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<document id="1E4C1EE0EDECA2140CC6FA47F1A4DD13" ID-DOI="10.11646/zootaxa.5109.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6336940" ID-ZooBank="7B6C0651-0935-4C77-B157-3ABF53E3AD81" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646727632931" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Ślipiński, Adam" docDate="2022" docId="038B11419C427023D4D8FD5AEBABF840" docLanguage="en" docName="zootaxa.5109.1.1.pdf.imf" docOrigin="Zootaxa 5109 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5109.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Botrodus dufaui Grouvelle" docType="treatment" docVersion="5" lastPageNumber="16" masterDocId="FFB269399C4D702CD44FFFD9E928FFB2" masterDocTitle="Revision of the family Murmidiidae (Coleoptera: Coccinelloidea)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="16" updateTime="1699303175474" updateUser="plazi">
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<mods:title id="1C62F5BADD7B62EFB79298453C998F80">Revision of the family Murmidiidae (Coleoptera: Coccinelloidea)</mods:title>
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<mods:namePart id="97C237354B2BDEE928EE5D11A98C7240">Jałoszyński, Paweł</mods:namePart>
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<mods:namePart id="F130549F4FC40701C387C5348DFEE1B7">Ślipiński, Adam</mods:namePart>
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<subSubSection id="C338F3DC9C427023D4D8FD5AE8C8FD2C" box="[151,480,643,670]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8B9DA0579C427023D4D8FD5AE8C8FD2C" blockId="15.[151,480,643,706]" box="[151,480,643,670]" pageId="15" pageNumber="16">
<heading id="D0D5173B9C427023D4D8FD5AE8C8FD2C" bold="true" box="[151,480,643,670]" fontSize="11" level="2" pageId="15" pageNumber="16" reason="4">
<taxonomicName id="4C22DBD49C427023D4D8FD5AE8C8FD2C" authority="Grouvelle" authorityName="Grouvelle" box="[151,480,643,670]" class="Insecta" family="Murmidiidae" genus="Botrodus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dufaui">
<emphasis id="B9567C459C427023D4D8FD5AE8C8FD2C" bold="true" box="[151,480,643,670]" pageId="15" pageNumber="16">
<emphasis id="B9567C459C427023D4D8FD5AE875FD2C" bold="true" box="[151,349,643,670]" italics="true" pageId="15" pageNumber="16">Botrodus dufaui</emphasis>
Grouvelle
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338F3DC9C427023D4D8FD71E80FFD70" box="[151,295,680,706]" pageId="15" pageNumber="16" type="description">
<paragraph id="8B9DA0579C427023D4D8FD71E80FFD70" blockId="15.[151,480,643,706]" box="[151,295,680,706]" pageId="15" pageNumber="16">
(
<figureCitation id="1319BCD29C427023D4D0FD71E837FD70" box="[159,287,680,706]" captionStart="FIGURES 2834" captionStartId="16.[151,264,1878,1902]" captionTargetBox="[151,1436,181,1854]" captionTargetId="figure-19@16.[151,1436,181,1854]" captionTargetPageId="16" captionText="FIGURES 2834. Morphological structures of Botrodus dufaui Grouvelle. Body in dorsal (2829), ventral (30) and lateral (31) views; pterothorax in ventral view (32); and aedeagus (3334)." figureDoi="http://doi.org/10.5281/zenodo.6336951" httpUri="https://zenodo.org/record/6336951/files/figure.png" pageId="15" pageNumber="16">Figs 2834</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C338F3DC9C427023D4D8FD35EA39FCB7" box="[151,785,748,773]" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="8B9DA0579C427023D4D8FD35EA39FCB7" blockId="15.[151,785,748,773]" box="[151,785,748,773]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB3287799C427023D4D8FD35EA39FCB7" box="[151,785,748,773]" pageId="15" pageNumber="16">
<taxonomicName id="4C22DBD49C427023D4D8FD35E88EFCB6" authority="Grouvelle" authorityName="Grouvelle" box="[151,422,748,772]" class="Insecta" family="Murmidiidae" genus="Botrodus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dufaui">
<emphasis id="B9567C459C427023D4D8FD35E812FCB6" box="[151,314,748,772]" italics="true" pageId="15" pageNumber="16">Botrodus dufaui</emphasis>
Grouvelle
</taxonomicName>
in
<treatmentCitation id="0A8386469C427023D589FD35EA25FCB7" author="Grouvelle, A. &amp; Raffray, A." box="[454,781,748,773]" page="297" pageId="15" pageNumber="16" year="1912">
<bibRefCitation id="EFB3DDA69C427023D589FD35EA25FCB7" author="Grouvelle, A. &amp; Raffray, A." box="[454,781,748,773]" pageId="15" pageNumber="16" pagination="290 - 312" refId="ref61305" refString="Grouvelle, A. &amp; Raffray, A. (1912) Supplement a la liste des Coleopteres de la Guadeloupe. 3 e supplement. Annales de la So- ciete Entomologique de France, 81, 290 - 312." type="journal article" year="1912">Grouvelle &amp; Raffray, 1912: 297</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C338F3DC9C427023D4D8FCEAED51FC27" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8B9DA0579C427023D4D8FCEAED51FC27" blockId="15.[151,1437,818,2034]" pageId="15" pageNumber="16">
<emphasis id="B9567C459C427023D4D8FCEAE895FCFE" bold="true" box="[151,445,818,845]" pageId="15" pageNumber="16">Type material examined.</emphasis>
<materialsCitation id="3B4AAA0A9C427023D58CFCEAEBE2FCC3" collectionCode="MNHN" collectorName="A. Slipinski" country="Guadeloupe" location="Guadeloupe" pageId="15" pageNumber="16" specimenCount="1" typeStatus="lectotype">
<emphasis id="B9567C459C427023D58CFCEAEBCBFCFE" bold="true" box="[451,739,818,845]" pageId="15" pageNumber="16">
<typeStatus id="54991EF59C427023D58CFCEAEB11FCFF" box="[451,569,819,845]" pageId="15" pageNumber="16" type="lectotype">Lectotype</typeStatus>
(
<collectingCountry id="F335E0C79C427023D608FCEBEBF3FCFE" box="[583,731,818,844]" name="Guadeloupe" pageId="15" pageNumber="16">Guadeloupe</collectingCountry>
)
</emphasis>
[here designated by
<collectorName id="26D7C5819C427023D789FCEAED78FCFD" box="[966,1104,819,847]" pageId="15" pageNumber="16">A. Ślipiński</collectorName>
], sex unknown, “Guadeloupe, Dufau”, “Type”, “
<taxonomicName id="4C22DBD49C427023D52AFC8EEB64FCC3" authority="Ty Grouv." authorityName="Ty Grouv." box="[357,588,855,881]" class="Insecta" family="Murmidiidae" genus="Botrodus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dufaui">B. dufaui Ty Grouv.</taxonomicName>
” (
<collectionCode id="ED3338929C427023D629FC8EEBEAFCC3" box="[614,706,855,881]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="16">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4AAA0A9C427023D699FC8EED5DFC27" collectionCode="MNHN" collectorName="A. Slipinski" country="Guadeloupe" location="Guadeloupe" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" typeStatus="paralectotype">
<emphasis id="B9567C459C427023D699FC8EEA89FCC1" bold="true" box="[726,929,855,883]" pageId="15" pageNumber="16">
<typeStatus id="54991EF59C427023D699FC8EEA57FCC3" box="[726,895,855,881]" pageId="15" pageNumber="16" type="paralectotype">Paralectotype</typeStatus>
:
<specimenCount id="9D246BDE9C427023D7C3FC81EA89FCC1" box="[908,929,856,883]" pageId="15" pageNumber="16" type="male"></specimenCount>
</emphasis>
, same data as for lectotype [specimen disarticulated, mounted in Canada balsam and placed on the same pin as
<typeStatus id="54991EF59C427023D7DDFCA3EAD4FC27" box="[914,1020,890,917]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
] (
<collectionCode id="ED3338929C427023D05EFCA2ED45FC27" box="[1041,1133,891,917]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="16">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C338F3DC9C427023D488FC47EC68FB97" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8B9DA0579C427023D488FC47EC68FB97" blockId="15.[151,1437,818,2034]" pageId="15" pageNumber="16">
<emphasis id="B9567C459C427023D488FC47E86AFC0A" bold="true" box="[199,322,926,952]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
Body moderately dark brown; elytra in lateral view evenly rounded; clypeus convex laterally, with anteromedian subtriangular impression; lateral elytral margin visible in dorsal view near humerus and in posterior third; mesoventrite with lateral longitudinal carinae along mesal margins of mesocoxae; mesoventral plate with anterior margin evenly rounded; each paramere ~3.5 times as long as broad at base, with only apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C338F3DC9C427023D488FBF7EB1AF87F" pageId="15" pageNumber="16" type="description">
<paragraph id="8B9DA0579C427023D488FBF7EB51FB6B" blockId="15.[151,1437,818,2034]" pageId="15" pageNumber="16">
<emphasis id="B9567C459C427023D488FBF7E871FBFA" bold="true" box="[199,345,1070,1096]" pageId="15" pageNumber="16">Description.</emphasis>
BL
<quantity id="4CDA0DB29C427023D5C2FBF6E8DDFBFB" box="[397,501,1071,1097]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="16" unit="mm" value="1.2">1.20 mm</quantity>
; BL/EW 1.92. Body elongate (
<figureCitation id="1319BCD29C427023D718FBF6EAF0FBFB" box="[855,984,1071,1097]" captionStart="FIGURES 2834" captionStartId="16.[151,264,1878,1902]" captionTargetBox="[151,1436,181,1854]" captionTargetId="figure-19@16.[151,1436,181,1854]" captionTargetPageId="16" captionText="FIGURES 2834. Morphological structures of Botrodus dufaui Grouvelle. Body in dorsal (2829), ventral (30) and lateral (31) views; pterothorax in ventral view (32); and aedeagus (3334)." figureDoi="http://doi.org/10.5281/zenodo.6336951" httpUri="https://zenodo.org/record/6336951/files/figure.png" pageId="15" pageNumber="16">Figs 2829</figureCitation>
) and in lateral view evenly, moderately convex (
<figureCitation id="1319BCD29C427023D4B6FB8AE866FBDF" box="[249,334,1107,1133]" captionStart="FIGURES 2834" captionStartId="16.[151,264,1878,1902]" captionTargetBox="[151,1436,181,1854]" captionTargetId="figure-19@16.[151,1436,181,1854]" captionTargetPageId="16" captionText="FIGURES 2834. Morphological structures of Botrodus dufaui Grouvelle. Body in dorsal (2829), ventral (30) and lateral (31) views; pterothorax in ventral view (32); and aedeagus (3334)." figureDoi="http://doi.org/10.5281/zenodo.6336951" httpUri="https://zenodo.org/record/6336951/files/figure.png" pageId="15" pageNumber="16">Fig. 31</figureCitation>
); pigmentation moderately dark brown, head and pronotum slightly lighter than elytra, legs light brown; dorsum covered with moderately sparse, extremely short yellowish setae barely visible under magnification 40 ×; cuticle glossy with very fine and superficial microreticulation distinct on head, on pronotum and elytra traces of microreticulation are poorly discernible.
</paragraph>
<paragraph id="8B9DA0579C427023D488FB3AECA1FAF7" blockId="15.[151,1437,818,2034]" pageId="15" pageNumber="16">
Head
<quantity id="4CDA0DB29C427023D542FB3AE85CFB4F" box="[269,372,1251,1277]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="16" unit="mm" value="0.3">0.30 mm</quantity>
wide; clypeus with convex sides and anteromedian subtriangular impression; frons weakly, evenly convex, both covered with dense and sharply marked but shallow and fine punctures separated by spaces subequal to their diameters (best discernible on frons behind eyes); eyes large, strongly convex, coarsely faceted.
</paragraph>
<paragraph id="8B9DA0579C427023D488FA96ED60FA4B" blockId="15.[151,1437,818,2034]" pageId="15" pageNumber="16">
Pronotum (
<figureCitation id="1319BCD29C427023D508FA96E8E1FADB" box="[327,457,1359,1385]" captionStart="FIGURES 2834" captionStartId="16.[151,264,1878,1902]" captionTargetBox="[151,1436,181,1854]" captionTargetId="figure-19@16.[151,1436,181,1854]" captionTargetPageId="16" captionText="FIGURES 2834. Morphological structures of Botrodus dufaui Grouvelle. Body in dorsal (2829), ventral (30) and lateral (31) views; pterothorax in ventral view (32); and aedeagus (3334)." figureDoi="http://doi.org/10.5281/zenodo.6336951" httpUri="https://zenodo.org/record/6336951/files/figure.png" pageId="15" pageNumber="16">Figs 2829</figureCitation>
) strongly transverse, widest at base; PL
<quantity id="4CDA0DB29C427023D7D4FA97ED2BFADB" box="[923,1027,1358,1385]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="16" unit="mm" value="0.25">0.25 mm</quantity>
, PW
<quantity id="4CDA0DB29C427023D00BFA97ED84FADB" box="[1092,1196,1358,1385]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="15" pageNumber="16" unit="mm" value="0.55">0.55 mm</quantity>
, PL/PW 0.45; lateral margins distinctly rounded in anterior half, posteriorly nearly straight, weakly convergent anterad, with narrow lateral carinae; disc along lateral carinae not flattened. Punctures on disc fine but distinct, sharply marked, slightly sparser than those on frons, punctures on anteromedian region separated by spaces slightly wider than their diameters, gradually reducing in diameter and depth towards sides and base of pronotum.
</paragraph>
<paragraph id="8B9DA0579C427023D488F9DAE893F9F3" blockId="15.[151,1437,818,2034]" pageId="15" pageNumber="16">
Prosternum (
<figureCitation id="1319BCD29C427023D514F9DAE898F9AF" box="[347,432,1539,1565]" captionStart="FIGURES 2834" captionStartId="16.[151,264,1878,1902]" captionTargetBox="[151,1436,181,1854]" captionTargetId="figure-19@16.[151,1436,181,1854]" captionTargetPageId="16" captionText="FIGURES 2834. Morphological structures of Botrodus dufaui Grouvelle. Body in dorsal (2829), ventral (30) and lateral (31) views; pterothorax in ventral view (32); and aedeagus (3334)." figureDoi="http://doi.org/10.5281/zenodo.6336951" httpUri="https://zenodo.org/record/6336951/files/figure.png" pageId="15" pageNumber="16">Fig. 30</figureCitation>
) smooth, not microreticulate and with very fine, sparse punctures; notosternal carinae strongly curved outwards.
</paragraph>
<paragraph id="8B9DA0579C427023D488F992E824F963" blockId="15.[151,1437,818,2034]" pageId="15" pageNumber="16">
Elytra (
<figureCitation id="1319BCD29C427023D554F992E8EFF9D7" box="[283,455,1611,1637]" captionStart="FIGURES 2834" captionStartId="16.[151,264,1878,1902]" captionTargetBox="[151,1436,181,1854]" captionTargetId="figure-19@16.[151,1436,181,1854]" captionTargetPageId="16" captionText="FIGURES 2834. Morphological structures of Botrodus dufaui Grouvelle. Body in dorsal (2829), ventral (30) and lateral (31) views; pterothorax in ventral view (32); and aedeagus (3334)." figureDoi="http://doi.org/10.5281/zenodo.6336951" httpUri="https://zenodo.org/record/6336951/files/figure.png" pageId="15" pageNumber="16">Figs 2829, 31</figureCitation>
) together oval, indistinctly rounded at sides, EL
<quantity id="4CDA0DB29C427023D7A1F992ED7DF9D7" box="[1006,1109,1611,1637]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="16" unit="mm" value="0.9">0.90 mm</quantity>
, EW
<quantity id="4CDA0DB29C427023D0DBF992EDD3F9D7" box="[1172,1275,1611,1637]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="15" pageNumber="16" unit="mm" value="0.63">0.63 mm</quantity>
, EL/EW 1.44. Lateral elytral margin visible in dorsal view near humerus and distinctly also in posterior third; elytral base ante- riorly convex; humerus with distinct, bulging callus; disc covered with punctures slightly sparser than those on pronotum.
</paragraph>
<paragraph id="8B9DA0579C427023D488F902EB2FF947" blockId="15.[151,1437,818,2034]" box="[199,519,1755,1781]" pageId="15" pageNumber="16">Hind wings fully developed.</paragraph>
<paragraph id="8B9DA0579C427023D488F926E880F88F" blockId="15.[151,1437,818,2034]" pageId="15" pageNumber="16">
Mesoventral plate (
<figureCitation id="1319BCD29C427023D5EFF926E8DAF8AB" box="[416,498,1791,1817]" captionStart="FIGURES 2834" captionStartId="16.[151,264,1878,1902]" captionTargetBox="[151,1436,181,1854]" captionTargetId="figure-19@16.[151,1436,181,1854]" captionTargetPageId="16" captionText="FIGURES 2834. Morphological structures of Botrodus dufaui Grouvelle. Body in dorsal (2829), ventral (30) and lateral (31) views; pterothorax in ventral view (32); and aedeagus (3334)." figureDoi="http://doi.org/10.5281/zenodo.6336951" httpUri="https://zenodo.org/record/6336951/files/figure.png" pageId="15" pageNumber="16">Fig. 32</figureCitation>
) with lateral longitudinal carinae and complete submarginal carina, anterior margin of plate evenly rounded.
</paragraph>
<paragraph id="8B9DA0579C427023D488F89EEC36F8D3" blockId="15.[151,1437,818,2034]" box="[199,1310,1863,1889]" pageId="15" pageNumber="16">
Metaventrite (
<figureCitation id="1319BCD29C427023D529F89EE89FF8D3" box="[358,439,1863,1889]" captionStart="FIGURES 2834" captionStartId="16.[151,264,1878,1902]" captionTargetBox="[151,1436,181,1854]" captionTargetId="figure-19@16.[151,1436,181,1854]" captionTargetPageId="16" captionText="FIGURES 2834. Morphological structures of Botrodus dufaui Grouvelle. Body in dorsal (2829), ventral (30) and lateral (31) views; pterothorax in ventral view (32); and aedeagus (3334)." figureDoi="http://doi.org/10.5281/zenodo.6336951" httpUri="https://zenodo.org/record/6336951/files/figure.png" pageId="15" pageNumber="16">Fig. 32</figureCitation>
) extremely finely and uniformly punctate; discrimen externally indiscernible.
</paragraph>
<paragraph id="8B9DA0579C427023D488F8B2EB1AF87F" blockId="15.[151,1437,818,2034]" pageId="15" pageNumber="16">
Aedeagus (
<figureCitation id="1319BCD29C427023D50BF8B2E8ECF837" box="[324,452,1899,1925]" captionStart="FIGURES 2834" captionStartId="16.[151,264,1878,1902]" captionTargetBox="[151,1436,181,1854]" captionTargetId="figure-19@16.[151,1436,181,1854]" captionTargetPageId="16" captionText="FIGURES 2834. Morphological structures of Botrodus dufaui Grouvelle. Body in dorsal (2829), ventral (30) and lateral (31) views; pterothorax in ventral view (32); and aedeagus (3334)." figureDoi="http://doi.org/10.5281/zenodo.6336951" httpUri="https://zenodo.org/record/6336951/files/figure.png" pageId="15" pageNumber="16">Figs 3334</figureCitation>
)
<quantity id="4CDA0DB29C427023D59DF8B2EB1DF837" box="[466,565,1899,1925]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="15" pageNumber="16" unit="mm" value="0.48">0.48 mm</quantity>
long, with parameres in lateral view straight, each ~3.5 times as long as broad at base, with three long apical setae, lacking ventral and sub-basal setae; penis inconspicuous, slender, nearly tubular and in distal half indistinctly curved.
</paragraph>
</subSubSection>
<subSubSection id="C338F3DC9C427023D488F80FEBABF840" box="[199,643,2006,2034]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8B9DA0579C427023D488F80FEBABF840" blockId="15.[151,1437,818,2034]" box="[199,643,2006,2034]" pageId="15" pageNumber="16">
<emphasis id="B9567C459C427023D488F80FE849F842" bold="true" box="[199,353,2006,2032]" pageId="15" pageNumber="16">Distribution.</emphasis>
Caribbean (
<collectingCountry id="F335E0C79C427023D5A6F80EEB5FF840" box="[489,631,2007,2034]" name="Guadeloupe" pageId="15" pageNumber="16">Guadeloupe</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>