174 lines
18 KiB
XML
174 lines
18 KiB
XML
<document id="4A027F66248B457B168BBCE1BA9C65AD" ID-DOI="10.3897/zookeys.1209.125897" ID-publisher-id="125897" URI-arpha="6ED9C06E-4DBB-5A79-A98B-FE427E12ED58" URI-zoobank="http://zoobank.org/CEE78803-61EB-40CC-8D63-46142E6383A" article-type="research-article" checkinTime="1723117912200" checkinUser="pensoft" docAuthor="Harden, Curt W. & Caterino, Michael S." docDate="2024" docId="DFBBD7522F2E5B979AC2FE02A8F5D5C2" docLanguage="en" docName="ZooKeys 1209: 69-197" docOrigin="ZooKeys 1209" docSource="https://zookeys.pensoft.net/article/125897/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Anillinus montrex Harden & Caterino 2024" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="197" masterDocId="6ED9C06E4DBB5A79A98BFE427E12ED58" masterDocTitle="Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)" masterLastPageNumber="197" masterPageNumber="69" pageNumber="69" updateTime="1723118642100" updateUser="ExternalLinkService">
|
||
<mods:mods id="61E4C26C79566557772835BC4656C541" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="EA74393F51E1AEDFD95F2CD522A4BA9C">
|
||
<mods:title id="6C2585812B13377FF3C1A3B2AC9E8053">Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="905679FBA3A94D2FC42260C2F745AA9F" type="personal">
|
||
<mods:role id="1237B1C1E38BB6E2613FBFA822F9F31B">
|
||
<mods:roleTerm id="7258ADAC7517A6AC36943FB9E4D2E875">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="12EE59E161C390735F8DE319835AE678">Harden, Curt W.</mods:namePart>
|
||
<mods:nameIdentifier id="5328A8456DBFF51F8B86AFF3FC6C2C0C" type="ORCID">0000-0002-5226-8725</mods:nameIdentifier>
|
||
<mods:affiliation id="F828E0056075777BD0478C98FE363DF7">Department of Plant and Environmental Sciences, Clemson University, 277 Poole Agricultural Center, Clemson, South Carolina, 29634, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="B2645DAC61FE3D4714F63E8BDBAA0691" type="personal">
|
||
<mods:role id="D692C263980BB4354BAB8BC9AAD54BAC">
|
||
<mods:roleTerm id="7162FC4056BF4FCD841D5CDF49E4255C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B5A524E66C7ABA6BFE13457C1DD4196C">Caterino, Michael S.</mods:namePart>
|
||
<mods:nameIdentifier id="64ACEEC749A940D8C69308B2FC1C60A9" type="ORCID">0000-0002-2597-5707</mods:nameIdentifier>
|
||
<mods:affiliation id="38EC3CBD07FEF53616F88507D6BFA44E">Department of Plant and Environmental Sciences, Clemson University, 277 Poole Agricultural Center, Clemson, South Carolina, 29634, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="8B4A8AE59EB32332A16BF3FB7098332F">text</mods:typeOfResource>
|
||
<mods:relatedItem id="638A02E2E3BE34C9326B8FCC95DB478A" type="host">
|
||
<mods:titleInfo id="E4CB70F3541413FD935581681311777F">
|
||
<mods:title id="8F0AD28EFF576B5466226FFEEB0AD3A4">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="1F14E2555DA1E400C6C92A951A90883D">
|
||
<mods:date id="EE6BD8AD4840A5F9DAFCAB8D8ED70EF4">2024</mods:date>
|
||
<mods:detail id="94F39342C888EB851D2F24FED1EC2033" type="pubDate">
|
||
<mods:number id="367B793DF2216ED61C14DC5256AA9C75">2024-08-08</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="5675BCFF8870D5D19C125A983BED9729" type="volume">
|
||
<mods:number id="44690FA1C0FE0522844FDE62D51785FA">1209</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="3400758D0EFF7127C5530B2B441058A3" unit="page">
|
||
<mods:start id="DBA7AEED86561860737089A4F75AE6A2">69</mods:start>
|
||
<mods:end id="053391CD0EC4D068A35CAB58ECF1B161">197</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="D405C7225217F3A633B62B8E30F272B3">journal article</mods:classification>
|
||
<mods:identifier id="A48897190A824B409262D6552D5C50C2" type="DOI">10.3897/zookeys.1209.125897</mods:identifier>
|
||
<mods:identifier id="511CD460188245F4B8F7361F065ED9BE" type="ZooBank">CEE78803-61EB-40CC-8D63-46142E6383A1</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="DFBBD7522F2E5B979AC2FE02A8F5D5C2" ID-DOI="http://doi.org/10.5281/zenodo.13270395" ID-Zenodo-Dep="13270395" ID-arpha="DFBBD752-2F2E-5B97-9AC2-FE02A8F5D5C2" LSID="urn:lsid:plazi:treatment:DFBBD7522F2E5B979AC2FE02A8F5D5C2" httpUri="http://treatment.plazi.org/id/DFBBD7522F2E5B979AC2FE02A8F5D5C2">
|
||
<subSubSection id="6AE8BA2B85493682E0FBF41FC9C687B1" type="nomenclature">
|
||
<paragraph id="3733FFE339DEC78DBB2277A821302839">
|
||
<taxonomicName id="48DB03C1BCBB23D26692D753FCBF71DB" authority="Harden & Caterino, 2024" authorityName="Harden & Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="montrex">
|
||
<emphasis id="3F0B16ACFE86BAA29B97BE67971E0FBE" italics="true">Anillinus montrex</emphasis>
|
||
<bibRefCitation id="3216EC113050E3260B3A744D68D8F42F" DOI="10.1093/isd/ixae009" author="Caterino MS & Harden CW" refId="B15" refString="Caterino MS, Harden CW (2024) A review of Appalachian Dasycerus Brongniart and the recognition of cryptic diversity within Dasycerus carolinensis (Coleoptera: Staphylinidae: Dasycerinae). Insect Systematics and Diversity 8 (2) [ixae 009]: 1 - 22. https: // doi. org / 10.1093 / isd / ixae 009" year="2024">Harden & Caterino, 2024</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4DF2CE82333FFF3812F622FBF18F2F2A" type="description">
|
||
<paragraph id="C9E210B7138F55E887059BD9F1020111">
|
||
<figureCitation id="C6531DE189E4A41C02DDF7649D6453EB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of eastern Nearctic Anillini. A Serranillus dunavani (South Carolina, Oconee Co.) B Anillinus moseleyae (Tennessee, Sevier Co.) C Anillinus dentatus (South Carolina, Abbeville Co.) D Serranillus jeanneli (South Carolina, Oconee Co.) E Anillinus folkertsioides (Alabama, Blount Co.) F Anillinus pecki (North Carolina, Avery Co.) G Anillinus sp. “ Georgia, Barnes Creek sp. 1 ” H Anillinus montrex (South Carolina, York Co.) I Anillinus sp. “ Tennessee, Kings Saltpeter Cave ” J Anillinus virginiae (Virginia, Bath Co.) K Anillinus erwini (Virginia, Grayson Co.) L Serranillus sp. (North Carolina, Graham Co.) M Anillinus sp. “ South Carolina, Waldrop Stone ” (South Carolina, Pickens Co.) N Anillinus sp. “ Alabama, Horseshoe Cave sp. 1 ” O Anillinus loweae (North Carolina, Haywood Co.) P Anillinus sp. “ Kentucky, Hestand sp. 1 ” Q Anillinus indianae (Indiana, Lawrence Co.) R Anillinus cf. barberi (West Virginia, Greenbrier Co.) S Anillinus hirsutus (Alabama, Madison Co.). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1209.125897.figure2" httpUri="https://binary.pensoft.net/fig/1110341">Figs 2 H</figureCitation>
|
||
,
|
||
<figureCitation id="99B73DCB0A8B90AB29BFA64A4F282816" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. SEM micrographs of male protarsi A Anillinus jancae, left protarsus, dorsal aspect B Anillinus mica, left protarsus, dorsal aspect C Anillinus montrex left protarsus, dorsal aspect D A. jancae, right protarsus, ventral aspect E Anillinus choestoea, right protarsus, ventral aspect F Anillinus dentatus, right protarsus, ventral aspect G Serranillus dunavani (Jeannel), right protarsus, ventral aspect H Anillinus murrayae Sokolov & Carlton, right protarsus, ventral aspect I A. montrex, right protarsus, ventral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure10" httpUri="https://binary.pensoft.net/fig/1110349">10 C, I</figureCitation>
|
||
,
|
||
<figureCitation id="166ED84AC9F72D550D71576EBB1CEA4B" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure21" httpUri="https://binary.pensoft.net/fig/1110360">21 L</figureCitation>
|
||
,
|
||
<figureCitation id="11084587587A0D83FFC78F640F926335" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1209.125897.figure24" httpUri="https://binary.pensoft.net/fig/1110363">24 L</figureCitation>
|
||
,
|
||
<figureCitation id="450F9E9A90F5D61879F1595DFDFFB70B" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Dorsal habitus of Anillinus species in the elongatus group and sinuaticollis group A Anillinus montrex B Anillinus arenicollis C Anillinus choestoea sp. nov. D Anillinus seneca sp. nov. E Anillinus mica sp. nov. F Anillinus micamicus sp. nov. Abdomens removed in B – F for DNA extraction. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1209.125897.figure34" httpUri="https://binary.pensoft.net/fig/1110373">34 A</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5CE911C364DB0B08BB13C7CA95152EAB" type="reference_group">
|
||
<paragraph id="9E6D376FB425B2636D3472B17D999135">
|
||
<treatmentCitationGroup id="CEEECF4CC084154375C5B2851353055C">
|
||
<treatmentCitation id="744F1FA50FA81BBFC546F10325132EB7" author="Caterino MS & Harden CW" page="25" year="2024">
|
||
<taxonomicName id="D7824F3FF6B09F07698628940BDA0F24" authority="Harden & Caterino, 2024: 25" authorityName="Harden & Caterino" authorityPageNumber="25" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="montrex">
|
||
<emphasis id="C6780DC192DCEF55D677C307E8E284D5" italics="true">Anillinus montrex</emphasis>
|
||
<comment id="6E7B4B1A69834DC6107124DF545C3E24">
|
||
<bibRefCitation id="4C66D214F717753F08D5FFF898EBDC60" DOI="10.1093/isd/ixae009" author="Caterino MS & Harden CW" refId="B15" refString="Caterino MS, Harden CW (2024) A review of Appalachian Dasycerus Brongniart and the recognition of cryptic diversity within Dasycerus carolinensis (Coleoptera: Staphylinidae: Dasycerinae). Insect Systematics and Diversity 8 (2) [ixae 009]: 1 - 22. https: // doi. org / 10.1093 / isd / ixae 009" year="2024">Harden & Caterino, 2024: 25</bibRefCitation>
|
||
.
|
||
</comment>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EJRDM" type="GenBank">
|
||
<paragraph id="09BC062BBB4F60BCB77456B54395DBC4">
|
||
<heading id="F9BFBC43FD0E2250EB1035E00B6FC9DE" reason="title">GenBank.</heading>
|
||
</paragraph>
|
||
<paragraph id="E272C313C096D4749FBD53FB77FED468">
|
||
<ext-link id="52FBFD481AE1AF0D3D41C49D2EB4EF28" ext-link-type="gen" xlink_href="OR853113" xlink_type="simple">OR 853113</ext-link>
|
||
,
|
||
<ext-link id="F16D0F607A46D0A9992CD617930D29D9" ext-link-type="gen" xlink_href="OR853294" xlink_type="simple">OR 853294</ext-link>
|
||
,
|
||
<ext-link id="B89F8321715216F1EDD45846A1A56127" ext-link-type="gen" xlink_href="OR839300" xlink_type="simple">OR 839300</ext-link>
|
||
,
|
||
<ext-link id="65E3C5438A050EBA72483D61FC489996" ext-link-type="gen" xlink_href="OR839565" xlink_type="simple">OR 839565</ext-link>
|
||
,
|
||
<ext-link id="0F8ADB3E6AA6916F48BBBB69F5A7005D" ext-link-type="gen" xlink_href="OR837884" xlink_type="simple">OR 837884</ext-link>
|
||
,
|
||
<ext-link id="881AE35FE4B9872B23C2EF093DC26034" ext-link-type="gen" xlink_href="OR838029" xlink_type="simple">OR 838029</ext-link>
|
||
,
|
||
<ext-link id="42FD7FE17EE26CDDC9523E55D94DEAB5" ext-link-type="gen" xlink_href="OR838218" xlink_type="simple">OR 838218</ext-link>
|
||
,
|
||
<ext-link id="45F02FC9269DE28DE7D5456939823532" ext-link-type="gen" xlink_href="OR838127" xlink_type="simple">OR 838127</ext-link>
|
||
,
|
||
<ext-link id="BDE46FE7E89CF40EC18D2587DBF6B9D5" ext-link-type="gen" xlink_href="OR839566" xlink_type="simple">OR 839566</ext-link>
|
||
,
|
||
<ext-link id="3211397FD4B9FDCF7BC69F7B1F629530" ext-link-type="gen" xlink_href="OR839299" xlink_type="simple">OR 839299</ext-link>
|
||
,
|
||
<ext-link id="57E5E47D886AD5FC177DA45752D9495F" ext-link-type="gen" xlink_href="OR839567" xlink_type="simple">OR 839567</ext-link>
|
||
,
|
||
<ext-link id="46E73ADA0CFD3FC8738710F6783400EA" ext-link-type="gen" xlink_href="OR837885" xlink_type="simple">OR 837885</ext-link>
|
||
,
|
||
<ext-link id="91C714E7B5C03038AA8F546C59B4E2B1" ext-link-type="gen" xlink_href="OR838030" xlink_type="simple">OR 838030</ext-link>
|
||
,
|
||
<ext-link id="B79C5A049298AF715BEA112A4946C7A1" ext-link-type="gen" xlink_href="OR838219" xlink_type="simple">OR 838219</ext-link>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EVTDM" type="diagnosis">
|
||
<paragraph id="AF331961DE67C97BB60378FF796E2EFB">
|
||
<heading id="B263868A86EB44B10B30F6280D4F44DD" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="71CC9B754CD8DF32A36F0A6811EB6122">
|
||
Members of this species are strongly flattened dorsoventrally, narrow, and parallel sided (Fig.
|
||
<figureCitation id="2B757C5FB9E51107840EC687D2AC248E" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Dorsal habitus of Anillinus species in the elongatus group and sinuaticollis group A Anillinus montrex B Anillinus arenicollis C Anillinus choestoea sp. nov. D Anillinus seneca sp. nov. E Anillinus mica sp. nov. F Anillinus micamicus sp. nov. Abdomens removed in B – F for DNA extraction. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1209.125897.figure34" httpUri="https://binary.pensoft.net/fig/1110373">34 A</figureCitation>
|
||
). Males have the first and second protarsomeres expanded and spinose on inner margin with ventral adhesive setae; the male second protarsomeres of this species are more enlarged than any other known
|
||
<taxonomicName id="7B356A92E3D610DA1142D1EF81E24DA4">
|
||
<emphasis id="940255E0AB4D3C3A780631D8B15AA859" italics="true">Anillinus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="8904F97A34A3FA92232D1EBB73048403" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. SEM micrographs of male protarsi A Anillinus jancae, left protarsus, dorsal aspect B Anillinus mica, left protarsus, dorsal aspect C Anillinus montrex left protarsus, dorsal aspect D A. jancae, right protarsus, ventral aspect E Anillinus choestoea, right protarsus, ventral aspect F Anillinus dentatus, right protarsus, ventral aspect G Serranillus dunavani (Jeannel), right protarsus, ventral aspect H Anillinus murrayae Sokolov & Carlton, right protarsus, ventral aspect I A. montrex, right protarsus, ventral aspect. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure10" httpUri="https://binary.pensoft.net/fig/1110349">10 C, I</figureCitation>
|
||
). The male genitalia are also distinctive, with a small apex that is abruptly bisected by the membranous dorsal margin. Female spermatheca (Fig.
|
||
<figureCitation id="41F3A46EDD10F80168F800AA7FB1176A" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure21" httpUri="https://binary.pensoft.net/fig/1110360">21 L</figureCitation>
|
||
) with stem coiled proximally, abruptly enlarged distally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EOUDM" type="distribution">
|
||
<paragraph id="88B5C655B33827368F10BF8D8F422514">
|
||
<heading id="9B7652A53971B1C110A67BCF506F4F91" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="404BFE477BDDD85B32B024CA0895FED9">
|
||
Known for certain only from a single hillside above a small stream in Kings Mountain State Park,
|
||
<collectingRegion id="CF8988A0A9FEF4D4F9A86D976C38D358" country="United Kingdom" name="York">York</collectingRegion>
|
||
Co., SC.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ETUDM" type="Sympatry">
|
||
<paragraph id="C5189B6136786EB0A3FA95758A4F2626">
|
||
<heading id="48FEF35DB8B26CE1EFCDF5C28187EC4F" reason="title">Sympatry.</heading>
|
||
</paragraph>
|
||
<paragraph id="958AB41913810CA6ABC4A05C675DEAF3">
|
||
Members of this species have been collected under rocks with
|
||
<taxonomicName id="6EFD0237929C53644B2666703FEF21B3" authorityName="Harden & Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="simplex" status="sp. nov.">
|
||
<emphasis id="EDE07F69EBE1D8DE10FBE9E5D50499A9" italics="true">A. simplex</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DAC0C3CD8E0AD8710CFADCCA0C92371D" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EEVDM" type="natural history">
|
||
<paragraph id="058CBF6CD238B11ACAAA2E99597EAE9A">
|
||
<heading id="D557FC14EAAEE4B6EBA51F442814BE57" reason="title">Natural history.</heading>
|
||
</paragraph>
|
||
<paragraph id="D9BD5C9A15F7A1C3DF8F12516543ACE0">Members of this species are endogean in habit, occurring under deeply embedded rocks at cold times of year. Specimens have been collected in December.</paragraph>
|
||
<paragraph id="711BDF4FF78E6B0F2F8ED89AD74F8701">
|
||
<title id="85D72956404B053407A98D0C56E07CB3">
|
||
<taxonomicName id="237D27D9CA9E9D2CF2EB2B4AE0D1837A" authorityName="Jeannel" authorityYear="1963" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="sinuaticollis">
|
||
‘
|
||
<emphasis id="A3B1A4204C1FB9B0DCECCFAF95F15B01" italics="true">sinuaticollis</emphasis>
|
||
</taxonomicName>
|
||
group’
|
||
</title>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |