422 lines
52 KiB
XML
422 lines
52 KiB
XML
<document id="39594EEDE7A8EA7BDF61A6484F61BEA7" ID-DOI="10.26028/cybium/2019-434-002" ID-ISSN="2101-0315" ID-Zenodo-Dep="12711559" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720644388693" checkinUser="felipe" docAuthor="Moritz, Timo, Straube, Nicolas & Neumann, Dirk" docDate="2019" docId="03915B38EF0CFFCDFC2717EE5DB88352" docLanguage="en" docName="Cybium.43.4.311-329.pdf" docOrigin="Cybium 43 (4)" docStyle="DocumentStyle:EAB95795642928CB986F0923DE58F0B7.1:Cybium.2019-.journal_article" docStyleId="EAB95795642928CB986F0923DE58F0B7" docStyleName="Cybium.2019-.journal_article" docStyleVersion="1" docTitle="Garra napata Moritz, Straube & Neumann, 2019, new species" docType="treatment" docVersion="2" lastPageNumber="319" masterDocId="FFA82340EF0AFFC5FFFB13345E168120" masterDocTitle="The Garra species (Cyprinidae) of the Main Nile basin with description of three new species" masterLastPageNumber="329" masterPageNumber="311" pageNumber="317" updateTime="1720649965938" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="CBA857D20B4461D00F59191EA5F74503">
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<mods:title id="53C3E4B2F3A6230D89997C5D15979C18">The Garra species (Cyprinidae) of the Main Nile basin with description of three new species</mods:title>
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</mods:titleInfo>
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<mods:name id="2975204889CAB7BAB2E6A59D09F04218" type="personal">
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<mods:role id="E5CC88B7B8632D0A50C5805B858D25A4">
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<mods:roleTerm id="17A7E1588E0464FCC92C8D7FC952665C">Author</mods:roleTerm>
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<mods:namePart id="243685AA667F0787889F9924C4BE7797">Moritz, Timo</mods:namePart>
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<mods:roleTerm id="8056AE072E4BB107B3FF63E105E4ACDC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9C0029F8613A9254DCD08E6AE3228FFF">Straube, Nicolas</mods:namePart>
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</mods:name>
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<mods:name id="DF4EB20F80463486B5BF987B55F07E38" type="personal">
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<mods:roleTerm id="8358A993898FCEB4120EB6F406268B70">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="72F0DC23F4E8A6DF113F981B39D49690">Neumann, Dirk</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4928E431B0868F4139495C82E278D874">text</mods:typeOfResource>
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<mods:titleInfo id="09C82D6891EC210BD69460608162FB84">
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<mods:title id="226E266B0ABF406C6B82965BC7DA7921">Cybium</mods:title>
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<mods:part id="46D61F0A2F93532F0D55CB3E66A40A67">
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<mods:date id="B869605292CE708615BF4BDC75F202D0">2019</mods:date>
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<mods:detail id="A8984302770D4CFE51C6CE3989357CC1" type="volume">
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<mods:number id="17138D59C04B6BF25CF25F4276FCD81E">43</mods:number>
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</mods:detail>
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<mods:detail id="6AE55D4C0546EDDFC99EA25271609490" type="issue">
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<mods:number id="AA367EF2B997FCCB1DE12D6E440B38B6">4</mods:number>
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<mods:start id="B744BB552091DB0618DDD184FD8DA58C">311</mods:start>
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<mods:end id="1F9B85107ECBF51F0633E9D650A19630">329</mods:end>
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<mods:classification id="323B7588EC78D25F0776491181CF170F">journal article</mods:classification>
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<mods:identifier id="5354BEA9EA954BCBA8B07D3D750C0E15" type="DOI">10.26028/cybium/2019-434-002</mods:identifier>
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<mods:identifier id="BECDDD32BA3E8FB7CFC9282C6D376D09" type="ISSN">2101-0315</mods:identifier>
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<mods:identifier id="1BD404ADA004AD1503EC4C49EF6B538C" type="Zenodo-Dep">12711559</mods:identifier>
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<treatment id="03915B38EF0CFFCDFC2717EE5DB88352" LSID="urn:lsid:plazi:treatment:03915B38EF0CFFCDFC2717EE5DB88352" httpUri="http://treatment.plazi.org/id/03915B38EF0CFFCDFC2717EE5DB88352" lastPageId="8" lastPageNumber="319" pageId="6" pageNumber="317">
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<subSubSection id="C322B9A5EF0CFFC3FC2717EE5B1685D2" box="[988,1280,1242,1266]" pageId="6" pageNumber="317" type="nomenclature">
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<paragraph id="8B87EA2EEF0CFFC3FC2717EE5B1685D2" blockId="6.[988,1280,1242,1266]" box="[988,1280,1242,1266]" pageId="6" pageNumber="317">
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<heading id="D0CF5D42EF0CFFC3FC2717EE5B1685D2" bold="true" box="[988,1280,1242,1266]" centered="true" fontSize="9" level="2" pageId="6" pageNumber="317" reason="2">
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<emphasis id="B94C363CEF0CFFC3FC2717EE5B1685D2" bold="true" box="[988,1280,1242,1266]" pageId="6" pageNumber="317">
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<taxonomicName id="4C3891ADEF0CFFC3FC2717EE5A6785D2" authorityName="Moritz & Straube & Neumann" authorityYear="2019" box="[988,1137,1242,1266]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="317" phylum="Chordata" rank="species" species="napata" status="sp. nov.">
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<emphasis id="B94C363CEF0CFFC3FC2717EE5A6785D2" bold="true" box="[988,1137,1242,1266]" italics="true" pageId="6" pageNumber="317">Garra napata</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27F8B47EF0CFFC3FB8617EE5B1685D2" box="[1149,1280,1242,1266]" pageId="6" pageNumber="317" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C322B9A5EF0CFFC3FC0617CE5B5F8472" pageId="6" pageNumber="317" type="description">
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||
<paragraph id="8B87EA2EEF0CFFC3FC0617CE5AC98432" blockId="6.[1021,1247,1274,1298]" box="[1021,1247,1274,1298]" pageId="6" pageNumber="317">
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||
(
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<figureCitation id="1303F6ABEF0CFFC3FBFF17CE5A498432" box="[1028,1119,1274,1298]" captionStart-0="Figure 8" captionStart-1="Figure 9" captionStartId-0="7.[813,876,685,707]" captionStartId-1="7.[813,877,1511,1533]" captionTargetBox-0="[813,1456,215,668]" captionTargetBox-1="[813,1456,787,1494]" captionTargetId-0="figure-601@7.[813,1456,215,668]" captionTargetId-1="figure-630@7.[813,1456,787,1494]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="Figure 8. – Garra napata sp. nov., holotype, ZSM 47244 (ex ZSM 39815#3), 39.9 mm SL, Sudan: Nile: Tarag Island, 10 Apr. 2008. Scale bar = 10 mm." captionText-1="Figure 9. – Live specimens of Garra napata sp. nov. All from the area of Tarag Island, 4th cataract, Nile, Sudan. A-B: ~38 mm SL, 9 Apr. 2008; C: ~50 mm SL, 10 Apr. 2008." figureDoi-0="http://doi.org/10.5281/zenodo.12711579" figureDoi-1="http://doi.org/10.5281/zenodo.12711583" httpUri-0="https://zenodo.org/record/12711579/files/figure.png" httpUri-1="https://zenodo.org/record/12711583/files/figure.png" pageId="6" pageNumber="317">Figs 8-9</figureCitation>
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, Tabs II-III)
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</paragraph>
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<paragraph id="8B87EA2EEF0CFFC3FCA9160F5B5F8472" blockId="6.[850,1353,1338,1362]" box="[850,1353,1338,1362]" pageId="6" pageNumber="317">
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<taxonomicName id="4C3891ADEF0CFFC3FCA9160F5D838472" authorityName="Moritz & Straube & Neumann" authorityYear="2019" box="[850,917,1339,1362]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="317" phylum="Chordata" rank="genus" status="sp. nov.">
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<emphasis id="B94C363CEF0CFFC3FCA9160F5D838472" box="[850,917,1339,1362]" italics="true" pageId="6" pageNumber="317">Garra</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27F8B47EF0CFFC3FC60160F5DFB8472" box="[923,1005,1339,1362]" pageId="6" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
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“flathead”
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<bibRefCitation id="EFA997DFEF0CFFC3FB9E160F5B5F8472" author="NEUMANN D. & OBERMAIER H. & MORITZ T." box="[1125,1353,1338,1362]" pageId="6" pageNumber="317" pagination="287 - 317" refId="ref15302" refString="NEUMANN D., OBERMAIER H. & MORITZ T., 2016. - Annotated check list for fishes of the Main Nile Basin in the Sudan and Egypt based on recent specimens records (2006 - 2015). Cybium, 40 (4): 287 - 317." type="journal article" year="2016">
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Neumann
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<emphasis id="B94C363CEF0CFFC3FB29160F5B168472" box="[1234,1280,1338,1362]" italics="true" pageId="6" pageNumber="317">et al</emphasis>
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||
., 2016
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</bibRefCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C322B9A5EF0CFFC2FCD6164E5F8E80D2" lastPageId="7" lastPageNumber="318" pageId="6" pageNumber="317" type="materials_examined">
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||
<paragraph id="8B87EA2EEF0CFFC3FCD6164E5D8684B2" blockId="6.[812,1456,1402,1907]" box="[813,912,1402,1426]" pageId="6" pageNumber="317">
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||
<emphasis id="B94C363CEF0CFFC3FCD6164E5D8684B2" bold="true" box="[813,912,1402,1426]" pageId="6" pageNumber="317">Material</emphasis>
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||
</paragraph>
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<paragraph id="8B87EA2EEF0CFFC3FCA916AE5A878713" blockId="6.[812,1456,1402,1907]" pageId="6" pageNumber="317">
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<materialsCitation id="3B50E073EF0CFFC3FCA916AE5A878713" collectingDate="2008-04-10" collectionCode="ZSM" collectorName="T. Moritz & Moritz, D & N. Pollath" country="Sudan" county="Merowe" latitude="18.59" location="Nile" longLatPrecision="765" longitude="31.97" municipality="Takar village" pageId="6" pageNumber="317" specimenCode="ZSM 47244, ZSM 39815, 3" specimenCount="1" typeStatus="holotype">
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<emphasis id="B94C363CEF0CFFC3FCA916AE5DA88492" box="[850,958,1434,1458]" italics="true" pageId="6" pageNumber="317">
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||
<typeStatus id="5483548CEF0CFFC3FCA916AE5DAC8492" box="[850,954,1434,1458]" pageId="6" pageNumber="317" type="holotype">Holotype</typeStatus>
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||
.
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||
</emphasis>
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–
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<specimenCode id="DB9E4255EF0CFFC3FC2216AF5A4D8493" box="[985,1115,1435,1459]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="6" pageNumber="317" type="Museum">ZSM 47244</specimenCode>
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(ex
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<specimenCode id="DB9E4255EF0CFFC3FB7716AF5B048492" box="[1164,1298,1434,1459]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="6" pageNumber="317" type="Museum">ZSM 39815</specimenCode>
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||
, #
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||
<specimenCode id="DB9E4255EF0CFFC3FAD616AF5B2C8493" box="[1325,1338,1435,1459]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="6" pageNumber="317" type="Museum">3</specimenCode>
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||
),
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||
<quantity id="4CC047CBEF0CFFC3FAB416AF5BB88492" box="[1359,1454,1435,1459]" metricMagnitude="-2" metricUnit="m" metricValue="3.9899999999999998" pageId="6" pageNumber="317" unit="mm" value="39.9">39.9 mm</quantity>
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SL,
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<collectingCountry id="F32FAABEEF0CFFC3FCA0168E5DB284F2" box="[859,932,1466,1490]" name="Sudan" pageId="6" pageNumber="317">Sudan</collectingCountry>
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||
:
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||
<location id="8EE7BCF5EF0CFFC3FC55168E5DF784F2" LSID="urn:lsid:plazi:treatment:03915B38EF0CFFCDFC2717EE5DB88352:8EE7BCF5EF0CFFC3FC55168E5DF784F2" box="[942,993,1466,1490]" country="Sudan" county="Merowe" latitude="18.59" longLatPrecision="765" longitude="31.97" municipality="Takar village" name="Nile" pageId="6" pageNumber="317">Nile</location>
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||
: rapids and riffles at small northern island opposite of
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||
<location id="8EE7BCF5EF0CFFC3FC5716EF5DFF84D3" LSID="urn:lsid:plazi:treatment:03915B38EF0CFFCDFC2717EE5DB88352:8EE7BCF5EF0CFFC3FC5716EF5DFF84D3" box="[940,1001,1499,1523]" country="Sudan" county="Merowe" latitude="18.59" longLatPrecision="765" longitude="31.97" municipality="Takar village" name="Tarag" pageId="6" pageNumber="317">Tarag</location>
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main island, close to
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<collectingMunicipality id="6BE37054EF0CFFC3FB2816EE5B7584D2" box="[1235,1379,1498,1522]" pageId="6" pageNumber="317">Takar village</collectingMunicipality>
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, north-east of
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||
<collectingCounty id="62E692A2EF0CFFC3FC8116CF5DC58732" box="[890,979,1531,1554]" pageId="6" pageNumber="317">Merowe</collectingCounty>
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(
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<geoCoordinate id="EE0C8CE9EF0CFFC3FC1A16CE5A288732" box="[993,1086,1530,1554]" degrees="18.59" direction="north" orientation="latitude" pageId="6" pageNumber="317" precision="555" value="18.59">N18.59°</geoCoordinate>
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<geoCoordinate id="EE0C8CE9EF0CFFC3FBBF16CF5AB68733" box="[1092,1184,1531,1555]" degrees="31.97" direction="east" orientation="longitude" pageId="6" pageNumber="317" precision="555" value="31.97">E31.97°</geoCoordinate>
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), coll.
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||
<collectorName id="26CD8FF8EF0CFFC3FB1C16CF5B588732" box="[1255,1358,1530,1554]" pageId="6" pageNumber="317">T. Moritz</collectorName>
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||
, D. Neumann &
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||
<collectorName id="26CD8FF8EF0CFFC3FC72152F5DE18713" box="[905,1015,1563,1587]" pageId="6" pageNumber="317">N. Pöllath</collectorName>
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,
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<date id="FF86CCEEEF0CFFC3FBFA152F5A988713" box="[1025,1166,1563,1587]" pageId="6" pageNumber="317" value="2008-04-10">
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<collectingDate id="EFC23506EF0CFFC3FBFA152F5A988713" box="[1025,1166,1563,1587]" pageId="6" pageNumber="317" value="2008-04-10">10 Apr. 2008</collectingDate>
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||
</date>
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||
.
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||
</materialsCitation>
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</paragraph>
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||
<paragraph id="8B87EA2EEF0CFFC2FCA9150F5F8E80D2" blockId="6.[812,1456,1402,1907]" lastBlockId="7.[132,775,217,498]" lastPageId="7" lastPageNumber="318" pageId="6" pageNumber="317">
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||
<materialsCitation id="3B50E073EF0CFFC3FCA9150F5A0F8753" collectingDate="2008-04-10" collectionCode="ZSM" collectorName="T. Moritz & Moritz, D & N. Pollath" country="Sudan" county="Merowe" latitude="18.59" location="Nile" longLatPrecision="765" longitude="31.97" municipality="Takar village" pageId="6" pageNumber="317" specimenCode="ZSM 39815" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="B94C363CEF0CFFC3FCA9150F5DDF8772" box="[850,969,1595,1618]" italics="true" pageId="6" pageNumber="317">
|
||
<typeStatus id="5483548CEF0CFFC3FCA9150F5DD38772" box="[850,965,1595,1618]" pageId="6" pageNumber="317" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
–
|
||
<specimenCode id="DB9E4255EF0CFFC3FC1F150F5A718772" box="[996,1127,1594,1619]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="6" pageNumber="317" type="Museum">ZSM 39815</specimenCode>
|
||
(6 now 3),
|
||
<quantity id="4CC047CBEF0CFFC3FB1C150E5B948772" box="[1255,1410,1594,1618]" metricMagnitude="-2" metricUnit="m" metricValue="3.2399999999999998" metricValueMax="3.85" metricValueMin="2.63" pageId="6" pageNumber="317" unit="mm" value="32.4" valueMax="38.5" valueMin="26.3">26.3-38.5 mm</quantity>
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||
SL, same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B50E073EF0CFFC3FBD9156F5AF787B3" collectingDate="2008-04-10" collectionCode="DMM" collectorName="T. Moritz & Moritz, D & N. Pollath" country="Sudan" county="Merowe" latitude="18.59" location="Nile" longLatPrecision="765" longitude="31.97" municipality="Takar village" pageId="6" pageNumber="317" specimenCode="ZSM 39815" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED2972EBEF0CFFC3FBD9156F5A738752" box="[1058,1125,1627,1650]" pageId="6" pageNumber="317">DMM</collectionCode>
|
||
IE/15073 (2) (ex
|
||
<specimenCode id="DB9E4255EF0CFFC3FADA156F5BB28752" box="[1313,1444,1626,1651]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="6" pageNumber="317" type="Museum">ZSM 39815</specimenCode>
|
||
),
|
||
<quantity id="4CC047CBEF0CFFC3FCD6154F5DD487B2" box="[813,962,1659,1683]" metricMagnitude="-2" metricUnit="m" metricValue="3.26" metricValueMax="3.85" metricValueMin="2.67" pageId="6" pageNumber="317" unit="mm" value="32.6" valueMax="38.5" valueMin="26.7">26.7-38.5 mm</quantity>
|
||
SL, same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B50E073EF0CFFC3FB10154F5A3187D3" collectingDate="1983-09" collectionCode="MRAC" collectorName="W. van Neer & R. Lauwers" country="Egypt" latitude="24.08" location="Elephantine Island" longLatPrecision="752" longitude="32.88" municipality="Nile" pageId="6" pageNumber="317" specimenCode="MRAC 1983-030" specimenCount="1" stateProvince="Aswan" typeStatus="paratype">
|
||
<specimenCode id="DB9E4255EF0CFFC3FB10154F5BBE87B3" box="[1259,1448,1659,1683]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="6" pageNumber="317" type="Museum">MRAC 1983-030</specimenCode>
|
||
- P-00887-0088 (2),
|
||
<quantity id="4CC047CBEF0CFFC3FC0F15AF5A9E8792" box="[1012,1160,1691,1715]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="7.029999999999999" metricValueMin="6.97" pageId="6" pageNumber="317" unit="mm" value="70.0" valueMax="70.3" valueMin="69.7">69.7-70.3 mm</quantity>
|
||
SL,
|
||
<collectingCountry id="F32FAABEEF0CFFC3FB4315AF5AEA8793" box="[1208,1276,1691,1715]" name="Egypt" pageId="6" pageNumber="317">Egypt</collectingCountry>
|
||
:
|
||
<collectingMunicipality id="6BE37054EF0CFFC3FAFF15AF5B248793" box="[1284,1330,1691,1715]" pageId="6" pageNumber="317">Nile</collectingMunicipality>
|
||
at
|
||
<location id="8EE7BCF5EF0CFFC3FAB415AF5DB787F3" LSID="urn:lsid:plazi:treatment:03915B38EF0CFFCDFC2717EE5DB88352:8EE7BCF5EF0CFFC3FAB415AF5DB787F3" country="Egypt" latitude="24.08" longLatPrecision="752" longitude="32.88" municipality="Nile" name="Elephantine Island" pageId="6" pageNumber="317" stateProvince="Aswan">Elephantine Island</location>
|
||
,
|
||
<collectingRegion id="49FC24CCEF0CFFC3FC50158F5DE087F3" box="[939,1014,1723,1747]" country="Egypt" name="Aswan" pageId="6" pageNumber="317">Aswan</collectingRegion>
|
||
(
|
||
<geoCoordinate id="EE0C8CE9EF0CFFC3FBFF158F5A7787F3" box="[1028,1121,1723,1747]" degrees="24.08" direction="north" orientation="latitude" pageId="6" pageNumber="317" precision="555" value="24.08">N24.08°</geoCoordinate>
|
||
<geoCoordinate id="EE0C8CE9EF0CFFC3FB93158F5AD287F3" box="[1128,1220,1723,1747]" degrees="32.88" direction="east" orientation="longitude" pageId="6" pageNumber="317" precision="555" value="32.88">E32.88°</geoCoordinate>
|
||
), coll.
|
||
<collectorName id="26CD8FF8EF0CFFC3FAF6158F5B8287F2" box="[1293,1428,1723,1746]" pageId="6" pageNumber="317">W. van Neer</collectorName>
|
||
&
|
||
<collectorName id="26CD8FF8EF0CFFC3FCD615EF5DBA87D2" box="[813,940,1755,1778]" pageId="6" pageNumber="317">R. Lauwers</collectorName>
|
||
,
|
||
<date id="FF86CCEEEF0CFFC3FC4315EF5A3187D3" box="[952,1063,1755,1779]" pageId="6" pageNumber="317" value="1983-09">
|
||
<collectingDate id="EFC23506EF0CFFC3FC4315EF5A3187D3" box="[952,1063,1755,1779]" pageId="6" pageNumber="317" value="1983-09">Sep. 1983</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B50E073EF0CFFC3FBC815EF5B798673" collectingDate="2008-04-09" collectionCode="ZSM" collectorName="T. Moritz & D. Neumann & N. Pollath" country="Sudan" county="Nile" latitude="18.57" location="Takar village" longLatPrecision="765" longitude="31.96" municipality="Tarag Island" pageId="6" pageNumber="317" specimenCode="ZSM 39814" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9E4255EF0CFFC3FBC815EF5AA387D3" box="[1075,1205,1755,1779]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="6" pageNumber="317" type="Museum">ZSM 39814</specimenCode>
|
||
(3),
|
||
<quantity id="4CC047CBEF0CFFC3FB1215EF5B9487D2" box="[1257,1410,1755,1779]" metricMagnitude="-2" metricUnit="m" metricValue="3.315" metricValueMax="4.08" metricValueMin="2.55" pageId="6" pageNumber="317" unit="mm" value="33.15" valueMax="40.8" valueMin="25.5">25.5-40.8 mm</quantity>
|
||
SL,
|
||
<collectingCountry id="F32FAABEEF0CFFC3FCD615CF5D608633" box="[813,886,1787,1811]" name="Sudan" pageId="6" pageNumber="317">Sudan</collectingCountry>
|
||
:
|
||
<collectingCounty id="62E692A2EF0CFFC3FC7B15CF5DB98633" box="[896,943,1787,1811]" pageId="6" pageNumber="317">Nile</collectingCounty>
|
||
at a smaller island close to
|
||
<collectingMunicipality id="6BE37054EF0CFFC3FB1B15CF5B7E8633" box="[1248,1384,1787,1811]" pageId="6" pageNumber="317">Tarag Island</collectingMunicipality>
|
||
opposite
|
||
<location id="8EE7BCF5EF0CFFC3FCA3142F5DF18613" LSID="urn:lsid:plazi:treatment:03915B38EF0CFFCDFC2717EE5DB88352:8EE7BCF5EF0CFFC3FCA3142F5DF18613" box="[856,999,1819,1843]" country="Sudan" county="Nile" latitude="18.57" longLatPrecision="765" longitude="31.96" municipality="Tarag Island" name="Takar village" pageId="6" pageNumber="317">Takar village</location>
|
||
, north-east of
|
||
<location id="8EE7BCF5EF0CFFC3FB84142F5ACE8612" LSID="urn:lsid:plazi:treatment:03915B38EF0CFFCDFC2717EE5DB88352:8EE7BCF5EF0CFFC3FB84142F5ACE8612" box="[1151,1240,1819,1842]" country="Sudan" county="Nile" latitude="18.57" longLatPrecision="765" longitude="31.96" municipality="Tarag Island" name="Merowe" pageId="6" pageNumber="317">Merowe</location>
|
||
(
|
||
<geoCoordinate id="EE0C8CE9EF0CFFC3FB1D142F5B548613" box="[1254,1346,1819,1843]" degrees="18.57" direction="north" orientation="latitude" pageId="6" pageNumber="317" precision="555" value="18.57">N18.57°</geoCoordinate>
|
||
<geoCoordinate id="EE0C8CE9EF0CFFC3FAB3142F5BB28613" box="[1352,1444,1819,1843]" degrees="31.96" direction="east" orientation="longitude" pageId="6" pageNumber="317" precision="555" value="31.96">E31.96°</geoCoordinate>
|
||
), coll.
|
||
<collectorName id="26CD8FF8EF0CFFC3FC9A140F5DD08673" box="[865,966,1851,1875]" pageId="6" pageNumber="317">T. Moritz</collectorName>
|
||
,
|
||
<collectorName id="26CD8FF8EF0CFFC3FC2B140F5A4E8672" box="[976,1112,1851,1874]" pageId="6" pageNumber="317">D. Neumann</collectorName>
|
||
&
|
||
<collectorName id="26CD8FF8EF0CFFC3FB83140F5AF08673" box="[1144,1254,1851,1875]" pageId="6" pageNumber="317">N. Pöllath</collectorName>
|
||
,
|
||
<date id="FF86CCEEEF0CFFC3FB0B140F5B798673" box="[1264,1391,1851,1875]" pageId="6" pageNumber="317" value="2008-04-09">
|
||
<collectingDate id="EFC23506EF0CFFC3FB0B140F5B798673" box="[1264,1391,1851,1875]" pageId="6" pageNumber="317" value="2008-04-09">9 Apr. 2008</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B50E073EF0CFFC2FA83140F5F298011" collectingDate="2008-04-09" collectionCode="ZSM" collectorName="T. Moritz & D. Neumann & N. Pollath" country="Sudan" county="Nile" lastPageId="7" lastPageNumber="318" latitude="18.58" location="Takar village" longLatPrecision="765" longitude="31.94" municipality="Tarag" pageId="6" pageNumber="317" specimenCode="ZSM 39816" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9E4255EF0CFFC3FA83140F5D648653" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="6" pageNumber="317" type="Museum">ZSM 39816</specimenCode>
|
||
(2),
|
||
<quantity id="4CC047CBEF0CFFC3FC50146F5A5C8652" box="[939,1098,1883,1907]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" metricValueMax="2.38" metricValueMin="1.88" pageId="6" pageNumber="317" unit="mm" value="21.3" valueMax="23.8" valueMin="18.8">18.8-23.8 mm</quantity>
|
||
SL,
|
||
<collectingCountry id="F32FAABEEF0CFFC3FB78146F5AD88653" box="[1155,1230,1883,1907]" name="Sudan" pageId="6" pageNumber="317">Sudan</collectingCountry>
|
||
:
|
||
<collectingCounty id="62E692A2EF0CFFC3FB22146F5B1F8653" box="[1241,1289,1883,1907]" pageId="6" pageNumber="317">Nile</collectingCounty>
|
||
at small rapid at right bank of
|
||
<collectingMunicipality id="6BE37054EF0DFFC2FEC113EE5F6F81D2" box="[314,377,218,242]" pageId="7" pageNumber="318">Tarag</collectingMunicipality>
|
||
main island, close to
|
||
<location id="8EE7BCF5EF0DFFC2FD8813ED5D1081D1" LSID="urn:lsid:plazi:treatment:03915B38EF0CFFCDFC2717EE5DB88352:8EE7BCF5EF0DFFC2FD8813ED5D1081D1" box="[627,774,217,241]" country="Sudan" county="Nile" latitude="18.58" longLatPrecision="765" longitude="31.94" municipality="Tarag" name="Takar village" pageId="7" pageNumber="318">Takar village</location>
|
||
(
|
||
<geoCoordinate id="EE0C8CE9EF0DFFC2FF7613CD5EFC8031" box="[141,234,249,273]" degrees="18.58" direction="north" orientation="latitude" pageId="7" pageNumber="318" precision="555" value="18.58">N18.58°</geoCoordinate>
|
||
<geoCoordinate id="EE0C8CE9EF0DFFC2FF0A13CD5F588031" box="[241,334,249,273]" degrees="31.94" direction="east" orientation="longitude" pageId="7" pageNumber="318" precision="555" value="31.94">E31.94°</geoCoordinate>
|
||
), coll.
|
||
<collectorName id="26CD8FF8EF0DFFC2FE6C13CE5C168031" box="[407,512,249,273]" pageId="7" pageNumber="318">T. Moritz</collectorName>
|
||
,
|
||
<collectorName id="26CD8FF8EF0DFFC2FDF013CE5C808031" box="[523,662,250,273]" pageId="7" pageNumber="318">D. Neumann</collectorName>
|
||
&
|
||
<collectorName id="26CD8FF8EF0DFFC2FD4213CE5EA68011" pageId="7" pageNumber="318">N. Pöllath</collectorName>
|
||
,
|
||
<date id="FF86CCEEEF0DFFC2FF40122D5F298011" box="[187,319,281,306]" pageId="7" pageNumber="318" value="2008-04-09">
|
||
<collectingDate id="EFC23506EF0DFFC2FF40122D5F298011" box="[187,319,281,306]" pageId="7" pageNumber="318" value="2008-04-09">9 Apr. 2008</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B50E073EF0DFFC2FEB1122D5F198071" collectingDate="2008-04-09" collectionCode="ZSM" collectorName="T. Moritz & D. Neumann & N. Pollath" country="Sudan" county="Nile" latitude="18.58" location="Takar village" longLatPrecision="765" longitude="31.94" municipality="Tarag" pageId="7" pageNumber="318" specimenCode="ZSM 39817" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9E4255EF0DFFC2FEB1122D5FDA8011" box="[330,460,281,305]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="7" pageNumber="318" type="Museum">ZSM 39817</specimenCode>
|
||
(5), 36.0-
|
||
<quantity id="4CC047CBEF0DFFC2FDC2122D5C8D8011" box="[569,667,281,305]" metricMagnitude="-2" metricUnit="m" metricValue="3.82" pageId="7" pageNumber="318" unit="mm" value="38.2">38.2 mm</quantity>
|
||
SL, same data as ZSM
|
||
</materialsCitation>
|
||
39816;
|
||
<materialsCitation id="3B50E073EF0DFFC2FE91120D5F6E8052" collectingDate="2008-04-09" collectionCode="ZSM" collectorName="T. Moritz & D. Neumann & N. Pollath" country="Sudan" county="Nile" latitude="18.58" location="Takar village" longLatPrecision="765" longitude="31.94" municipality="Tarag" pageId="7" pageNumber="318" specimenCode="ZSM 39818" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9E4255EF0DFFC2FE91120D5FFA8071" box="[362,492,313,337]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="7" pageNumber="318" type="Museum">ZSM 39818</specimenCode>
|
||
(6 now 2), 18.3-36.0 mm SL, same data as ZSM
|
||
</materialsCitation>
|
||
39816;
|
||
<materialsCitation id="3B50E073EF0DFFC2FE34126E5C7480B2" collectingDate="2008-04-09" collectionCode="AMNH, ZSM" collectorName="T. Moritz & D. Neumann & N. Pollath" country="Sudan" county="Nile" latitude="18.58" location="Takar village" longLatPrecision="765" longitude="31.94" municipality="Tarag" pageId="7" pageNumber="318" specimenCode="AMNH 267145, ZSM 39818" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9E4255EF0DFFC2FE34126E5C6F8051" box="[463,633,345,370]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="7" pageNumber="318">AMNH 267145</specimenCode>
|
||
(2) (ex
|
||
<specimenCode id="DB9E4255EF0DFFC2FD34126E5EDD80B2" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="7" pageNumber="318" type="Museum">ZSM 39818</specimenCode>
|
||
),
|
||
<quantity id="4CC047CBEF0DFFC2FF27124D5F6680B1" box="[220,368,377,401]" metricMagnitude="-2" metricUnit="m" metricValue="2.405" metricValueMax="2.66" metricValueMin="2.15" pageId="7" pageNumber="318" unit="mm" value="24.05" valueMax="26.6" valueMin="21.5">21.5-26.6 mm</quantity>
|
||
SL, same data as ZSM
|
||
</materialsCitation>
|
||
39816;
|
||
<materialsCitation id="3B50E073EF0DFFC2FD4C124E5F0380F2" collectingDate="2008-04-09" collectionCode="BMNH" collectorName="T. Moritz & D. Neumann & N. Pollath" country="Sudan" county="Nile" latitude="18.58" location="Takar village" longLatPrecision="765" longitude="31.94" municipality="Tarag" pageId="7" pageNumber="318" specimenCode="ZSM 39818" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED2972EBEF0DFFC2FD4C124E5D1180B1" box="[695,775,378,401]" pageId="7" pageNumber="318">BMNH</collectionCode>
|
||
2018.9.3.1-2 (2) (ex
|
||
<specimenCode id="DB9E4255EF0DFFC2FE9C12AE5FFB8092" box="[359,493,410,434]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="7" pageNumber="318" type="Museum">ZSM 39818</specimenCode>
|
||
),
|
||
<quantity id="4CC047CBEF0DFFC2FDFA12AD5C8D8091" box="[513,667,409,433]" metricMagnitude="-2" metricUnit="m" metricValue="2.565" metricValueMax="2.98" metricValueMin="2.15" pageId="7" pageNumber="318" unit="mm" value="25.65" valueMax="29.8" valueMin="21.5">21.5-29.8 mm</quantity>
|
||
SL, same data as ZSM
|
||
</materialsCitation>
|
||
39816;
|
||
<materialsCitation id="3B50E073EF0DFFC2FE8E128E5F5E80D2" collectingDate="2008-04-09" collectionCode="ZSM" collectorName="T. Moritz & D. Neumann & N. Pollath" country="Sudan" county="Nile" latitude="18.58" location="Takar village" longLatPrecision="765" longitude="31.94" municipality="Tarag" pageId="7" pageNumber="318" specimenCode="ZSM 39820" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DB9E4255EF0DFFC2FE8E128E5FEA80F2" box="[373,508,442,466]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="7" pageNumber="318" type="Museum">ZSM 39820</specimenCode>
|
||
(2),
|
||
<quantity id="4CC047CBEF0DFFC2FDCD128D5CC080F1" box="[566,726,441,465]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" metricValueMax="2.19" metricValueMin="1.35" pageId="7" pageNumber="318" unit="mm" value="17.7" valueMax="21.9" valueMin="13.5">13.5-21.9 mm</quantity>
|
||
SL, same data as ZSM
|
||
</materialsCitation>
|
||
39814.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C322B9A5EF0DFFC2FF7F112D5F728713" pageId="7" pageNumber="318" type="diagnosis">
|
||
<paragraph id="8B87EA2EEF0DFFC2FF7F112D5EE48311" blockId="7.[131,775,537,1587]" box="[132,242,537,561]" pageId="7" pageNumber="318">
|
||
<emphasis id="B94C363CEF0DFFC2FF7F112D5EE48311" bold="true" box="[132,242,537,561]" pageId="7" pageNumber="318">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B87EA2EEF0DFFC2FF51110E5F728713" blockId="7.[131,775,537,1587]" pageId="7" pageNumber="318">
|
||
<taxonomicName id="4C3891ADEF0DFFC2FF51110E5F578371" authorityName="Moritz & Straube & Neumann" authorityYear="2019" box="[170,321,570,593]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="318" phylum="Chordata" rank="species" species="napata">
|
||
<emphasis id="B94C363CEF0DFFC2FF51110E5F578371" box="[170,321,570,593]" italics="true" pageId="7" pageNumber="318">Garra napata</emphasis>
|
||
</taxonomicName>
|
||
differs from all other African
|
||
<taxonomicName id="4C3891ADEF0DFFC2FD75110E5EA58352" pageId="7" pageNumber="318">
|
||
<emphasis id="B94C363CEF0DFFC2FD75110E5CC48371" box="[654,722,570,593]" italics="true" pageId="7" pageNumber="318">Garra</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, except
|
||
<taxonomicName id="4C3891ADEF0DFFC2FEE9116E5F6A8352" box="[274,380,602,626]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="318" phylum="Chordata" rank="species" species="jamila">
|
||
<emphasis id="B94C363CEF0DFFC2FEE9116E5F6A8352" box="[274,380,602,626]" italics="true" pageId="7" pageNumber="318">G. jamila</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3891ADEF0DFFC2FE4F116E5C758352" authorityName="Menon" authorityYear="1958" box="[436,611,602,626]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="318" phylum="Chordata" rank="species" species="ethelwynnae">
|
||
<emphasis id="B94C363CEF0DFFC2FE4F116E5C758352" box="[436,611,602,626]" italics="true" pageId="7" pageNumber="318">G. ethelwynnae</emphasis>
|
||
</taxonomicName>
|
||
, in a scaleless area on the dorsal body area clearly exceeding beyond dorsal fin origin (
|
||
<figureCitation id="1303F6ABEF0DFFC2FF0D11AE5F5C8392" box="[246,330,666,690]" captionStart="Figure 6" captionStartId="6.[133,198,562,584]" captionTargetBox="[142,764,227,533]" captionTargetId="figure-693@6.[132,778,215,544]" captionTargetPageId="6" captionText="Figure 6. – Scaled (dark) and unscaled (light grey) areas in dorsal view. A: Garra vinciguerrae; B: G. napata; C: G. jamila; D: G. sannarensis." figureDoi="http://doi.org/10.5281/zenodo.12711575" httpUri="https://zenodo.org/record/12711575/files/figure.png" pageId="7" pageNumber="318">Fig. 6B</figureCitation>
|
||
) and thus in an incomplete count of only 1 to 2.5 (vs. 3 or more) scale rows between the origin of the dorsal fin and lateral line series and dorsal fin. It is distinguished from
|
||
<taxonomicName id="4C3891ADEF0DFFC2FEE511CE5FDC8232" authorityName="Menon" authorityYear="1958" box="[286,458,762,786]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="318" phylum="Chordata" rank="species" species="ethelwynnae">
|
||
<emphasis id="B94C363CEF0DFFC2FEE511CE5FDC8232" box="[286,458,762,786]" italics="true" pageId="7" pageNumber="318">G. ethelwynnae</emphasis>
|
||
</taxonomicName>
|
||
from the Damas River basin (
|
||
<collectingCountry id="F32FAABEEF0DFFC2FF77102E5ECB8212" box="[140,221,794,818]" name="Eritrea" pageId="7" pageNumber="318">Eritrea</collectingCountry>
|
||
) by its intermediate [
|
||
<typeStatus id="5483548CEF0DFFC2FE34102E5C148212" box="[463,514,794,818]" pageId="7" pageNumber="318">type</typeStatus>
|
||
B] developed disc and only minute free posterior flap (vs. prominent disc [
|
||
<typeStatus id="5483548CEF0DFFC2FD4F100E5CF38272" box="[692,741,826,850]" pageId="7" pageNumber="318">type</typeStatus>
|
||
C] with well-developed posterior flap) and an asquamate postpelvic area (vs. squamate).
|
||
<taxonomicName id="4C3891ADEF0DFFC2FE55104E5C0C82B2" authorityName="Moritz & Straube & Neumann" authorityYear="2019" box="[430,538,890,914]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="318" phylum="Chordata" rank="species" species="napata">
|
||
<emphasis id="B94C363CEF0DFFC2FE55104E5C0C82B2" box="[430,538,890,914]" italics="true" pageId="7" pageNumber="318">G. napata</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C3891ADEF0DFFC2FF7F10AE5EFE8292" box="[132,232,922,946]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="318" phylum="Chordata" rank="species" species="jamila">
|
||
<emphasis id="B94C363CEF0DFFC2FF7F10AE5EFE8292" box="[132,232,922,946]" italics="true" pageId="7" pageNumber="318">G. jamila</emphasis>
|
||
</taxonomicName>
|
||
of the Atbara (Main Nile basin) by its intermediate [
|
||
<typeStatus id="5483548CEF0DFFC2FF71108F5EAD82F3" box="[138,187,955,979]" pageId="7" pageNumber="318">type</typeStatus>
|
||
B] developed disc with a very small free posterior flap (vs. prominent disc [
|
||
<typeStatus id="5483548CEF0DFFC2FE9F10EF5F8082D3" box="[356,406,987,1011]" pageId="7" pageNumber="318">type</typeStatus>
|
||
C] with well-developed posterior flap), a smooth upper lip (vs. crenate) and, in live specimens, by an uncoloured anal fin (vs. red anal fin); dark humeral spot large and nearly twice as big as red postopercular spot (vs. equally sized small humeral and postopercular spots); fixed specimens usually display a pronounced lateral band and pale back (vs. dark back strongly and contrasting light belly). It is distinguished from
|
||
<taxonomicName id="4C3891ADEF0DFFC2FE2D178F5C9485F2" box="[470,642,1211,1234]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="318" phylum="Chordata" rank="species" species="vinciguerrae">
|
||
<emphasis id="B94C363CEF0DFFC2FE2D178F5C9485F2" box="[470,642,1211,1234]" italics="true" pageId="7" pageNumber="318">G. vinciguerrae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3891ADEF0DFFC2FD4C178F5EC885D2" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="318" phylum="Chordata" rank="species" species="sannarensis">
|
||
<emphasis id="B94C363CEF0DFFC2FD4C178F5EC885D2" italics="true" pageId="7" pageNumber="318">G. sannarensis</emphasis>
|
||
</taxonomicName>
|
||
by its narrow disc (width much smaller than mouth vs. as large as mouth width), virtually absent lateroposteri- or flap of the disc (vs. clearly developed) (
|
||
<figureCitation id="1303F6ABEF0DFFC2FDA4162E5CA08412" box="[607,694,1306,1330]" captionStart="Figure 5" captionStartId="5.[132,198,1741,1763]" captionTargetBox="[550,1453,1303,1899]" captionTargetId="figure-637@5.[548,1455,1301,1901]" captionTargetPageId="5" captionText="Figure 5. – Underside of head show- ing the different types of sucking-discs. A: Garra vinciguerrae; B: G. napa- ta; C: G. sannarensis; D: G. jamila, holotype; E: G. jamila, large speci- men; F: G. sp. “White Nile”. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12711571" httpUri="https://zenodo.org/record/12711571/files/figure.png" pageId="7" pageNumber="318">Fig. 5B</figureCitation>
|
||
), and a relatively broad head, i.e. cheeks visible when viewed from dorsal (vs. cheeks not visible); from
|
||
<taxonomicName id="4C3891ADEF0DFFC2FDED166F5CD28452" box="[534,708,1371,1394]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="318" phylum="Chordata" rank="species" species="vinciguerrae">
|
||
<emphasis id="B94C363CEF0DFFC2FDED166F5CD28452" box="[534,708,1371,1394]" italics="true" pageId="7" pageNumber="318">G. vinciguerrae</emphasis>
|
||
</taxonomicName>
|
||
by its prominent dark colour markings in the centre of the dorsal-fin membrane near its base (vs. no such pattern) (
|
||
<figureCitation id="1303F6ABEF0DFFC2FD5916AE5CEF8492" box="[674,761,1434,1458]" captionStart="Figure 6" captionStartId="6.[133,198,562,584]" captionTargetBox="[142,764,227,533]" captionTargetId="figure-693@6.[132,778,215,544]" captionTargetPageId="6" captionText="Figure 6. – Scaled (dark) and unscaled (light grey) areas in dorsal view. A: Garra vinciguerrae; B: G. napata; C: G. jamila; D: G. sannarensis." figureDoi="http://doi.org/10.5281/zenodo.12711575" httpUri="https://zenodo.org/record/12711575/files/figure.png" pageId="7" pageNumber="318">Fig. 6B</figureCitation>
|
||
); from
|
||
<emphasis id="B94C363CEF0DFFC2FF44168F5ECF84F2" box="[191,217,1467,1490]" italics="true" pageId="7" pageNumber="318">G.</emphasis>
|
||
sp. “White Nile” by its visible cheeks when viewed from dorsal (vs. not visible from above) and by its short barbels barely reaching margin of lower-lip pad (vs. almost reaching each other).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF47BAA6EF0DFFC2FCD611995DF183D3" ID-DOI="http://doi.org/10.5281/zenodo.12711579" ID-Zenodo-Dep="12711579" httpUri="https://zenodo.org/record/12711579/files/figure.png" pageId="7" pageNumber="318" startId="7.[813,876,685,707]" targetBox="[813,1456,215,668]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8B87EA2EEF0DFFC2FCD611995DF183D3" blockId="7.[812,1455,685,755]" pageId="7" pageNumber="318">
|
||
Figure 8. –
|
||
<taxonomicName id="4C3891ADEF0DFFC2FC67119A5A3783E3" authorityName="Moritz & Straube & Neumann" authorityYear="2019" box="[924,1057,686,707]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="318" phylum="Chordata" rank="species" species="napata" status="sp. nov.">
|
||
<emphasis id="B94C363CEF0DFFC2FC67119A5A3783E3" box="[924,1057,686,707]" italics="true" pageId="7" pageNumber="318">Garra napata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27F8B47EF0DFFC2FBDC119A5A6383E3" box="[1063,1141,686,707]" pageId="7" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ZSM 47244 (ex ZSM 39815#3), 39.9 mm SL, Sudan: Nile: Tarag Island, 10 Apr. 2008. Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF47BAA6EF0DFFC2FCD616D35AA3870D" ID-DOI="http://doi.org/10.5281/zenodo.12711583" ID-Zenodo-Dep="12711583" httpUri="https://zenodo.org/record/12711583/files/figure.png" pageId="7" pageNumber="318" startId="7.[813,877,1511,1533]" targetBox="[813,1456,787,1494]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8B87EA2EEF0DFFC2FCD616D35AA3870D" blockId="7.[812,1455,1511,1581]" pageId="7" pageNumber="318">
|
||
Figure 9. – Live specimens of
|
||
<taxonomicName id="4C3891ADEF0DFFC2FBA316DC5AC984DD" authorityName="Moritz & Straube & Neumann" authorityYear="2019" box="[1112,1247,1512,1533]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="318" phylum="Chordata" rank="species" species="napata" status="sp. nov.">
|
||
<emphasis id="B94C363CEF0DFFC2FBA316DC5AC984DD" box="[1112,1247,1512,1533]" italics="true" pageId="7" pageNumber="318">Garra napata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27F8B47EF0DFFC2FB1D16DC5B2484DD" box="[1254,1330,1512,1533]" pageId="7" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
|
||
All from the area of Tarag Island, 4
|
||
<superScript id="7C4D4766EF0DFFC2FBFE16C95A04872B" attach="left" box="[1029,1042,1533,1547]" fontSize="6" pageId="7" pageNumber="318">th</superScript>
|
||
cataract, Nile, Sudan.
|
||
<emphasis id="B94C363CEF0DFFC2FB1516CB5B0E8734" bold="true" box="[1262,1304,1535,1556]" pageId="7" pageNumber="318">A-B</emphasis>
|
||
: ~38 mm SL, 9 Apr. 2008;
|
||
<emphasis id="B94C363CEF0DFFC2FC6C15235DBE870D" bold="true" box="[919,936,1559,1581]" pageId="7" pageNumber="318">C</emphasis>
|
||
: ~50 mm SL, 10 Apr. 2008.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C322B9A5EF0DFFCDFF7F156E5D6280F1" lastPageId="8" lastPageNumber="319" pageId="7" pageNumber="318" type="description">
|
||
<paragraph id="8B87EA2EEF0DFFC2FF7F156E5F1E8752" blockId="7.[132,775,1626,1907]" box="[132,264,1626,1650]" pageId="7" pageNumber="318">
|
||
<emphasis id="B94C363CEF0DFFC2FF7F156E5F1E8752" bold="true" box="[132,264,1626,1650]" pageId="7" pageNumber="318">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B87EA2EEF0DFFCDFF51154F5C4D85A2" blockId="7.[132,775,1626,1907]" lastBlockId="8.[131,776,770,1907]" lastPageId="8" lastPageNumber="319" pageId="7" pageNumber="318">
|
||
Based on
|
||
<typeStatus id="5483548CEF0DFFC2FEE1154F5F6A87B3" box="[282,380,1659,1683]" pageId="7" pageNumber="318" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="1303F6ABEF0DFFC2FE76154F5FC787B3" box="[397,465,1659,1683]" captionStart="Figure 8" captionStartId="7.[813,876,685,707]" captionTargetBox="[813,1456,215,668]" captionTargetId="figure-601@7.[813,1456,215,668]" captionTargetPageId="7" captionText="Figure 8. – Garra napata sp. nov., holotype, ZSM 47244 (ex ZSM 39815#3), 39.9 mm SL, Sudan: Nile: Tarag Island, 10 Apr. 2008. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.12711579" httpUri="https://zenodo.org/record/12711579/files/figure.png" pageId="7" pageNumber="318">Fig. 8</figureCitation>
|
||
) and
|
||
<specimenCount id="9D3E21A7EF0DFFC2FDE9154F5CB087B3" box="[530,678,1659,1683]" count="24" pageId="7" pageNumber="318" type="generic" typeStatus="paratype">24 paratypes</specimenCount>
|
||
. A small
|
||
<taxonomicName id="4C3891ADEF0DFFC2FF7E15AF5F098793" box="[133,287,1691,1715]" pageId="7" pageNumber="318">
|
||
<emphasis id="B94C363CEF0DFFC2FF7E15AF5EDF8792" box="[133,201,1691,1714]" italics="true" pageId="7" pageNumber="318">Garra</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
with a maximum recorded size
|
||
<quantity id="4CC047CBEF0DFFC2FD7B15AF5CF78792" box="[640,737,1691,1715]" metricMagnitude="-2" metricUnit="m" metricValue="7.029999999999999" pageId="7" pageNumber="318" unit="mm" value="70.3">70.3 mm</quantity>
|
||
SL (84.0 mm TL). Counts and proportional measurements are given in tables II and III. Head and anterior body region dorsoventrally flattened; posterior half of body laterally compressed. Body dorsoventrally compressed, thinner than most other
|
||
<taxonomicName id="4C3891ADEF0DFFC2FF3B140F5F438673" box="[192,341,1851,1875]" pageId="7" pageNumber="318">
|
||
<emphasis id="B94C363CEF0DFFC2FF3B140F5F158672" box="[192,259,1851,1874]" italics="true" pageId="7" pageNumber="318">Garra</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in the Main Nile. Head relatively flat and broad, cheeks visible when viewed from dorsal. Disc intermediate (type B), with very small lateroposterior flap without papillae; central callus of disc smaller than mouth and usually longer than wide (
|
||
<figureCitation id="1303F6ABEF0DFFC2FBC415AF5A878793" box="[1087,1169,1691,1715]" captionStart="Figure 5" captionStartId="5.[132,198,1741,1763]" captionTargetBox="[550,1453,1303,1899]" captionTargetId="figure-637@5.[548,1455,1301,1901]" captionTargetPageId="5" captionText="Figure 5. – Underside of head show- ing the different types of sucking-discs. A: Garra vinciguerrae; B: G. napa- ta; C: G. sannarensis; D: G. jamila, holotype; E: G. jamila, large speci- men; F: G. sp. “White Nile”. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12711571" httpUri="https://zenodo.org/record/12711571/files/figure.png" pageId="7" pageNumber="318">Fig. 5B</figureCitation>
|
||
). Rostral cap poorly developed, its margin being only weakly fringed if fringed at all. Two well-developed pairs of maxillary barbels. Dorsal fin originates midway on body, its distal margin slightly convex or straight with one large and one smaller simple ray, often supported by an additional minute small ray deeply embedded in the anterior dorsal fin tissue and visible only in X-ray images; unbranched rays followed by 7 branched. Pectoral fins inserting ventrally low on body; pelvic fins reaching beyond vent, but ending in front of base of anal fin. Anal fin with one large and one small simple ray, often supported by additional minute rays deeply embedded into the anterior fin tissue and visible only in X-ray images; 5 branched rays, distal fin membrane straight. Caudal fin emarginated, upper and lower lobe rounded. Body covered with large scales, except for chest, belly and postpelvic area; predorsal area naked, asquamate area expands laterally onto body flanks (
|
||
<figureCitation id="1303F6ABEF02FFCDFD56171D5CE88561" box="[685,766,1065,1089]" captionStart="Figure 6" captionStartId="6.[133,198,562,584]" captionTargetBox="[142,764,227,533]" captionTargetId="figure-693@6.[132,778,215,544]" captionTargetPageId="6" captionText="Figure 6. – Scaled (dark) and unscaled (light grey) areas in dorsal view. A: Garra vinciguerrae; B: G. napata; C: G. jamila; D: G. sannarensis." figureDoi="http://doi.org/10.5281/zenodo.12711575" httpUri="https://zenodo.org/record/12711575/files/figure.png" pageId="8" pageNumber="319">Fig. 6B</figureCitation>
|
||
) consequently reducing the number of scale rows between the dorsal fin origin and lateral line series to 2.5.
|
||
</paragraph>
|
||
<caption id="DF47BAA6EF02FFCDFF7F11B95FD5839A" ID-DOI="http://doi.org/10.5281/zenodo.12711585" ID-Zenodo-Dep="12711585" httpUri="https://zenodo.org/record/12711585/files/figure.png" pageId="8" pageNumber="319" startId="8.[132,195,653,675]" targetBox="[134,773,217,633]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8B87EA2EEF02FFCDFF7F11B95FD5839A" blockId="8.[132,776,653,699]" pageId="8" pageNumber="319">
|
||
Figure 10. – Small rapid at right bank of Tarag main island (13 Apr. 2008), type locality of
|
||
<taxonomicName id="4C3891ADEF02FFCDFEA611915FAA839A" authorityName="Moritz & Straube & Neumann" authorityYear="2019" box="[349,444,677,698]" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="319" phylum="Chordata" rank="species" species="napata">
|
||
<emphasis id="B94C363CEF02FFCDFEA611915FAA839A" box="[349,444,677,698]" italics="true" pageId="8" pageNumber="319">G. napata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B87EA2EEF02FFCDFF5117BF5C638612" blockId="8.[131,776,770,1907]" pageId="8" pageNumber="319">
|
||
<emphasis id="B94C363CEF02FFCDFF5117BF5F228583" box="[170,308,1163,1187]" italics="true" pageId="8" pageNumber="319">Colouration.</emphasis>
|
||
– Head and flanks in life deeply olive-green, body sides gradually getting darker towards mid-dorsal line, lower third of flanks and belly whitish (
|
||
<figureCitation id="1303F6ABEF02FFCDFDD617F85C7985C5" box="[557,623,1228,1253]" captionStart="Figure 9" captionStartId="7.[813,877,1511,1533]" captionTargetBox="[813,1456,787,1494]" captionTargetId="figure-630@7.[813,1456,787,1494]" captionTargetPageId="7" captionText="Figure 9. – Live specimens of Garra napata sp. nov. All from the area of Tarag Island, 4th cataract, Nile, Sudan. A-B: ~38 mm SL, 9 Apr. 2008; C: ~50 mm SL, 10 Apr. 2008." figureDoi="http://doi.org/10.5281/zenodo.12711583" httpUri="https://zenodo.org/record/12711583/files/figure.png" pageId="8" pageNumber="319">Fig. 9</figureCitation>
|
||
). Flank scales with dark anterior centres and light distal part resulting in reticulate pattern on flanks, towards tail additional, usually 7 irregular, vertically merged dark green to blackish blotches. Caudal peduncle with prominent and broad, slightly rectangular marking, contrasting the flank colouration. Iris orange. Prominent red spot at posterior dorsal corner of opercle merges with dark-green humeral spot, nearly twice as large as blotch on the first lateral line series scale. Pectoral, pelvic and anal fins hyaline to slightly orange with few scattered melanophores along fin rays. Caudal fin membrane hyaline, rays bordered by dark grey melanophores in proximal twothirds, often fused to a dark spot at the base of the inner-most caudal rays; distal third of caudal fin membrane slightly orange, border of ventral and dorsal lobes orange. Dorsal-fin base with four prominent dark spots at branched rays 3 to 6; middle of fin membrane with horizontal dark band with very dark to black zone in anterior half (
|
||
<figureCitation id="1303F6ABEF02FFCDFDFC15CD5C488631" box="[519,606,1785,1809]" captionStart="Figure 7" captionStartId="6.[132,195,796,818]" captionTargetBox="[133,774,667,773]" captionTargetId="figure-729@6.[132,775,664,779]" captionTargetPageId="6" captionText="Figure 7. – Dorsal fin melanophore pattern. A: Garra vinciguerrae; B: G. napata; C: G. jamila; D: G. sannarensis." figureDoi="http://doi.org/10.5281/zenodo.12711577" httpUri="https://zenodo.org/record/12711577/files/figure.png" pageId="8" pageNumber="319">Fig. 7B</figureCitation>
|
||
); distal third of dorsal fin membrane orange or slightly orange.
|
||
</paragraph>
|
||
<paragraph id="8B87EA2EEF02FFCDFF51140E5D6280F1" blockId="8.[131,776,770,1907]" lastBlockId="8.[812,1456,217,465]" pageId="8" pageNumber="319">Preserved specimens dark brown on flanks and dorsum; lower third of flanks and ventral side pale whitish. Posterior half of flanks darker; faint dark band along lateral line occa- sionally visible in specimens, the irregular colour pattern on tail faintly recognisable. First scale of lateral line series with dark blotch. Dark bar at end of caudal peduncle and innermost caudal rays with dark spots at insertion. Pectoral pelvic and anal fin whitish-hyaline with single melanophore along rays, dark dorsal fin markings also visible in preserved specimens.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C322B9A5EF02FFCDFCD612CD5DB88352" pageId="8" pageNumber="319" type="distribution">
|
||
<paragraph id="8B87EA2EEF02FFCDFCD612CD5DAE8331" blockId="8.[813,1455,505,626]" box="[813,952,505,529]" pageId="8" pageNumber="319">
|
||
<emphasis id="B94C363CEF02FFCDFCD612CD5DAE8331" bold="true" box="[813,952,505,529]" pageId="8" pageNumber="319">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B87EA2EEF02FFCDFCA9112E5DB88352" blockId="8.[813,1455,505,626]" pageId="8" pageNumber="319">
|
||
Known from the area around Elephantine Island at the 1
|
||
<superScript id="7C4D4766EF02FFCDFCC111035D518367" attach="left" box="[826,839,567,583]" fontSize="7" pageId="8" pageNumber="319">st</superScript>
|
||
cataract and Tarag Island below the (former) 4
|
||
<superScript id="7C4D4766EF02FFCDFABE11025B428367" attach="left" box="[1349,1364,566,583]" fontSize="7" pageId="8" pageNumber="319">th</superScript>
|
||
cataract (
|
||
<figureCitation id="1303F6ABEF02FFCDFCCF116E5D608352" box="[820,886,602,626]" captionStart="Figure 1" captionStartId="1.[813,879,1813,1835]" captionTargetBox="[830,1438,215,1796]" captionTargetId="figure-464@1.[830,1438,215,1796]" captionTargetPageId="1" captionText="Figure 1. – Records of Garra species from the Main Nile basin (dotted line). Star – Garra vinciguerrae; diamond – G. napata; square – G. jamila; triangle – G. sannarensis; cross – G. sp. “White Nile”; numbers indicate the cataracts of the Nile river." figureDoi="http://doi.org/10.5281/zenodo.12711561" httpUri="https://zenodo.org/record/12711561/files/figure.png" pageId="8" pageNumber="319">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="1303F6ABEF02FFCDFC78116E5DB78352" box="[899,929,602,626]" captionStart="Figure 10" captionStartId="8.[132,195,653,675]" captionTargetBox="[134,773,217,633]" captionTargetId="figure-710@8.[132,775,215,635]" captionTargetPageId="8" captionText="Figure 10. – Small rapid at right bank of Tarag main island (13 Apr. 2008), type locality of G. napata." figureDoi="http://doi.org/10.5281/zenodo.12711585" httpUri="https://zenodo.org/record/12711585/files/figure.png" pageId="8" pageNumber="319">10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |