292 lines
27 KiB
XML
292 lines
27 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.594.7539" ID-GBIF-Dataset="1c3bcc53-d564-4464-95ed-dc7b7f04c61a" ID-PMC="PMC4926698" ID-Pensoft-Pub="1313-2970-594-11" ID-PubMed="27408557" ID-ZBK="FDFE14E46C7C4E7DBA415DDBD8F62E2A" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-594-11" ModsDocOrigin="ZooKeys 594" ModsDocTitle="A new family Lepidocharontidae with description of Lepidocharon gen. n., from the Great Barrier Reef, Australia, and redefinition of the Microparasellidae (Isopoda, Asellota)" checkinTime="1464725666723" checkinUser="pensoft" docAuthor="Galassi, Diana M. P., Bruce, Niel L., Fiasca, Barbara & Dole-Olivier, Marie-Jose" docDate="2016" docId="4D20587EC58B57E99665B5F7E0F15B77" docLanguage="en" docName="ZooKeys 594: 11-50" docOrigin="ZooKeys 594" docSource="http://dx.doi.org/10.3897/zookeys.594.7539" docTitle="Lepidocharontidae Galassi & Bruce, fam. n." docType="treatment" docUuid="B21B701E-73AE-4433-84C3-1EA34618814B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="15" masterDocId="FFE1FFBDFFB7FFF0277A375A6652FFF7" masterDocTitle="A new family Lepidocharontidae with description of Lepidocharon gen. n., from the Great Barrier Reef, Australia, and redefinition of the Microparasellidae (Isopoda, Asellota)" masterLastPageNumber="50" masterPageNumber="11" pageNumber="12" updateTime="1668163155426" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new family Lepidocharontidae with description of Lepidocharon gen. n., from the Great Barrier Reef, Australia, and redefinition of the Microparasellidae (Isopoda, Asellota)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Galassi, Diana M. P.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bruce, Niel L.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fiasca, Barbara</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dole-Olivier, Marie-Jose</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="volume">
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<mods:number>594</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>11</mods:start>
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<mods:end>50</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.594.7539</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.594.7539</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-594-11</mods:identifier>
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<mods:identifier type="ZBK">FDFE14E46C7C4E7DBA415DDBD8F62E2A</mods:identifier>
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<mods:identifier type="ZooBank">FDFE14E46C7C4E7DBA415DDBD8F62E2A</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127903061" LSID="urn:lsid:zoobank.org:act:B21B701E-73AE-4433-84C3-1EA34618814B" httpUri="http://treatment.plazi.org/id/4D20587EC58B57E99665B5F7E0F15B77" lastPageId="4" lastPageNumber="15" pageId="1" pageNumber="12">
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<subSubSection pageId="1" pageNumber="12" type="multiple">
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<paragraph pageId="1" pageNumber="12">Taxon classification Animalia Isopoda Lepidocharontidae</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="12" type="nomenclature">
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<paragraph pageId="1" pageNumber="12">
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<taxonomicName LSID="http://zoobank.org/B21B701E-73AE-4433-84C3-1EA34618814B" authority="Galassi & Bruce" family="Lepidocharontidae" lsidName="" pageId="1" pageNumber="12" rank="family">Lepidocharontidae Galassi & Bruce</taxonomicName>
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<taxonomicNameLabel pageId="1" pageNumber="12">fam. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="13" pageId="1" pageNumber="12" type="diagnosis">
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<paragraph pageId="1" pageNumber="12">Diagnosis.</paragraph>
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<paragraph lastPageId="2" lastPageNumber="13" pageId="1" pageNumber="12">
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Male. Body dorsally flat, slender, ~4
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<normalizedToken originalValue="–10×">-10x</normalizedToken>
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long as wide, without chromatophores; somites all subsimilar in width, somites sub-rectangular or trapezoidal, lateral margins of head and pereionites sub-parallel. Pleon of one segment, with free lateral margins. Head with weak or absent rostrum, without pseudorostrum. Eyes absent. Antennula with maximally 4 flagellar articles. Antenna flagellum longer than podomeres. Antennal scale (rudimentary exopod) present, even if more or less developed among genera. Mandible incisor with 2 to 8 cusps; molar process subconical, without grinding surface, with apical unequal smooth and pinnate setae; spine row and lacinia mobilis
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<pageBreakToken pageId="2" pageNumber="13" start="start">present</pageBreakToken>
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, the latter only on left mandible. Maxilliped slender, covering entire mouthpart field, endite distal margin narrowly rounded; epipod slender, quadrate or distally acute; palp composed of 5 articles; 2 stiff pectinate setae always present on maxilliped distal article. Pereiopods 1-7 subsimilar, always without subchela; all pereiopods with 2 dactylar claws; pereiopods articulating dorso-laterally or laterally, and projecting ventrally (in
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<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Janinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Janinella" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Janinella</taxonomicName>
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and
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<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
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) or outwards (in
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<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Lepidocharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lepidocharon" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Lepidocharon</taxonomicName>
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). Penial processes with openings coalescent and medial. Male pleopods 1 and 2 not operculate; male pleopod 1 distally rounded or subtruncate, with or without acute distolateral lobes; proximal part of the pleopod with or without scale-like elements on postero-lateral margins; stylet-guiding grooves running parallel to the lateral free distal margin of pleopod 1 and folded by a hyaline lamella (transversal and unfolded in
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<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Janinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Janinella" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Janinella</taxonomicName>
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); pleopod 3 endopod with 3 plumose setae (marine taxa) or without plumose setae (freshwater taxa), exopod slender. Pleopod 4 globular, pleopod 5 absent. Uropods biramous, ventrally inserted on pleotelson, protopod large, c. 0.5-1.3 as long as pleotelson; protopod length/width ratio c. 2.5-4.5; rami slender with exopod articulating anteriorly to endopod. Anus terminal, not covered by pleopods. Anus outside pleopodal chamber, between bases of uropodal protopods.
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</paragraph>
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<paragraph pageId="2" pageNumber="13">Female. Operculum (pleopod 2) from sub-quadrate (as long as wide) to more than 2 times longer than wide, with free distal margin deeply incised medially, faintly incised, or without medial incision, armed with 4 or 2 setae, or unarmed.</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="13" type="genera included">
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<paragraph pageId="2" pageNumber="13">Genera included.</paragraph>
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<paragraph pageId="2" pageNumber="13">
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<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
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Karaman, 1934;
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<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Janinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Janinella" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Janinella</taxonomicName>
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Albuquerque, Boulanouar & Coineau, 2014;
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<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Lepidocharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lepidocharon" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Lepidocharon</taxonomicName>
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Galassi & Bruce, gen. n.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="13" type="genus incertae sedis">
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<paragraph pageId="2" pageNumber="13">Genus incertae sedis.</paragraph>
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<paragraph pageId="2" pageNumber="13">
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<taxonomicName class="Malacostraca" genus="Angeliera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angeliera" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Angeliera</taxonomicName>
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Chappuis & Delamare Deboutteville, 1952.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="15" pageId="2" pageNumber="13" type="remarks">
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<paragraph pageId="2" pageNumber="13">Remarks.</paragraph>
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<paragraph pageId="2" pageNumber="13">
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<bibRefCitation author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="683 - 747" title="A systematic review of the family Janiridae (Crustacea: Isopoda: Asellota)." url="10.1071/IT9940683" volume="8" year="1994">
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Wilson and
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<normalizedToken originalValue="Wägele">Waegele</normalizedToken>
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(1994)
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</bibRefCitation>
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and
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<bibRefCitation author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="749 - 766" title="A phylogenetic analysis of the isopod family Janiridae (Crustacea)." url="10.1071/IT9940749" volume="8" year="1994">Wilson (1994)</bibRefCitation>
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critically discussed the status of the
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<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
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in relation to the
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<taxonomicName family="Janiridae" lsidName="" pageId="2" pageNumber="13" rank="family">Janiridae</taxonomicName>
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on a cladistic phylogenetic basis.
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<bibRefCitation author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="683 - 747" title="A systematic review of the family Janiridae (Crustacea: Isopoda: Asellota)." url="10.1071/IT9940683" volume="8" year="1994">
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Wilson and
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<normalizedToken originalValue="Wägele">Waegele</normalizedToken>
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(1994)
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</bibRefCitation>
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in their review of the
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<taxonomicName family="Janiridae" lsidName="" pageId="2" pageNumber="13" rank="family">Janiridae</taxonomicName>
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analysed the status of the family
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<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
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in detail, reviewing the history of the debate over the status of the family.
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<bibRefCitation author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="749 - 766" title="A phylogenetic analysis of the isopod family Janiridae (Crustacea)." url="10.1071/IT9940749" volume="8" year="1994">Wilson (1994)</bibRefCitation>
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also included the
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<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
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in his cladistic analysis of the phylogeny of the
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<taxonomicName family="Janiridae" lsidName="" pageId="2" pageNumber="13" rank="family">Janiridae</taxonomicName>
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.
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<bibRefCitation author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="683 - 747" title="A systematic review of the family Janiridae (Crustacea: Isopoda: Asellota)." url="10.1071/IT9940683" volume="8" year="1994">
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Wilson and
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<normalizedToken originalValue="Wägele">Waegele</normalizedToken>
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(1994
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</bibRefCitation>
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: page 721) stated "The family concept of the
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<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
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may be open to challenge because
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<taxonomicName class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
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is distinct from the other three genera in these autapomorphies", these being the differences in the somatic and uropodal morphology. These authors went on to say "the composition of this family will require further study." In
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<bibRefCitation author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="749 - 766" title="A phylogenetic analysis of the isopod family Janiridae (Crustacea)." url="10.1071/IT9940749" volume="8" year="1994">
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<normalizedToken originalValue="Wilson’s">Wilson's</normalizedToken>
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(1994)
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</bibRefCitation>
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analysis, there were no supporting apomorphies for the
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<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
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as then constituted, but there were separate supporting apomorphies for the genus
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<taxonomicName class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
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and the clade holding the remaining genera, strongly suggesting that potentially these were two monophyletic clades, albeit the
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<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
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being monogeneric. The principle basis for this is that each group had unique and derived uropod morphology and substantial differences in body morphology.
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</paragraph>
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<paragraph lastPageId="3" lastPageNumber="14" pageId="2" pageNumber="13">
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The description of the new genus
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<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Lepidocharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lepidocharon" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Lepidocharon</taxonomicName>
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Galassi & Bruce, gen. n. led to a re-appraisal of the taxonomic status of the
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<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
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and its constituent genera. We conclude that the
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<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
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is a mono-generic family supported by a prominent acute or narrowly rounded rostrum, the antennal flagellum shorter than podomeres,
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<pageBreakToken pageId="3" pageNumber="14" start="start">all</pageBreakToken>
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somites with straight lateral margins that also have scales, an indisputable ventral position of the pereiopods, the unique uniramous and short uropods (see Appendix 1). The remaining genera are housed in the new family
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<taxonomicName family="Lepidocharontidae" lsidName="" pageId="3" pageNumber="14" rank="family">Lepidocharontidae</taxonomicName>
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fam. n., the diagnostic characters being the elongate body (up to 10
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as maximum width), a weak or absent rostrum, the antennal flagellum longer than podomeres, the pereionites rectangular or trapezoidal in dorsal view, with sub-parallel lateral margins, a lateral or dorso-lateral position of the pereiopods, a tendency to reduction of pereiopodal coxal plates, and the uropod with a large protopod with the exopod articulating anteriorly and separately to the endopod.
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</paragraph>
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<paragraph pageId="3" pageNumber="14">
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Within the family
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<taxonomicName family="Lepidocharontidae" lsidName="" pageId="3" pageNumber="14" rank="family">Lepidocharontidae</taxonomicName>
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there is great uniformity of the diagnostic characters among all the genera. The genus
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<taxonomicName class="Malacostraca" genus="Angeliera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angeliera" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Angeliera</taxonomicName>
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has been placed incertae sedis in the
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<taxonomicName lsidName="Asellota" pageId="3" pageNumber="14" rank="suborder" suborder="Asellota">Asellota</taxonomicName>
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on the basis of marked differences in several morphological features that set this genus far away the basic body plan observed in
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<taxonomicName family="Lepidocharontidae" lsidName="" pageId="3" pageNumber="14" rank="family">Lepidocharontidae</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="3" pageNumber="14">
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The largest genus in the
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<taxonomicName family="Lepidocharontidae" lsidName="" pageId="3" pageNumber="14" rank="family">Lepidocharontidae</taxonomicName>
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is
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<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
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with 77 species, both marine and freshwater. Many species lack full descriptions, and there are inconsistencies in the distribution of certain characters within the genus. A dorsal view of the head is not routinely figured; when figured, it can be seen that some species do have a rostral point or rostrum, while others have the anterior margin of the head weakly concave.
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</paragraph>
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<paragraph pageId="3" pageNumber="14">
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The generic name
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<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
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is unavailable under the
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<bibRefCitation author="Harrison, K" journalOrPublisher="The International Trust for Zoological Nomenclature, London, UK" pageId="22" pageNumber="33" title="International Code of Zoological Nomenclature" year="1999">ICZN (1999)</bibRefCitation>
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's rules because the genus was established by
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<bibRefCitation author="Karaman, S" journalOrPublisher="Mitteilungen ueber Hohlen- und Karstforschung" pageId="23" pageNumber="34" pagination="42 - 44" title="Beitrage zur Kenntnis des Isopoden-Familie Microparasellidae." volume="1934" year="1934">Karaman (1934)</bibRefCitation>
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without type species designation. This prevents the use of
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<normalizedToken originalValue="“Microcharontidae”">"Microcharontidae"</normalizedToken>
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(from the most speciose and well-known genus
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<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
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) as the family name because
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<bibRefCitation author="Harrison, K" journalOrPublisher="The International Trust for Zoological Nomenclature, London, UK" pageId="22" pageNumber="33" title="International Code of Zoological Nomenclature" year="1999">ICZN (1999)</bibRefCitation>
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' article 13.2 unambiguously states: "To be available, every new family-group name published after 1930 must satisfy the provisions of Article 13.1 and must be formed from an available genus-group name then used as valid by the author in the family-group taxon [Arts. 11.7.1.1, 29]". Therefore we here propose the name
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<taxonomicName family="Lepidocharontidae" lsidName="" pageId="3" pageNumber="14" rank="family">Lepidocharontidae</taxonomicName>
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fam. n.
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</paragraph>
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<paragraph pageId="3" pageNumber="14">
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The family name
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<taxonomicName family="Microparasellidae" lsidName="" pageId="3" pageNumber="14" rank="family">Microparasellidae</taxonomicName>
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was first introduced by
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<bibRefCitation author="Karaman, S" journalOrPublisher="Mitteilungen ueber Hohlen- und Karstforschung" pageId="23" pageNumber="34" pagination="42 - 44" title="Beitrage zur Kenntnis des Isopoden-Familie Microparasellidae." volume="1934" year="1934">Karaman (1934</bibRefCitation>
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: page 44) when describing the genus
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<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
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, although the family had been earlier diagnosed by
|
||
<bibRefCitation author="Karaman, S" journalOrPublisher="Zoologischer Anzeiger" pageId="23" pageNumber="34" pagination="16 - 22" title="Neue Isopoden aus unterirdischen Gewaessern Jugoslawiens." volume="102" year="1933">Karaman (1933</bibRefCitation>
|
||
: page 17) with the accompanying statement "
|
||
<taxonomicName class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
|
||
n. fam., n. gen.", but without type-species designation for the genus
|
||
<taxonomicName class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
|
||
(see
|
||
<bibRefCitation author="Karaman, S" journalOrPublisher="Zoologischer Anzeiger" pageId="23" pageNumber="34" pagination="16 - 22" title="Neue Isopoden aus unterirdischen Gewaessern Jugoslawiens." volume="102" year="1933">Karaman 1933</bibRefCitation>
|
||
). According to the
|
||
<bibRefCitation author="Harrison, K" journalOrPublisher="The International Trust for Zoological Nomenclature, London, UK" pageId="22" pageNumber="33" title="International Code of Zoological Nomenclature" year="1999">ICZN (1999</bibRefCitation>
|
||
: Article 13.2) the family name
|
||
<taxonomicName family="Microparasellidae" lsidName="" pageId="3" pageNumber="14" rank="family">Microparasellidae</taxonomicName>
|
||
proposed by
|
||
<bibRefCitation author="Karaman, S" journalOrPublisher="Mitteilungen ueber Hohlen- und Karstforschung" pageId="23" pageNumber="34" pagination="42 - 44" title="Beitrage zur Kenntnis des Isopoden-Familie Microparasellidae." volume="1934" year="1934">Karaman (1934)</bibRefCitation>
|
||
is thus a nomen nudum, because the family was erected on the unavailable generic name
|
||
<taxonomicName class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
|
||
Karaman, 1933 that lacked type-species designation. Nevertheless, Article 13.2.1 states that "A family group name first published after 1930 and before 1961 which does not satisfy the provisions of Article 13.1 is available from its original publication only if it was used as valid before 2000, and also was not rejected by an author who, after 1960 and before 2000, expressly applied Article 13 of the then current editions of the Code". The family name was considered valid until 2000, and for this reason it is an available name as
|
||
<taxonomicName family="Microparasellidae" lsidName="" pageId="3" pageNumber="14" rank="family">Microparasellidae</taxonomicName>
|
||
Karaman, 1934. The family name
|
||
<taxonomicName family="Microparasellidae" lsidName="" pageId="3" pageNumber="14" rank="family">Microparasellidae</taxonomicName>
|
||
is then valid.
|
||
</paragraph>
|
||
<paragraph lastPageId="4" lastPageNumber="15" pageId="3" pageNumber="14">
|
||
Conversely, we provisionally maintain current and common usage of the names
|
||
<taxonomicName class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
|
||
and
|
||
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
|
||
, and this is discussed in more detail together with a new diagnosis for the
|
||
<taxonomicName family="Microparasellidae" lsidName="" pageId="3" pageNumber="14" rank="family">Microparasellidae</taxonomicName>
|
||
(see Appendix 1). As the nomenclature within
|
||
<pageBreakToken pageId="4" pageNumber="15" start="start">the</pageBreakToken>
|
||
family
|
||
<taxonomicName family="Microparasellidae" lsidName="" pageId="4" pageNumber="15" rank="family">Microparasellidae</taxonomicName>
|
||
is well established and widely used, a proposition (Galassi and Bruce in preparation) will be submitted to the ICZN Commission for maintaining the stability of the current nomenclature and related authorities.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="15" type="key">
|
||
<paragraph pageId="4" pageNumber="15">
|
||
Key to
|
||
<taxonomicName family="Microparasellidae" lsidName="" pageId="4" pageNumber="15" rank="family">Microparasellidae</taxonomicName>
|
||
and genera of
|
||
<taxonomicName family="Lepidocharontidae" lsidName="" pageId="4" pageNumber="15" rank="family">Lepidocharontidae</taxonomicName>
|
||
fam. n.
|
||
</paragraph>
|
||
<paragraph pageId="4" pageNumber="15">
|
||
<table pageId="4" pageNumber="15">
|
||
<tr pageId="4" pageNumber="15">
|
||
<td colspan="1" pageId="4" pageNumber="15" rowspan="1">
|
||
<taxonomicName family="Microparasellidae" lsidName="" pageId="4" pageNumber="15" rank="family">Microparasellidae</taxonomicName>
|
||
(
|
||
<taxonomicName class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
|
||
)
|
||
</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="15">
|
||
<td colspan="1" pageId="4" pageNumber="15" rowspan="1">2</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="15">
|
||
<td colspan="1" pageId="4" pageNumber="15" rowspan="1">
|
||
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="15">
|
||
<td colspan="1" pageId="4" pageNumber="15" rowspan="1">4</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="15">
|
||
<td colspan="1" pageId="4" pageNumber="15" rowspan="1">
|
||
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Janinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Janinella" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">Janinella</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="15">
|
||
<td colspan="1" pageId="4" pageNumber="15" rowspan="1">
|
||
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Lepidocharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lepidocharon" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">Lepidocharon</taxonomicName>
|
||
gen. n.
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |