415 lines
39 KiB
XML
415 lines
39 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3790424" ID-GBIF-Dataset="5101dbd6-06ac-4b6e-991e-b66eec55ca52" ID-GBIF-Taxon="163750798" ID-ISSN="1313–2970" ID-Zenodo-Dep="3790424" ID-ZooBank="46085EC4-FB49-4272-803A-A0FF21EE52CD" LSID="urn:lsid:zoobank.org:act:C1F51397-8A1C-404B-AAF5-2B68F46C6A53" approvalRequired="22" approvalRequired_for_textStreams="22" checkinTime="1587986115529" checkinUser="plazi" docAuthor="Kraus, Fred & Allison, Allen" docDate="2009" docId="AF1DA83BD457FFE6FF3ADFB844537EBB" docLanguage="en" docName="ZK_article_2096.pdf" docOrigin="ZooKeys 26 (26)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@42210d27" docStyleName="zookeys.2008.journal_article" docTitle="Albericus murritus Kraus & Allison 2009, sp. n." docType="treatment" docUuid="C1F51397-8A1C-404B-AAF5-2B68F46C6A53" docUuidSource="ZooBank" docVersion="6" lastPageId="4" lastPageNumber="57" masterDocId="5324D043D455FFE2FFB6DC144745783C" masterDocTitle="New microhylid Frogs from the Muller Range, Papua New Guinea" masterLastPageNumber="76" masterPageNumber="53" pageId="2" pageNumber="55" updateTime="1643569204779" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New microhylid Frogs from the Muller Range, Papua New Guinea</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kraus, Fred</mods:namePart>
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<mods:affiliation>University of Michigan, Ann Arbor, United States of America</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Allison, Allen</mods:namePart>
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<mods:affiliation>Bishop Museum ,,</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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||
<mods:title>ZooKeys</mods:title>
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||
</mods:titleInfo>
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||
<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-10-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>26</mods:number>
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</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>26</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>53</mods:start>
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<mods:end>76</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.3897/zookeys.26.258</mods:identifier>
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<mods:identifier type="GBIF-Dataset">5101dbd6-06ac-4b6e-991e-b66eec55ca52</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576562</mods:identifier>
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||
<mods:identifier type="ZooBank">46085EC4-FB49-4272-803A-A0FF21EE52CD</mods:identifier>
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||
</mods:mods>
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||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790424" ID-GBIF-Taxon="163750798" ID-Zenodo-Dep="3790424" LSID="urn:lsid:zoobank.org:act:C1F51397-8A1C-404B-AAF5-2B68F46C6A53" httpUri="http://treatment.plazi.org/id/AF1DA83BD457FFE6FF3ADFB844537EBB" lastPageId="4" lastPageNumber="57" pageId="2" pageNumber="55">
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||
<subSubSection pageId="2" pageNumber="55" type="nomenclature">
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||
<paragraph blockId="2.[140,969,940,1038]" box="[140,434,940,967]" pageId="2" pageNumber="55">
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||
<heading box="[140,434,940,967]" fontSize="11" level="2" pageId="2" pageNumber="55" reason="3">
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||
<emphasis bold="true" box="[140,434,940,967]" pageId="2" pageNumber="55">
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<taxonomicName authority="Kraus & Allison, 2009" authorityName="Kraus & Allison" authorityYear="2009" box="[140,362,940,967]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="murritus" status="sp. n.">
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<emphasis bold="true" box="[140,362,940,967]" italics="true" pageId="2" pageNumber="55">
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<emphasis box="[140,249,940,967]" inLineHeading="true" pageId="2" pageNumber="55" reason="1">Albericus</emphasis>
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murritus
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</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[369,434,941,967]" pageId="2" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
<paragraph blockId="2.[140,969,940,1038]" box="[140,969,976,1003]" pageId="2" pageNumber="55">
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||
<uri box="[140,969,976,1003]" pageId="2" pageNumber="55">
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||
urn:lsid:zoobank.org:act:
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||
<uuid box="[421,969,976,1003]" pageId="2" pageNumber="55">C1F51397-8A1C-404B-AAF5-2B68F46C6A53</uuid>
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||
</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[140,222,1011,1038]" pageId="2" pageNumber="55" type="description">
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<paragraph blockId="2.[140,969,940,1038]" box="[140,222,1011,1038]" pageId="2" pageNumber="55">Fig. 1A</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="2" pageNumber="55" type="materials_examined">
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||
<paragraph blockId="2.[140,1108,1082,1672]" pageId="2" pageNumber="55">
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||
<materialsCitation ID-GBIF-Occurrence="2608940806" collectingDate="2009-04-02" collectionCode="BPBM, FK" collectorName="F. Kraus" country="Papua New Guinea" elevation="1700" latitude="-5.656703" location="Tumbutu River" longLatPrecision="1" longitude="142.63423" municipality="Muller Range" pageId="2" pageNumber="55" specimenCode="BPBM 33657, FK 13097" specimenCount="1" stateProvince="Southern Highlands" typeStatus="holotype">
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||
<emphasis bold="true" box="[140,260,1082,1108]" pageId="2" pageNumber="55">
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||
<emphasis box="[140,256,1082,1108]" inLineHeading="true" pageId="2" pageNumber="55" reason="1">
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<typeStatus box="[140,256,1082,1108]" pageId="2" pageNumber="55">Holotype</typeStatus>
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</emphasis>
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.
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</emphasis>
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<specimenCode box="[269,430,1082,1109]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="55" type="Museum">BPBM 33657</specimenCode>
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(field tag
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<specimenCode box="[549,670,1082,1108]" collectionCode="FK" pageId="2" pageNumber="55">FK 13097</specimenCode>
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), collected by
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<collectorName box="[837,933,1082,1108]" pageId="2" pageNumber="55">F. Kraus</collectorName>
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||
, S of
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<location LSID="urn:lsid:plazi:treatment:AF1DA83BD457FFE6FF3ADFB844537EBB:226B4FF6D457FFE0FC5ED82E478F7C4B" country="Papua New Guinea" latitude="-5.656703" longLatPrecision="1" longitude="142.63423" municipality="Muller Range" name="Tumbutu River" pageId="2" pageNumber="55" stateProvince="Southern Highlands">Tumbutu River</location>
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,
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<collectingMunicipality box="[217,380,1117,1144]" pageId="2" pageNumber="55">Muller Range</collectingMunicipality>
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||
,
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||
<geoCoordinate box="[396,547,1117,1144]" degrees="5.6567028" direction="south" orientation="latitude" pageId="2" pageNumber="55" precision="1" value="-5.656703">5.6567028°S</geoCoordinate>
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||
,
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||
<geoCoordinate box="[562,746,1117,1143]" degrees="142.6342342" direction="east" orientation="longitude" pageId="2" pageNumber="55" precision="1" value="142.63423">142.6342342°E</geoCoordinate>
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||
,
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||
<quantity box="[761,858,1117,1144]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="55" unit="m" value="1700.0">
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||
<elevation box="[761,858,1117,1144]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="55" unit="m" value="1700.0">1700 m</elevation>
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||
</quantity>
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||
,
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<collectingRegion country="Papua New Guinea" name="Southern Highlands" pageId="2" pageNumber="55">Southern Highlands Province</collectingRegion>
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||
,
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<collectingCountry box="[252,474,1152,1179]" name="Papua New Guinea" pageId="2" pageNumber="55">Papua New Guinea</collectingCountry>
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||
,
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||
<date box="[486,635,1152,1178]" pageId="2" pageNumber="55" value="2009-04-02">
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||
<collectingDate box="[486,635,1152,1178]" pageId="2" pageNumber="55" value="2009-04-02">2 April 2009</collectingDate>
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||
</date>
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||
.
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||
</materialsCitation>
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</paragraph>
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||
<paragraph blockId="2.[140,1108,1082,1672]" pageId="2" pageNumber="55">
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<materialsCitation ID-GBIF-Occurrence="2608940809" collectingDate="2009-03-21" collectingDateMax="2009-03-22" collectingDateMin="2009-03-21" collectionCode="BPBM" elevation="1700" latitude="-5.643116" location="Kunida" longLatPrecision="1" longitude="142.63423" municipality="Muller Range" pageId="2" pageNumber="55" specimenCode="BPBM 33636-40" specimenCount="1" typeStatus="paratype">
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<emphasis bold="true" box="[185,427,1187,1213]" pageId="2" pageNumber="55">
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<emphasis box="[185,300,1187,1213]" inLineHeading="true" pageId="2" pageNumber="55" reason="1">
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<typeStatus box="[185,300,1187,1213]" pageId="2" pageNumber="55">Paratypes</typeStatus>
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</emphasis>
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(n = 31).
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</emphasis>
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<specimenCode box="[439,651,1187,1214]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="55" type="Museum">BPBM 33636–40</specimenCode>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD457FFE6FF3ADFB844537EBB:226B4FF6D457FFE0FD2AD8B745B67C81" box="[668,755,1187,1213]" latitude="-5.643116" longLatPrecision="1" longitude="142.63423" municipality="Muller Range" name="Kunida" pageId="2" pageNumber="55">Kunida</location>
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,
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<collectingMunicipality box="[772,935,1187,1214]" pageId="2" pageNumber="55">Muller Range</collectingMunicipality>
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,
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||
<geoCoordinate box="[952,1103,1187,1214]" degrees="5.6431159" direction="south" orientation="latitude" pageId="2" pageNumber="55" precision="1" value="-5.643116">5.6431159°S</geoCoordinate>
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||
,
|
||
<geoCoordinate box="[140,323,1223,1249]" degrees="142.6342342" direction="east" orientation="longitude" pageId="2" pageNumber="55" precision="1" value="142.63423">142.6342342°E</geoCoordinate>
|
||
,
|
||
<quantity box="[338,433,1223,1249]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="55" unit="m" value="1700.0">
|
||
<elevation box="[338,433,1223,1249]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="55" unit="m" value="1700.0">1700 m</elevation>
|
||
</quantity>
|
||
,
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||
<date box="[449,680,1223,1249]" pageId="2" pageNumber="55" value="2009-03-21" valueMax="2009-03-22" valueMin="2009-03-21">
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||
<collectingDate box="[449,680,1223,1249]" pageId="2" pageNumber="55" value="2009-03-21" valueMax="2009-03-22" valueMin="2009-03-21">21–22 March 2009</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
;
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||
<materialsCitation ID-GBIF-Occurrence="2608940803" collectingDate="2009-03-27" collectionCode="BPBM" elevation="2177" latitude="-5.66954" location="Mt. Itukua" longLatPrecision="1" longitude="142.62334" municipality="Muller Range" pageId="2" pageNumber="55" specimenCode="BPBM 33641" specimenCount="1" typeStatus="paratype">
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||
<specimenCode box="[695,860,1223,1249]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="55" type="Museum">BPBM 33641</specimenCode>
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||
, E slope
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||
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD457FFE6FF3ADFB844537EBB:226B4FF6D457FFE0FC7AD8D3430A7CDD" box="[972,1103,1223,1249]" latitude="-5.66954" longLatPrecision="1" longitude="142.62334" municipality="Muller Range" name="Mt. Itukua" pageId="2" pageNumber="55">Mt. Itukua</location>
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||
,
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||
<collectingMunicipality box="[140,301,1258,1284]" pageId="2" pageNumber="55">Muller Range</collectingMunicipality>
|
||
,
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||
<geoCoordinate box="[315,437,1258,1285]" degrees="5.66954" direction="south" orientation="latitude" pageId="2" pageNumber="55" precision="1" value="-5.66954">5.66954°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[452,605,1258,1284]" degrees="142.62334" direction="east" orientation="longitude" pageId="2" pageNumber="55" precision="1" value="142.62334">142.62334°E</geoCoordinate>
|
||
,
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||
<quantity box="[620,715,1258,1284]" metricMagnitude="3" metricUnit="m" metricValue="2.177" pageId="2" pageNumber="55" unit="m" value="2177.0">
|
||
<elevation box="[620,715,1258,1284]" metricMagnitude="3" metricUnit="m" metricValue="2.177" pageId="2" pageNumber="55" unit="m" value="2177.0">2177 m</elevation>
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||
</quantity>
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||
,
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||
<date box="[730,916,1258,1284]" pageId="2" pageNumber="55" value="2009-03-27">
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<collectingDate box="[730,916,1258,1284]" pageId="2" pageNumber="55" value="2009-03-27">27 March 2009</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
;
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||
<materialsCitation ID-GBIF-Occurrence="2608940802" collectingDate="2009-03-31" collectionCode="BPBM" latitude="-5.6503625" location="Tumbutu River" longLatPrecision="1" longitude="142.63963" municipality="Mt. Paramo" pageId="2" pageNumber="55" specimenCode="BPBM 33642- 53" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="55" type="Museum">BPBM 33642– 53</specimenCode>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD457FFE6FF3ADFB844537EBB:226B4FF6D457FFE0FF01D91946237D1B" box="[183,358,1293,1319]" latitude="-5.6503625" longLatPrecision="1" longitude="142.63963" municipality="Mt. Paramo" name="Tumbutu River" pageId="2" pageNumber="55">Tumbutu River</location>
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below
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<collectingMunicipality box="[441,581,1294,1320]" pageId="2" pageNumber="55">Mt. Paramo</collectingMunicipality>
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||
,
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||
<geoCoordinate box="[593,744,1293,1320]" degrees="5.6503623" direction="south" orientation="latitude" pageId="2" pageNumber="55" precision="1" value="-5.6503625">5.6503623°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[756,909,1293,1319]" degrees="142.63963" direction="east" orientation="longitude" pageId="2" pageNumber="55" precision="1" value="142.63963">142.63963°E</geoCoordinate>
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||
,
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||
<date box="[922,1103,1293,1319]" pageId="2" pageNumber="55" value="2009-03-31">
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||
<collectingDate box="[922,1103,1293,1319]" pageId="2" pageNumber="55" value="2009-03-31">31 March 2009</collectingDate>
|
||
</date>
|
||
</materialsCitation>
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||
;
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||
<materialsCitation ID-GBIF-Occurrence="2608940811" box="[140,1103,1328,1355]" collectingDate="2009-04-01" collectionCode="BPBM" latitude="-5.6503625" location="Tumbutu River" longLatPrecision="1" longitude="142.63963" municipality="Mt. Paramo" pageId="2" pageNumber="55" specimenCode="BPBM 33654-56" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode box="[140,351,1328,1355]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="55" type="Museum">BPBM 33654–56</specimenCode>
|
||
, same data as BPBM 33642–53 except collected
|
||
<date box="[945,1103,1328,1354]" pageId="2" pageNumber="55" value="2009-04-01">
|
||
<collectingDate box="[945,1103,1328,1354]" pageId="2" pageNumber="55" value="2009-04-01">1 April 2009</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2608940820" box="[140,811,1363,1390]" collectingDate="2009-04-02" collectionCode="BPBM, PNGNM" collectorName="F. Kraus" country="Papua New Guinea" elevation="1700" latitude="-5.656703" location="Tumbutu River" longLatPrecision="1" longitude="142.63423" municipality="Muller Range" pageId="2" pageNumber="55" specimenCode="BPBM 33658, PNGNM 24095-96" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
|
||
<specimenCode box="[140,301,1363,1390]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="55" type="Museum">BPBM 33658</specimenCode>
|
||
,
|
||
<specimenCode box="[313,551,1363,1390]" collectionCode="PNGNM" pageId="2" pageNumber="55">PNGNM 24095–96</specimenCode>
|
||
, same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2608940805" collectingDate="2009-04-02" collectionCode="BPBM, PNGNM" country="Papua New Guinea" elevation="1777" latitude="-5.64545" location="Mt. Paramo" longLatPrecision="1" longitude="142.63904" pageId="2" pageNumber="55" specimenCode="BPBM 33659, PNGNM 24093-94" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
|
||
<specimenCode box="[823,985,1363,1390]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="55" type="Museum">BPBM 33659</specimenCode>
|
||
,
|
||
<specimenCode collectionCode="PNGNM" pageId="2" pageNumber="55">PNGNM 24093–94</specimenCode>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD457FFE6FF3ADFB844537EBB:226B4FF6D457FFE0FEA2D96346E17DAD" box="[276,420,1399,1425]" country="Papua New Guinea" latitude="-5.64545" longLatPrecision="1" longitude="142.63904" name="Mt. Paramo" pageId="2" pageNumber="55" stateProvince="Southern Highlands">Mt. Paramo</location>
|
||
,
|
||
<geoCoordinate box="[436,557,1398,1425]" degrees="5.64545" direction="south" orientation="latitude" pageId="2" pageNumber="55" precision="1" value="-5.64545">5.64545°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[573,727,1399,1425]" degrees="142.63904" direction="east" orientation="longitude" pageId="2" pageNumber="55" precision="1" value="142.63904">142.63904°E</geoCoordinate>
|
||
,
|
||
<quantity box="[743,840,1399,1425]" metricMagnitude="3" metricUnit="m" metricValue="1.777" pageId="2" pageNumber="55" unit="m" value="1777.0">
|
||
<elevation box="[743,840,1399,1425]" metricMagnitude="3" metricUnit="m" metricValue="1.777" pageId="2" pageNumber="55" unit="m" value="1777.0">1777 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date box="[856,1013,1399,1425]" pageId="2" pageNumber="55" value="2009-04-02">
|
||
<collectingDate box="[856,1013,1399,1425]" pageId="2" pageNumber="55" value="2009-04-02">2 April 2009</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2608940813" collectingDate="2009-04-02" collectionCode="BPBM" country="Papua New Guinea" elevation="1777" latitude="-5.64545" location="Mt. Paramo" longLatPrecision="1" longitude="142.63904" pageId="2" pageNumber="55" specimenCode="BPBM 33660-63" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
|
||
<specimenCode collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="2" pageNumber="55" type="Museum">BPBM 33660–63</specimenCode>
|
||
, same data as BPBM 33642–53 except collected
|
||
<date box="[814,963,1434,1460]" pageId="2" pageNumber="55" value="2009-04-02">
|
||
<collectingDate box="[814,963,1434,1460]" pageId="2" pageNumber="55" value="2009-04-02">2 April 2009</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="3" lastPageNumber="56" pageId="2" pageNumber="55" type="diagnosis">
|
||
<paragraph blockId="2.[140,1108,1082,1672]" pageId="2" pageNumber="55">
|
||
<emphasis bold="true" box="[185,312,1469,1495]" pageId="2" pageNumber="55">
|
||
<emphasis box="[185,307,1469,1495]" inLineHeading="true" pageId="2" pageNumber="55" reason="1">Diagnosis</emphasis>
|
||
.
|
||
</emphasis>
|
||
A small species (adult SV =
|
||
<quantity box="[634,807,1469,1496]" metricMagnitude="-2" metricUnit="m" metricValue="1.645" metricValueMax="1.83" metricValueMin="1.46" pageId="2" pageNumber="55" unit="mm" value="16.45" valueMax="18.3" valueMin="14.6">14.6–18.3 mm</quantity>
|
||
) distinguished by its combination of oblique lores, distinct tympanum, relatively broad snout (IN/SV =
|
||
<geoCoordinate box="[1027,1092,1504,1530]" degrees="0.081" direction="north" orientation="latitude" pageId="2" pageNumber="55" precision="55" value="0.081">0.081</geoCoordinate>
|
||
<geoCoordinate degrees="0.096" direction="west" orientation="longitude" pageId="2" pageNumber="55" precision="55" value="-0.096">– 0.096</geoCoordinate>
|
||
, mean 0.086), relatively wide finger discs (3rdF/SV =
|
||
<geoCoordinate box="[818,887,1539,1566]" degrees="0.059" direction="north" orientation="latitude" pageId="2" pageNumber="55" precision="55" value="0.059">0.059</geoCoordinate>
|
||
<geoCoordinate box="[886,969,1539,1566]" degrees="0.081" direction="west" orientation="longitude" pageId="2" pageNumber="55" precision="55" value="-0.081">–0.081</geoCoordinate>
|
||
), and advertisement call consisting of a single peep uttered in a continuous series.
|
||
</paragraph>
|
||
<paragraph blockId="2.[140,1108,1082,1672]" pageId="2" pageNumber="55">
|
||
<emphasis bold="true" box="[185,589,1609,1636]" pageId="2" pageNumber="55">
|
||
<emphasis box="[185,344,1609,1635]" inLineHeading="true" pageId="2" pageNumber="55" reason="1">Comparisons</emphasis>
|
||
with other species.
|
||
</emphasis>
|
||
The new species differs from all congeners except
|
||
<taxonomicName authority="Menzies" authorityName="Menzies" authorityYear="1999" box="[221,480,1645,1671]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="swanhildae">
|
||
<emphasis box="[221,377,1645,1671]" italics="true" pageId="2" pageNumber="55">A. swanhildae</emphasis>
|
||
Menzies
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Kraus and Allison" authorityName="Kraus and Allison" box="[541,919,1645,1672]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="exclamitans">
|
||
<emphasis box="[541,702,1645,1671]" italics="true" pageId="2" pageNumber="55">A. exclamitans</emphasis>
|
||
Kraus and Allison
|
||
</taxonomicName>
|
||
in having a call
|
||
</paragraph>
|
||
<paragraph blockId="3.[140,1108,863,949]" pageId="3" pageNumber="56">
|
||
<emphasis bold="true" box="[140,238,863,885]" pageId="3" pageNumber="56">Figure |.</emphasis>
|
||
Photos in life of
|
||
<emphasis bold="true" box="[395,414,863,885]" pageId="3" pageNumber="56">A</emphasis>
|
||
<typeStatus box="[420,499,864,885]" pageId="3" pageNumber="56">paratype</typeStatus>
|
||
of
|
||
<taxonomicName authority="Kraus & Allison, 2009" authorityName="Kraus & Allison" authorityYear="2009" box="[529,692,864,885]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="murritus" status="sp. n.">
|
||
<emphasis box="[529,692,864,885]" italics="true" pageId="3" pageNumber="56">Albericus murritus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[698,748,864,885]" pageId="3" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
|
||
(BPBM 33656)
|
||
<emphasis bold="true" box="[904,921,863,885]" pageId="3" pageNumber="56">B</emphasis>
|
||
<typeStatus box="[927,1006,864,885]" pageId="3" pageNumber="56">paratype</typeStatus>
|
||
of
|
||
<taxonomicName authority="Kraus & Allison, 2009" authorityName="Kraus & Allison" authorityYear="2009" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="caverniphilus" status="sp. n.">
|
||
<emphasis italics="true" pageId="3" pageNumber="56">Cophixalus caverniphilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[305,356,896,917]" pageId="3" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
|
||
(BPBM 33711)
|
||
<emphasis bold="true" box="[514,533,895,917]" pageId="3" pageNumber="56">C</emphasis>
|
||
<typeStatus box="[539,618,896,917]" pageId="3" pageNumber="56">paratype</typeStatus>
|
||
of
|
||
<taxonomicName authority="Kraus & Allison, 2009" authorityName="Kraus & Allison" authorityYear="2009" box="[650,850,896,917]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="anamiatoi" status="sp. n.">
|
||
<emphasis box="[650,850,896,917]" italics="true" pageId="3" pageNumber="56">Oreophryne anamiatoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[857,908,896,917]" pageId="3" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
|
||
(BPBM 33764), and
|
||
<emphasis bold="true" box="[140,159,927,949]" pageId="3" pageNumber="56">D</emphasis>
|
||
<typeStatus box="[165,244,928,949]" pageId="3" pageNumber="56">paratype</typeStatus>
|
||
of
|
||
<taxonomicName authority="Kraus & Allison, 2009" authorityName="Kraus & Allison" authorityYear="2009" box="[275,475,928,949]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="anamiatoi" status="sp. n.">
|
||
<emphasis box="[275,475,928,949]" italics="true" pageId="3" pageNumber="56">Oreophryne anamiatoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[481,532,928,949]" pageId="3" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
|
||
(BPBM 33765).
|
||
</paragraph>
|
||
<paragraph blockId="3.[140,1108,1011,1672]" pageId="3" pageNumber="56">
|
||
consisting of a peep; all other
|
||
<taxonomicName authorityName="Burton & Zweifel" authorityYear="1995" box="[485,583,1011,1037]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis box="[485,583,1011,1037]" italics="true" pageId="3" pageNumber="56">Albericus</emphasis>
|
||
</taxonomicName>
|
||
have calls consisting of a single honk/buzz or a series of clicks. From
|
||
<taxonomicName authorityName="Menzies" authorityYear="1999" box="[414,570,1047,1073]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="swanhildae">
|
||
<emphasis box="[414,570,1047,1073]" italics="true" pageId="3" pageNumber="56">A. swanhildae</emphasis>
|
||
</taxonomicName>
|
||
the new species differs in its larger size (SV =
|
||
<quantity box="[140,311,1082,1109]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.54" metricValueMin="1.3599999999999999" pageId="3" pageNumber="56" unit="mm" value="14.5" valueMax="15.4" valueMin="13.6">13.6–15.4 mm</quantity>
|
||
in
|
||
<taxonomicName authorityName="Menzies" authorityYear="1999" box="[349,502,1082,1108]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="swanhildae">
|
||
<emphasis box="[349,502,1082,1108]" italics="true" pageId="3" pageNumber="56">A. swanhildae</emphasis>
|
||
</taxonomicName>
|
||
), oblique (vs. vertical) lores, wider finger discs (3rdF/ SV =
|
||
<quantity box="[201,377,1117,1144]" metricMagnitude="-3" metricUnit="m" metricValue="1.4731999999999998" metricValueMax="1.5493999999999999" metricValueMin="1.3969999999999998" pageId="3" pageNumber="56" unit="in" value="0.057999999999999996" valueMax="0.061" valueMin="0.055">
|
||
<geoCoordinate box="[201,267,1117,1144]" degrees="0.055" direction="north" orientation="latitude" pageId="3" pageNumber="56" precision="55" value="0.055">0.055</geoCoordinate>
|
||
<geoCoordinate box="[268,348,1117,1144]" degrees="0.061" direction="west" orientation="longitude" pageId="3" pageNumber="56" precision="55" value="-0.061">–0.061</geoCoordinate>
|
||
in
|
||
</quantity>
|
||
<taxonomicName authorityName="Menzies" authorityYear="1999" box="[384,536,1117,1143]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="swanhildae">
|
||
<emphasis box="[384,536,1117,1143]" italics="true" pageId="3" pageNumber="56">A. swanhildae</emphasis>
|
||
</taxonomicName>
|
||
), and in having a single broad dark band across the center of each shank (vs. three narrow dark bars across each shank in
|
||
<taxonomicName authorityName="Menzies" authorityYear="1999" box="[936,1091,1152,1179]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="swanhildae">
|
||
<emphasis box="[936,1091,1152,1179]" italics="true" pageId="3" pageNumber="56">A. swanhildae</emphasis>
|
||
</taxonomicName>
|
||
). From
|
||
<taxonomicName box="[209,368,1187,1214]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="exclamitans">
|
||
<emphasis box="[209,368,1187,1214]" italics="true" pageId="3" pageNumber="56">A. exclamitans</emphasis>
|
||
</taxonomicName>
|
||
the new species differs in having the tympanum evident (vs. hidden) in males and in having the call consist of an infrequently produced single peep (vs. rapid burst of 3–48 peeps in
|
||
<taxonomicName box="[540,701,1258,1284]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="exclamitans">
|
||
<emphasis box="[540,701,1258,1284]" italics="true" pageId="3" pageNumber="56">A. exclamitans</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName authorityName="Kraus & Allison" authorityYear="2009" box="[729,933,1258,1284]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="murritus">
|
||
<emphasis box="[729,933,1258,1284]" italics="true" pageId="3" pageNumber="56">Albericus murritus</emphasis>
|
||
</taxonomicName>
|
||
is also slightly smaller than
|
||
<taxonomicName box="[289,449,1293,1319]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="exclamitans">
|
||
<emphasis box="[289,449,1293,1319]" italics="true" pageId="3" pageNumber="56">A. exclamitans</emphasis>
|
||
</taxonomicName>
|
||
(male SV =
|
||
<quantity box="[597,772,1293,1320]" metricMagnitude="-2" metricUnit="m" metricValue="1.645" metricValueMax="1.83" metricValueMin="1.46" pageId="3" pageNumber="56" unit="mm" value="16.45" valueMax="18.3" valueMin="14.6">14.6–18.3 mm</quantity>
|
||
, female SV = 15.0–
|
||
<quantity box="[1002,1107,1293,1320]" metricMagnitude="-2" metricUnit="m" metricValue="1.83" pageId="3" pageNumber="56" unit="mm" value="18.3">18.3 mm</quantity>
|
||
in
|
||
<taxonomicName authorityName="Kraus & Allison" authorityYear="2009" box="[173,303,1329,1355]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="murritus">
|
||
<emphasis box="[173,303,1329,1355]" italics="true" pageId="3" pageNumber="56">A. murritus</emphasis>
|
||
</taxonomicName>
|
||
vs. 15.3–20.7 and 18.0–22.0 in
|
||
<taxonomicName box="[692,853,1328,1355]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="exclamitans">
|
||
<emphasis box="[692,853,1328,1355]" italics="true" pageId="3" pageNumber="56">A. exclamitans</emphasis>
|
||
</taxonomicName>
|
||
) and has a somewhat broader snout (IN/SV =
|
||
<geoCoordinate box="[420,488,1363,1390]" degrees="0.074" direction="north" orientation="latitude" pageId="3" pageNumber="56" precision="55" value="0.074">0.074</geoCoordinate>
|
||
<geoCoordinate box="[487,570,1363,1390]" degrees="0.085" direction="west" orientation="longitude" pageId="3" pageNumber="56" precision="55" value="-0.085">–0.085</geoCoordinate>
|
||
, mean
|
||
<quantity box="[650,747,1363,1390]" metricMagnitude="-3" metricUnit="m" metricValue="2.0066" pageId="3" pageNumber="56" unit="in" value="0.079">0.079 in</quantity>
|
||
<taxonomicName box="[754,912,1363,1390]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="exclamitans">
|
||
<emphasis box="[754,912,1363,1390]" italics="true" pageId="3" pageNumber="56">A. exclamitans</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56" type="description">
|
||
<paragraph blockId="3.[140,1108,1011,1672]" lastBlockId="4.[140,1108,167,1672]" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56">
|
||
<emphasis bold="true" box="[185,489,1398,1425]" pageId="3" pageNumber="56">
|
||
<emphasis box="[185,329,1398,1424]" inLineHeading="true" pageId="3" pageNumber="56" reason="1">Description</emphasis>
|
||
of
|
||
<typeStatus box="[374,484,1399,1425]" pageId="3" pageNumber="56">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male. Head relatively wide (HW/SV = 0.38), with oblique and shallowly concave loreal region; canthus rostralis broadly rounded; nostrils small, crescent-shaped, much closer to tip of snout than to eyes; distance from external naris to eye larger than internarial distance (EN/IN = 1.14, IN/SV = 0.083, EN/SV = 0.095); snout bluntly rounded when viewed from above, truncate when viewed from side; eyes moderately large (EY/SV = 0.13), eyelid approximately 2/3 width of interorbital distance; tympanum indistinct, partially hidden by surrounding skin. Dorsum pustulose on body and limbs; supratympanic fold absent but row of dorsal pustules occupy that area; ventral surfaces coarsely granular. Fingers unwebbed, bearing discs with terminal grooves; relative lengths 3>4>2>1. Finger discs approximately twice widths of penultimate phalanges. Subarticular tubercles not well developed; metacarpal tubercles absent. Toes unwebbed, bearing discs with terminal grooves; relative lengths 4>5>3>2>1. Toe discs smaller than those of fingers (3rdF/4thT = 1.24), approximately 1.5 times width of penultimate phalanges. Subarticular tubercles low; metatarsal tubercles lacking. Hind legs rather short (TL/SV = 0.40); arms rather long (ArmL/SV = 0.54).
|
||
</paragraph>
|
||
<paragraph blockId="4.[140,1108,167,1672]" pageId="4" pageNumber="57">In preservative, dorsal ground color an irregular mix of yellow-tan and brown, with the former predominating dorsolaterally and the latter mid-dorsally; parts of both fields suffused with russet. Irregular black flecks and markings scattered throughout, concentrated above shoulders, on rear of head, and above tympana. Traces of a short, lighter, yellow-brown bar on each scapula; similar-colored interocular bar and vestiges of lumbar ocelli. Sides dark gray flecked with black and light blue-gray. Face yellow-tan flecked with black and russet. Legs yellow-tan with one broad, dark, centrally placed band on each thigh and shank. Rear of thighs dark brown with narrow yellow-tan band proximally. Venter dirty light gray evenly and densely peppered with dark gray; palmar and plantar surfaces same. Front margin of mandible russet. Iris black flecked with silver.</paragraph>
|
||
<paragraph blockId="4.[140,1108,167,1672]" pageId="4" pageNumber="57">
|
||
<emphasis bold="true" box="[185,304,800,826]" pageId="4" pageNumber="57">
|
||
<emphasis box="[185,298,800,826]" inLineHeading="true" pageId="4" pageNumber="57" reason="1">Variation</emphasis>
|
||
.
|
||
</emphasis>
|
||
Mensural variation for the
|
||
<typeStatus box="[619,667,801,827]" pageId="4" pageNumber="57">type</typeStatus>
|
||
series is shown in Table 1. Th ere is no obvious sexual dimorphism in morphometric features, although there is slight evidence that females may average larger in body size. However, this difference is slight considering the normal pattern of larger female size in most Papuan frogs.
|
||
</paragraph>
|
||
<paragraph blockId="4.[140,1108,167,1672]" pageId="4" pageNumber="57">
|
||
Most specimens appear dark brown in preservative (darker than the
|
||
<typeStatus box="[991,1095,941,968]" pageId="4" pageNumber="57">holotype</typeStatus>
|
||
), with a few contrasting light-brown streaks or lines scattered on dorsum. These typically involve a single short line on each scapula, an interocular bar, and often traces of lumbar ocelli. Lighter specimens are similar but show more clearly the variable mottling seen in the
|
||
<typeStatus box="[323,426,1082,1108]" pageId="4" pageNumber="57">holotype</typeStatus>
|
||
. Density of dark ventral stippling varies from sparse to dense, making the overall appearance of the venter vary from light gray to black. Both ventral extremes appear in frogs with both light and dark dorsa, but venters of frogs that are lighter dorsally average somewhat paler than those of dark frogs.
|
||
</paragraph>
|
||
<paragraph blockId="4.[140,1108,167,1672]" pageId="4" pageNumber="57">
|
||
<emphasis bold="true" box="[185,339,1222,1248]" pageId="4" pageNumber="57">
|
||
<emphasis box="[185,253,1222,1248]" inLineHeading="true" pageId="4" pageNumber="57" reason="1">Color</emphasis>
|
||
in life.
|
||
</emphasis>
|
||
BPBM 33636: “Dorsum dark tan with irregular black blotches and tiny white or tan dots on some warts; fairly warty. Orange-brown interocular bar, suprascapular marks, and on arms and heels. Venter charcoal gray with tiny light-gray punctations. Light-tan patch from eye to rictus. Iris brown.” BPBM 33637 was yellow-brown with cream and black markings, venter densely punctated with light gray, and iris tan.
|
||
</paragraph>
|
||
<paragraph blockId="4.[140,1108,167,1672]" pageId="4" pageNumber="57">
|
||
<emphasis bold="true" box="[185,242,1398,1424]" pageId="4" pageNumber="57">
|
||
<emphasis box="[185,237,1398,1424]" inLineHeading="true" pageId="4" pageNumber="57" reason="1">Call</emphasis>
|
||
.
|
||
</emphasis>
|
||
This species begins calling at dusk and calls in highest numbers during the first few hours of darkness. Th e call consists of a single “peep” note uttered in a continuous series with occasional breaks (
|
||
<figureCitation box="[584,649,1469,1495]" captionStart="Figure 2" captionStartId="5.[140,212,1425,1447]" captionTargetBox="[147,1085,875,1315]" captionTargetId="figure@5.[140,1108,868,1404]" captionTargetPageId="5" captionText="Figure 2. A Waveform, B power spectrum, and C spectrogram of call “T” of Albericus murritus sp. n. (BPBM 33641) recorded on E slope Mt. Itukua, Muller Range, Southern Highlands Province, Papua New Guinea on 27 March 2009 at 2020 h. Air temperature 14.7 °C." figureDoi="http://doi.org/10.5281/zenodo.3769053" httpUri="https://zenodo.org/record/3769053/files/figure.png" pageId="4" pageNumber="57">Fig 2</figureCitation>
|
||
). We recorded calls of two individuals, BPBM 33641 and BPBM 33657 (
|
||
<tableCitation box="[528,616,1504,1530]" captionStart="Table 2" captionStartId="6.[140,200,166,188]" captionText="Table 2. Call statistics of Albericus murritus sp. n. recorded by Fred Kraus on E slope Mt. Itukua, Muller Range, Southern Highlands Province, Papua New Guinea. BPBM 33641 was recorded on 27 March 2009 at 2020 h, BPBM 33657 on 2 April 2009 at 2215 h. Air temperatures were 14.7 and 18.8 °C, respectively." pageId="4" pageNumber="57">Table 2</tableCitation>
|
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). The notes had a mean duration of
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<geoCoordinate box="[1025,1107,1504,1531]" degrees="0.151" direction="south" orientation="latitude" pageId="4" pageNumber="57" precision="55" value="-0.151">0.151 s</geoCoordinate>
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(range
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<geoCoordinate box="[218,286,1540,1566]" degrees="0.093" direction="north" orientation="latitude" pageId="4" pageNumber="57" precision="55" value="0.093">0.093</geoCoordinate>
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<geoCoordinate box="[286,370,1540,1566]" degrees="0.213" direction="west" orientation="longitude" pageId="4" pageNumber="57" precision="55" value="-0.213">–0.213</geoCoordinate>
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). The inter-note interval for BPBM 33641 ranged from 4.1–
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<geoCoordinate box="[1054,1107,1540,1566]" degrees="9.0" direction="south" orientation="latitude" pageId="4" pageNumber="57" precision="5555" value="-9.0">9.0 s</geoCoordinate>
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(n = 20), except for two instances of 31.6 and
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<geoCoordinate box="[648,717,1574,1601]" degrees="35.9" direction="south" orientation="latitude" pageId="4" pageNumber="57" precision="5555" value="-35.9">35.9 s</geoCoordinate>
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. Many species of
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<taxonomicName authorityName="Burton & Zweifel" authorityYear="1995" box="[911,1009,1575,1601]" class="Amphibia" family="Microhylidae" genus="Albericus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="57" phylum="Chordata" rank="genus">
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<emphasis box="[911,1009,1575,1601]" italics="true" pageId="4" pageNumber="57">Albericus</emphasis>
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</taxonomicName>
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produce calls in groups separated by periods of silence (A. Allison, pers. obs.), and our brief recording suggests that this is occurring in BPBM 33641.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |