346 lines
48 KiB
XML
346 lines
48 KiB
XML
<document ID-CLB-Dataset="128624" ID-DOI="http://dx.doi.org/10.3897/zookeys.1118.83156" ID-GBIF-Dataset="1ee2e89c-fb32-4486-8b69-3d8ab0845ea6" ID-PMC="PMC9848743" ID-Pensoft-Pub="1313-2970-1118-133" ID-Pensoft-UUID="2D72BBAAE1585D86B1DFD3CEC29E0E61" ID-PubMed="36761806" ID-ZooBank="43714075E8324248991638E5AEFA9F71" ModsDocID="1313-2970-1118-133" checkinTime="1661416745058" checkinUser="pensoft" docAuthor="Ha, Ngoc Linh, Truong, Xuan Lam, Ishikawa, Tadashi, Jaitrong, Weeyawat, Lee, Chi Feng, Chouangthavy, Bounsanong & Eguchi, Katsuyuki" docDate="2022" docId="EFED6E08BFEC5FD18C238DAFB14CF4EA" docLanguage="en" docName="ZooKeys 1118: 133-180" docOrigin="ZooKeys 1118" docPubDate="2022-08-24" docSource="http://dx.doi.org/10.3897/zookeys.1118.83156" docTitle="Biasticus griseocapillus Ha, Truong & Ishikawa 2022, sp. nov." docType="treatment" docUuid="BDAB0B60-64D5-4529-8715-2E1B010AD57B" docUuidSource="ZooBank" docVersion="6" id="2D72BBAAE1585D86B1DFD3CEC29E0E61" lastPageNumber="133" masterDocId="2D72BBAAE1585D86B1DFD3CEC29E0E61" masterDocTitle="Three new species of the genus Biasticus Stal, 1867 (Insecta, Heteroptera, Reduviidae, Harpactorinae) from Central Highlands, Vietnam" masterLastPageNumber="180" masterPageNumber="133" pageNumber="133" updateTime="1719497085934" updateUser="ExternalLinkService">
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<mods:title id="060C7DF72E2BDA05C45626F9913570B5">Three new species of the genus Biasticus Stal, 1867 (Insecta, Heteroptera, Reduviidae, Harpactorinae) from Central Highlands, Vietnam</mods:title>
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<mods:namePart id="9024BEA61F8E4AA48627E9557DC5E0C0">Ha, Ngoc Linh</mods:namePart>
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<mods:affiliation id="F0F636418E3D5E33B92EF333A0AC2C63">Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, Minami-osawa 1 - 1, Hachioji, Tokyo 192 - 0397, Japan & Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:nameIdentifier id="FF4FBC6045FEF4C27B3041BBB076CF73" type="email">linh.hangoc02@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D9B58568D559B1705957CB7904995EAE">Truong, Xuan Lam</mods:namePart>
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<mods:affiliation id="1B2EA3C4C58EFD5C37441C56941701F9">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="BD1150B42B71C7EAA9FC4EB872F76EBB">Ishikawa, Tadashi</mods:namePart>
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<mods:affiliation id="3F482F8F473EEBF63F14F55A5F3D596E">Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="946B7C4C58C9347E2B934241799EA31F">Jaitrong, Weeyawat</mods:namePart>
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<mods:affiliation id="732696D4AF4E2EE1373242823DC66B5E">Faculty of Agriculture, Tokyo University of Agriculture, Atsugi, Kanagawa, Japan</mods:affiliation>
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<mods:namePart id="A7887C5D0EF54D8315086AB50334FB7D">Lee, Chi Feng</mods:namePart>
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<mods:affiliation id="858136D40807DB83C0730DFDB149D039">Office of Natural Science Research, National Science Museum, 39 Moo 3, Khlong 5, Khlong Luang, Pathum Thani, 12120 Thailand</mods:affiliation>
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<mods:namePart id="7C66F67117DDEA8F08BEE737D5300AEE">Chouangthavy, Bounsanong</mods:namePart>
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<mods:affiliation id="BC31A69B4929637BBBE8D374F5A38F7B">Applied Zoology Division, Taiwan Agricultural Research Institute, Taichung 413, Taiwan</mods:affiliation>
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<mods:namePart id="4127910FF1C00431D568B9F685D05EB1">Eguchi, Katsuyuki</mods:namePart>
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<mods:affiliation id="6DAE8C25EC7B33FF80925905A4FBD900">Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, Minami-osawa 1 - 1, Hachioji, Tokyo 192 - 0397, Japan</mods:affiliation>
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<mods:date id="D17C22D363BA363583FFA3B4087C4DFC">2022</mods:date>
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<mods:number id="C97ECEEEF43E7807362F6E669EAF7E25">2022-08-24</mods:number>
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<treatment id="EFED6E08BFEC5FD18C238DAFB14CF4EA" ID-GBIF-Taxon="199821304" LSID="urn:lsid:zoobank.org:act:BDAB0B60-64D5-4529-8715-2E1B010AD57B" httpUri="http://treatment.plazi.org/id/EFED6E08BFEC5FD18C238DAFB14CF4EA" lastPageNumber="133" pageId="0" pageNumber="133">
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<subSubSection id="95B54E8617CF30EAB630CE5EE966AAFA" pageId="0" pageNumber="133" type="nomenclature">
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<paragraph id="7C6374B735150B8C9348C17D9DD3B572" pageId="0" pageNumber="133">
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<taxonomicName id="AFFA4059A11FDAAE5ADFFB531A4F240B" LSID="https://zoobank.org/BDAB0B60-64D5-4529-8715-2E1B010AD57B" authority="Ha, Truong & Ishikawa" authorityName="Ha, Truong & Ishikawa" authorityYear="2022" class="Insecta" family="Reduviidae" genus="Biasticus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Biasticus griseocapillus" order="Hemiptera" pageId="0" pageNumber="133" phylum="Arthropoda" rank="species" species="griseocapillus" status="sp. nov.">Biasticus griseocapillus Ha, Truong & Ishikawa</taxonomicName>
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<taxonomicNameLabel id="68CD959A02FB51EC6C435CC7087E1C20" pageId="0" pageNumber="133">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="C2164FACA280372C4DB30C0035FDE8A1" pageId="0" pageNumber="133" type="description">
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<paragraph id="3C9F05F0EF522328768CB02D2BC3E33E" pageId="0" pageNumber="133">
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<figureCitation id="50E6E3E47A2E06BBC5A116698A7D2863" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Female external genitalia of six morphospecies in ventral view A type fA, Biasticus taynguyenensis Ha, Truong & Ishikawa, sp. nov., holotype, ♀, HNL 2018 - 073 B type fB, B. griseocapillus Ha, Truong & Ishikawa, sp. nov., holotype, ♀, HNL 2018 - 038 C type fC, B. luteicollis Ha, Truong & Ishikawa, sp. nov., paratype, ♀, HNL 2018 - 024 D type fD, B. flavinotus (Matsumura, 1913), ♀, HNL 2018 - 117 E type fE, B. confusus Hsiao et al., 1979, ♀, VN-Hem- 1998 - 012 F type fF, B. flavus (Distant, 1903), ♀, LA-Redu- 2004 - 006." figureDoi="10.3897/zookeys.1118.83156.figure5" httpUri="https://binary.pensoft.net/fig/734748" pageId="0" pageNumber="133">Figs 5B</figureCitation>
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<figureCitation id="6290FDE5CC440B9E51DC9E3E9B889A35" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia of four morphospecies A-D type mA, Biasticus taynguyenensis Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 545 E-I type mB, B. griseocapillus Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 546 J-N type mC, B. luteicollis Ha, Truong & Ishikawa, sp. nov., holotype, ♂, HNL 2018 - 025 O-R mF, B. flavus (Distant, 1903), ♂, HEM-TH 2004 - 018. A, E, J, O pygophore with paramere (s) in dorsal view B, F, K, P phallus in dorsal view C, G, L, Q phallus in lateral view D, I, N, R articulatory apparatus H, M distal dorsal lobe of endosoma." figureDoi="10.3897/zookeys.1118.83156.figure6" httpUri="https://binary.pensoft.net/fig/734749" pageId="0" pageNumber="133">, 6E-I</figureCitation>
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<figureCitation id="F499D9D4682BF93966DE02EC6AC6A22F" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Biasticus griseocapillus Ha, Truong & Ishikawa, sp. nov., holotype, ♀, HNL 2018 - 038 A body in dorsal view B body in lateral view C head in dorsal view D head in lateral view." figureDoi="10.3897/zookeys.1118.83156.figure16" httpUri="https://binary.pensoft.net/fig/734759" pageId="0" pageNumber="133">, 16</figureCitation>
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<figureCitation id="0FF787A88777B454CB0441A8D2224DAA" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Biasticus griseocapillus Ha, Truong & Ishikawa, sp. nov., holotype, ♀, HNL 2018 - 038 A pronotum in dorsal view B anterior pronotal lobe in dorsal view C, D setae on posterior pronotum E right hemelytron F right hind wing." figureDoi="10.3897/zookeys.1118.83156.figure17" httpUri="https://binary.pensoft.net/fig/734760" pageId="0" pageNumber="133">, 17</figureCitation>
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<figureCitation id="FD650F34849DA71ED54DFC07F8DE0094" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Biasticus griseocapillus Ha, Truong & Ishikawa, sp. nov. A female genitalia in ventral view, holotype, ♀, HNL 2018 - 038 B-M male genitalia, paratype, ♂, TXL 2016 - 546 B-F pygophore with paramere B dorsal view C lateral view D ventral view E apical portion of pygophore, showing median process (mpp) and paramere F left paramere G-M phallus G dorsal view H lateral view I articulatory apparatus (aa) J aedeagus with endosoma semi-everted, dorsal view K aedeagus with endosoma semi-everted, ventral view L aedeagus with endosoma semi-everted, lateral view M distal dorsal lobe of endosoma (ddl)." figureDoi="10.3897/zookeys.1118.83156.figure18" httpUri="https://binary.pensoft.net/fig/734761" pageId="0" pageNumber="133">, 18</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="729979EA5921C1EDA22637AD8309F90D" pageId="0" pageNumber="133" type="type material">
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<paragraph id="475EA5AAF26CF2DF73CD6C6A2222C283" pageId="0" pageNumber="133">Type material.</paragraph>
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<paragraph id="58E62A0775904EB0BF4900858ACEE5B8" pageId="0" pageNumber="133">
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||
<emphasis id="00D287DA7BA15D3EDA20E4F198E79F67" bold="true" pageId="0" pageNumber="133">
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<emphasis id="ECCB5115297EFCADD8299DB298602B84" italics="true" pageId="0" pageNumber="133">Holotype</emphasis>
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.
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</emphasis>
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1♀; HNL2018-038; Vietnam, Dak Lak Province, Chu Yang Sin National Park; 09.v.2018; X. L. Truong leg.; NSMT.
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<emphasis id="329DF525C7151628EC34A4BA3EA93F35" bold="true" pageId="0" pageNumber="133">
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<emphasis id="343F645A2393EF978962BFEBDDA4A006" italics="true" pageId="0" pageNumber="133">Paratypes</emphasis>
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.
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</emphasis>
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1♀; HNL2018-007; Vietnam, Gia Lai Province, Kon Chu Rang Nature Reserve; 05.v.2018; X. L. Truong leg.; IEBR. 1♀; HNL2018-037; Vietnam, Dak Lak Province, Chu Yang Sin National Park; 09.v.2018; X. L. Truong leg.; VNMN. 1♂; TXL2016-546; Vietnam, Gia Lai Province, Kon Chu Rang Nature Reserve; 28.iv.2016; X. L. Truong leg.; NSMT.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="462ECB1DA5003DDB96629522BE4F6309" pageId="0" pageNumber="133" type="diagnosis">
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<paragraph id="12063BF05B383B683EFF81665EA942C5" pageId="0" pageNumber="133">Diagnosis.</paragraph>
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<paragraph id="5E483D8ACF50EF44539B7AA3838AA741" pageId="0" pageNumber="133">Body shiny blackish brown; anterior pronotal lobe black or blackish brown with some rows of long bent griseous setae; posterior pronotal lobe blackish brown or brown and densely covered with short bent griseous setae somewhat interspersed with long griseous setae; scutellum black in basal half and dark brown or brown in lateral margin and apical half; abdominal sternites shiny sanguineous; laterotergites II-VI luteous, segmentally suffused with dark brown spots or blackish brown spots; laterotergite VII sanguineous.</paragraph>
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<paragraph id="3BA8D87A91BC80C7B18F8D8EAC686F7C" pageId="0" pageNumber="133">
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In general appearance,
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<taxonomicName id="D99A5911AAD0B034FC9BD714181E5F5D" authorityName="Ha, Truong & Ishikawa" authorityYear="2022" class="Clitellata" family="Reduviidae" genus="Biasticus" kingdom="Animalia" lsidName="Biasticus griseocapillus" order="Hemiptera" pageId="0" pageNumber="133" phylum="Annelida" rank="species" species="griseocapillus">
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<emphasis id="976DEE23BADFECAAE2925B997C3F4822" italics="true" pageId="0" pageNumber="133">Biasticus griseocapillus</emphasis>
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</taxonomicName>
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sp. nov. resembles
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<taxonomicName id="A34BF758DD81C8C79A73C29A77D3796D" lsidName="B. confusus" pageId="0" pageNumber="133" rank="species" species="confusus">
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<emphasis id="803EA9F7D4A49D21B2D19C7901D5E5CD" italics="true" pageId="0" pageNumber="133">B. confusus</emphasis>
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</taxonomicName>
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Hsiao et al., 1979, especially in general colors of body, pronotum, and sterna. But the new species can be distinguished from
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<taxonomicName id="B663A933405BA8872B2C5CD1B86D29DB" lsidName="B. confusus" pageId="0" pageNumber="133" rank="species" species="confusus">
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<emphasis id="5DDD9085C4C8E44B1948142CBB9B4A84" italics="true" pageId="0" pageNumber="133">B. confusus</emphasis>
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</taxonomicName>
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by a combination of the following characters: antennal pedicel longer than first flagellomere (in
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<taxonomicName id="0DB520DCBC5C13A18D40D1E997DEF485" lsidName="B. confusus" pageId="0" pageNumber="133" rank="species" species="confusus">
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<emphasis id="1BD1EAA16BB6DB99C5985386BA003A14" italics="true" pageId="0" pageNumber="133">B. confusus</emphasis>
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</taxonomicName>
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pedicel as long as first flagellomere), second flagellomere of antenna as long as pedicel (in
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<taxonomicName id="F0AB55A2FF8D4B134AAC58CCB81C9373" lsidName="B. confusus" pageId="0" pageNumber="133" rank="species" species="confusus">
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<emphasis id="1DC2BF087FA314B8813DF4E3DCD24C90" italics="true" pageId="0" pageNumber="133">B. confusus</emphasis>
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</taxonomicName>
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second flagellomere longer than pedicel), proportional average length of first to third visible labial segments 1.1:1.4:0.3 (in
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<taxonomicName id="D95072233B1A59C595978383E4E1C30A" lsidName="B. confusus" pageId="0" pageNumber="133" rank="species" species="confusus">
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<emphasis id="D63A8261496ADE5D29000A4D43B96E42" italics="true" pageId="0" pageNumber="133">B. confusus</emphasis>
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</taxonomicName>
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0.9:1.2:0.3), anterior pronotal lobe with some rows of bent setae (in
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<taxonomicName id="27570B93F66A529AE1FE6896BE5F0F3A" lsidName="B. confusus" pageId="0" pageNumber="133" rank="species" species="confusus">
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<emphasis id="8A73F03ED6635A1CA359449D83B4409D" italics="true" pageId="0" pageNumber="133">B. confusus</emphasis>
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</taxonomicName>
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without row of bent setae), posterior pronotal lobe 2.6
|
||
<normalizedToken id="4591F30BC5AE58352C5F6EAAA8C32AB5" originalValue="×">x</normalizedToken>
|
||
as long as anterior pronotal lobe (in
|
||
<taxonomicName id="202821FF773DFF7E9C5D256D48FD21B5" lsidName="B. confusus" pageId="0" pageNumber="133" rank="species" species="confusus">
|
||
<emphasis id="C3238D221F40E33753DE539B218BDC31" italics="true" pageId="0" pageNumber="133">B. confusus</emphasis>
|
||
</taxonomicName>
|
||
2.0
|
||
<normalizedToken id="9B6113B1AE02609674ADCBA60C1E4856" originalValue="×">x</normalizedToken>
|
||
), and apical margin of median process of pygophore weakly convex and emarginate at middle (in
|
||
<taxonomicName id="BBE9698B3EC3817B7D5839B3E210111E" lsidName="B. confusus" pageId="0" pageNumber="133" rank="species" species="confusus">
|
||
<emphasis id="EA3F517DA3E346718BADFF25D1527399" italics="true" pageId="0" pageNumber="133">B. confusus</emphasis>
|
||
</taxonomicName>
|
||
weakly convex without such emargination).
|
||
</paragraph>
|
||
<paragraph id="95CEFDB5FC3E4AE176D2794AFADB5771" pageId="0" pageNumber="133">
|
||
Furthermore, this species is similar to
|
||
<taxonomicName id="BD2C21EC9EDAB4781DF78BA8C55DE795" lsidName="B. ventralis" pageId="0" pageNumber="133" rank="species" species="ventralis">
|
||
<emphasis id="BF885CB61BB961CBFA693BBB1995313C" italics="true" pageId="0" pageNumber="133">B. ventralis</emphasis>
|
||
</taxonomicName>
|
||
Hsiao et al., 1979 in general colors of body, pronotum, and sterna but the new species can be distinguished from
|
||
<taxonomicName id="304AEAA6EBD8D7F18F2A0CB5AD1E337F" lsidName="B. ventralis" pageId="0" pageNumber="133" rank="species" species="ventralis">
|
||
<emphasis id="A3C5979B517E93E9538465DCF0EDE42B" italics="true" pageId="0" pageNumber="133">B. ventralis</emphasis>
|
||
</taxonomicName>
|
||
by a combination of the following characters: antennal pedicel longer than first flagellomere (in
|
||
<taxonomicName id="86FD5ABF6A722FCA1D29A30132229B43" lsidName="B. ventralis" pageId="0" pageNumber="133" rank="species" species="ventralis">
|
||
<emphasis id="C93BFC375432E8A5BDC77CD8FE93EC8F" italics="true" pageId="0" pageNumber="133">B. ventralis</emphasis>
|
||
</taxonomicName>
|
||
pedicel ~ 1/2 as long as first flagellomere), second flagellomere as long as pedicel (in
|
||
<taxonomicName id="371FE6EC1BA9D923F39D5D12F7D6A2FC" lsidName="B. ventralis" pageId="0" pageNumber="133" rank="species" species="ventralis">
|
||
<emphasis id="796D1FA599CE72322B1E8F3AF6DAC0C8" italics="true" pageId="0" pageNumber="133">B. ventralis</emphasis>
|
||
</taxonomicName>
|
||
second flagellomere 2.5
|
||
<normalizedToken id="50164ED4E7B9B2B7BC0390AB3E130B68" originalValue="×">x</normalizedToken>
|
||
as long as pedicel), proportional average length of antennal scape, pedicel, first and second flagellomeres 3.0:1.6:1.4:1.6 (in
|
||
<taxonomicName id="1807793B4440257E18602E99676A16E4" lsidName="B. ventralis" pageId="0" pageNumber="133" rank="species" species="ventralis">
|
||
<emphasis id="34CB8063003F593D2EF40DBC19A55F1E" italics="true" pageId="0" pageNumber="133">B. ventralis</emphasis>
|
||
</taxonomicName>
|
||
3.3:1.1:2.0:2.8), and proportional average length of first to third visible labial segments 1.1:1.4:0.3 (in
|
||
<taxonomicName id="14CCD69A71155113B862EC195FD294C8" lsidName="B. ventralis" pageId="0" pageNumber="133" rank="species" species="ventralis">
|
||
<emphasis id="CDC10595487CA2FB7FD6BCA7052222F9" italics="true" pageId="0" pageNumber="133">B. ventralis</emphasis>
|
||
</taxonomicName>
|
||
1.0:1.3:0.3).
|
||
</paragraph>
|
||
<paragraph id="D0C0474626A6683D9622FE356D801B11" pageId="0" pageNumber="133">
|
||
Also,
|
||
<taxonomicName id="924B9EA1D01EA9A989129E0C4989BB75" authorityName="Ha, Truong & Ishikawa" authorityYear="2022" class="Clitellata" family="Reduviidae" genus="Biasticus" kingdom="Animalia" lsidName="Biasticus griseocapillus" order="Hemiptera" pageId="0" pageNumber="133" phylum="Annelida" rank="species" species="griseocapillus">
|
||
<emphasis id="CB98ED56B07CCA233194BF846ABBB67A" italics="true" pageId="0" pageNumber="133">Biasticus griseocapillus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is similar to
|
||
<taxonomicName id="A3E8606D82A6F968D8F91094202EDB73" lsidName="B. taynguyenensis" pageId="0" pageNumber="133" rank="species" species="taynguyenensis">
|
||
<emphasis id="99EB8C9D8098894959D495E2F6885D40" italics="true" pageId="0" pageNumber="133">B. taynguyenensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in the body color pattern, the proportion average length of the antenna segments, and the proportion average length of the visible labial segments. However,
|
||
<taxonomicName id="E185A22BB4DF530B9D87DB08C4184823" lsidName="B. griseocapillus" pageId="0" pageNumber="133" rank="species" species="griseocapillus">
|
||
<emphasis id="95E704FFF2B7FACB82D605F015499E4A" italics="true" pageId="0" pageNumber="133">B. griseocapillus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from the latter by a combination of the following characters: collar, anterior pronotal lobe, and posterior pronotal lobe covered with long bent griseous setae (in
|
||
<taxonomicName id="A49EE106F2ED481F609B9220568E040D" lsidName="B. taynguyenensis" pageId="0" pageNumber="133" rank="species" species="taynguyenensis">
|
||
<emphasis id="F49E31F483A9F8EEBB0B2FC140BAD023" italics="true" pageId="0" pageNumber="133">B. taynguyenensis</emphasis>
|
||
</taxonomicName>
|
||
short bent cream-yellow setae), and median process of pygophore (mpp) 0.2
|
||
<normalizedToken id="23A98BB5506F44222DAF493EA7B4B5F5" originalValue="×">x</normalizedToken>
|
||
as long as wide, with apical margin weakly convex and emarginate medially, and with apicolateral corner slightly produced laterad and pointed (in
|
||
<taxonomicName id="95BF6F986074D40126BEACA08384AD4B" lsidName="B. taynguyenensis" pageId="0" pageNumber="133" rank="species" species="taynguyenensis">
|
||
<emphasis id="5D536C5A291C5747CB921564A3373921" italics="true" pageId="0" pageNumber="133">B. taynguyenensis</emphasis>
|
||
</taxonomicName>
|
||
, 0.3
|
||
<normalizedToken id="F896E715896F06B3AB5AEE4F789B7020" originalValue="×">x</normalizedToken>
|
||
as long as wide, with apical margin weakly and continuously concave, and with apicolateral corner distinctly produced posterolaterad and pointed).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A9CEE5F913F67F0D731AEC8265BADED7" pageId="0" pageNumber="133" type="description">
|
||
<paragraph id="6C12FFAEAC2A706A5D4FA739D4D10939" pageId="0" pageNumber="133">Description.</paragraph>
|
||
<paragraph id="BC8CDDA0A6DBC5E2024E9C933A4B8750" pageId="0" pageNumber="133">
|
||
<emphasis id="3B8A7AC07BE761978B619EAD805A5535" bold="true" pageId="0" pageNumber="133">
|
||
Female description.
|
||
<emphasis id="939FEE03A7AE9C6A5DD1E90282D3F68B" italics="true" pageId="0" pageNumber="133">Coloration</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body shiny blackish brown. Head dorsum (except anteclypeus, maxillary plate, and gena) black; anteclypeus, labrum, maxillary plate, and anterior region of gena brown; central fascia to head venter, and posterior region of gena pale luteous; first visible labial segment brown; second and third visible labial segments blackish brown; tips of first and second visible labial segments yellowish brown; area around lateral ocellus with reddish brown suffusion; a longitudinally elongated yellowish brown spot present between lateral ocelli. Base of scape brownish black; remaining of scape and pedicel, first and second flagellomeres blackish brown. Collar, anterolateral angles, anterior pronotal lobe, and anterior acetabulum black; posterior pronotal lobe, pleura, and thoracic sterna blackish brown; scutellum black in basal half and dark brown in lateral margin and apical half; stridulatory sulcus luteous; coxae and trochanters yellowish brown; intersecting area of fore coxae and fore trochanters luteous; femora, tibiae, and tarsi blackish brown. Corium and clavus darkish brown, apically brownish yellow; membrane bronzy brown, semi-hyaline. Hind wings faintly semi-hyaline. Abdominal mediotergites I+II to V reddish brown or sanguineous, somewhat suffused with blackish brown; mediotergite VI suffused with reddish brown and irregularly suffused with sanguineous; mediotergite VII and laterotergite VII to apex of abdomen sanguineous; abdominal sternite shiny sanguineous; laterotergites II-VI luteous, segmentally suffused with dark brown spots; laterotergite VII sanguineous. External genitalia sanguineous.
|
||
</paragraph>
|
||
<paragraph id="39E23507D877DB0525AFC1692A665127" pageId="0" pageNumber="133">
|
||
<emphasis id="31A35DB76A2A497A2D12D9BFA7F0FD22" bold="true" pageId="0" pageNumber="133">
|
||
<emphasis id="D87294F70B33C81C473D16843E54B036" italics="true" pageId="0" pageNumber="133">Structure</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body medium-sized (BL = 10.61-10.84 mm), elongate, and somewhat robust. Head subelongate and robust (HL/PoW = 2.37-2.45), shorter than pronotum (HL/PnL = 0.81-0.82); postocular area of head sub-globose (PoL/PoW = 0.85-1.03), distinctly wider than anteocular area (PoW/AoW = 1.44-1.47), slightly longer than anteocular area (PoL/AoL = 1.05-1.27), constricted behind compound eyes, with a wide and deep interocular sulcus; neck short. Compound eyes protruding laterally, nearly globose, with posterior margin sub-straight; lateral ocelli produced, elevated behind interocular sulcus, widely separated from each other (OCD/PoW = 0.43-0.46); interspace between lateral ocelli wider than distance between compound eye and lateral ocellus (OCD/COD = 1.93-2.17). First visible labial segment shorter than second segment (R1L/R2L = 0.80-0.84), longer than anteocular area of head (R1L/AoL = 1.58-1.63), extending beyond level of middle of compound eye when labium laid backward; proportional average length of first to third visible labial segments 1.1:1.3:0.3. Scape ~ 1.5
|
||
<normalizedToken id="17CA0FA1C73C0BB1944EF6A64E825461" originalValue="×">x</normalizedToken>
|
||
as long as head; pedicel slightly longer than first flagellomere and nearly equal in length to second flagellomere; proportional average length of scape, pedicel, first and second flagellomeres 3.0:1.6:1.4:1.6. Collar very short in dorsal view, with anterolateral angle weakly and roundly produced; anterior pronotal lobe round and bulged, slightly rough, with middle longitudinal sulcus deep and narrow posteriorly; posterior pronotal lobe with slightly swollen anteromedial elevation; humerus roughly triangular, with round apex; posterior margin of pronotum straight; posterior angles round, not exceeded to posterior margin of pronotum. Scutellum triangular, somewhat triangularly depressed basally, apically produced and sloping downward; posterior apex round; anterior margin slightly convex. Femora thick, apically moderately nodulose; fore femora very slightly incrassated, thicker than mid and hind femora. Hemelytra surpassing apex of abdomen when fully closed, 0.8
|
||
<normalizedToken id="1B796B72AB9F41433C371BA044A6CA7A" originalValue="×">x</normalizedToken>
|
||
as long as body length; discal cell nearly parallelogram-shaped, twice as long as width; Sc 0.7
|
||
<normalizedToken id="7CBAB7B3541A7802BFBE5163186BAA1E" originalValue="×">x</normalizedToken>
|
||
as long as hemelytron length, 1.4
|
||
<normalizedToken id="AEED31A267D586D288A373F4860E0A64" originalValue="×">x</normalizedToken>
|
||
as long as R + M. Hind wing ~ 3.2
|
||
<normalizedToken id="4B2E3E77796853429E5434BEDD4A579B" originalValue="×">x</normalizedToken>
|
||
as long as maximum width. Connexivum slightly dilated and ascending with segmental incisures; abdominal laterotergite VIII (AL8) with thin posterior margin (0.01 mm); abdominal sternite VII (AS7) forming a wide subpentagonal posteromedian concavity, with inner posterolateral margin almost straight; gonocoxa VIII (Gc8) broad, ~ 1.4
|
||
<normalizedToken id="0D6AF5F082143D0DD310E67F1336450E" originalValue="×">x</normalizedToken>
|
||
wider than length, slightly opened inward for gonapophyses VIII (Gp8), with posterior margin gently slanting anteromesad, and with apical inner corner not produced; Gp8 small and subtriangular, 3.9
|
||
<normalizedToken id="4BD8E97F9541EB8BF917DFAE43D974C6" originalValue="×">x</normalizedToken>
|
||
longer than width; gonoplac (Gpl) V-shaped, thin, with maximum thick ~ 0.025 mm.
|
||
</paragraph>
|
||
<paragraph id="CE6D9E9926A7DCCE5A4D00AD9EA476A4" pageId="0" pageNumber="133">
|
||
<emphasis id="C62CC11A590D0C007D68D35CD790C00D" bold="true" pageId="0" pageNumber="133">
|
||
<emphasis id="66D63DF2C65E54ECB485C8633E50E2F3" italics="true" pageId="0" pageNumber="133">Vestiture</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body clothed with griseous setation. Head, except interocular sulcus and posterior margin of clypeus, covered with long erect setae; posterior margin of clypeus, head venter, anterolateral area of neck with short bent setae and sparsely with long erect setae; anteclypeus, labium, scape with a few short setae; pedicel, first and second flagellomeres covered with short erect to recumbent setae; dorsum of neck without setae. Collar, anterior margin of anterior pronotal lobe with dense bent pubescence; anterior pronotal lobe with some rows of long bent griseous setae; lateral area of anterior pronotal lobe with long erect setae; posterior pronotal lobe densely covered with short bent griseous setae somewhat interspersed with long griseous setae; lateral slope area and posterior apex of scutellum with dense short to long setae; posterior area of posterior pronotal lobe with long erect setae; pleura, thoracic sterna, and coxae with short bent setae; trochanters, femora, and tibiae with long erect setae; corium with bent griseous pubescence. Anterior margin of abdominal mediotergite I+II with some long slender erect yellowish brown setae; mediotergites and dorsal laterotergites sparsely covered with a few short vertical setae; abdominal sternites interspersed with long vertical setae; posterior margin of abdominal laterotergite VIII (AL8) with long erect slender setae; gonocoxa VIII (Gc8) with long slender bent setae; apex of external genitalia with long thick erect setae.
|
||
</paragraph>
|
||
<paragraph id="721D056AA11BACFA8E4200A584889DD6" pageId="0" pageNumber="133">
|
||
<emphasis id="0F39734A938A3E444B7627C963D9491D" bold="true" pageId="0" pageNumber="133">Male description.</emphasis>
|
||
General external morphology similar to that of the female.
|
||
</paragraph>
|
||
<paragraph id="732487D40582DD9BC359BCD5E52AE13A" pageId="0" pageNumber="133">
|
||
<emphasis id="1DA23ED86A55A49CEA7DBB974ECBCCC3" bold="true" pageId="0" pageNumber="133">
|
||
<emphasis id="89DF5CDADBB0DCA6BDF93CFE32004665" italics="true" pageId="0" pageNumber="133">Coloration</emphasis>
|
||
.
|
||
</emphasis>
|
||
Almost similar to female but brighter than female. Gena brown in anterior half and luteous in posterior half; labium blackish brown; tips of first and second visible labial segments luteous. Collar, anterolateral angles, anterior pronotal lobe, and anterior acetabulum blackish brown; posterior pronotal lobe, pleura, and thoracic sterna brown; scutellum black in basal half and brown in lateral margin and apical half; stridulatory sulcus pale gray. Laterotergites II-VI pale luteous, segmentally suffused with blackish brown spots; laterotergite VII sanguineous. Pygophore ventrally orpiment-orange; paramere semi-hyaline, pale orange.
|
||
</paragraph>
|
||
<paragraph id="80B14897479F6FBD576F86C3E5A39EF8" pageId="0" pageNumber="133">
|
||
<emphasis id="31C44D5D60EBE4BF19DEED06F8DB200B" bold="true" pageId="0" pageNumber="133">
|
||
<emphasis id="D835F104529684613A64DAC5CE2133B1" italics="true" pageId="0" pageNumber="133">Structure</emphasis>
|
||
.
|
||
</emphasis>
|
||
Almost similar to female except for the following characters. Scape 1.8
|
||
<normalizedToken id="20E400652C6FC4258E844DC8AB1C3B66" originalValue="×">x</normalizedToken>
|
||
as long as head and more than twice as long as pedicel; pedicel slightly longer than first flagellomere; second flagellomere missing; proportional length of scape, pedicel, and first flagellomere 3.5:1.6:1.4. Hemelytra surpassing apex of abdomen when fully closed, 0.9
|
||
<normalizedToken id="B6D4473694D76121B9BF7E1D9A2FBAA4" originalValue="×">x</normalizedToken>
|
||
as long as body; Sc 0.8
|
||
<normalizedToken id="03B31F8C1971384E03B5D306D37EA90A" originalValue="×">x</normalizedToken>
|
||
as long as hemelytron, and 1.5
|
||
<normalizedToken id="CB4A52E27D284174F832D69C04CD81EA" originalValue="×">x</normalizedToken>
|
||
as long as R + M. Pygophore ovoid; median process of pygophore (mpp) broad and low, 0.2
|
||
<normalizedToken id="EC3129E73A59D482B81D1E61CC1684CA" originalValue="×">x</normalizedToken>
|
||
as long as wide, with apical margin weakly convex and emarginate medially, with apicolateral corner slightly produced laterad and pointed; paramere long, slender, clavate, somewhat curved medially, and apically subnodulose and round (Figs
|
||
<figureCitation id="F676DE86F030A4E0635C50439C109A76" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia of four morphospecies A-D type mA, Biasticus taynguyenensis Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 545 E-I type mB, B. griseocapillus Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 546 J-N type mC, B. luteicollis Ha, Truong & Ishikawa, sp. nov., holotype, ♂, HNL 2018 - 025 O-R mF, B. flavus (Distant, 1903), ♂, HEM-TH 2004 - 018. A, E, J, O pygophore with paramere (s) in dorsal view B, F, K, P phallus in dorsal view C, G, L, Q phallus in lateral view D, I, N, R articulatory apparatus H, M distal dorsal lobe of endosoma." figureDoi="10.3897/zookeys.1118.83156.figure6" httpUri="https://binary.pensoft.net/fig/734749" pageId="0" pageNumber="133">6E</figureCitation>
|
||
,
|
||
<figureCitation id="24744476BE264269D38028CD86B2E4AE" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Biasticus griseocapillus Ha, Truong & Ishikawa, sp. nov. A female genitalia in ventral view, holotype, ♀, HNL 2018 - 038 B-M male genitalia, paratype, ♂, TXL 2016 - 546 B-F pygophore with paramere B dorsal view C lateral view D ventral view E apical portion of pygophore, showing median process (mpp) and paramere F left paramere G-M phallus G dorsal view H lateral view I articulatory apparatus (aa) J aedeagus with endosoma semi-everted, dorsal view K aedeagus with endosoma semi-everted, ventral view L aedeagus with endosoma semi-everted, lateral view M distal dorsal lobe of endosoma (ddl)." figureDoi="10.3897/zookeys.1118.83156.figure18" httpUri="https://binary.pensoft.net/fig/734761" pageId="0" pageNumber="133">18B-F</figureCitation>
|
||
). Aedeagus in dorsal view ovoid, dorsally sclerotized (Figs
|
||
<figureCitation id="024E6CD9B672FA8C25FC837B61501896" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia of four morphospecies A-D type mA, Biasticus taynguyenensis Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 545 E-I type mB, B. griseocapillus Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 546 J-N type mC, B. luteicollis Ha, Truong & Ishikawa, sp. nov., holotype, ♂, HNL 2018 - 025 O-R mF, B. flavus (Distant, 1903), ♂, HEM-TH 2004 - 018. A, E, J, O pygophore with paramere (s) in dorsal view B, F, K, P phallus in dorsal view C, G, L, Q phallus in lateral view D, I, N, R articulatory apparatus H, M distal dorsal lobe of endosoma." figureDoi="10.3897/zookeys.1118.83156.figure6" httpUri="https://binary.pensoft.net/fig/734749" pageId="0" pageNumber="133">6F</figureCitation>
|
||
,
|
||
<figureCitation id="2130C2251F343955E39EBCCFEDF06AEE" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Biasticus griseocapillus Ha, Truong & Ishikawa, sp. nov. A female genitalia in ventral view, holotype, ♀, HNL 2018 - 038 B-M male genitalia, paratype, ♂, TXL 2016 - 546 B-F pygophore with paramere B dorsal view C lateral view D ventral view E apical portion of pygophore, showing median process (mpp) and paramere F left paramere G-M phallus G dorsal view H lateral view I articulatory apparatus (aa) J aedeagus with endosoma semi-everted, dorsal view K aedeagus with endosoma semi-everted, ventral view L aedeagus with endosoma semi-everted, lateral view M distal dorsal lobe of endosoma (ddl)." figureDoi="10.3897/zookeys.1118.83156.figure18" httpUri="https://binary.pensoft.net/fig/734761" pageId="0" pageNumber="133">18G, J</figureCitation>
|
||
) and in lateral view long and narrow (Figs
|
||
<figureCitation id="9B116B67221CE6CD1886814D1D0C3411" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia of four morphospecies A-D type mA, Biasticus taynguyenensis Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 545 E-I type mB, B. griseocapillus Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 546 J-N type mC, B. luteicollis Ha, Truong & Ishikawa, sp. nov., holotype, ♂, HNL 2018 - 025 O-R mF, B. flavus (Distant, 1903), ♂, HEM-TH 2004 - 018. A, E, J, O pygophore with paramere (s) in dorsal view B, F, K, P phallus in dorsal view C, G, L, Q phallus in lateral view D, I, N, R articulatory apparatus H, M distal dorsal lobe of endosoma." figureDoi="10.3897/zookeys.1118.83156.figure6" httpUri="https://binary.pensoft.net/fig/734749" pageId="0" pageNumber="133">6G</figureCitation>
|
||
,
|
||
<figureCitation id="A15B27CF14B72C036D02A380E530D7A4" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Biasticus griseocapillus Ha, Truong & Ishikawa, sp. nov. A female genitalia in ventral view, holotype, ♀, HNL 2018 - 038 B-M male genitalia, paratype, ♂, TXL 2016 - 546 B-F pygophore with paramere B dorsal view C lateral view D ventral view E apical portion of pygophore, showing median process (mpp) and paramere F left paramere G-M phallus G dorsal view H lateral view I articulatory apparatus (aa) J aedeagus with endosoma semi-everted, dorsal view K aedeagus with endosoma semi-everted, ventral view L aedeagus with endosoma semi-everted, lateral view M distal dorsal lobe of endosoma (ddl)." figureDoi="10.3897/zookeys.1118.83156.figure18" httpUri="https://binary.pensoft.net/fig/734761" pageId="0" pageNumber="133">18H, L</figureCitation>
|
||
); articulatory apparatus (aa) in ventral view with basal plate arms relatively slender and jointly forming a U-shape, and in lateral view arched strongly (Figs
|
||
<figureCitation id="81DF18298AF7F6EEAB8A89F6C7752961" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia of four morphospecies A-D type mA, Biasticus taynguyenensis Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 545 E-I type mB, B. griseocapillus Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 546 J-N type mC, B. luteicollis Ha, Truong & Ishikawa, sp. nov., holotype, ♂, HNL 2018 - 025 O-R mF, B. flavus (Distant, 1903), ♂, HEM-TH 2004 - 018. A, E, J, O pygophore with paramere (s) in dorsal view B, F, K, P phallus in dorsal view C, G, L, Q phallus in lateral view D, I, N, R articulatory apparatus H, M distal dorsal lobe of endosoma." figureDoi="10.3897/zookeys.1118.83156.figure6" httpUri="https://binary.pensoft.net/fig/734749" pageId="0" pageNumber="133">6G, I</figureCitation>
|
||
,
|
||
<figureCitation id="807BB3015E9EBCAC30F23C2EFB2AF5AF" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Biasticus griseocapillus Ha, Truong & Ishikawa, sp. nov. A female genitalia in ventral view, holotype, ♀, HNL 2018 - 038 B-M male genitalia, paratype, ♂, TXL 2016 - 546 B-F pygophore with paramere B dorsal view C lateral view D ventral view E apical portion of pygophore, showing median process (mpp) and paramere F left paramere G-M phallus G dorsal view H lateral view I articulatory apparatus (aa) J aedeagus with endosoma semi-everted, dorsal view K aedeagus with endosoma semi-everted, ventral view L aedeagus with endosoma semi-everted, lateral view M distal dorsal lobe of endosoma (ddl)." figureDoi="10.3897/zookeys.1118.83156.figure18" httpUri="https://binary.pensoft.net/fig/734761" pageId="0" pageNumber="133">18H, I</figureCitation>
|
||
); dorsal phallothecal sclerite (dps) in lateral view with posteromedian part weakly produced posterodorsad (Figs
|
||
<figureCitation id="6B5A3677CF569A037F544F818807B8E8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia of four morphospecies A-D type mA, Biasticus taynguyenensis Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 545 E-I type mB, B. griseocapillus Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 546 J-N type mC, B. luteicollis Ha, Truong & Ishikawa, sp. nov., holotype, ♂, HNL 2018 - 025 O-R mF, B. flavus (Distant, 1903), ♂, HEM-TH 2004 - 018. A, E, J, O pygophore with paramere (s) in dorsal view B, F, K, P phallus in dorsal view C, G, L, Q phallus in lateral view D, I, N, R articulatory apparatus H, M distal dorsal lobe of endosoma." figureDoi="10.3897/zookeys.1118.83156.figure6" httpUri="https://binary.pensoft.net/fig/734749" pageId="0" pageNumber="133">6G</figureCitation>
|
||
,
|
||
<figureCitation id="FECA8904C0925A2A0F4837804BB63C99" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Biasticus griseocapillus Ha, Truong & Ishikawa, sp. nov. A female genitalia in ventral view, holotype, ♀, HNL 2018 - 038 B-M male genitalia, paratype, ♂, TXL 2016 - 546 B-F pygophore with paramere B dorsal view C lateral view D ventral view E apical portion of pygophore, showing median process (mpp) and paramere F left paramere G-M phallus G dorsal view H lateral view I articulatory apparatus (aa) J aedeagus with endosoma semi-everted, dorsal view K aedeagus with endosoma semi-everted, ventral view L aedeagus with endosoma semi-everted, lateral view M distal dorsal lobe of endosoma (ddl)." figureDoi="10.3897/zookeys.1118.83156.figure18" httpUri="https://binary.pensoft.net/fig/734761" pageId="0" pageNumber="133">18L</figureCitation>
|
||
); spoon-like sclerites (sps) hyaline and glabrous; both membranous sac-like lobes posterolaterally produced; distal dorsal lobe of endosoma (ddl) round, with membranous surface covered with large hyaline prickles (Figs
|
||
<figureCitation id="04F4A0AEE3A31C2C865BC75504B0C615" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Male genitalia of four morphospecies A-D type mA, Biasticus taynguyenensis Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 545 E-I type mB, B. griseocapillus Ha, Truong & Ishikawa, sp. nov., paratype, ♂, TXL 2016 - 546 J-N type mC, B. luteicollis Ha, Truong & Ishikawa, sp. nov., holotype, ♂, HNL 2018 - 025 O-R mF, B. flavus (Distant, 1903), ♂, HEM-TH 2004 - 018. A, E, J, O pygophore with paramere (s) in dorsal view B, F, K, P phallus in dorsal view C, G, L, Q phallus in lateral view D, I, N, R articulatory apparatus H, M distal dorsal lobe of endosoma." figureDoi="10.3897/zookeys.1118.83156.figure6" httpUri="https://binary.pensoft.net/fig/734749" pageId="0" pageNumber="133">6H</figureCitation>
|
||
,
|
||
<figureCitation id="9D0536744D814781ACF704180193BE1B" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Biasticus griseocapillus Ha, Truong & Ishikawa, sp. nov. A female genitalia in ventral view, holotype, ♀, HNL 2018 - 038 B-M male genitalia, paratype, ♂, TXL 2016 - 546 B-F pygophore with paramere B dorsal view C lateral view D ventral view E apical portion of pygophore, showing median process (mpp) and paramere F left paramere G-M phallus G dorsal view H lateral view I articulatory apparatus (aa) J aedeagus with endosoma semi-everted, dorsal view K aedeagus with endosoma semi-everted, ventral view L aedeagus with endosoma semi-everted, lateral view M distal dorsal lobe of endosoma (ddl)." figureDoi="10.3897/zookeys.1118.83156.figure18" httpUri="https://binary.pensoft.net/fig/734761" pageId="0" pageNumber="133">18M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="3C82C1F0913740A0D91AC22D01873D85" pageId="0" pageNumber="133">
|
||
<emphasis id="CCF17771F904D6AB0687EA1B274EB71E" bold="true" pageId="0" pageNumber="133">
|
||
<emphasis id="BEF5490DE187B37C8069AB6F9E180B2B" italics="true" pageId="0" pageNumber="133">Vestiture</emphasis>
|
||
.
|
||
</emphasis>
|
||
Almost similar to female except for the following characters. Posterior margin of clypeus with short bent pubescence; anteclypeus and scape without setae. Collar and anterior margin of anterior pronotal lobe with short bent setae and some long bent setae; anterior pronotal lobe with some rows of long bent griseous setae; posterior pronotal lobe with bent griseous setae; scutellum with long slender erect setae abundantly, especially in lateral slopes; posterior apex of scutellum with a pinch of long slender erect and bent setae; pleura, thoracic sterna, and coxae with short bent setae, with a few long slender setae; trochanters sparsely with short bent pubescence and a few long setae; corium with bent pubescence. Pygophore ventrally covered with a few bent pubescence, more abundant laterally; paramere with a few long erect thick setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0B93B9798B9B2FAAD37E569CB437AD45" pageId="0" pageNumber="133" type="description">
|
||
<paragraph id="E34CA590F2ACCC57BB4310ECBD45297D" pageId="0" pageNumber="133">Measurements.</paragraph>
|
||
<paragraph id="3A1D07EA56B73CDADFC417ED026841FF" pageId="0" pageNumber="133">
|
||
All dimensions are given in mm.
|
||
<emphasis id="76BD1FEC32345A9684FED3FD64B512A2" bold="true" italics="true" pageId="0" pageNumber="133">Holotype</emphasis>
|
||
(♀): BL 10.84; HL 1.99; AoL 0.67; AoW 0.56; PoL 0.85; PoW 0.82; OE 1.05; IE 0.56; ED 0.60; OD 0.17; OCD 0.36; COD 0.18; R1L 1.09; R2L 1.31; R3L 0.32; A1L 3.01; A2L 1.59; A3L 1.52; A4L 1.56; PnL 2.45; PnW 2.74; APL 0.79; PPL 1.67; HeL 8.56; HeW 2.67; Sc 6.69; R+M 4.67; HWL 5.68; HWW 1.68; AFL 3.37; ATL 4.00; MFL 2.77; MTL 3.47; PFL 3.94; PTL 5.44.
|
||
<emphasis id="1F109DD73B5074876100250A20E31CA5" bold="true" italics="true" pageId="0" pageNumber="133">Paratype</emphasis>
|
||
(♂): BL 9.77; HL 1.91; AoL 0.64; AoW 0.54; PoL 0.78; PoW 0.79; OE 1.05; IE 0.53; ED 0.63; OD 0.14; OCD 0.38; COD 0.19; R1L 1.05; R2L 1.31; R3L 0.29; A1L 3.48; A2L 1.62; A3L 1.49; A4L n/a; PnL 2.12; PnW 2.57; APL 0.59; PPL 1.52; HeL 8.45; HeW 2.79; Sc 6.78; R+M 4.52; AFL 3.44; ATL 4.06; MFL 2.90; MTL 3.52; PFL 3.99; PTL 5.36.
|
||
<emphasis id="482963B9D5538242B0DDAA75BAB389AF" bold="true" italics="true" pageId="0" pageNumber="133">Paratypes</emphasis>
|
||
(♀). BL 10.61-10.75; HL 1.94-1.95; AoL 0.67; AoW 0.54-0.57; PoL 0.70-0.76; PoW 0.79-0.82; OE 1.05; IE 0.54-0.55; ED 0.58-0.62; OD 0.16; OCD 0.36-0.38; R1L 1.05-1.06; R2L 1.27-1.31; R3L 0.31-0.34; A1L 2.92-2.97; A2L 1.57-1.61; A3L 1.36-1.43; A4L 1.60-1.64; PnL 2.36-2.38; PnW 2.58-2.74; APL 0.72-0.84; PPL 1.52-1.66; HeL 8.65-8.88; HeW 2.75-2.98; Sc 6.30-6.49; R+M 4.40-4.48; HWL 5.78-5.89; HWW 1.79-1.91; AFL 3.31-3.33; ATL 3.78-3.93; MFL 2.70-2.78; MTL 3.42-3.51; PFL 3.69-3.87; PTL 4.91-5.35.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1CF644FCF84FD4819F273E3CBA85540" pageId="0" pageNumber="133" type="distribution">
|
||
<paragraph id="BBF827D0FA95420E1AC4CAFAF8B96203" pageId="0" pageNumber="133">Distribution.</paragraph>
|
||
<paragraph id="F29DEF67B7449DEDBF6F9AF0FDA10771" pageId="0" pageNumber="133">Vietnam, Central Highlands (Gia Lai, Dak Lak).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E52DCD4DCECE1AB4783876353F57803A" pageId="0" pageNumber="133" type="type locality">
|
||
<paragraph id="0534D0C9A58AEF8155D4B24503391734" pageId="0" pageNumber="133">Type locality.</paragraph>
|
||
<paragraph id="F95DAAE3D92B3D2D0FCD83A4D49BAB07" pageId="0" pageNumber="133">Vietnam, Central Highlands, Dak Lak Province, Chu Yang Sin National Park.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="34913037A0B2843D54E9640D95AAA4A1" pageId="0" pageNumber="133" type="etymology">
|
||
<paragraph id="1DD7AA9A96B0F9E590CADFB893D70D23" pageId="0" pageNumber="133">Etymology.</paragraph>
|
||
<paragraph id="3CA5C3A24FA041F8CD22E0D60553943A" pageId="0" pageNumber="133">This new species is named after griseous pubescence on the pronotum.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |