165 lines
20 KiB
XML
165 lines
20 KiB
XML
<document id="D44D4D728B32522C37F8BEDEAB791817" ID-DOI="10.11646/zootaxa.1957.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5241137" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629590285433" checkinUser="felipe" docAuthor="Sartori, Michel, Peters, Janice G. & Hubbard, Michael D." docDate="2008" docId="039AE86B4402FFFBCE8E72C8FB41FE6F" docLanguage="en" docName="zt01957p051.pdf" docOrigin="Zootaxa 1957 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1957.1.1" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Teloganodes dentatus Navas 1931" docType="treatment" docVersion="3" lastPageNumber="11" masterDocId="FFA390134408FFF0CE197632FFA3FFCD" masterDocTitle="A revision of Oriental Teloganodidae (Insecta, Ephemeroptera, Ephemerelloidea)" masterLastPageNumber="51" masterPageNumber="1" pageNumber="11" updateTime="1699082080968" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="2CA7497F7DDA71DCAC1CF1E17B4BAAC3">
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<mods:title id="424B739A45326F1CA42D202843807980">A revision of Oriental Teloganodidae (Insecta, Ephemeroptera, Ephemerelloidea)</mods:title>
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<mods:name id="F6B7109D226D9523FAEC8262309AA4C5" type="personal">
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<mods:namePart id="AB1C8842622797E14A22573E08652A72">Sartori, Michel</mods:namePart>
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<mods:affiliation id="C5BD1874E3D54E4F609FDEDA5901ABCE">Museum of zoology, Palais de Rumine, Place Riponne 6, CH- 1014 Lausanne, Switzerland. E-mail: michel. sartori @ vd. ch Laboratory of Aquatic Entomology, Florida A & M University, Tallahassee, FL 32307, USA</mods:affiliation>
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<mods:namePart id="D0107EB9EEC4F01A1781ED1BC6A8B1B3">Peters, Janice G.</mods:namePart>
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<mods:namePart id="9EA230A96EA43AE036B7F040C235A39B">Hubbard, Michael D.</mods:namePart>
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<mods:title id="67DA2CACB08F1C13F2900ACDDA4C7829">Zootaxa</mods:title>
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<mods:part id="D31E897C463CDF49419CA7AD676212B6">
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<mods:date id="3738D4A789DACF8F55C98BA30BD977B8">2008</mods:date>
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<mods:number id="A81C711217E72FCEF3B6178647C4F82E">2008-12-05</mods:number>
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<mods:number id="364BE0F967D0D98AE649EBEA1E66CFB2">1957</mods:number>
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<mods:url id="CF488F17AB55AD32B78923A4DC4AD4F0">https://biotaxa.org/Zootaxa/article/view/zootaxa.1957.1.1</mods:url>
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<treatment id="039AE86B4402FFFBCE8E72C8FB41FE6F" LSID="urn:lsid:plazi:treatment:039AE86B4402FFFBCE8E72C8FB41FE6F" httpUri="http://treatment.plazi.org/id/039AE86B4402FFFBCE8E72C8FB41FE6F" lastPageId="11" lastPageNumber="11" pageId="10" pageNumber="11">
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<subSubSection id="C3290AF64402FFFACE8E72C8FD94FAD9" box="[151,567,1274,1300]" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="8B8C597D4402FFFACE8E72C8FD94FAD9" blockId="10.[151,567,1274,1335]" box="[151,567,1274,1300]" pageId="10" pageNumber="11">
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<heading id="D0C4EE114402FFFACE8E72C8FD94FAD9" bold="true" box="[151,567,1274,1300]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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<taxonomicName id="4C3322FE4402FFFACE8E72C8FD94FAD9" ID-CoL="558P4" authority="Navas, 1931" authorityName="Navas" authorityYear="1931" box="[151,567,1274,1300]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dentatus">
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<emphasis id="B947856F4402FFFACE8E72C8FD94FAD9" bold="true" box="[151,567,1274,1300]" pageId="10" pageNumber="11">
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<emphasis id="B947856F4402FFFACE8E72C8FE39FAD9" bold="true" box="[151,410,1274,1300]" italics="true" pageId="10" pageNumber="11">Teloganodes dentatus</emphasis>
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<bibRefCitation id="EFA2248C4402FFFACFBB72C8FD94FAD9" author="Navas, L." box="[418,567,1274,1300]" pageId="10" pageNumber="11" pagination="16 - 23" refId="ref26191" refString="Navas, L. (1931) Insectos de la India. 4 a serie. Revista de la Academia de Ciencias de Zaragoza, 15, 16 - 23." type="journal article" year="1931">Navás, 1931</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3290AF64402FFFACE8E732FFEB4FAFA" box="[151,279,1309,1335]" pageId="10" pageNumber="11" type="description">
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<paragraph id="8B8C597D4402FFFACE8E732FFEB4FAFA" blockId="10.[151,567,1274,1335]" box="[151,279,1309,1335]" pageId="10" pageNumber="11">
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(
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<figureCitation id="130845F84402FFFACE86732FFEADFAFA" box="[159,270,1309,1335]" captionStart="FIGURES 1–6" captionStartId="9.[151,269,1864,1888]" captionTargetBox="[251,1344,188,1838]" captionTargetId="figure-98@9.[240,1345,185,1848]" captionTargetPageId="9" captionText="FIGURES 1–6. Teloganodes spp. Fig. 1: Forewing of the female subimago lectotype of Teloganodes tristis (Hagen) [MCZ]; Fig. 2: Forewing of a male subimago syntype of Teloganodes major Eaton [MCZ]. Fig. 3: Hindwing of the female lectotype of Teloganodes tristis (Hagen) [MCZ]; Fig. 4: Hindwing of the male holotype of Teloganodes dentatus Navás [MNHN]; Fig. 5: Genitalia of a male subimago syntype of Teloganodes major Eaton [MCZ]; Fig. 6: Genitalia in ventral view of the male holotype of T. dentatus Navás [MNHN]." figureDoi="http://doi.org/10.5281/zenodo.5241139" httpUri="https://zenodo.org/record/5241139/files/figure.png" pageId="10" pageNumber="11">Figs. 4, 6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3290AF64402FFFACE8E735FFD8BFA49" box="[151,552,1389,1412]" pageId="10" pageNumber="11" type="reference_group">
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<paragraph id="8B8C597D4402FFFACE8E735FFD8BFA49" blockId="10.[151,552,1389,1412]" box="[151,552,1389,1412]" pageId="10" pageNumber="11">
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<treatmentCitationGroup id="AB237E534402FFFACE8E735FFD8BFA49" box="[151,552,1389,1412]" pageId="10" pageNumber="11">
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<treatmentCitation id="0A927F6C4402FFFACE8E735FFD87FA49" author="Navas, L." box="[151,548,1389,1412]" page="19" pageId="10" pageNumber="11" year="1931">
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<taxonomicName id="4C3322FE4402FFFACE8E735FFD87FA49" authority="Navas, 1931: 19" authorityName="Navas" authorityPageNumber="19" authorityYear="1931" box="[151,548,1389,1412]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dentata">
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<emphasis id="B947856F4402FFFACE8E735FFED0FA49" box="[151,371,1389,1412]" italics="true" pageId="10" pageNumber="11">Teloganodes dentata</emphasis>
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<bibRefCitation id="EFA2248C4402FFFACF63735FFD87FA49" author="Navas, L." box="[378,548,1389,1412]" pageId="10" pageNumber="11" pagination="16 - 23" refId="ref26191" refString="Navas, L. (1931) Insectos de la India. 4 a serie. Revista de la Academia de Ciencias de Zaragoza, 15, 16 - 23." type="journal article" year="1931">Navás, 1931:19</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3290AF64402FFFACE8E7387FC15F84F" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph id="8B8C597D4402FFFACE8E7387FF47F972" blockId="10.[151,1436,1461,1727]" pageId="10" pageNumber="11">
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<emphasis id="B947856F4402FFFACE8E7387FE31FA02" bold="true" box="[151,402,1461,1487]" pageId="10" pageNumber="11">Material examined.</emphasis>
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In the collections of the
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<collectionCode id="ED22C1B84402FFFACCC67387FC9AFA02" box="[735,825,1461,1487]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="11">MNHN</collectionCode>
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is housed the
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<typeStatus id="5488E7DF4402FFFACDE57387FBC1FA02" box="[1020,1122,1461,1487]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="4C3322FE4402FFFACA8E7384FA3FFA02" authorityName="Navas" authorityYear="1931" box="[1175,1436,1462,1487]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dentatus">
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<emphasis id="B947856F4402FFFACA8E7384FA3FFA02" box="[1175,1436,1462,1487]" italics="true" pageId="10" pageNumber="11">Teloganodes dentatus</emphasis>
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</taxonomicName>
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described by Navás in 1931. According to the ICZN, the ending –
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<emphasis id="B947856F4402FFFACDB673ECFC46FA3A" box="[943,997,1502,1527]" italics="true" pageId="10" pageNumber="11">odes</emphasis>
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is masculine; therefore the adjective used by Navás to name his species should be spelled
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<taxonomicName id="4C3322FE4402FFFACD067034FC21F9D2" authorityName="Navas" authorityYear="1931" box="[799,898,1542,1567]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dentatus">
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<emphasis id="B947856F4402FFFACD067034FC21F9D2" box="[799,898,1542,1567]" italics="true" pageId="10" pageNumber="11">dentatus</emphasis>
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</taxonomicName>
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and not
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<emphasis id="B947856F4402FFFACDF67034FBECF9D2" box="[1007,1103,1542,1567]" italics="true" pageId="10" pageNumber="11">
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<taxonomicName id="4C3322FE4402FFFACDF67034FBE8F9D2" authorityName="Navas" authorityYear="1931" box="[1007,1099,1542,1567]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dentata">dentata</taxonomicName>
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.
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</emphasis>
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The species is described in detail and illustrated. Although Navás is supposed to have seen several specimens (“varios ejemplares”) collected in Khandala (
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<collectingCountry id="F32419ED4402FFFACF9B7067FE62F9A2" box="[386,449,1621,1647]" name="India" pageId="10" pageNumber="11">India</collectingCountry>
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)
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<date id="FF8D7FBD4402FFFACFC87067FD3BF9A2" box="[465,664,1621,1647]" pageId="10" pageNumber="11" value="1928-05-20" valueMax="1928-05-22" valueMin="1928-05-20">20–22 May 1928</date>
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and
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<date id="FF8D7FBD4402FFFACCCB7067FC39F9A2" box="[722,922,1621,1647]" pageId="10" pageNumber="11" value="1927-10-14">14 October 1927</date>
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, a single specimen is housed in Paris. It is a pinned male imago in perfect condition; no piece broken except the tip of the cerci. It bears the following labels:
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</paragraph>
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<paragraph id="8B8C597D4402FFFACE8E70C2FCCFF8C7" blockId="10.[151,1046,1776,1922]" box="[151,876,1776,1802]" pageId="10" pageNumber="11">
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i) Khandala (
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<collectingCountry id="F32419ED4402FFFACF4170C2FE3BF8C7" box="[344,408,1776,1802]" name="India" pageId="10" pageNumber="11">India</collectingCountry>
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) 14.X. [19]27 [white label handwritten]
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</paragraph>
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<paragraph id="8B8C597D4402FFFACE8E712AFBB5F8FF" blockId="10.[151,1046,1776,1922]" box="[151,1046,1816,1842]" pageId="10" pageNumber="11">
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ii)
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<taxonomicName id="4C3322FE4402FFFACEC1712AFDA5F8FF" authority="Nav." authorityName="Navas" authorityYear="1931" box="[216,518,1816,1842]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dentata">Teloganodes dentata Nav.</taxonomicName>
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[handwritten] P. Navás S.J. det [typewritten]
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</paragraph>
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<paragraph id="8B8C597D4402FFFACE8E7172FD90F897" blockId="10.[151,1046,1776,1922]" box="[151,563,1856,1882]" pageId="10" pageNumber="11">
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iii)
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<typeStatus id="5488E7DF4402FFFACEC17172FEBCF897" box="[216,287,1856,1882]" pageId="10" pageNumber="11">Typus</typeStatus>
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[red label handwritten]
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</paragraph>
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<paragraph id="8B8C597D4402FFFACE8E715AFC15F84F" blockId="10.[151,1046,1776,1922]" box="[151,950,1896,1922]" pageId="10" pageNumber="11">iv) Museum Paris Longin Navas legit 19 [white label typewritten]</paragraph>
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</subSubSection>
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<subSubSection id="C3290AF64402FFFBCEDF7182FB41FE6F" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
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<paragraph id="8B8C597D4402FFFBCEDF7182FB41FE6F" blockId="10.[151,1436,1968,2038]" lastBlockId="11.[151,1436,152,418]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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<emphasis id="B947856F4402FFFACEDF7182FEFDF807" bold="true" box="[198,350,1968,1994]" pageId="10" pageNumber="11">Description.</emphasis>
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This specimen shares most of the characters found in
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<taxonomicName id="4C3322FE4402FFFACDED7182FBF5F804" baseAuthorityName="Hagen" baseAuthorityYear="1858" box="[1012,1110,1968,1993]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="tristis">
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<emphasis id="B947856F4402FFFACDED7182FBF5F804" box="[1012,1110,1968,1993]" italics="true" pageId="10" pageNumber="11">T. tristis</emphasis>
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</taxonomicName>
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, particularly: upper part of compound eyes reddish brown, vein MP
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<subScript id="17B75B384402FFFACC6771DAFD25F83B" attach="left" box="[638,646,2024,2038]" fontSize="6" pageId="10" pageNumber="11">2</subScript>
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of forewing long,
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<specimenCount id="9D3592F44402FFFACD7271EAFBA1F83F" box="[875,1026,2008,2034]" pageId="10" pageNumber="11" type="intercalary">3 intercalary</specimenCount>
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veins in cubital area of forewing, hindwing broad (
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<figureCitation id="130845F84403FFFBCF4776AAFE05FF7F" box="[350,422,152,178]" captionStart="FIGURES 1–6" captionStartId="9.[151,269,1864,1888]" captionTargetBox="[251,1344,188,1838]" captionTargetId="figure-98@9.[240,1345,185,1848]" captionTargetPageId="9" captionText="FIGURES 1–6. Teloganodes spp. Fig. 1: Forewing of the female subimago lectotype of Teloganodes tristis (Hagen) [MCZ]; Fig. 2: Forewing of a male subimago syntype of Teloganodes major Eaton [MCZ]. Fig. 3: Hindwing of the female lectotype of Teloganodes tristis (Hagen) [MCZ]; Fig. 4: Hindwing of the male holotype of Teloganodes dentatus Navás [MNHN]; Fig. 5: Genitalia of a male subimago syntype of Teloganodes major Eaton [MCZ]; Fig. 6: Genitalia in ventral view of the male holotype of T. dentatus Navás [MNHN]." figureDoi="http://doi.org/10.5281/zenodo.5241139" httpUri="https://zenodo.org/record/5241139/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
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), no posterior processes on mesoscutellum, gill socket vestiges on segments II–VI, terminal filament absent, cerci whitish with purple rings. The forelegs have the following proportion: tibia ca.1.8 times femur length and 1.3 times tarsi length. It mainly differs from
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<taxonomicName id="4C3322FE4403FFFBCDAC76DAFBB7FECC" baseAuthorityName="Hagen" baseAuthorityYear="1858" box="[949,1044,232,257]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tristis">
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<emphasis id="B947856F4403FFFBCDAC76DAFBB7FECC" box="[949,1044,232,257]" italics="true" pageId="11" pageNumber="12">T. tristis</emphasis>
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</taxonomicName>
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by the darker colour of the abdomen, and shape of the costal process of the hind wing. Genitalia (
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<figureCitation id="130845F84403FFFBCD897722FC75FEE7" box="[912,982,272,298]" captionStart="FIGURES 1–6" captionStartId="9.[151,269,1864,1888]" captionTargetBox="[251,1344,188,1838]" captionTargetId="figure-98@9.[240,1345,185,1848]" captionTargetPageId="9" captionText="FIGURES 1–6. Teloganodes spp. Fig. 1: Forewing of the female subimago lectotype of Teloganodes tristis (Hagen) [MCZ]; Fig. 2: Forewing of a male subimago syntype of Teloganodes major Eaton [MCZ]. Fig. 3: Hindwing of the female lectotype of Teloganodes tristis (Hagen) [MCZ]; Fig. 4: Hindwing of the male holotype of Teloganodes dentatus Navás [MNHN]; Fig. 5: Genitalia of a male subimago syntype of Teloganodes major Eaton [MCZ]; Fig. 6: Genitalia in ventral view of the male holotype of T. dentatus Navás [MNHN]." figureDoi="http://doi.org/10.5281/zenodo.5241139" httpUri="https://zenodo.org/record/5241139/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
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) resemble the subimaginal genitalia of
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<taxonomicName id="4C3322FE4403FFFBCE8E770AFF54FE9C" baseAuthorityName="Hagen" baseAuthorityYear="1858" box="[151,247,312,337]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tristis">
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<emphasis id="B947856F4403FFFBCE8E770AFF54FE9C" box="[151,247,312,337]" italics="true" pageId="11" pageNumber="12">T. tristis</emphasis>
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</taxonomicName>
|
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(as in
|
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<figureCitation id="130845F84403FFFBCF51770AFE2CFE9F" box="[328,399,312,338]" captionStart="FIGURES 1–6" captionStartId="9.[151,269,1864,1888]" captionTargetBox="[251,1344,188,1838]" captionTargetId="figure-98@9.[240,1345,185,1848]" captionTargetPageId="9" captionText="FIGURES 1–6. Teloganodes spp. Fig. 1: Forewing of the female subimago lectotype of Teloganodes tristis (Hagen) [MCZ]; Fig. 2: Forewing of a male subimago syntype of Teloganodes major Eaton [MCZ]. Fig. 3: Hindwing of the female lectotype of Teloganodes tristis (Hagen) [MCZ]; Fig. 4: Hindwing of the male holotype of Teloganodes dentatus Navás [MNHN]; Fig. 5: Genitalia of a male subimago syntype of Teloganodes major Eaton [MCZ]; Fig. 6: Genitalia in ventral view of the male holotype of T. dentatus Navás [MNHN]." figureDoi="http://doi.org/10.5281/zenodo.5241139" httpUri="https://zenodo.org/record/5241139/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
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), i.e. gonopods 3-segmented, the first segment slightly longer than the second, the third one about two times as long as wide. Penis lobes fused for entire length except the apex “U” shaped. Ventrally a groove is visible that ends at the middle of the penes. The styliger plate is markedly convex.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |