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<document id="3F24E207A00D3E264FE78BF4C53710FD" ID-CLB-Dataset="45928" ID-DOI="10.11646/zootaxa.3793.2.3" ID-GBIF-Dataset="cec96d21-19fa-4e66-8305-9e70166f04f7" ID-ISSN="1175-5326" ID-Zenodo-Dep="230581" ID-ZooBank="5E6DB00C-E98F-4D9D-958B-95B634E6D9E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460412815762" checkinUser="plazi" docAuthor="Baca, Stephen M., Gustafson, Grey T., Toledo, Mario &amp; Miller, Kelly B." docDate="2014" docId="03D9AD22FFB6FFF675A8FA70FCEA97D7" docLanguage="en" docName="zt03793p246.pdf" docOrigin="Zootaxa 3793 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liocanthydrus angustus Guignot 1957" docType="treatment" docVersion="9" lastPageNumber="238" masterDocId="FFE0D55AFFB2FFF1753FFF9FFF65972F" masterDocTitle="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species" masterLastPageNumber="246" masterPageNumber="231" pageNumber="235" updateTime="1698301202102" updateUser="ExternalLinkService">
<mods:mods id="B216449FEF52A2614EBBA08150AC1FD0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DDF9670EAAC1FB6260628E7E8D4BA3CD">
<mods:title id="1CD74CC9260CDFFC0AC9004E86FE993F">Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species</mods:title>
</mods:titleInfo>
<mods:name id="1155C14C0381305159D5A85C22137177" type="personal">
<mods:role id="2CC9694D3F367763C30F7971867B6F39">
<mods:roleTerm id="7C94145C7306558FB60E56FF28CCE990">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="42799ED4EE475BBC6FE7EDF11309AEDC">Baca, Stephen M.</mods:namePart>
</mods:name>
<mods:name id="A33855790D2685D878872A4F6DC3BEE0" type="personal">
<mods:role id="081CE43E0EDEF59056411A8D1156E8C8">
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</mods:role>
<mods:namePart id="D7D974309A68665913E7E30B733E848A">Gustafson, Grey T.</mods:namePart>
</mods:name>
<mods:name id="285B51BF381C31405935609D75A15CFF" type="personal">
<mods:role id="8EEAECF288AE0DF691DF8D074A691036">
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</mods:role>
<mods:namePart id="AA9157894063930489DC924BF614DD17">Toledo, Mario</mods:namePart>
</mods:name>
<mods:name id="66E02E5FAF4262BC40D5BADE2A3BFDBB" type="personal">
<mods:role id="CBE46F187CA3EF148BC4F3AAA1167A93">
<mods:roleTerm id="DFEA5672A7EA2EC1BEFEFCE9DF28C2F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FF63C185CC5E3FD0B04835578519E026">Miller, Kelly B.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3BABAF111CAE4A6D2AD098623DDF5D6B">text</mods:typeOfResource>
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<mods:titleInfo id="F78E6486F74C8E8FE4A7C2BBD7D6FA3A">
<mods:title id="188E29149F997B54030F1038087E77F6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D202EB917BD0504BA03707E6AF6ED6D8">
<mods:date id="6E7E334FEE87486D11335B4E6F02C39B">2014</mods:date>
<mods:detail id="A7E1CB5DF23CAF41834676E052BFFA04" type="volume">
<mods:number id="F12AA4C1EC5B634D727495D057A2B823">3793</mods:number>
</mods:detail>
<mods:detail id="B020093548A4E504B4044842821B2040" type="issue">
<mods:number id="AC671D0C6D3D100ADA428171AA60636C">2</mods:number>
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<mods:start id="AD869D064EDA7C3F3F15AD4D369BCD63">231</mods:start>
<mods:end id="18D3257C8BD76BDCA44E8A1C223DB156">246</mods:end>
</mods:extent>
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<mods:classification id="D1C35436397D0E66349890E426ABC7FA">journal article</mods:classification>
<mods:identifier id="6FD65B48006BE4F1914B3711F119D99C" type="CLB-Dataset">45928</mods:identifier>
<mods:identifier id="407C0F9C232682185AE09897E0BFF64F" type="DOI">10.11646/zootaxa.3793.2.3</mods:identifier>
<mods:identifier id="4215F1A243F1BD92A087729EF7185248" type="GBIF-Dataset">cec96d21-19fa-4e66-8305-9e70166f04f7</mods:identifier>
<mods:identifier id="2E4522B6F98BD2304FBB8CAA09DE1BE8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9C54E7F95846906620602C5908C2B5C3" type="Zenodo-Dep">230581</mods:identifier>
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</mods:mods>
<treatment id="03D9AD22FFB6FFF675A8FA70FCEA97D7" ID-DOI="http://doi.org/10.5281/zenodo.6131923" ID-GBIF-Taxon="119423984" ID-Zenodo-Dep="6131923" LSID="urn:lsid:plazi:treatment:03D9AD22FFB6FFF675A8FA70FCEA97D7" httpUri="http://treatment.plazi.org/id/03D9AD22FFB6FFF675A8FA70FCEA97D7" lastPageId="7" lastPageNumber="238" pageId="4" pageNumber="235">
<subSubSection id="C36A4FBFFFB6FFF575A8FA70FDE09125" box="[151,645,1519,1546]" pageId="4" pageNumber="235" type="nomenclature">
<paragraph id="8BCF1C34FFB6FFF575A8FA70FDE09125" blockId="4.[151,645,1519,1546]" box="[151,645,1519,1546]" pageId="4" pageNumber="235">
<heading id="D087AB58FFB6FFF575A8FA70FDE09125" bold="true" box="[151,645,1519,1546]" fontSize="11" level="1" pageId="4" pageNumber="235" reason="1">
<taxonomicName id="4C7067B7FFB6FFF575A8FA70FDE09125" authority="Guignot, 1957" authorityName="Guignot" authorityYear="1957" box="[151,645,1519,1546]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B904C026FFB6FFF575A8FA70FDE09125" bold="true" box="[151,645,1519,1546]" pageId="4" pageNumber="235">
<emphasis id="B904C026FFB6FFF575A8FA70FEDE9126" bold="true" box="[151,443,1519,1545]" italics="true" pageId="4" pageNumber="235">Liocanthydrus angustus</emphasis>
(
<bibRefCitation id="EFE161C5FFB6FFF574F5FA6FFD189125" author="Guignot" box="[458,637,1520,1546]" pageId="4" pageNumber="235" refString="Guignot, F. (1957) Contribution a la connaissance des dytiscides Sud-Americains. Revue Francaise d'Entomologie, 24, 33 - 45." type="journal article" year="1957">Guignot, 1957</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36A4FBFFFB6FFF575A8F9A9FD989144" pageId="4" pageNumber="235" type="reference_group">
<paragraph id="8BCF1C34FFB6FFF575A8F9A9FD989144" blockId="4.[151,1153,1590,1643]" pageId="4" pageNumber="235">
<treatmentCitationGroup id="AB603B1AFFB6FFF575A8F9A9FD989144" pageId="4" pageNumber="235">
<taxonomicName id="4C7067B7FFB6FFF575A8F9A9FD859163" authority="Guignot (1957: 42)" authorityName="Guignot" authorityPageNumber="42" authorityYear="1957" box="[151,736,1590,1613]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="angustus" subGenus="Liocanthydrus">
<emphasis id="B904C026FFB6FFF575A8F9A9FE689163" box="[151,269,1590,1612]" italics="true" pageId="4" pageNumber="235">Canthydrus</emphasis>
(
<emphasis id="B904C026FFB6FFF57423F9A9FECB9163" box="[284,430,1590,1612]" italics="true" pageId="4" pageNumber="235">Liocanthydrus</emphasis>
)
<emphasis id="B904C026FFB6FFF57482F9A7FD729162" box="[445,535,1592,1613]" italics="true" pageId="4" pageNumber="235">angustus</emphasis>
<treatmentCitation id="0AD13A25FFB6FFF57722F9A9FD859163" author="Guignot" box="[541,736,1590,1612]" page="42" pageId="4" pageNumber="235" year="1957">
<bibRefCitation id="EFE161C5FFB6FFF57722F9A9FD859163" author="Guignot" box="[541,736,1590,1612]" pageId="4" pageNumber="235" refString="Guignot, F. (1957) Contribution a la connaissance des dytiscides Sud-Americains. Revue Francaise d'Entomologie, 24, 33 - 45." type="journal article" year="1957">Guignot (1957: 42)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(orig. descr.);
<treatmentCitation id="0AD13A25FFB6FFF5764BF9A9FB5A9163" author="Nilsson" box="[884,1087,1590,1612]" page="109" pageId="4" pageNumber="235" year="2005">
<bibRefCitation id="EFE161C5FFB6FFF5764BF9A9FB5A9163" author="Nilsson" box="[884,1087,1590,1612]" pageId="4" pageNumber="235" refString="Nilsson, A. N. (2005) Family Noteridae (Coleoptera, Adephaga). In: Nilsson, A. N. &amp; van Vondel, B. J., (Eds.), Amphizoidae, Aspidytidae, Haliplidae, Noteridae and Paelobiidae (Coleoptera, Adephaga). World Catalogue of Insects. Vol. 7. Apollo Books, Stenstrup, 87 - 153. http: // dx. doi. org / 10.1649 / 0010 - 065 x (2006) 60 [305: aahnap] 2.0. co; 2" type="book chapter" year="2005">Nilsson (2005: 109)</bibRefCitation>
</treatmentCitation>
(cat.).
<taxonomicName id="4C7067B7FFB6FFF575A8F9CAFD539144" authority="Guignot, 1957" authorityName="Guignot" authorityYear="1957" box="[151,566,1621,1643]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B904C026FFB6FFF575A8F9CAFEEC9144" box="[151,393,1621,1643]" italics="true" pageId="4" pageNumber="235">Liocanthydrus angustus</emphasis>
(
<bibRefCitation id="EFE161C5FFB6FFF574A8F9CAFD4A9144" author="Guignot" box="[407,559,1621,1643]" pageId="4" pageNumber="235" refString="Guignot, F. (1957) Contribution a la connaissance des dytiscides Sud-Americains. Revue Francaise d'Entomologie, 24, 33 - 45." type="journal article" year="1957">Guignot, 1957</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="0AD13A25FFB6FFF5777FF9CAFD989144" author="Nilsson" box="[576,765,1621,1643]" page="27" pageId="4" pageNumber="235" year="2011">
<bibRefCitation id="EFE161C5FFB6FFF5777FF9CAFD989144" author="Nilsson" box="[576,765,1621,1643]" pageId="4" pageNumber="235" refString="Nilsson, A. N. (2011) A world catalogue of the family Noteridae, or the burrowing water beetles (Coleoptera, Adephaga). Version 16. VIII. 2011, 1 - 54 pp. Available from: http: // www 2. emg. umu. se / projects / biginst / andersn / WCN / WCN _ 20110816. pdf (accessed 9 January 2014)" type="journal article" year="2011">Nilsson (2011: 27)</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36A4FBFFFB6FFF575A8F908FA7B90A7" pageId="4" pageNumber="235" type="materials_examined">
<paragraph id="8BCF1C34FFB6FFF575A8F908FD90919F" blockId="4.[151,1436,1687,1928]" box="[151,757,1687,1712]" pageId="4" pageNumber="235">
<emphasis id="B904C026FFB6FFF575A8F908FE4A919F" bold="true" box="[151,303,1687,1712]" pageId="4" pageNumber="235">
<typeStatus id="54CBA296FFB6FFF575A8F908FFB7919F" box="[151,210,1687,1712]" pageId="4" pageNumber="235">Type</typeStatus>
locality
</emphasis>
:
<collectingCountry id="F3675CA4FFB6FFF57401F908FEE0919F" box="[318,389,1687,1712]" name="Brazil" pageId="4" pageNumber="235">Brazil</collectingCountry>
, Pará State, Sierra de Cachimbo.
</paragraph>
<paragraph id="8BCF1C34FFB6FFF575F8F923FCAB9032" blockId="4.[151,1436,1687,1928]" pageId="4" pageNumber="235">
<emphasis id="B904C026FFB6FFF575F8F923FE9191FC" bold="true" box="[199,500,1724,1749]" pageId="4" pageNumber="235">
<typeStatus id="54CBA296FFB6FFF575F8F923FE6791FA" box="[199,258,1724,1749]" pageId="4" pageNumber="235">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="54CBA296FFB6FFF574C2F922FD3A91FA" box="[509,607,1725,1749]" pageId="4" pageNumber="235" type="paratype">Paratype</typeStatus>
(female): “
<collectingCountry id="F3675CA4FFB6FFF577DCF923FC1291FA" box="[739,887,1724,1749]" name="Brazil" pageId="4" pageNumber="235">BRASILIEN</collectingCountry>
/Para/Cachimbo/
<date id="FFCE3AF4FFB6FFF57105F923FBE391FA" box="[1082,1158,1724,1749]" pageId="4" pageNumber="235" value="1955-10">X.1955</date>
/Pereira” [printed], “♀”, “
<typeStatus id="54CBA296FFB6FFF5759EF97FFE6D91D7" box="[161,264,1760,1784]" pageId="4" pageNumber="235" type="paratype">Paratype</typeStatus>
” [red; printed], “Museum Paris/1960/coll. F. Guignot” [printed] (1 ex.,
<collectionCode id="ED6184F1FFB6FFF57179F97FFBC491D7" LSID="urn:lsid:biocol.org:col:34988" box="[1094,1185,1760,1784]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="235">MNHN</collectionCode>
).
<typeStatus id="54CBA296FFB6FFF57189F940FA7B91D7" box="[1206,1310,1759,1784]" pageId="4" pageNumber="235" type="holotype">Holotype</typeStatus>
(male, not studied) in
<collectionCode id="ED6184F1FFB6FFF5742AF89BFE169033" box="[277,371,1796,1820]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="235">MZUSP</collectionCode>
.
<emphasis id="B904C026FFB6FFF57442F89BFEDB9032" bold="true" box="[381,446,1796,1821]" pageId="4" pageNumber="235">Note:</emphasis>
All label descriptions are included in brackets.
</paragraph>
<paragraph id="8BCF1C34FFB6FFF575F8F8B8FA7B90A7" blockId="4.[151,1436,1687,1928]" pageId="4" pageNumber="235">
<emphasis id="B904C026FFB6FFF575F8F8B8FDD4906F" bold="true" box="[199,689,1831,1856]" pageId="4" pageNumber="235">
Other material examined.
<collectingCountry id="F3675CA4FFB6FFF57730F8B8FD39906F" box="[527,604,1831,1856]" name="Brazil" pageId="4" pageNumber="235">Brazil</collectingCountry>
: Goiás
</emphasis>
, Caldas Novas, at light,
<date id="FFCE3AF4FFB6FFF576E5F8B8FBE6906F" box="[986,1155,1831,1856]" pageId="4" pageNumber="235" value="1993-04-13" valueMax="1993-05-01" valueMin="1993-04-13">13.iv–1.v.1993</date>
, J. Sár (1 ex.,
<collectionCode id="ED6184F1FFB6FFF57008F8B7FAF5906F" box="[1335,1424,1832,1856]" pageId="4" pageNumber="235">HNMB</collectionCode>
);
<emphasis id="B904C026FFB6FFF575A8F8D3FE51904A" bold="true" box="[151,308,1868,1893]" pageId="4" pageNumber="235">Mato Grosso</emphasis>
, Jacaré, PNXingu,
<date id="FFCE3AF4FFB6FFF5772BF8D3FD0E904A" box="[532,619,1868,1893]" pageId="4" pageNumber="235" value="1965-11">xi.1965</date>
, MALvarenga &amp; WCABokermann,
<collectionCode id="ED6184F1FFB6FFF57133F8D2FB5B904B" LSID="urn:lsid:biocol.org:col:12647" box="[1036,1086,1869,1892]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12647" name="Belfast Natural History and Philosophical Society" pageId="4" pageNumber="235">BLT</collectionCode>
(1 ex.,
<collectionCode id="ED6184F1FFB6FFF571A5F8D3FB81904B" box="[1178,1252,1868,1892]" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="235">FSCA</collectionCode>
; 1 ex.,
<collectionCode id="ED6184F1FFB6FFF57005F8D3FAF5904B" LSID="urn:lsid:biocol.org:col:34871" box="[1338,1424,1868,1892]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="235">USNM</collectionCode>
);
<emphasis id="B904C026FFB6FFF575A8F8F0FE3090A7" bold="true" box="[151,341,1903,1928]" pageId="4" pageNumber="235">
<collectingCountry id="F3675CA4FFB6FFF575A8F8F0FE3590A7" box="[151,336,1903,1928]" name="French Guiana" pageId="4" pageNumber="235">French Guiana</collectingCountry>
:
</emphasis>
Sinnamary, Piste Sante Elie, Crique Toissant,
<date id="FFCE3AF4FFB6FFF57662F8F0FCBD90A7" box="[861,984,1903,1928]" pageId="4" pageNumber="235" value="2007-11-26">26.xi.2007</date>
, D. Lohez leg. (1 ex., MTC).
</paragraph>
</subSubSection>
<caption id="DF0F4CBCFFB7FFF475A8FB29FCBB93C4" httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="5" pageNumber="236" targetBox="[192,1397,212,1170]" targetPageId="5">
<paragraph id="8BCF1C34FFB7FFF475A8FB29FCBB93C4" blockId="5.[151,1436,1206,1259]" pageId="5" pageNumber="236">
<emphasis id="B904C026FFB7FFF475A8FB29FE5193E3" bold="true" box="[151,308,1206,1228]" pageId="5" pageNumber="236">FIGURES 3–9</emphasis>
.
<taxonomicName id="4C7067B7FFB7FFF4747EFB29FEB693E3" box="[321,467,1206,1228]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis id="B904C026FFB7FFF4747EFB29FEB693E3" box="[321,467,1206,1228]" italics="true" pageId="5" pageNumber="236">Liocanthydrus</emphasis>
</taxonomicName>
species, dorsal habitus: 3)
<taxonomicName id="4C7067B7FFB7FFF477D9FB27FC3993E2" box="[742,860,1208,1229]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B904C026FFB7FFF477D9FB27FC3993E2" box="[742,860,1208,1229]" italics="true" pageId="5" pageNumber="236">L. angustus</emphasis>
</taxonomicName>
; 4)
<taxonomicName id="4C7067B7FFB7FFF476BAFB27FB6493E3" box="[901,1025,1206,1229]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="B904C026FFB7FFF476BAFB27FB6493E3" box="[901,1025,1206,1229]" italics="true" pageId="5" pageNumber="236">L. uniformis</emphasis>
</taxonomicName>
; 5)
<taxonomicName id="4C7067B7FFB7FFF47115FB27FBEB93E3" box="[1066,1166,1206,1229]" class="Insecta" family="Noteridae" genus="Siolius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B904C026FFB7FFF47115FB27FBEB93E3" box="[1066,1166,1206,1229]" italics="true" pageId="5" pageNumber="236">L. bicolor</emphasis>
</taxonomicName>
; 6)
<taxonomicName id="4C7067B7FFB7FFF47187FB27FA5F93E3" box="[1208,1338,1206,1229]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="armulatus" status="sp. nov.">
<emphasis id="B904C026FFB7FFF47187FB27FA5F93E3" box="[1208,1338,1206,1229]" italics="true" pageId="5" pageNumber="236">L. armulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377D5DFFB7FFF4707FFB29FA1C93E3" box="[1344,1401,1206,1228]" pageId="5" pageNumber="236" rank="species">
<emphasis id="B904C026FFB7FFF4707FFB29FA1C93E3" bold="true" box="[1344,1401,1206,1228]" pageId="5" pageNumber="236">sp. n.</emphasis>
</taxonomicNameLabel>
; 7)
<taxonomicName id="4C7067B7FFB7FFF475A8FB49FF9193C4" box="[151,244,1237,1259]" class="Insecta" family="Noteridae" genus="Siolius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="clayae">
<emphasis id="B904C026FFB7FFF475A8FB49FFC093C4" box="[151,165,1238,1259]" italics="true" pageId="5" pageNumber="236">L</emphasis>
.
<emphasis id="B904C026FFB7FFF4758DFB4AFF9193C4" box="[178,244,1237,1259]" italics="true" pageId="5" pageNumber="236">clayae</emphasis>
</taxonomicName>
; 8)
<taxonomicName id="4C7067B7FFB7FFF47423FB49FEE593C4" box="[284,384,1238,1259]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="nanops" status="sp. nov.">
<emphasis id="B904C026FFB7FFF47423FB49FEE593C4" box="[284,384,1238,1259]" italics="true" pageId="5" pageNumber="236">L. nanops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377D5DFFB7FFF474B9FB4AFEDA93C4" box="[390,447,1237,1259]" pageId="5" pageNumber="236" rank="species">
<emphasis id="B904C026FFB7FFF474B9FB4AFEDA93C4" bold="true" box="[390,447,1237,1259]" pageId="5" pageNumber="236">sp. n.</emphasis>
</taxonomicNameLabel>
; 9)
<taxonomicName id="4C7067B7FFB7FFF474D7FB49FDE593C4" box="[488,640,1238,1259]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="octoguttatus">
<emphasis id="B904C026FFB7FFF474D7FB49FDE593C4" box="[488,640,1238,1259]" italics="true" pageId="5" pageNumber="236">L. octoguttatus</emphasis>
</taxonomicName>
(illustration). Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="DF0F4CBCFFB7FFF475A8F82DFD8C90C8" httpUri="https://zenodo.org/record/230584/files/figure.png" pageId="5" pageNumber="236" targetBox="[164,1430,1309,1937]" targetPageId="5">
<paragraph id="8BCF1C34FFB7FFF475A8F82DFD8C90C8" blockId="5.[151,1436,1970,2023]" pageId="5" pageNumber="236">
<emphasis id="B904C026FFB7FFF475A8F82DFE3690E7" bold="true" box="[151,339,1970,1992]" pageId="5" pageNumber="236">FIGURES 10–15.</emphasis>
<taxonomicName id="4C7067B7FFB7FFF47465F82DFE8990E7" box="[346,492,1970,1992]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis id="B904C026FFB7FFF47465F82DFE8990E7" box="[346,492,1970,1992]" italics="true" pageId="5" pageNumber="236">Liocanthydrus</emphasis>
</taxonomicName>
species, lateral aspect: 10)
<taxonomicName id="4C7067B7FFB7FFF4763DF82CFC1290E7" box="[770,887,1971,1992]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B904C026FFB7FFF4763DF82CFC1290E7" box="[770,887,1971,1992]" italics="true" pageId="5" pageNumber="236">L. angustus</emphasis>
</taxonomicName>
; 11)
<taxonomicName id="4C7067B7FFB7FFF47694F82CFB4290E7" box="[939,1063,1970,1992]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="B904C026FFB7FFF47694F82CFB4290E7" box="[939,1063,1970,1992]" italics="true" pageId="5" pageNumber="236">L. uniformis</emphasis>
</taxonomicName>
; 12)
<taxonomicName id="4C7067B7FFB7FFF47163F82CFBA590E7" box="[1116,1216,1970,1992]" class="Insecta" family="Noteridae" genus="Siolius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B904C026FFB7FFF47163F82CFBA590E7" box="[1116,1216,1970,1992]" italics="true" pageId="5" pageNumber="236">L. bicolor</emphasis>
</taxonomicName>
; 13)
<taxonomicName id="4C7067B7FFB7FFF471C9F82CFA1D90E7" box="[1270,1400,1970,1992]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="armulatus" status="sp. nov.">
<emphasis id="B904C026FFB7FFF471C9F82CFA1D90E7" box="[1270,1400,1970,1992]" italics="true" pageId="5" pageNumber="236">L. armulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377D5DFFB7FFF47042F82DFFCE90C9" pageId="5" pageNumber="236" rank="species">
<emphasis id="B904C026FFB7FFF47042F82DFFCE90C9" bold="true" pageId="5" pageNumber="236">sp. n.</emphasis>
</taxonomicNameLabel>
; 14)
<taxonomicName id="4C7067B7FFB7FFF475DEF84DFE5890C8" box="[225,317,2001,2023]" class="Insecta" family="Noteridae" genus="Siolius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="clayae">
<emphasis id="B904C026FFB7FFF475DEF84DFE5890C8" box="[225,317,2001,2023]" italics="true" pageId="5" pageNumber="236">L. clayae</emphasis>
</taxonomicName>
; 15)
<taxonomicName id="4C7067B7FFB7FFF4744CF84DFEB390C8" box="[371,470,2002,2023]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="nanops" status="sp. nov.">
<emphasis id="B904C026FFB7FFF4744CF84DFEB390C8" box="[371,470,2002,2023]" italics="true" pageId="5" pageNumber="236">L. nanops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377D5DFFB7FFF474E2F84EFD7390C9" box="[477,534,2001,2023]" pageId="5" pageNumber="236" rank="species">
<emphasis id="B904C026FFB7FFF474E2F84EFD7390C9" bold="true" box="[477,534,2001,2023]" pageId="5" pageNumber="236">sp. n.</emphasis>
</taxonomicNameLabel>
Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="DF0F4CBCFFB4FFF775A8FC5AFAF29318" httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="6" pageNumber="237" targetBox="[163,1436,193,935]" targetPageId="6">
<paragraph id="8BCF1C34FFB4FFF775A8FC5AFAF29318" blockId="6.[151,1436,965,1079]" pageId="6" pageNumber="237">
<emphasis id="B904C026FFB4FFF775A8FC5AFE3594F4" bold="true" box="[151,336,965,987]" pageId="6" pageNumber="237">FIGURES 16–22</emphasis>
. Aedeagi of
<taxonomicName id="4C7067B7FFB4FFF774E6FC5AFD0E94F4" box="[473,619,965,987]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="genus">
<emphasis id="B904C026FFB4FFF774E6FC5AFD0E94F4" box="[473,619,965,987]" italics="true" pageId="6" pageNumber="237">Liocanthydrus</emphasis>
</taxonomicName>
species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16)
<taxonomicName id="4C7067B7FFB4FFF77020FC7AFAF094D5" box="[1311,1429,997,1018]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B904C026FFB4FFF77020FC7AFAF094D5" box="[1311,1429,997,1018]" italics="true" pageId="6" pageNumber="237">L. angustus</emphasis>
</taxonomicName>
; 17)
<taxonomicName id="4C7067B7FFB4FFF77580FB9CFE599337" box="[191,316,1026,1048]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="B904C026FFB4FFF77580FB9CFE599337" box="[191,316,1026,1048]" italics="true" pageId="6" pageNumber="237">L. uniformis</emphasis>
</taxonomicName>
; 18)
<taxonomicName id="4C7067B7FFB4FFF7744CFB9CFEB29337" box="[371,471,1026,1048]" class="Insecta" family="Noteridae" genus="Siolius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B904C026FFB4FFF7744CFB9CFEB29337" box="[371,471,1026,1048]" italics="true" pageId="6" pageNumber="237">L. bicolor</emphasis>
</taxonomicName>
; 19)
<taxonomicName id="4C7067B7FFB4FFF77731FB9CFDF59337" box="[526,656,1026,1048]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="armulatus" status="sp. nov.">
<emphasis id="B904C026FFB4FFF77731FB9CFDF59337" box="[526,656,1026,1048]" italics="true" pageId="6" pageNumber="237">L. armulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377D5DFFB4FFF777A8FB9DFDB49337" box="[663,721,1026,1048]" pageId="6" pageNumber="237" rank="species">
<emphasis id="B904C026FFB4FFF777A8FB9DFDB49337" bold="true" box="[663,721,1026,1048]" pageId="6" pageNumber="237">sp. n.</emphasis>
</taxonomicNameLabel>
; 20)
<taxonomicName id="4C7067B7FFB4FFF77637FB9CFC009337" box="[776,869,1026,1048]" class="Insecta" family="Noteridae" genus="Siolius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="clayae">
<emphasis id="B904C026FFB4FFF77637FB9CFC009337" box="[776,869,1026,1048]" italics="true" pageId="6" pageNumber="237">L. clayae</emphasis>
</taxonomicName>
; 21)
<taxonomicName id="4C7067B7FFB4FFF776A3FB9CFB649337" box="[924,1025,1027,1048]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="nanops" status="sp. nov.">
<emphasis id="B904C026FFB4FFF776A3FB9CFB649337" box="[924,1025,1027,1048]" italics="true" pageId="6" pageNumber="237">L. nanops</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377D5DFFB4FFF77137FB9DFB249337" box="[1032,1089,1026,1048]" pageId="6" pageNumber="237" rank="species">
<emphasis id="B904C026FFB4FFF77137FB9DFB249337" bold="true" box="[1032,1089,1026,1048]" pageId="6" pageNumber="237">sp. n.</emphasis>
</taxonomicNameLabel>
; 22)
<taxonomicName id="4C7067B7FFB4FFF77146FB9CFA749337" box="[1145,1297,1027,1048]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="octoguttatus">
<emphasis id="B904C026FFB4FFF77146FB9CFA749337" box="[1145,1297,1027,1048]" italics="true" pageId="6" pageNumber="237">L. octoguttatus</emphasis>
</taxonomicName>
. Scale bars = 0.2 mm.
<emphasis id="B904C026FFB4FFF775CFFBBEFE499318" bold="true" box="[240,300,1057,1079]" pageId="6" pageNumber="237">Notes</emphasis>
: Terminology follows K.B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces.
</paragraph>
</caption>
<subSubSection id="C36A4FBFFFB4FFF775F8FBC1FCEB9261" pageId="6" pageNumber="237" type="diagnosis">
<paragraph id="8BCF1C34FFB4FFF775F8FBC1FD6C93CC" blockId="6.[151,1437,1118,1971]" pageId="6" pageNumber="237">
<emphasis id="B904C026FFB4FFF775F8FBC1FE249358" bold="true" box="[199,321,1118,1143]" pageId="6" pageNumber="237">Diagnosis.</emphasis>
Elytra without maculation, distinguished from other immaculate members by the following combination of characters: a) large eye size, EW/HW = 1.63–1.73 (
<figureCitation id="134B00B1FFB4FFF776B7FB1DFCAE93B4" box="[904,971,1154,1179]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="6" pageNumber="237">Fig. 3</figureCitation>
); b) body outline as in
<figureCitation id="134B00B1FFB4FFF771F2FB1DFA7593B4" box="[1229,1296,1154,1179]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="6" pageNumber="237">Fig. 3</figureCitation>
, slender and gradually attenuate posteriorly; c) aedeagus as in
<figureCitation id="134B00B1FFB4FFF777F1FB39FC469390" box="[718,803,1190,1215]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="6" pageNumber="237">Figs 16</figureCitation>
a–e, with left lateral lobe slender, distinctly subequal in length to median lobe (
<figureCitation id="134B00B1FFB4FFF774A5FB55FE8993CC" box="[410,492,1226,1251]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="6" pageNumber="237">Fig. 16</figureCitation>
d).
</paragraph>
<paragraph id="8BCF1C34FFB4FFF775F8FB71FCEB9261" blockId="6.[151,1437,1118,1971]" pageId="6" pageNumber="237">
Externally
<taxonomicName id="4C7067B7FFB4FFF7747DFB70FEA79229" box="[322,450,1263,1286]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B904C026FFB4FFF7747DFB70FEA79229" box="[322,450,1263,1286]" italics="true" pageId="6" pageNumber="237">L. angustus</emphasis>
</taxonomicName>
appears very similar to
<taxonomicName id="4C7067B7FFB4FFF777F2FB70FC309228" box="[717,853,1263,1287]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="uniformis">
<emphasis id="B904C026FFB4FFF777F2FB70FC309228" box="[717,853,1263,1287]" italics="true" pageId="6" pageNumber="237">L. uniformis</emphasis>
</taxonomicName>
. However,
<taxonomicName id="4C7067B7FFB4FFF776EAFB70FB339229" box="[981,1110,1263,1286]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="B904C026FFB4FFF776EAFB70FB339229" box="[981,1110,1263,1286]" italics="true" pageId="6" pageNumber="237">L. angustus</emphasis>
</taxonomicName>
has a larger relative eye size, EW/HW = 1.63–1.73 (fig. 3), a body outline that is less rounded and posteriorly more attenuate (
<figureCitation id="134B00B1FFB4FFF771F1FA8DFA749204" box="[1230,1297,1298,1323]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="6" pageNumber="237">Fig. 3</figureCitation>
), and has an aedeagus with a distinctly longer and more slender left lateral lobe.
</paragraph>
</subSubSection>
<subSubSection id="C36A4FBFFFB4FFF775F8FAC5FE87909C" pageId="6" pageNumber="237" type="description">
<paragraph id="8BCF1C34FFB4FFF775F8FAC5FB589294" blockId="6.[151,1437,1118,1971]" pageId="6" pageNumber="237">
<emphasis id="B904C026FFB4FFF775F8FAC5FE3C925C" bold="true" box="[199,345,1370,1395]" pageId="6" pageNumber="237">Description.</emphasis>
Measurements. TL =
<quantity id="4C88B1D1FFB4FFF7776AFAC5FD88925D" box="[597,749,1370,1395]" metricMagnitude="-3" metricUnit="m" metricValue="2.9749999999999996" metricValueMax="3.05" metricValueMin="2.9" pageId="6" pageNumber="237" unit="mm" value="2.9749999999999996" valueMax="3.05" valueMin="2.9">2.9–3.05 mm</quantity>
, male max. = 3.0 mm, female max. =
<quantity id="4C88B1D1FFB4FFF7719FFAC5FA62925D" box="[1184,1287,1370,1395]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" pageId="6" pageNumber="237" unit="mm" value="3.05">3.05 mm</quantity>
, male min. =
<quantity id="4C88B1D1FFB4FFF775A8FAE0FF8A92B9" box="[151,239,1407,1431]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="6" pageNumber="237" unit="mm" value="2.9">2.9 mm</quantity>
, GW =
<quantity id="4C88B1D1FFB4FFF77476FAE1FE8492B9" box="[329,481,1406,1431]" metricMagnitude="-3" metricUnit="m" metricValue="1.4249999999999998" metricValueMax="1.45" metricValueMin="1.4" pageId="6" pageNumber="237" unit="mm" value="1.4249999999999998" valueMax="1.45" valueMin="1.4">1.4–1.45 mm</quantity>
, L/GW = 2.07–2.18, HW =
<quantity id="4C88B1D1FFB4FFF7761FFAE0FCCF92B9" box="[800,938,1407,1431]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="6" pageNumber="237" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.8–0.9 mm</quantity>
, EW =
<quantity id="4C88B1D1FFB4FFF7713FFAE1FBFD92B9" box="[1024,1176,1406,1431]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="6" pageNumber="237" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.5–0.55 mm</quantity>
, HW/EW = 1.63–1.73, PNWpost =
<quantity id="4C88B1D1FFB4FFF7741DFA3CFECE9295" box="[290,427,1443,1467]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="6" pageNumber="237" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3–1.4 mm</quantity>
, PNWant =
<quantity id="4C88B1D1FFB4FFF7770DFA3DFDAC9295" box="[562,713,1442,1467]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="9.0" metricValueMin="8.5" pageId="6" pageNumber="237" unit="mm" value="0.875" valueMax="0.9" valueMin="0.85">0.85–0.9 mm</quantity>
, PNWpost/PNWant = 1.51–1.55.
</paragraph>
<paragraph id="8BCF1C34FFB4FFF775F8FA59FE289108" blockId="6.[151,1437,1118,1971]" pageId="6" pageNumber="237">
<emphasis id="B904C026FFB4FFF775F8FA59FE2F92F0" box="[199,330,1478,1503]" italics="true" pageId="6" pageNumber="237">Coloration.</emphasis>
Elytra without maculation; color of elytra brown to reddish brown; color of head and pronotum yellowish red to reddish brown, slightly lighter in color than elytra (
<figureCitation id="134B00B1FFB4FFF776A9FA75FCBC912C" box="[918,985,1514,1539]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="6" pageNumber="237">Fig. 3</figureCitation>
); color of venter reddish brown, similar in color to head.
</paragraph>
<paragraph id="8BCF1C34FFB4FFF775F8F9ADFE289140" blockId="6.[151,1437,1118,1971]" pageId="6" pageNumber="237">
<emphasis id="B904C026FFB4FFF775F8F9ADFE639164" box="[199,262,1586,1611]" italics="true" pageId="6" pageNumber="237">Body.</emphasis>
Body outline as in
<figureCitation id="134B00B1FFB4FFF774D8F9ADFD499164" box="[487,556,1586,1611]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="6" pageNumber="237">Fig. 3</figureCitation>
; oval, narrow, attenuate posteriorly to apex; narrow, weakly convex in lateral aspect (
<figureCitation id="134B00B1FFB4FFF775D3F9C9FE5A9140" box="[236,319,1622,1647]" captionStart="FIGURES 10 – 15" captionStartId="5.[151,264,1970,1992]" captionTargetBox="[164,1430,1309,1937]" captionTargetId="figure@5.[151,1436,1309,1949]" captionTargetPageId="5" captionText="FIGURES 10 – 15. Liocanthydrus species, lateral aspect: 10) L. angustus; 11) L. uniformis; 12) L. bicolor; 13) L. armulatus sp. n.; 14) L. clayae; 15) L. nanops sp. n. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230584/files/figure.png" pageId="6" pageNumber="237">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8BCF1C34FFB4FFF775F8F9E5FB4791BC" blockId="6.[151,1437,1118,1971]" box="[199,1058,1658,1683]" pageId="6" pageNumber="237">
<emphasis id="B904C026FFB4FFF775F8F9E5FE6E91BC" box="[199,267,1658,1683]" italics="true" pageId="6" pageNumber="237">Head.</emphasis>
Eyes large relative to head capsule, HW/EW = 1.63–1.73 (
<figureCitation id="134B00B1FFB4FFF7769EF9E5FC8C91BC" box="[929,1001,1658,1683]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="6" pageNumber="237">Figs 3</figureCitation>
,
<figureCitation id="134B00B1FFB4FFF776C9F9E4FB7091BC" box="[1014,1045,1659,1683]" captionStart="FIGURES 10 – 15" captionStartId="5.[151,264,1970,1992]" captionTargetBox="[164,1430,1309,1937]" captionTargetId="figure@5.[151,1436,1309,1949]" captionTargetPageId="5" captionText="FIGURES 10 – 15. Liocanthydrus species, lateral aspect: 10) L. angustus; 11) L. uniformis; 12) L. bicolor; 13) L. armulatus sp. n.; 14) L. clayae; 15) L. nanops sp. n. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230584/files/figure.png" pageId="6" pageNumber="237">10</figureCitation>
).
</paragraph>
<paragraph id="8BCF1C34FFB4FFF775F8F901FBD49198" blockId="6.[151,1437,1118,1971]" box="[199,1201,1694,1719]" pageId="6" pageNumber="237">
<emphasis id="B904C026FFB4FFF775F8F901FE7D9198" box="[199,280,1694,1719]" italics="true" pageId="6" pageNumber="237">Thorax</emphasis>
. Elytra with lateral margins slightly ascendant medially in lateral aspect (
<figureCitation id="134B00B1FFB4FFF7716FF901FBC69198" box="[1104,1187,1694,1719]" captionStart="FIGURES 10 – 15" captionStartId="5.[151,264,1970,1992]" captionTargetBox="[164,1430,1309,1937]" captionTargetId="figure@5.[151,1436,1309,1949]" captionTargetPageId="5" captionText="FIGURES 10 – 15. Liocanthydrus species, lateral aspect: 10) L. angustus; 11) L. uniformis; 12) L. bicolor; 13) L. armulatus sp. n.; 14) L. clayae; 15) L. nanops sp. n. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230584/files/figure.png" pageId="6" pageNumber="237">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8BCF1C34FFB4FFF775F8F95DFD12900C" blockId="6.[151,1437,1118,1971]" pageId="6" pageNumber="237">
<emphasis id="B904C026FFB4FFF775F8F95DFE5D91F4" box="[199,312,1730,1755]" italics="true" pageId="6" pageNumber="237">Abdomen.</emphasis>
Aedeagus as in
<figureCitation id="134B00B1FFB4FFF774D0F95DFD2791F4" box="[495,578,1730,1755]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="6" pageNumber="237">Figs 16</figureCitation>
a–e; median lobe narrow and attenuate (
<figureCitation id="134B00B1FFB4FFF7713CF95DFB3D91F4" box="[1027,1112,1730,1755]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="6" pageNumber="237">Figs 16</figureCitation>
a–c); left lateral lobe narrow, subequal in length to median lobe, with setal fringe limited to distal third (
<figureCitation id="134B00B1FFB4FFF776D4F979FB5A91D0" box="[1003,1087,1766,1791]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="6" pageNumber="237">Fig. 16</figureCitation>
d); right lateral lobe triangular, distal angle acute, not projected (
<figureCitation id="134B00B1FFB4FFF77736F895FD39900C" box="[521,604,1802,1827]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller &amp; Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="6" pageNumber="237">Fig. 16</figureCitation>
e).
</paragraph>
<paragraph id="8BCF1C34FFB4FFF775F8F8B1FE87909C" blockId="6.[151,1437,1118,1971]" pageId="6" pageNumber="237">
<emphasis id="B904C026FFB4FFF775F8F8B1FE5E9068" bold="true" box="[199,315,1838,1863]" pageId="6" pageNumber="237">Variation.</emphasis>
The variation exhibited by this species was difficult to assess due to the limited number of specimens examined for this study. The examined individuals differed slightly in overall color, with some specimens slightly darker and more reddish brown than others. There was also some very slight variation in the overall size of the specimens.
</paragraph>
</subSubSection>
<subSubSection id="C36A4FBFFFB5FFF675F8FF08FC6697FA" pageId="7" pageNumber="238" type="biology_ecology">
<paragraph id="8BCF1C34FFB5FFF675F8FF08FC6697FA" blockId="7.[151,1436,151,248]" pageId="7" pageNumber="238">
<emphasis id="B904C026FFB5FFF675F8FF08FE42979F" bold="true" box="[199,295,151,176]" pageId="7" pageNumber="238">Biology.</emphasis>
Available collecting data were very limited for this species. One specimen was labeled as taken from a light and another named a creek as the collection site.
</paragraph>
</subSubSection>
<subSubSection id="C36A4FBFFFB5FFF675F8FF40FCEA97D7" box="[199,911,223,248]" pageId="7" pageNumber="238" type="distribution">
<paragraph id="8BCF1C34FFB5FFF675F8FF40FCEA97D7" blockId="7.[151,1436,151,248]" box="[199,911,223,248]" pageId="7" pageNumber="238">
<emphasis id="B904C026FFB5FFF675F8FF40FE0597D7" bold="true" box="[199,352,223,248]" pageId="7" pageNumber="238">Distribution.</emphasis>
Known from
<collectingCountry id="F3675CA4FFB5FFF674C1FF40FDC197D7" box="[510,676,223,248]" name="French Guiana" pageId="7" pageNumber="238">French Guiana</collectingCountry>
and
<collectingCountry id="F3675CA4FFB5FFF677E5FF40FC7B97D7" box="[730,798,223,248]" name="Brazil" pageId="7" pageNumber="238">Brazil</collectingCountry>
(
<figureCitation id="134B00B1FFB5FFF67611FF40FCE797D7" box="[814,898,223,248]" captionStart="FIGURE 23" captionStartId="7.[151,250,1491,1513]" captionText="FIGURE 23. Distribution map of Liocanthydrus species." pageId="7" pageNumber="238">Fig. 23</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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