" direction="north" orientation="latitude" pageId="39" pageNumber="40" precision="1" value="29.598223">29°35'53.6''N</geoCoordinate>
<geoCoordinate id="EE0050155C38FF9C4CDEFA79FA92FA42" box="[1215,1372,1446,1471]" direction="west" orientation="longitude" pageId="39" pageNumber="40" precision="1" value="-82.36522">82°21'54.8''W</geoCoordinate>
,
<collectingDate id="EFCEE9FA5C38FF9C4D08FA78FF1CFA1E" pageId="39" pageNumber="40" value="2004-05">
<date id="FF8A10125C38FF9C4D08FA78FF1CFA1E" pageId="39" pageNumber="40" value="2004-05">May 2004</date>
</collectingDate>
,
<collectingMethod id="52754EC55C38FF9C48BCFA15FEA5FA1F" box="[221,363,1482,1507]" pageId="39" pageNumber="40">Malaise trap</collectingMethod>
,
<collectorName id="26C153045C38FF9C4917FA14FE22FA1E" box="[374,492,1482,1507]" pageId="39" pageNumber="40">D.B. Wahl</collectorName>
(
<collectionCode id="ED25AE175C38FF9C499AFA14FD99FA1E" box="[507,599,1483,1507]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="39" pageNumber="40">AMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E65595C38FF9C48A6FA31FDB0F9FA" box="[199,638,1518,1543]" pageId="39" pageNumber="40" type="distribution">
<paragraph id="8B8B36D25C38FF9C48A6FA31FDB0F9FA" blockId="39.[151,1437,798,1651]" box="[199,638,1518,1543]" pageId="39" pageNumber="40">
<emphasis id="B940EAC05C38FF9C48A6FA31FEAEF9FA" bold="true" box="[199,352,1518,1543]" pageId="39" pageNumber="40">Distribution.</emphasis>
<collectingCountry id="F32376425C38FF9C4906FA30FE56F9FA" box="[359,408,1519,1543]" name="United States of America" pageId="39" pageNumber="40">U.S.</collectingCountry>
A:
<collectingRegion id="49F0F8305C38FF9C49DDFA31FDD4F9FA" box="[444,538,1518,1543]" country="United States of America" name="Virginia" pageId="39" pageNumber="40">Virginia</collectingRegion>
,
<collectingRegion id="49F0F8305C38FF9C4A44FA31FDB4F9FA" box="[549,634,1518,1543]" country="United States of America" name="Florida" pageId="39" pageNumber="40">Florida</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E65595C38FF9C48A6F9CDFAAAF98E" pageId="39" pageNumber="40" type="discussion">
<paragraph id="8B8B36D25C38FF9C48A6F9CDFAAAF98E" blockId="39.[151,1437,798,1651]" pageId="39" pageNumber="40">
<emphasis id="B940EAC05C38FF9C48A6F9CDFEF5F9D6" bold="true" box="[199,315,1554,1579]" pageId="39" pageNumber="40">Remarks.</emphasis>
This species differs from
<taxonomicName id="4C344D515C38FF9C4AE8F9CCFC76F9D6" authority="Evans, 1966" authorityName="Evans" authorityYear="1966" box="[649,952,1554,1579]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="jocundus">
<emphasis id="B940EAC05C38FF9C4AE8F9CCFCDBF9D6" box="[649,789,1554,1579]" italics="true" pageId="39" pageNumber="40">A. jocundus</emphasis>
<bibRefCitation id="EFA54B235C38FF9C4B44F9CCFC76F9D6" box="[805,952,1554,1579]" pageId="39" pageNumber="40" refId="ref280374" refString="Evans, H. E. (1966) A revision of the genus Anisepyris Kieffer (Hymenoptera. Bethylidae). Studia Entomologica, 9, 1 - 120." type="journal article">Evans, 1966</bibRefCitation>
</taxonomicName>
by having the frons without a median impression; the ocelli very small; and the mesopleuron with all foveae closed, whereas
<taxonomicName id="4C344D515C38FF9C4C1DF9E8FACFF9B2" authorityName="Evans" authorityYear="1966" box="[1148,1281,1590,1615]" class="Insecta" family="Bethylidae" genus="Anisepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="jocundus">
<emphasis id="B940EAC05C38FF9C4C1DF9E8FACFF9B2" box="[1148,1281,1590,1615]" italics="true" pageId="39" pageNumber="40">A. jocundus</emphasis>
</taxonomicName>
has the frons with a median impression; the mesopleuron with the lower foveae closed; and the mesopleural suture opened.
</paragraph>
</subSubSection>
</treatment>
</document>