197 lines
31 KiB
XML
197 lines
31 KiB
XML
<document id="D5FE0D6F39A9F802D5883E7CDDFC13A5" ID-CLB-Dataset="27575" ID-DOI="10.11646/zootaxa.4554.1.9" ID-GBIF-Dataset="ff7707f0-058f-4f40-8eb8-2b6c9c75a173" ID-ISSN="1175-5326" ID-Zenodo-Dep="2623565" ID-ZooBank="A804E949-109A-468D-B58B-CF7C8BCB3059" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="diego" checkinTime="1554271225461" checkinUser="plazi" docAuthor="Vandenberg, Natalia J. & Hanson, Paul E." docDate="2019" docId="DF18F971FFD50B3CFF4BFCBF69C8BAFB" docLanguage="en" docName="zootaxa.4554.1.9.pdf" docOrigin="Zootaxa 4554 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Moiradiomus Vandenberg & Hanson 2019" docType="key" docVersion="10" lastPageNumber="278" masterDocId="23218109FFC20B2BFFDCFFF96C54BC27" masterDocTitle="Overview of the lady beetle tribe Diomini (Coleoptera: Coccinellidae) and description of a new phytophagous, silk-spinning genus from Costa Rica that induces food bodies on leaves of Piper (Piperaceae)" masterLastPageNumber="285" masterPageNumber="255" pageNumber="278" updateTime="1728643685052" updateUser="ExternalLinkService">
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<mods:title id="FFD984457EDA3038A5D5E580195595E9">Overview of the lady beetle tribe Diomini (Coleoptera: Coccinellidae) and description of a new phytophagous, silk-spinning genus from Costa Rica that induces food bodies on leaves of Piper (Piperaceae)</mods:title>
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<mods:namePart id="048276B38A919FF363B423F6C9ABDE56">Vandenberg, Natalia J.</mods:namePart>
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<mods:namePart id="DF065223A0E5E772C56110AB7E9E9D42">Hanson, Paul E.</mods:namePart>
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<mods:title id="107F407863A885A93E548AC61E3FD554">Zootaxa</mods:title>
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<mods:part id="6EA914F601E7C8F86703960C23A6FF0D">
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<mods:date id="53BB7415C93D8BC97B8A8FF15E00ED42">2019</mods:date>
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<mods:number id="048907B629D9AC10C16C511B54BE4B83">2019-02-08</mods:number>
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<mods:number id="6A1043197BD871E5C83208C6B0A65F7A">4554</mods:number>
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<mods:identifier id="DFED6D2587C5CD6D86B88E3B0FCB709D" type="DOI">10.11646/zootaxa.4554.1.9</mods:identifier>
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<treatment id="DF18F971FFD50B3CFF4BFCBF69C8BAFB" ID-DOI="http://doi.org/10.5281/zenodo.5921894" ID-GBIF-Taxon="154535007" ID-Zenodo-Dep="5921894" LSID="urn:lsid:plazi:treatment:DF18F971FFD50B3CFF4BFCBF69C8BAFB" httpUri="http://treatment.plazi.org/id/DF18F971FFD50B3CFF4BFCBF69C8BAFB" lastPageNumber="278" pageId="23" pageNumber="278">
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<subSubSection id="1FAB1BECFFD50B3CFF4BFCBF6F8EB835" pageId="23" pageNumber="278" type="nomenclature">
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<paragraph id="570E4867FFD50B3CFF4BFCBF6E44BF47" blockId="23.[151,528,838,864]" box="[151,528,838,864]" pageId="23" pageNumber="278">
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<heading id="0C46FF0BFFD50B3CFF4BFCBF6E44BF47" bold="true" box="[151,528,838,864]" fontSize="11" level="1" pageId="23" pageNumber="278" reason="1">
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<emphasis id="65C59475FFD50B3CFF4BFCBF6E44BF47" bold="true" box="[151,528,838,864]" pageId="23" pageNumber="278">
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Key to species of
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<taxonomicName id="90B133E4FFD50B3CFEB1FCBF6E44BF47" ID-CoL="CG2R4" authorityName="Vandenberg & Hanson" authorityYear="2019" box="[365,528,838,864]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="278" phylum="Arthropoda" rank="genus">
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<emphasis id="65C59475FFD50B3CFEB1FCBF6E44BF47" bold="true" box="[365,528,838,864]" italics="true" pageId="23" pageNumber="278">Moiradiomus</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="570E4867FFD50B3CFF4BFC746F8EB835" blockId="23.[151,1436,909,1042]" pageId="23" pageNumber="278">
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The species of
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<taxonomicName id="90B133E4FFD50B3CFE9BFC746D8ABF81" authorityName="Vandenberg & Hanson" authorityYear="2019" box="[327,478,909,934]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="278" phylum="Arthropoda" rank="genus">
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<emphasis id="65C59475FFD50B3CFE9BFC746D8ABF81" box="[327,478,909,934]" italics="true" pageId="23" pageNumber="278">Moiradiomus</emphasis>
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</taxonomicName>
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can be readily separated by characteristics of the male genitalia and the species of larval host plant. Additional morphological and color differences are added to the key as confidence characters, but we project that these will prove insufficient for identification purposes, particularly considering our small sample sizes and the potentially large number of undescribed species in the genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="1FAB1BECFFD50B3CFF4BFBB869C8BAFB" pageId="23" pageNumber="278" type="key">
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<keyStep id="EC4550C2FFD50B3CFF4BFBB869CFB904" pageId="23" pageNumber="278">
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<paragraph id="570E4867FFD50B3CFF4BFBB869C8B888" blockId="23.[151,1436,1089,1756]" pageId="23" pageNumber="278">
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<keyLead id="EC40EB52FFD50B3CFF4BFBB869C8B888" pageId="23" pageNumber="278">
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1. Male genitalia with short, broad paramere (length only 2× width) (
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<figureCitation id="CF8A54E2FFD50B3CFCA8FBB86F9DB871" box="[884,969,1089,1110]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="23" pageNumber="278">Fig. 32a</figureCitation>
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); basal lobe in ventral view (
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<figureCitation id="CF8A54E2FFD50B3CFB36FBB8696AB871" box="[1258,1342,1089,1110]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="23" pageNumber="278">Fig. 32b</figureCitation>
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) strongly tapered toward apex with rounded apical knob. Penis tube (
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<figureCitation id="CF8A54E2FFD50B3CFCFFFBA76F20B854" box="[803,884,1118,1139]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="23" pageNumber="278">Fig. 32c</figureCitation>
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) with slight angulation and irregular swelling near distal ½. Larvae develop on food bodies of
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<taxonomicName id="90B133E4FFD50B3CFD98FB856E9CB8B6" box="[580,712,1147,1169]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="23" pageNumber="278" phylum="Tracheophyta" rank="species" species="reticulatum">
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<emphasis id="65C59475FFD50B3CFD98FB856E9CB8B6" box="[580,712,1147,1169]" italics="true" pageId="23" pageNumber="278">P. reticulatum</emphasis>
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</taxonomicName>
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. Adults proportionally shorter (
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<figureCitation id="CF8A54E2FFD50B3CFC2AFB85683CB8B6" box="[1014,1128,1148,1169]" captionStart="FIGURES 17–20" captionStartId="14.[151,264,1965,1988]" captionTargetBox="[151,1436,260,1911]" captionTargetId="figure@14.[151,1436,260,1911]" captionTargetPageId="14" captionText="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.2623573" httpUri="https://zenodo.org/record/2623573/files/figure.png" pageId="23" pageNumber="278">Figs. 19, 20</figureCitation>
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) than other known species, with prosternal carinae (
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<figureCitation id="CF8A54E2FFD50B3CFE45FB606D8BB889" box="[409,479,1177,1198]" captionStart="FIGURES 25–28" captionStartId="15.[151,264,1975,1998]" captionTargetBox="[151,1436,1466,1918]" captionTargetId="figure@15.[151,1436,1461,1924]" captionTargetPageId="15" captionText="FIGURES 25–28. Diagrammatic ventral view of Moiradiomus prosterna (setae, color patterns, and surface punctation not indicated): 25, M. clotho; 26, M. lachesis; 27, M. atopos; 28, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623578" httpUri="https://zenodo.org/record/2623578/files/figure.png" pageId="23" pageNumber="278">Fig. 28</figureCitation>
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) nearly parallel, framing subtrapezoidal depression.........................
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<emphasis id="65C59475FFD50B3CFB25FB606934B889" box="[1273,1376,1177,1198]" italics="true" pageId="23" pageNumber="278">
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<taxonomicName id="90B133E4FFD50B3CFB25FB606908B889" authorityName="Vandenberg & Hanson" authorityYear="2019" box="[1273,1372,1177,1198]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="278" phylum="Arthropoda" rank="species" species="nanita" status="n. sp.">M. nanita</taxonomicName>
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,
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</emphasis>
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<emphasis id="65C59475FFD50B3CFABAFB6069C8B888" bold="true" box="[1382,1436,1177,1199]" pageId="23" pageNumber="278">
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<taxonomicNameLabel id="7EF6290EFFD50B3CFABAFB6069C8B888" box="[1382,1436,1177,1199]" pageId="23" pageNumber="278" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="570E4867FFD50B3CFF4BFB4F69CFB904" blockId="23.[151,1436,1089,1756]" pageId="23" pageNumber="278">
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<keyLead id="EC40EB52FFD50B3CFF4BFB4F69CFB904" pageId="23" pageNumber="278">
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1’. Male genitalia with paramere longer (length approximately 2.5–3.0× width) (
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<figureCitation id="CF8A54E2FFD50B3CFC06FB4F68CDB8EC" box="[986,1177,1206,1227]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="23" pageNumber="278">Figs. 29a, 30a, 31a</figureCitation>
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); basal lobe not as above. Penis tube without angulation and swelling near distal ½ (
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<figureCitation id="CF8A54E2FFD50B3CFCC3FB2D6F8AB8CE" box="[799,990,1236,1257]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="23" pageNumber="278">Figs. 29c, 30c, 31c</figureCitation>
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). Larvae develop on other
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<taxonomicName id="90B133E4FFD50B3CFB35FB2D694AB8CE" box="[1257,1310,1236,1257]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="23" pageNumber="278" phylum="Tracheophyta" rank="genus">
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<emphasis id="65C59475FFD50B3CFB35FB2D694AB8CE" box="[1257,1310,1236,1257]" italics="true" pageId="23" pageNumber="278">Piper</emphasis>
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</taxonomicName>
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hosts. Adult form elongate oval (
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<figureCitation id="CF8A54E2FFD50B3CFE79FB086E4EB921" box="[421,538,1265,1286]" captionStart-0="FIGURES 13–16" captionStart-1="FIGURES 17–20" captionStartId-0="13.[151,264,1957,1980]" captionStartId-1="14.[151,264,1965,1988]" captionTargetBox-0="[151,1436,245,1896]" captionTargetBox-1="[151,1436,260,1911]" captionTargetId-0="figure@13.[151,1436,245,1896]" captionTargetId-1="figure@14.[151,1436,260,1911]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 13–16. Moiradiomus species habitus illustrations. 13, M. clotho: holotype, male; 14, M. clotho: paratype, female. 15, M. lachesis: holotype, male; 16, M. lachesis: paratype, female. Scale bar = 0.5 mm." captionText-1="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.2623571" figureDoi-1="http://doi.org/10.5281/zenodo.2623573" httpUri-0="https://zenodo.org/record/2623571/files/figure.png" httpUri-1="https://zenodo.org/record/2623573/files/figure.png" pageId="23" pageNumber="278">Figs. 13–18</figureCitation>
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), prosternal carinae more or less distinctly converging toward apex, framing subtriangular to ovoid depression (
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<figureCitation id="CF8A54E2FFD50B3CFE4DFAF76E50B904" box="[401,516,1294,1315]" captionStart="FIGURES 25–28" captionStartId="15.[151,264,1975,1998]" captionTargetBox="[151,1436,1466,1918]" captionTargetId="figure@15.[151,1436,1461,1924]" captionTargetPageId="15" captionText="FIGURES 25–28. Diagrammatic ventral view of Moiradiomus prosterna (setae, color patterns, and surface punctation not indicated): 25, M. clotho; 26, M. lachesis; 27, M. atopos; 28, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623578" httpUri="https://zenodo.org/record/2623578/files/figure.png" pageId="23" pageNumber="278">Figs. 25–27</figureCitation>
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)........................................................................... 2
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="EC4550C2FFD50B3CFF4BFAD569CFB991" pageId="23" pageNumber="278">
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<paragraph id="570E4867FFD50B3CFF4BFAD569C8B95B" blockId="23.[151,1436,1089,1756]" pageId="23" pageNumber="278">
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<keyLead id="EC40EB52FFD50B3CFF4BFAD569C8B95B" pageId="23" pageNumber="278">
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2. Male genitalia with basal lobe strongly unevenly tapered in apical half (
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<figureCitation id="CF8A54E2FFD50B3CFC4BFAD56FBFB966" box="[919,1003,1324,1345]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="23" pageNumber="278">Fig. 29b</figureCitation>
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), terminating in hook-shaped process resembling a vulture’s head (
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<figureCitation id="CF8A54E2FFD50B3CFE62FAB06E47B979" box="[446,531,1353,1374]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="23" pageNumber="278">Fig. 29d</figureCitation>
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). Larvae develop on food bodies of
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<taxonomicName id="90B133E4FFD50B3CFCB3FAB06810B979" authorityName="W. C. Burger" authorityYear="1971" box="[879,1092,1352,1374]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="23" pageNumber="278" phylum="Tracheophyta" rank="species" species="holdridgeianum">
|
||
<emphasis id="65C59475FFD50B3CFCB3FAB06810B979" box="[879,1092,1352,1374]" italics="true" pageId="23" pageNumber="278">Piper holdridgeianum</emphasis>
|
||
</taxonomicName>
|
||
. Adult with terminal maxillary palpomere elongate, moderately expanded distally (
|
||
<figureCitation id="CF8A54E2FFD50B3CFD6FFA9F6F71B95C" box="[691,805,1382,1403]" captionStart="FIGURES 13–16" captionStartId="13.[151,264,1957,1980]" captionTargetBox="[151,1436,245,1896]" captionTargetId="figure@13.[151,1436,245,1896]" captionTargetPageId="13" captionText="FIGURES 13–16. Moiradiomus species habitus illustrations. 13, M. clotho: holotype, male; 14, M. clotho: paratype, female. 15, M. lachesis: holotype, male; 16, M. lachesis: paratype, female. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.2623571" httpUri="https://zenodo.org/record/2623571/files/figure.png" pageId="23" pageNumber="278">Figs. 13, 14</figureCitation>
|
||
).......................................
|
||
<taxonomicName id="90B133E4FFD50B3CFB24FA9F690DB95C" authorityName="Vandenberg & Hanson" authorityYear="2019" box="[1272,1369,1381,1403]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="278" phylum="Arthropoda" rank="species" species="clotho" status="n. sp.">
|
||
<emphasis id="65C59475FFD50B3CFB24FA9F690DB95C" box="[1272,1369,1381,1403]" italics="true" pageId="23" pageNumber="278">M. clotho</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="65C59475FFD50B3CFABAFA9F69C8B95B" bold="true" box="[1382,1436,1382,1404]" pageId="23" pageNumber="278">
|
||
<taxonomicNameLabel id="7EF6290EFFD50B3CFABAFA9F69C8B95B" box="[1382,1436,1382,1404]" pageId="23" pageNumber="278" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="570E4867FFD50B3CFF4BFA7D69CFB991" blockId="23.[151,1436,1089,1756]" pageId="23" pageNumber="278">
|
||
<keyLead id="EC40EB52FFD50B3CFF4BFA7D69CFB991" pageId="23" pageNumber="278">
|
||
2.’ Male genitalia with basal lobe nearly parallel-sided in apical half, terminating in oblique apex (
|
||
<figureCitation id="CF8A54E2FFD50B3CFBB6FA7D68A3B9BE" box="[1130,1271,1412,1433]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="23" pageNumber="278">Figs. 30b, 31b</figureCitation>
|
||
). Larvae develop on other
|
||
<taxonomicName id="90B133E4FFD50B3CFEEAFA586D3FB991" box="[310,363,1441,1462]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="23" pageNumber="278" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="65C59475FFD50B3CFEEAFA586D3FB991" box="[310,363,1441,1462]" italics="true" pageId="23" pageNumber="278">Piper</emphasis>
|
||
</taxonomicName>
|
||
hosts. Adult with terminal maxillary palpomere short, strongly expanded distally (
|
||
<figureCitation id="CF8A54E2FFD50B3CFB58FA5868A3B991" box="[1156,1271,1441,1462]" captionStart-0="FIGURES 13–16" captionStart-1="FIGURES 17–20" captionStartId-0="13.[151,264,1957,1980]" captionStartId-1="14.[151,264,1965,1988]" captionTargetBox-0="[151,1436,245,1896]" captionTargetBox-1="[151,1436,260,1911]" captionTargetId-0="figure@13.[151,1436,245,1896]" captionTargetId-1="figure@14.[151,1436,260,1911]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 13–16. Moiradiomus species habitus illustrations. 13, M. clotho: holotype, male; 14, M. clotho: paratype, female. 15, M. lachesis: holotype, male; 16, M. lachesis: paratype, female. Scale bar = 0.5 mm." captionText-1="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.2623571" figureDoi-1="http://doi.org/10.5281/zenodo.2623573" httpUri-0="https://zenodo.org/record/2623571/files/figure.png" httpUri-1="https://zenodo.org/record/2623573/files/figure.png" pageId="23" pageNumber="278">Figs. 15–18</figureCitation>
|
||
)............ 3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="EC4550C2FFD50B3CFF4BFA4769C8BAFB" pageId="23" pageNumber="278">
|
||
<paragraph id="570E4867FFD50B3CFF4BFA4769C8BA6E" blockId="23.[151,1436,1089,1756]" pageId="23" pageNumber="278">
|
||
<keyLead id="EC40EB52FFD50B3CFF4BFA4769C8BA6E" pageId="23" pageNumber="278">
|
||
3. Basal lobe of male genitalia with pronounced dorsal hump at basal 1/3, forming a conspicuous subangulation in lateral view (
|
||
<figureCitation id="CF8A54E2FFD50B3CFF34FA256D68B9D6" box="[232,316,1500,1521]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="23" pageNumber="278">Fig. 31a</figureCitation>
|
||
). Prosternum with strongly convergent carinae framing subtriangular depression with attenuate, narrowly rounded apex (
|
||
<figureCitation id="CF8A54E2FFD50B3CFEC1FA006D32BA29" box="[285,358,1529,1550]" captionStart="FIGURES 25–28" captionStartId="15.[151,264,1975,1998]" captionTargetBox="[151,1436,1466,1918]" captionTargetId="figure@15.[151,1436,1461,1924]" captionTargetPageId="15" captionText="FIGURES 25–28. Diagrammatic ventral view of Moiradiomus prosterna (setae, color patterns, and surface punctation not indicated): 25, M. clotho; 26, M. lachesis; 27, M. atopos; 28, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623578" httpUri="https://zenodo.org/record/2623578/files/figure.png" pageId="23" pageNumber="278">Fig. 27</figureCitation>
|
||
). Larvae develop on food bodies of
|
||
<taxonomicName id="90B133E4FFD50B3CFD16FA006FC3BA29" box="[714,919,1528,1550]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="23" pageNumber="278" phylum="Tracheophyta" rank="species" species="friedrichsthalii">
|
||
<emphasis id="65C59475FFD50B3CFD16FA006FC3BA29" box="[714,919,1528,1550]" italics="true" pageId="23" pageNumber="278">Piper friedrichsthalii</emphasis>
|
||
</taxonomicName>
|
||
. Adult form narrow, tapered toward apex; male with scutellar shield same color as elytron, and base of pronotum in front of shield with light brown transverse mark (
|
||
<figureCitation id="CF8A54E2FFD50B3CFACBF9EF6909BA0C" box="[1303,1373,1558,1579]" captionStart="FIGURES 17–20" captionStartId="14.[151,264,1965,1988]" captionTargetBox="[151,1436,260,1911]" captionTargetId="figure@14.[151,1436,260,1911]" captionTargetPageId="14" captionText="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.2623573" httpUri="https://zenodo.org/record/2623573/files/figure.png" pageId="23" pageNumber="278">Fig. 17</figureCitation>
|
||
)...........................................................................................
|
||
<taxonomicName id="90B133E4FFD50B3CFB31F9CD690DBA6E" authorityName="Vandenberg & Hanson" authorityYear="2019" box="[1261,1369,1588,1609]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="278" phylum="Arthropoda" rank="species" species="atropos" status="n. sp.">
|
||
<emphasis id="65C59475FFD50B3CFB31F9CD690DBA6E" box="[1261,1369,1588,1609]" italics="true" pageId="23" pageNumber="278">M. atropos</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="65C59475FFD50B3CFABAF9CA69C8BA6E" bold="true" box="[1382,1436,1587,1609]" pageId="23" pageNumber="278">
|
||
<taxonomicNameLabel id="7EF6290EFFD50B3CFABAF9CA69C8BA6E" box="[1382,1436,1587,1609]" pageId="23" pageNumber="278" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="570E4867FFD50B3CFF4BF9A869C8BAFB" blockId="23.[151,1436,1089,1756]" pageId="23" pageNumber="278">
|
||
<keyLead id="EC40EB52FFD50B3CFF4BF9A869C8BAFB" pageId="23" pageNumber="278">
|
||
3’. Basal lobe of male genitalia with dorsal surface nearly flat, weakly sinuous in lateral view (
|
||
<figureCitation id="CF8A54E2FFD50B3CFB91F9A868F5BA41" box="[1101,1185,1617,1638]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="23" pageNumber="278">Fig. 30a</figureCitation>
|
||
). Prosternum with weakly convergent carinae framing elongate ovoid depression (
|
||
<figureCitation id="CF8A54E2FFD50B3CFCD4F9976F07BAA4" box="[776,851,1646,1667]" captionStart="FIGURES 25–28" captionStartId="15.[151,264,1975,1998]" captionTargetBox="[151,1436,1466,1918]" captionTargetId="figure@15.[151,1436,1461,1924]" captionTargetPageId="15" captionText="FIGURES 25–28. Diagrammatic ventral view of Moiradiomus prosterna (setae, color patterns, and surface punctation not indicated): 25, M. clotho; 26, M. lachesis; 27, M. atopos; 28, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623578" httpUri="https://zenodo.org/record/2623578/files/figure.png" pageId="23" pageNumber="278">Fig. 26</figureCitation>
|
||
). Larvae develop on food bodies of
|
||
<taxonomicName id="90B133E4FFD50B3CFB1EF99769C1BAA4" box="[1218,1429,1645,1667]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="23" pageNumber="278" phylum="Tracheophyta" rank="species" species="lanosibracteum">
|
||
<emphasis id="65C59475FFD50B3CFB1EF99769C1BAA4" box="[1218,1429,1645,1667]" italics="true" pageId="23" pageNumber="278">Piper lanosibracteum</emphasis>
|
||
</taxonomicName>
|
||
. Adult form less narrow, oval with lateral margins nearly parallel-sided, then rounded off in apical 2/5; male scutellar shield may be paler than elytron and base of pronotum in front of shield without brown transverse mark (
|
||
<figureCitation id="CF8A54E2FFD50B3CFB4CF950688CBA99" box="[1168,1240,1705,1726]" captionStart="FIGURES 13–16" captionStartId="13.[151,264,1957,1980]" captionTargetBox="[151,1436,245,1896]" captionTargetId="figure@13.[151,1436,245,1896]" captionTargetPageId="13" captionText="FIGURES 13–16. Moiradiomus species habitus illustrations. 13, M. clotho: holotype, male; 14, M. clotho: paratype, female. 15, M. lachesis: holotype, male; 16, M. lachesis: paratype, female. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.2623571" httpUri="https://zenodo.org/record/2623571/files/figure.png" pageId="23" pageNumber="278">Fig. 15</figureCitation>
|
||
)......................................................................................................
|
||
<emphasis id="65C59475FFD50B3CFB3AF93F690BBAFC" box="[1254,1375,1733,1755]" italics="true" pageId="23" pageNumber="278">
|
||
<taxonomicName id="90B133E4FFD50B3CFB3AF93F690CBAFC" authorityName="Vandenberg & Hanson" authorityYear="2019" box="[1254,1368,1733,1755]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="278" phylum="Arthropoda" rank="species" species="lachesis">M. lachesis</taxonomicName>
|
||
.,
|
||
</emphasis>
|
||
<emphasis id="65C59475FFD50B3CFABAF93F69C8BAFB" bold="true" box="[1382,1436,1734,1756]" pageId="23" pageNumber="278">n. sp.</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |