344 lines
58 KiB
XML
344 lines
58 KiB
XML
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<mods:title id="1C3ADB990AAF17DDEDBA7A04A4BEF3E6">Five new species of the genus Philydrodes Bernhauer, 1929 from China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<taxonomicName id="4C234D3EFFE8B936FF41595684A49D70" ID-CoL="8XNVT" authority="Bernhauer, 1929" authorityName="Bernhauer" authorityYear="1929" box="[151,507,1588,1614]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAAFFFE8B936FF41595684799D70" bold="true" box="[151,294,1588,1614]" italics="true" pageId="1" pageNumber="170">Philydrodes</emphasis>
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<bibRefCitation id="EFB24B4CFFE8B936FEFB595684A49D70" author="Bernhauer" box="[301,507,1588,1614]" pageId="1" pageNumber="170" refString="Bernhauer, M. (1929) Neue Kurzflugler des palaarktischen Gebietes. Koleopterologische Rundschau, 14, 177 - 195." type="journal article" year="1929">Bernhauer, 1929</bibRefCitation>
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<subSubSection id="C3396536FFE8B936FF41591886249D91" pageId="1" pageNumber="170" type="reference_group">
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<paragraph id="8B9C36BDFFE8B936FF41591884AE9DAE" blockId="1.[151,891,1658,1711]" box="[151,497,1658,1680]" pageId="1" pageNumber="170">
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<taxonomicName id="4C234D3EFFE8B936FF41591884AE9DAE" ID-CoL="8XNVT" authority="Bernhauer, 1929: 179" authorityName="Bernhauer" authorityPageNumber="179" authorityYear="1929" box="[151,497,1658,1680]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAAFFFE8B936FF415918844E9DAE" box="[151,273,1658,1680]" italics="true" pageId="1" pageNumber="170">Philydrodes</emphasis>
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<treatmentCitation id="0A8210ACFFE8B936FEC0591884AE9DAE" author="Bernhauer" box="[278,497,1658,1680]" page="179" pageId="1" pageNumber="170" year="1929">
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<bibRefCitation id="EFB24B4CFFE8B936FEC05918849E9DAE" author="Bernhauer" box="[278,449,1658,1680]" pageId="1" pageNumber="170" refString="Bernhauer, M. (1929) Neue Kurzflugler des palaarktischen Gebietes. Koleopterologische Rundschau, 14, 177 - 195." type="journal article" year="1929">Bernhauer, 1929</bibRefCitation>
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: 179
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<paragraph id="8B9C36BDFFE8B936FF4159FB86249D91" blockId="1.[151,891,1658,1711]" box="[151,891,1689,1711]" pageId="1" pageNumber="170">
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<taxonomicName id="4C234D3EFFE8B936FF4159FB87169D91" ID-CoL="8XNVT" authority="Nakane & Sawada 1956: 179" authorityName="Nakane & Sawada" authorityPageNumber="179" authorityYear="1956" box="[151,585,1689,1711]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus">
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<emphasis id="B957EAAFFFE8B936FF4159FB844F9D91" box="[151,272,1689,1711]" italics="true" pageId="1" pageNumber="170">Philydrodes</emphasis>
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:
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<treatmentCitation id="0A8210ACFFE8B936FECB59FB87169D91" author="Nakane" box="[285,585,1689,1711]" page="179" pageId="1" pageNumber="170" year="1956">
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<bibRefCitation id="EFB24B4CFFE8B936FECB59FB87499D91" author="Nakane" box="[285,534,1689,1711]" pageId="1" pageNumber="170" refString="Nakane, T. & Sawada, K. (1956) On the genus Philydrodes Bernhauer in Japan with descriptions of a new subgenus and several new species. The Scientific Reports of the Saikyo University (A), 2, 179 - 186." type="journal article" year="1956">Nakane & Sawada 1956</bibRefCitation>
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: 179
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;
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<treatmentCitation id="0A8210ACFFE8B936FD8559FB86479D91" author="Watanabe" box="[595,792,1689,1711]" page="61" pageId="1" pageNumber="170" year="1990">
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<bibRefCitation id="EFB24B4CFFE8B936FD8559FB87AC9D91" author="Watanabe" box="[595,755,1689,1711]" pageId="1" pageNumber="170" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 55 - 391." type="journal article" year="1990">Watanabe 1990</bibRefCitation>
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: 61
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, 265, 309
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<emphasis id="B957EAAFFFE8B936FF4159BE84769DCB" bold="true" box="[151,297,1756,1781]" pageId="1" pageNumber="170">Description.</emphasis>
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Body more or less flattened, shiny, with coriaceous microsculpture on head and pronotum; length 3.0–8.0 mm. Coloration varying from reddish brown to black, sometimes with paler pronotum and/or elytra. Punctation variable: small and sometimes indistinct on head, deeper and denser on pronotum and elytra. Head subtrapezoidal (Fig.
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<bibRefCitation id="EFB24B4CFFE8B936FE1C582587289C5E" author="Watanabe" box="[458,631,1863,1888]" pageId="1" pageNumber="170" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 55 - 391." type="journal article" year="1990">Watanabe 1990</bibRefCitation>
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) to subquadrate (
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<figureCitation id="13182A38FFE8B936FCEE582586D69C5E" box="[824,905,1863,1888]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="1" pageNumber="170">Fig. 11</figureCitation>
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), with postocular parts significantly widened in male or not widened in both sexes, with wide and deep trapezoidal interocellar impression and moderately deep furrows (grooves) beginning above ocelli and obliquely extending posteriad; gular sutures united or slightly separated at narrowest point (
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<figureCitation id="13182A38FFE8B936FE3458D6876C9CF2" box="[482,563,1972,1997]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="1" pageNumber="170">Fig. 11</figureCitation>
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). Eyes convex, small to medium-sized. Ocelli moderately large, very distinct. Labrum transverse (
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<figureCitation id="13182A38FFE8B936FEA858BA84999CCF" box="[382,454,2008,2033]" captionStart="FIGURES 6 – 10" captionStartId="5.[151,264,949,972]" captionTargetBox="[160,1427,199,923]" captionTargetId="figure@5.[151,1436,193,929]" captionTargetPageId="5" captionText="FIGURES 6 – 10. Mouthparts (dorsal view) of Philydrodes tridentatus: 6 — labrum, 7 — mandibles (left and right), 8 — maxilla (right), 9 — labium, 10 — submentum. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291398/files/figure.png" pageId="1" pageNumber="170">Fig. 6</figureCitation>
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). Mandibles slender, each mandible with two elongated teeth on inner margin, right mandible with additional distinct smoothed tooth (
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<figureCitation id="13182A38FFEBB935FD375FF586769B8E" box="[737,809,151,176]" captionStart="FIGURES 6 – 10" captionStartId="5.[151,264,949,972]" captionTargetBox="[160,1427,199,923]" captionTargetId="figure@5.[151,1436,193,929]" captionTargetPageId="5" captionText="FIGURES 6 – 10. Mouthparts (dorsal view) of Philydrodes tridentatus: 6 — labrum, 7 — mandibles (left and right), 8 — maxilla (right), 9 — labium, 10 — submentum. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291398/files/figure.png" pageId="2" pageNumber="171">Fig. 7</figureCitation>
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). Labial palpomeres distinctly longer than their width (
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<figureCitation id="13182A38FFEBB935FF495FDE85B89BEA" box="[159,231,188,213]" captionStart="FIGURES 6 – 10" captionStartId="5.[151,264,949,972]" captionTargetBox="[160,1427,199,923]" captionTargetId="figure@5.[151,1436,193,929]" captionTargetPageId="5" captionText="FIGURES 6 – 10. Mouthparts (dorsal view) of Philydrodes tridentatus: 6 — labrum, 7 — mandibles (left and right), 8 — maxilla (right), 9 — labium, 10 — submentum. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291398/files/figure.png" pageId="2" pageNumber="171">Fig. 9</figureCitation>
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). Maxillae narrow, with elongated galea and maxillary palpomeres; apical maxillary palpomere narrow, indistinctly widened in median part and distinctly shorter than preceeding palpomere. Antennae reaching apical third or fourth of elytra, with very elongated antennomeres. Pronotum convex, moderately small, distinctly transverse or narrow, subequal to head; prothorax with elongated intercoxal process and wide triangular mesosternal processes (Fig.
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<bibRefCitation id="EFB24B4CFFEBB935FDFB5E2E87819A5A" author="Watanabe" box="[557,734,332,357]" pageId="2" pageNumber="171" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 55 - 391." type="journal article" year="1990">Watanabe 1990</bibRefCitation>
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) or near basal portion (
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<figureCitation id="13182A38FFEBB935FC265E2E81199A5A" box="[1008,1094,332,357]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 12</figureCitation>
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). Mesothorax elongated, with acute intercoxal process reaching posterior margin of mesocoxae (
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<figureCitation id="13182A38FFEBB935FC595E0D86BA9AB6" box="[911,997,367,392]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 14</figureCitation>
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). Scutellum large, with tongue-shaped apical portion (
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<figureCitation id="13182A38FFEBB935FE855EF684F19A92" box="[339,430,404,429]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 13</figureCitation>
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). Metathorax subtrapezoidal, with moderately deep intercoxal cavities and angular intercoxal process (
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<figureCitation id="13182A38FFEBB935FEAC5ED5848E9AEE" box="[378,465,439,464]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 14</figureCitation>
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). Metendosternite with elongated basal stalk and furcal arms, as in
|
||
<figureCitation id="13182A38FFEBB935FB245ED580169AEE" box="[1266,1353,439,464]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 16</figureCitation>
|
||
. Elytra moderately short, distinctly dilated posteriad. Hind wings fully developed. Legs slender, with transverse procoxae (
|
||
<figureCitation id="13182A38FFEBB935FF495E9D85AF9926" box="[159,240,511,536]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 18</figureCitation>
|
||
) and metacoxae (
|
||
<figureCitation id="13182A38FFEBB935FE605E9D87589926" box="[438,519,511,536]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 20</figureCitation>
|
||
) and suboval mesocoxae (
|
||
<figureCitation id="13182A38FFEBB935FCF95E9D86DC9926" box="[815,899,511,536]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 19</figureCitation>
|
||
); protarsi of males with dilated tarsomeres (
|
||
<figureCitation id="13182A38FFEBB935FAB95E9D85E99901" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 21</figureCitation>
|
||
); 4th segment of protarsi concaved apically (
|
||
<figureCitation id="13182A38FFEBB935FD765D4487AD9901" box="[672,754,550,575]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 21</figureCitation>
|
||
), 4th segments of meso- (
|
||
<figureCitation id="13182A38FFEBB935FBD05D4481089901" box="[1030,1111,550,575]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 22</figureCitation>
|
||
) and metatarsi (
|
||
<figureCitation id="13182A38FFEBB935FADC5D4480049901" box="[1290,1371,550,575]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 23</figureCitation>
|
||
) with elongated membranous apex lobed beneath; apical tarsal segments distinctly longer than three preceeding segments. Abdomen with two moderately wide transverse tomentose spots on abdominal tergite IV and with palisade fridge on apical margin of abdominal tergite VII. Aedeagus with large or small phallobase; parameres short and moderately wide, or long and narrow. Female genital segment with moderately wide and long gonocoxites, with very small transverse styli (
|
||
<figureCitation id="13182A38FFEBB935FD4F5DB887B199CD" box="[665,750,730,755]" captionStart="FIGURES 24 – 29" captionStartId="7.[151,264,1952,1975]" captionTargetBox="[165,1428,1097,1924]" captionTargetId="figure@7.[151,1436,1095,1931]" captionTargetPageId="7" captionText="FIGURES 24 – 29. Abdominal segments of Philydrodes tridentatus: 24 — male tergite VIII, 25 — male sternite VIII, 26 — male genital segment, 27 — female tergite VIII, 28 — female sternite VIII, 29 — female genital segment. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291274/files/figure.png" pageId="2" pageNumber="171">Fig. 29</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9C36BDFFEBB935FF115D9C87DA9EA1" blockId="2.[151,1437,151,2016]" pageId="2" pageNumber="171">
|
||
<emphasis id="B957EAAFFFEBB935FF115D9C844E9829" bold="true" box="[199,273,766,791]" pageId="2" pageNumber="171">Notes.</emphasis>
|
||
<bibRefCitation id="EFB24B4CFFEBB935FECA5D9C84BF9829" author="Watanabe" box="[284,480,766,791]" pageId="2" pageNumber="171" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 55 - 391." type="journal article" year="1990">Watanabe (1990)</bibRefCitation>
|
||
divided genus into two subgenera: nominative and
|
||
<emphasis id="B957EAAFFFEBB935FB935D9C80559829" box="[1093,1290,766,791]" italics="true" pageId="2" pageNumber="171">
|
||
<taxonomicName id="4C234D3EFFEBB935FB935D9C80599829" box="[1093,1286,766,791]" class="Insecta" family="Staphylinidae" genus="Minyphilydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">Minyphilydrodes</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The species from the nominative subgenus characterized by the larger body length (4.5–8.0 mm), by the distinctly pronounced dimorphism in males having considerably widened postocular parts of head (see Fig.
|
||
<quantity id="4CDB9B58FFEBB935FBBE5C2581EF9861" box="[1128,1200,838,863]" metricMagnitude="1" metricUnit="m" metricValue="1.07442" pageId="2" pageNumber="171" unit="in" value="423.0">423 in</quantity>
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<bibRefCitation id="EFB24B4CFFEBB935FB6F5C2480349861" author="Watanabe" box="[1209,1387,838,863]" pageId="2" pageNumber="171" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 55 - 391." type="journal article" year="1990">Watanabe 1990</bibRefCitation>
|
||
), by the slender penultimate segment of maxillary palpi three times longer than broad and by the more flattened pronotum with clearly visible lateral margins when viewed dorsally.
|
||
<taxonomicName id="4C234D3EFFEBB935FC625CEC812E9899" box="[948,1137,910,935]" class="Insecta" family="Staphylinidae" genus="Minyphilydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FC625CEC812E9899" box="[948,1137,910,935]" italics="true" pageId="2" pageNumber="171">Minyphilydrodes</emphasis>
|
||
</taxonomicName>
|
||
contains species with the smaller body length (3.0–
|
||
<quantity id="4CDB9B58FFEBB935FE6F5CD1874B98F5" box="[441,532,947,971]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="171" unit="mm" value="4.3">4.3 mm</quantity>
|
||
), without modified postocular parts of head (
|
||
<figureCitation id="13182A38FFEBB935FBC75CD0813C98F5" box="[1041,1123,946,971]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 11</figureCitation>
|
||
), with penultimate segment of maxillary palpi distinctly widened apicad and less than three times longer than broad, with more convex pronotum and lateral margins of pronotum indistinctly visible from the dorsal side. The Chinese species have some sufficient characters similar to that of
|
||
<taxonomicName id="4C234D3EFFEBB935FD835B7C864E9F09" box="[597,785,1054,1079]" class="Insecta" family="Staphylinidae" genus="Minyphilydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FD835B7C864E9F09" box="[597,785,1054,1079]" italics="true" pageId="2" pageNumber="171">Minyphilydrodes</emphasis>
|
||
</taxonomicName>
|
||
: proportions of maxillary palpi, not widened postocular parts of head (head of males is slightly wider than pronotum and distinctly wider in females). At the same time, the body length of Chinese species varies from
|
||
<quantity id="4CDB9B58FFEBB935FD465B04866A9F41" box="[656,821,1126,1151]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="5.7" metricValueMin="3.5" pageId="2" pageNumber="171" unit="mm" value="4.6" valueMax="5.7" valueMin="3.5">3.5 to 5.7 mm</quantity>
|
||
, the pronotum is distinctly smaller, narrower and less transverse, with clearly visible lateral margins if viewed dorsally, the prothorax with triangular mesosternal processes shifted to the basal portion (compare Fig.
|
||
<quantity id="4CDB9B58FFEBB935FD0D5BCD867E9FF9" box="[731,801,1198,1223]" metricMagnitude="1" metricUnit="m" metricValue="1.07696" pageId="2" pageNumber="171" unit="in" value="424.0">424 in</quantity>
|
||
<bibRefCitation id="EFB24B4CFFEBB935FCFE5BCC868B9FF9" author="Watanabe" box="[808,980,1198,1223]" pageId="2" pageNumber="171" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 55 - 391." type="journal article" year="1990">Watanabe 1990</bibRefCitation>
|
||
and
|
||
<figureCitation id="13182A38FFEBB935FBDF5BCC81049FF9" box="[1033,1115,1198,1223]" captionStart="FIGURES 11 – 23" captionStartId="6.[151,264,1628,1651]" captionTargetBox="[168,1426,347,1600]" captionTargetId="figure@6.[151,1436,332,1608]" captionTargetPageId="6" captionText="FIGURES 11 – 23. Body parts of Philydrodes tridentatus (paratype, male): 11 — ventral aspect of head, 12 — ventral aspect of prothorax, 13 — scutellum (dorsal view), 14 — ventral aspect of mesothorax, 15 — ventral aspect of metathorax, 16 — metendosternite (ventral view), 17 — ventral aspect of elytron, 18 — fore leg (anterior view), 19 — middle leg (anterior view), 20 — hind leg (anterior view), 21 — protarsus (dorsal view), 22 — mesotarsus (dorsal view), 23 — metatarsus (dorsal view). Scale bars 0.1 mm." httpUri="https://zenodo.org/record/291273/files/figure.png" pageId="2" pageNumber="171">Fig. 12</figureCitation>
|
||
), and aedeagus with small to large phallobase and long parameres. Three species (
|
||
<taxonomicName id="4C234D3EFFEBB935FD355BB086289FD5" box="[739,887,1234,1259]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="tibetanus" status="sp. nov.">
|
||
<emphasis id="B957EAAFFFEBB935FD355BB0865D9FD5" box="[739,770,1234,1259]" italics="true" pageId="2" pageNumber="171">Ph</emphasis>
|
||
.
|
||
<emphasis id="B957EAAFFFEBB935FCC65BB086289FD5" box="[784,887,1234,1259]" italics="true" pageId="2" pageNumber="171">tibetanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26457D4FFEBB935FCAB5BB086EA9FD5" box="[893,949,1234,1259]" pageId="2" pageNumber="171" rank="species">
|
||
<emphasis id="B957EAAFFFEBB935FCAB5BB086EA9FD5" bold="true" box="[893,949,1234,1259]" pageId="2" pageNumber="171">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C234D3EFFEBB935FC155BB081379FD5" box="[963,1128,1234,1259]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="tridentatus" status="sp. nov.">
|
||
<emphasis id="B957EAAFFFEBB935FC155BB086BD9FD5" box="[963,994,1234,1259]" italics="true" pageId="2" pageNumber="171">Ph</emphasis>
|
||
.
|
||
<emphasis id="B957EAAFFFEBB935FC395BB081379FD5" box="[1007,1128,1234,1259]" italics="true" pageId="2" pageNumber="171">tridentatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26457D4FFEBB935FBB95BB081F89FD5" box="[1135,1191,1234,1259]" pageId="2" pageNumber="171" rank="species">
|
||
<emphasis id="B957EAAFFFEBB935FBB95BB081F89FD5" bold="true" box="[1135,1191,1234,1259]" pageId="2" pageNumber="171">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C234D3EFFEBB935FB0B5BB0800B9FD5" box="[1245,1364,1234,1259]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="wrasei" status="sp. nov.">
|
||
<emphasis id="B957EAAFFFEBB935FB0B5BB081A39FD5" box="[1245,1276,1234,1259]" italics="true" pageId="2" pageNumber="171">Ph</emphasis>
|
||
.
|
||
<emphasis id="B957EAAFFFEBB935FADF5BB6800B9FD5" box="[1289,1364,1236,1259]" italics="true" pageId="2" pageNumber="171">wrasei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26457D4FFEBB935FA8D5BB080CD9FD5" box="[1371,1426,1234,1259]" pageId="2" pageNumber="171" rank="species">
|
||
<emphasis id="B957EAAFFFEBB935FA8D5BB080CD9FD5" bold="true" box="[1371,1426,1234,1259]" pageId="2" pageNumber="171">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
) have latero-apical and apical projections of the median lobe of the aedeagus similar to that of
|
||
<emphasis id="B957EAAFFFEBB935FAF45B94801E9E31" box="[1314,1345,1270,1295]" italics="true" pageId="2" pageNumber="171">Ph</emphasis>
|
||
. (s.str.)
|
||
<emphasis id="B957EAAFFFEBB935FF415A7E84799E0D" box="[151,294,1308,1331]" italics="true" pageId="2" pageNumber="171">tsurugisanus</emphasis>
|
||
<bibRefCitation id="EFB24B4CFFEBB935FEE65A7884B99E0D" author="Watanabe" box="[304,486,1306,1331]" pageId="2" pageNumber="171" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 55 - 391." type="journal article" year="1990">Watanabe, 1990</bibRefCitation>
|
||
from Shikoku (see Fig.
|
||
<quantity id="4CDB9B58FFEBB935FCD25A7986119E0D" box="[772,846,1306,1331]" metricMagnitude="1" metricUnit="m" metricValue="1.12268" pageId="2" pageNumber="171" unit="in" value="442.0">442 in</quantity>
|
||
<bibRefCitation id="EFB24B4CFFEBB935FC815A7881549E0D" author="Watanabe" box="[855,1035,1306,1331]" pageId="2" pageNumber="171" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 55 - 391." type="journal article" year="1990">Watanabe 1990</bibRefCitation>
|
||
). Thus, the described species from China were tentatively placed by me within
|
||
<taxonomicName id="4C234D3EFFEBB935FD465A5C86129E69" box="[656,845,1342,1367]" class="Insecta" family="Staphylinidae" genus="Minyphilydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FD465A5C86129E69" box="[656,845,1342,1367]" italics="true" pageId="2" pageNumber="171">Minyphilydrodes</emphasis>
|
||
</taxonomicName>
|
||
as a group of related species, and real phylogenetic relations between taxa of
|
||
<taxonomicName id="4C234D3EFFEBB935FE6A5A00871D9E45" box="[444,578,1378,1403]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FE6A5A00871D9E45" box="[444,578,1378,1403]" italics="true" pageId="2" pageNumber="171">Philydrodes</emphasis>
|
||
</taxonomicName>
|
||
must be verified in the future after the detailed examination of the internal structure of the aedeagi of Japanese species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3396536FFEBB934FF115AC884BA9929" lastPageId="3" lastPageNumber="172" pageId="2" pageNumber="171" type="diagnosis">
|
||
<paragraph id="8B9C36BDFFEBB934FF115AC881539A7E" blockId="2.[151,1437,151,2016]" lastBlockId="3.[151,1437,151,536]" lastPageId="3" lastPageNumber="172" pageId="2" pageNumber="171">
|
||
<emphasis id="B957EAAFFFEBB935FF115AC8848F9EFD" bold="true" box="[199,464,1450,1475]" pageId="2" pageNumber="171">Differential diagnosis.</emphasis>
|
||
Based on the shape of mandibles, the presence and arrangment of two teeth on the inner margin, the short apical segment of the maxillary palp distinctly smaller and narrower than preceeding segment, by the densely pubescent antennes, and by the general shape of the aedeagus,
|
||
<taxonomicName id="4C234D3EFFEBB935FB525A9080559D35" box="[1156,1290,1522,1547]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FB525A9080559D35" box="[1156,1290,1522,1547]" italics="true" pageId="2" pageNumber="171">Philydrodes</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C234D3EFFEBB935FF41597484A79D11" authority="Tronquet, 1981" authorityName="Tronquet" authorityYear="1981" box="[151,504,1558,1583]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FF415974841A9D11" box="[151,325,1558,1583]" italics="true" pageId="2" pageNumber="171">Hygrodromicus</emphasis>
|
||
Tronquet, 1981
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C234D3EFFEBB935FDD2597486119D11" authority="Nakane, 1983" authorityName="Nakane" authorityYear="1983" box="[516,846,1558,1583]" class="Insecta" family="Staphylinidae" genus="Liophilydrodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FDD2597487F39D11" box="[516,684,1558,1583]" italics="true" pageId="2" pageNumber="171">Liophilydrodes</emphasis>
|
||
<bibRefCitation id="EFB24B4CFFEBB935FD65597486119D11" author="Nakane" box="[691,846,1558,1583]" pageId="2" pageNumber="171" refString="Nakane, T. (1983) New name for genus Lioplax Nakane et K. Sawada. Fragmenta Coleopterologica, 35 / 37, 148." type="journal article" year="1983">Nakane, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C234D3EFFEBB935FC55597481F69D11" authority="LeConte, 1874" authorityName="LeConte" authorityYear="1874" box="[899,1193,1558,1583]" class="Insecta" family="Staphylinidae" genus="Microedus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FC55597486A59D11" box="[899,1018,1558,1583]" italics="true" pageId="2" pageNumber="171">Microedus</emphasis>
|
||
LeConte, 1874
|
||
</taxonomicName>
|
||
, and, by the character of microsculpture of the head and pronotum and by the very narrow apical maxillary palpomere it is more similar to the latter two genera. It can be distinguished from
|
||
<taxonomicName id="4C234D3EFFEBB935FD31593C86019D49" box="[743,862,1630,1655]" class="Insecta" family="Staphylinidae" genus="Microedus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FD31593C86019D49" box="[743,862,1630,1655]" italics="true" pageId="2" pageNumber="171">Microedus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B957EAAFFFEBB935FC42593C81639D49" box="[916,1084,1630,1655]" italics="true" pageId="2" pageNumber="171">Liophylidrodes</emphasis>
|
||
by the slightly widened medial part of the apical maxillary palpomere (compared to genera with apical maxillary palpomere narrower, parallelsided and awl-shaped, and with wider preapical palpomere), and by the less transverse pronotum. From
|
||
<taxonomicName id="4C234D3EFFEBB935FF4159A8841A9DDD" box="[151,325,1738,1763]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FF4159A8841A9DDD" box="[151,325,1738,1763]" italics="true" pageId="2" pageNumber="171">Hygrodromicus</emphasis>
|
||
</taxonomicName>
|
||
it differs by the narrower body and by the distinctly wider preapical maxillar palpomere, as well as by the well developed ocelli (ocelli of some species of
|
||
<taxonomicName id="4C234D3EFFEBB935FCC9598C86929C39" box="[799,973,1774,1799]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FCC9598C86929C39" box="[799,973,1774,1799]" italics="true" pageId="2" pageNumber="171">Hygrodromicus</emphasis>
|
||
</taxonomicName>
|
||
can be reduced). From all these genera
|
||
<taxonomicName id="4C234D3EFFEBB935FF41587084429C15" box="[151,285,1810,1835]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FF41587084429C15" box="[151,285,1810,1835]" italics="true" pageId="2" pageNumber="171">Philydrodes</emphasis>
|
||
</taxonomicName>
|
||
differs by the larger head, sometimes distinctly wider than pronotum as in
|
||
<taxonomicName id="4C234D3EFFEBB935FBAA587080C39C15" authority="LeConte, 1852" authorityName="LeConte" authorityYear="1852" box="[1148,1436,1810,1835]" class="Insecta" family="Staphylinidae" genus="Brathinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FBAA587081B59C15" box="[1148,1258,1810,1835]" italics="true" pageId="2" pageNumber="171">Brathinus</emphasis>
|
||
LeConte, 1852
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C234D3EFFEBB935FF1D5854871B9C71" authority="Gravenhorst, 1802" authorityName="Gravenhorst" authorityYear="1802" box="[203,580,1846,1871]" class="Insecta" family="Staphylinidae" genus="Anthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FF1D585484009C71" box="[203,351,1846,1871]" italics="true" pageId="2" pageNumber="171">Anthophagus</emphasis>
|
||
Gravenhorst, 1802
|
||
</taxonomicName>
|
||
, with significantly or insignificantly pronounced sexual dimorphism (for details see above), by the more slender mandibles with narrow basal part, by the narrower maxillary palpi, and labial palpomeres. The shape of phallobase of some Chinese species of
|
||
<taxonomicName id="4C234D3EFFEBB935FBDE581C81D19CA9" box="[1032,1166,1918,1943]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FBDE581C81D19CA9" box="[1032,1166,1918,1943]" italics="true" pageId="2" pageNumber="171">Philydrodes</emphasis>
|
||
</taxonomicName>
|
||
are similar to that of
|
||
<taxonomicName id="4C234D3EFFEBB935FF4158C084AD9C85" authority="Zerche 1992" authorityName="Zerche" authorityYear="1992" box="[151,498,1954,1979]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FF4158C0841A9C85" box="[151,325,1954,1979]" italics="true" pageId="2" pageNumber="171">Hygrodromicus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB24B4CFFEBB935FE8058C084B49C85" author="Zerche" box="[342,491,1954,1979]" pageId="2" pageNumber="171" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche 1992</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
which also can be 90° rotated inside the abdomen in lateral position (if viewed dorsally). In habitus,
|
||
<taxonomicName id="4C234D3EFFEBB935FE5F58A487509CE1" box="[393,527,1990,2015]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEBB935FE5F58A487509CE1" box="[393,527,1990,2015]" italics="true" pageId="2" pageNumber="171">Philydrodes</emphasis>
|
||
</taxonomicName>
|
||
is related to the Holarctic group of genera of the tribe
|
||
<taxonomicName id="4C234D3EFFEBB935FB5658A480429CE1" box="[1152,1309,1990,2015]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="tribe" tribe="Anthophagini">Anthophagini</taxonomicName>
|
||
, all taxa of which have deep interocellar impressions between eyes.
|
||
<bibRefCitation id="EFB24B4CFFEAB934FCCE5FF586ED9B8E" author="Zerche" box="[792,946,151,176]" pageId="3" pageNumber="172" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche (1992</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB24B4CFFEAB934FC685FFA86A79B8E" author="Zerche" box="[958,1016,152,176]" pageId="3" pageNumber="172" refString="Zerche, L. (2003) Arten der Hygrogeus - Gruppe aus dem Fernen Osten Russlands, aus Japan und China. Beitrage zur Entomologie, 53 (2), 277 - 295." type="journal article" year="2003">2003</bibRefCitation>
|
||
) named these genera as “
|
||
<taxonomicName id="4C234D3EFFEAB934FACF5FFB80CC9B8E" box="[1305,1427,153,176]" class="Insecta" family="Staphylinidae" genus="Hygrogeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEAB934FACF5FFB80CC9B8E" box="[1305,1427,153,176]" italics="true" pageId="3" pageNumber="172">Hygrogeus</emphasis>
|
||
</taxonomicName>
|
||
- Gruppe” and included several palaearctic genera (
|
||
<taxonomicName id="4C234D3EFFEAB934FD135FDE81519BEA" authority="Zerche, 1992" authorityName="Zerche" authorityYear="1992" box="[709,1038,188,213]" class="Insecta" family="Staphylinidae" genus="Altaiodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEAB934FD135FDE862E9BEB" box="[709,881,188,213]" italics="true" pageId="3" pageNumber="172">Altaiodromicus</emphasis>
|
||
<bibRefCitation id="EFB24B4CFFEAB934FCAE5FDE81519BEA" author="Zerche" box="[888,1038,188,213]" pageId="3" pageNumber="172" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C234D3EFFEAB934FBCC5FDE80C79BEB" authority="Redtenbacher, 1857" authorityName="Redtenbacher" authorityYear="1857" box="[1050,1432,188,213]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEAB934FBCC5FDE81EF9BEB" box="[1050,1200,188,213]" italics="true" pageId="3" pageNumber="172">Geodromicus</emphasis>
|
||
Redtenbacher, 1857
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C234D3EFFEAB934FF415FBD87429BC6" authority="Tronquet, 1981" authorityName="Tronquet" authorityYear="1981" box="[151,541,223,248]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEAB934FF415FBD841A9BC6" box="[151,325,223,248]" italics="true" pageId="3" pageNumber="172">Hygrodromicus</emphasis>
|
||
Tronquet, 1981
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C234D3EFFEAB934FDED5F8381569BC6" authority="Mulsant and Rey, 1880" authorityName="Mulsant and Rey" authorityYear="1880" box="[571,1033,223,248]" class="Insecta" family="Staphylinidae" genus="Hygrogeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEAB934FDED5F8387EA9BC6" box="[571,693,225,248]" italics="true" pageId="3" pageNumber="172">Hygrogeus</emphasis>
|
||
Mulsant and Rey, 1880
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C234D3EFFEAB934FBF05FBD80C79BC6" authority="Nakane, 1983" authorityName="Nakane" authorityYear="1983" box="[1062,1432,223,248]" class="Insecta" family="Staphylinidae" genus="Liophilydrodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEAB934FBF05FBD81909BC6" box="[1062,1231,223,248]" italics="true" pageId="3" pageNumber="172">Liophilydrodes</emphasis>
|
||
<bibRefCitation id="EFB24B4CFFEAB934FB3E5FBD80C79BC6" author="Nakane" box="[1256,1432,223,248]" pageId="3" pageNumber="172" refString="Nakane, T. (1983) New name for genus Lioplax Nakane et K. Sawada. Fragmenta Coleopterologica, 35 / 37, 148." type="journal article" year="1983">Nakane, 1983</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C234D3EFFEAB934FF415E6687489A22" authority="Zerche, 1992" authorityName="Zerche" authorityYear="1992" box="[151,535,260,285]" class="Insecta" family="Staphylinidae" genus="Paratrichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEAB934FF415E66842E9A23" box="[151,369,260,285]" italics="true" pageId="3" pageNumber="172">Paratrichodromeus</emphasis>
|
||
<bibRefCitation id="EFB24B4CFFEAB934FEAA5E6687489A22" author="Zerche" box="[380,535,260,285]" pageId="3" pageNumber="172" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C234D3EFFEAB934FDF05E6686009A22" authority="Luze, 1903" authorityName="Luze" authorityYear="1903" box="[550,863,260,285]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEAB934FDF05E6687909A23" box="[550,719,260,285]" italics="true" pageId="3" pageNumber="172">Trichodromeus</emphasis>
|
||
Luze, 1903
|
||
</taxonomicName>
|
||
), and to which
|
||
<taxonomicName id="4C234D3EFFEAB934FBCA5E6681CC9A23" box="[1052,1171,260,285]" class="Insecta" family="Staphylinidae" genus="Microedus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEAB934FBCA5E6681CC9A23" box="[1052,1171,260,285]" italics="true" pageId="3" pageNumber="172">Microedus</emphasis>
|
||
</taxonomicName>
|
||
, with several Nearctic (
|
||
<bibRefCitation id="EFB24B4CFFEAB934FF485E45847C9A7E" author="Hatch" box="[158,291,295,320]" pageId="3" pageNumber="172" refString="Hatch, M. H. (1957) The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, 16 (2), i - ix + 1 - 384." type="journal article" year="1957">Hatch 1957</bibRefCitation>
|
||
) and Palaearctic (
|
||
<bibRefCitation id="EFB24B4CFFEAB934FE3D5E4A87949A7E" author="Newton" box="[491,715,295,320]" pageId="3" pageNumber="172" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. & Chandler, D. S. (2000) 22. Staphylinidae Latreille, 1802. In: Arnett, R. H. & Thomas, M. C. (Ed.), American Beetles. Fol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 272 - 418." type="journal article" year="2000">Newton et al., 2000</bibRefCitation>
|
||
) species, should be referred.
|
||
</paragraph>
|
||
<paragraph id="8B9C36BDFFEAB934FF115E2E84BA9929" blockId="3.[151,1437,151,536]" pageId="3" pageNumber="172">
|
||
<emphasis id="B957EAAFFFEAB934FF115E2E84159A5B" bold="true" box="[199,330,332,357]" pageId="3" pageNumber="172">Bionomics.</emphasis>
|
||
The detailed bionomics data for the most Japanese
|
||
<taxonomicName id="4C234D3EFFEAB934FC735E2E81749A5B" box="[933,1067,332,357]" class="Insecta" family="Staphylinidae" genus="Philydrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="172" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B957EAAFFFEAB934FC735E2E81749A5B" box="[933,1067,332,357]" italics="true" pageId="3" pageNumber="172">Philydrodes</emphasis>
|
||
</taxonomicName>
|
||
are unknown:
|
||
<bibRefCitation id="EFB24B4CFFEAB934FB0C5E2E80C39A5A" author="Watanabe" box="[1242,1436,332,357]" pageId="3" pageNumber="172" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 55 - 391." type="journal article" year="1990">Watanabe (1990)</bibRefCitation>
|
||
provided only a comment that these species can be “[...] found in damp dead leaves accumulated along mountain streams or under stones at the stream side”. Two species described from the Eastern Shikoku were collected in high altitudes near streams in wet litter (
|
||
<bibRefCitation id="EFB24B4CFFEAB934FDE45ED587B79AEE" author="Watanabe" box="[562,744,439,464]" pageId="3" pageNumber="172" refString="Watanabe, Y. (2012) Two new species of the subgenus Minyphilhydrodes (Coleoptera, Staphylinidae) from Eastern Shikoku, Japan. Japanese Journal of Systematic Entomology, 18 (1), 17 - 22." type="journal article" year="2012">Watanabe 2012</bibRefCitation>
|
||
). All Chinese species have been collected in high mountain habitats with elevations between
|
||
<quantity id="4CDB9B58FFEAB934FDDF5EBE87019ACA" box="[521,606,476,501]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="172" unit="m" value="1550.0">1550 m</quantity>
|
||
a.s.l. to
|
||
<quantity id="4CDB9B58FFEAB934FD6A5EBE864E9ACA" box="[700,785,476,500]" metricMagnitude="3" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="172" unit="m" value="4200.0">4200 m</quantity>
|
||
a.s.l. from wet mosses, debris, leaf litter, under stones etc. near rivers and small streams.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |