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<document ID-DOI="10.5733/afin.054.0213" ID-ISSN="2305-2562" ID-Zenodo-Dep="7918661" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1683583268648" checkinUser="felipe" docAuthor="Niekerk, P. van &amp; Dippenaar-Schoeman, A. S." docDate="2013" docId="03D8D254FFE5A93F21A5FB3BFD47FA7C" docLanguage="en" docName="AfrInvertebr.54.2.447-476.pdf" docOrigin="African Invertebrates 54 (2)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.054.0213" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.4:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="4" docTitle="Heriaeus foordi Niekerk &amp; Dippenaar-Schoeman 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="463" masterDocId="FFE1AA2CFFEBA92F2055FF99FFE4FFE6" masterDocTitle="A revision of the crab spider genus Heriaeus Simon, 1875 (Araneae: Thomisidae) in the Afrotropical Region" masterLastPageNumber="447" masterPageNumber="447" pageNumber="461" updateTime="1683701492856" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>A revision of the crab spider genus Heriaeus Simon, 1875 (Araneae: Thomisidae) in the Afrotropical Region</mods:title>
</mods:titleInfo>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Niekerk, P. van</mods:namePart>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Dippenaar-Schoeman, A. S.</mods:namePart>
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<mods:title>African Invertebrates</mods:title>
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<mods:date>2013</mods:date>
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<mods:number>2013-11-14</mods:number>
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<mods:number>54</mods:number>
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<mods:number>2</mods:number>
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<mods:identifier type="ISSN">2305-2562</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7917981" ID-Zenodo-Dep="7917981" LSID="urn:lsid:plazi:treatment:03D8D254FFE5A93F21A5FB3BFD47FA7C" httpUri="http://treatment.plazi.org/id/03D8D254FFE5A93F21A5FB3BFD47FA7C" lastPageId="16" lastPageNumber="463" pageId="14" pageNumber="461">
<subSubSection box="[496,751,1186,1213]" pageId="14" pageNumber="461" type="nomenclature">
<paragraph blockId="14.[144,1104,1186,1677]" box="[496,751,1186,1213]" pageId="14" pageNumber="461">
<heading box="[496,751,1186,1213]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="461" reason="2">
<taxonomicName ID-CoL="3KYV5" authority="Niekerk &amp; Dippenaar-Schoeman, 2013" authorityName="Niekerk &amp; Dippenaar-Schoeman" authorityYear="2013" box="[496,684,1186,1212]" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="461" phylum="Arthropoda" rank="species" species="foordi" status="sp. n.">
<emphasis bold="true" box="[496,684,1186,1212]" pageId="14" pageNumber="461">Heriaeus foordi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[691,751,1187,1213]" pageId="14" pageNumber="461" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="463" pageId="14" pageNumber="461" type="description">
<paragraph blockId="14.[144,1104,1186,1677]" box="[524,723,1226,1253]" pageId="14" pageNumber="461">
<figureCitation box="[524,595,1226,1253]" captionStart="Figs 19" captionStartId="3.[181,220,1090,1112]" captionTargetBox="[187,1132,204,1065]" captionTargetId="figure-241@3.[187,1132,204,1065]" captionTargetPageId="3" captionText="Figs 19. Heriaeus spp., habitus:(1) Heriaeus sp. long spiniform abdominal setae; (2) H.allenjonesi antoni sp. n. female; (3) H.antoni sp. n. female; (4) H.copricola sp. n. male; (5) H. crassispinus Lawrence, 1942 male; (6) H. foordi sp. n. male; (7) H. latifrons Lessert, 1919 female; (8) H.madagascar sp. n. male; (9) H. muizenberg sp. n. female." figureDoi="http://doi.org/10.5281/zenodo.7918663" httpUri="https://zenodo.org/record/7918663/files/figure.png" pageId="14" pageNumber="461">Figs 6</figureCitation>
,
<figureCitation box="[607,683,1227,1253]" captionStart="Figs 3036" captionStartId="15.[181,220,1193,1215]" captionTargetBox="[189,1133,212,1162]" captionTargetId="figure-164@15.[181,1141,204,1174]" captionTargetPageId="15" captionText="Figs 3036.(3033) Heriaeus foordi sp. n.: (30) male palp, ventral view; (31) male palp, lateral view; (32) epigyne, ventral view; (33) epigyne, dorsal view; (34) Heriaeus latifrons Lessert, 1919, epigyne, ventral view; (35, 36) Heriaeus madagascar sp. n.: (35) male palp, ventral view; (36) male palp, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7918671" httpUri="https://zenodo.org/record/7918671/files/figure.png" pageId="14" pageNumber="461">3033</figureCitation>
,
<figureCitation box="[695,723,1226,1253]" captionStart="Figs 5759" captionStartId="27.[347,386,1654,1676]" captionTargetBox="[181,1141,609,1633]" captionTargetId="figure-176@27.[132,1190,602,1636]" captionTargetPageId="27" captionText="Figs 5759. Distribution of Heriaeus spp. in the Afrotropical Region." figureDoi="http://doi.org/10.5281/zenodo.7918679" httpUri="https://zenodo.org/record/7918679/files/figure.png" pageId="14" pageNumber="461">58</figureCitation>
</paragraph>
<paragraph blockId="14.[144,1104,1186,1677]" pageId="14" pageNumber="461">
Etymology: Named for Stefan Foord of the University of Venda, who collected the
<typeStatus box="[144,246,1299,1325]" pageId="14" pageNumber="461">holotype</typeStatus>
. He is recognized for his contribution in recording spider diversity in the African savannah.
</paragraph>
<paragraph blockId="14.[144,1104,1186,1677]" pageId="14" pageNumber="461">
Diagnosis: The species is recognized by a combination of long, dark brown, spiniform abdominal setae (
<figureCitation box="[340,408,1403,1429]" captionStart="Figs 19" captionStartId="3.[181,220,1090,1112]" captionTargetBox="[187,1132,204,1065]" captionTargetId="figure-241@3.[187,1132,204,1065]" captionTargetPageId="3" captionText="Figs 19. Heriaeus spp., habitus:(1) Heriaeus sp. long spiniform abdominal setae; (2) H.allenjonesi antoni sp. n. female; (3) H.antoni sp. n. female; (4) H.copricola sp. n. male; (5) H. crassispinus Lawrence, 1942 male; (6) H. foordi sp. n. male; (7) H. latifrons Lessert, 1919 female; (8) H.madagascar sp. n. male; (9) H. muizenberg sp. n. female." figureDoi="http://doi.org/10.5281/zenodo.7918663" httpUri="https://zenodo.org/record/7918663/files/figure.png" pageId="14" pageNumber="461">Fig. 6</figureCitation>
), the small dome-shaped hood of the female epigyne (
<figureCitation box="[1014,1095,1403,1429]" captionStart="Figs 3036" captionStartId="15.[181,220,1193,1215]" captionTargetBox="[189,1133,212,1162]" captionTargetId="figure-164@15.[181,1141,204,1174]" captionTargetPageId="15" captionText="Figs 3036.(3033) Heriaeus foordi sp. n.: (30) male palp, ventral view; (31) male palp, lateral view; (32) epigyne, ventral view; (33) epigyne, dorsal view; (34) Heriaeus latifrons Lessert, 1919, epigyne, ventral view; (35, 36) Heriaeus madagascar sp. n.: (35) male palp, ventral view; (36) male palp, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7918671" httpUri="https://zenodo.org/record/7918671/files/figure.png" pageId="14" pageNumber="461">Fig. 32</figureCitation>
) and the long embolus with straight tip in the male (
<figureCitation box="[726,811,1435,1461]" captionStart="Figs 3036" captionStartId="15.[181,220,1193,1215]" captionTargetBox="[189,1133,212,1162]" captionTargetId="figure-164@15.[181,1141,204,1174]" captionTargetPageId="15" captionText="Figs 3036.(3033) Heriaeus foordi sp. n.: (30) male palp, ventral view; (31) male palp, lateral view; (32) epigyne, ventral view; (33) epigyne, dorsal view; (34) Heriaeus latifrons Lessert, 1919, epigyne, ventral view; (35, 36) Heriaeus madagascar sp. n.: (35) male palp, ventral view; (36) male palp, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7918671" httpUri="https://zenodo.org/record/7918671/files/figure.png" pageId="14" pageNumber="461">Fig. 30</figureCitation>
). The RTA is long, with a prolaterally curved distal tooth and a basal lobe (
<figureCitation box="[690,773,1467,1493]" captionStart="Figs 3036" captionStartId="15.[181,220,1193,1215]" captionTargetBox="[189,1133,212,1162]" captionTargetId="figure-164@15.[181,1141,204,1174]" captionTargetPageId="15" captionText="Figs 3036.(3033) Heriaeus foordi sp. n.: (30) male palp, ventral view; (31) male palp, lateral view; (32) epigyne, ventral view; (33) epigyne, dorsal view; (34) Heriaeus latifrons Lessert, 1919, epigyne, ventral view; (35, 36) Heriaeus madagascar sp. n.: (35) male palp, ventral view; (36) male palp, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7918671" httpUri="https://zenodo.org/record/7918671/files/figure.png" pageId="14" pageNumber="461">Fig. 31</figureCitation>
).
</paragraph>
<paragraph blockId="14.[144,1104,1186,1677]" box="[144,283,1507,1533]" pageId="14" pageNumber="461">Description:</paragraph>
<paragraph blockId="14.[144,1104,1186,1677]" box="[144,351,1547,1573]" pageId="14" pageNumber="461">
<emphasis box="[144,228,1547,1573]" italics="true" pageId="14" pageNumber="461">Female</emphasis>
(
<typeStatus box="[243,339,1547,1573]" pageId="14" pageNumber="461">allotype</typeStatus>
).
</paragraph>
<paragraph blockId="14.[144,1104,1186,1677]" lastBlockId="16.[144,1104,203,1434]" lastPageId="16" lastPageNumber="463" pageId="14" pageNumber="461">
<emphasis box="[144,189,1587,1613]" italics="true" pageId="14" pageNumber="461">Size</emphasis>
: TL 4.61, CL 2.25, CW 2.18.
<emphasis box="[525,604,1587,1613]" italics="true" pageId="14" pageNumber="461">Colour</emphasis>
: Carapace yellow-brown, with two broad irregularly shaped longitudinal dark brown stripes; setae dark brown; sternum with white markings; clypeus brown; chelicerae yellow-brown, with brown marks proximally and white marks distally and laterally; eye region white between anterior eyes, brown between posterior eyes; eye tubercles white or greyish white.Abdomen brown dorsally, mottled with white, with dark brown circular marks usually forming a distinct pattern; venter lined with white; laterally with white striae with brown spots. Legs yellow-brown, with white marks on all segments except metatarsi and tarsi. Palp segments all with white markings except tibia with brown markings.
<emphasis box="[802,912,1491,1517]" italics="true" pageId="15" pageNumber="462">Carapace</emphasis>
:
<emphasis box="[926,1134,1491,1516]" italics="true" pageId="15" pageNumber="462">Eye measurements</emphasis>
: AMEAME 0.25, ALEAME 0.14, PMEPME 0.19, PLEPME 0.30, AMEPME 0.27, MOQL AMEPME 0.34, MOQAW AMEAME 0.31, MOQPW PMEPME 0.27.
<emphasis box="[244,451,1587,1612]" italics="true" pageId="15" pageNumber="462">Leg measurements</emphasis>
: leg I Fe 1.76, Pat 0.91, Tib 1.75, Mt 1.66, Ta 0.95, total 7.03; leg II Fe 1.41, Pat 0.74, Tib 1.27, Mt 1.25, Ta 0.83, total 5.50; leg III Fe 1.06, Pat 0.53, Tib 1.00, Mt 0.67, Ta 0.66, total 3.92; leg IV Fe 1.15, Pat 0.65, Tib 1.02, Mt 0.75, Ta 0.62, total 4.19.
<emphasis box="[328,434,204,230]" italics="true" pageId="16" pageNumber="463">Abdomen</emphasis>
: Round, truncated anteriorly; setae long and dark brown or short and transparent.
<emphasis box="[392,484,236,261]" italics="true" pageId="16" pageNumber="463">Epigyne</emphasis>
: With small dome­shaped hood; internal organs visible as two indistinct bean-shaped structures lying diagonally below hood (
<figureCitation box="[949,1035,268,294]" captionStart="Figs 3036" captionStartId="15.[181,220,1193,1215]" captionTargetBox="[189,1133,212,1162]" captionTargetId="figure-164@15.[181,1141,204,1174]" captionTargetPageId="15" captionText="Figs 3036.(3033) Heriaeus foordi sp. n.: (30) male palp, ventral view; (31) male palp, lateral view; (32) epigyne, ventral view; (33) epigyne, dorsal view; (34) Heriaeus latifrons Lessert, 1919, epigyne, ventral view; (35, 36) Heriaeus madagascar sp. n.: (35) male palp, ventral view; (36) male palp, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7918671" httpUri="https://zenodo.org/record/7918671/files/figure.png" pageId="16" pageNumber="463">Fig. 32</figureCitation>
); four small dark posterior spots indicate position of fertilization ducts; copulatory ducts two tubes lying diagonally below hood (
<figureCitation box="[546,630,332,358]" captionStart="Figs 3036" captionStartId="15.[181,220,1193,1215]" captionTargetBox="[189,1133,212,1162]" captionTargetId="figure-164@15.[181,1141,204,1174]" captionTargetPageId="15" captionText="Figs 3036.(3033) Heriaeus foordi sp. n.: (30) male palp, ventral view; (31) male palp, lateral view; (32) epigyne, ventral view; (33) epigyne, dorsal view; (34) Heriaeus latifrons Lessert, 1919, epigyne, ventral view; (35, 36) Heriaeus madagascar sp. n.: (35) male palp, ventral view; (36) male palp, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7918671" httpUri="https://zenodo.org/record/7918671/files/figure.png" pageId="16" pageNumber="463">Fig. 33</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7918671" ID-Zenodo-Dep="7918671" httpUri="https://zenodo.org/record/7918671/files/figure.png" pageId="15" pageNumber="462" startId="15.[181,220,1193,1215]" targetBox="[189,1133,212,1162]" targetPageId="15" targetType="figure">
<paragraph blockId="15.[181,1142,1193,1287]" pageId="15" pageNumber="462">
Figs 3036. (3033)
<taxonomicName authority="Niekerk &amp; Dippenaar-Schoeman, 2013" authorityName="Niekerk &amp; Dippenaar-Schoeman" authorityYear="2013" box="[375,518,1193,1215]" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="462" phylum="Arthropoda" rank="species" species="foordi" status="sp. n.">
<emphasis box="[375,518,1193,1215]" italics="true" pageId="15" pageNumber="462">Heriaeus foordi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[522,573,1194,1215]" pageId="15" pageNumber="462" rank="species">sp. n.</taxonomicNameLabel>
: (30) male palp, ventral view; (31) male palp, lateral view; (32) epigyne, ventral view; (33) epigyne, dorsal view; (34)
<taxonomicName authority="Lessert, 1919" authorityName=": Lessert" authorityYear="1919" box="[754,1053,1217,1239]" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="462" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis box="[754,920,1217,1239]" italics="true" pageId="15" pageNumber="462">Heriaeus latifrons</emphasis>
<bibRefCitation author="LESSERT, R. DE" box="[927,1053,1217,1239]" pageId="15" pageNumber="462" pagination="99 - 234" refId="ref19984" refString="LESSERT, R. DE. 1919. Araignees du Kilimandjaro et du Merou (suite). 3. Thomisidae. Revue Suisse de Zoologie 27: 99 - 234." type="journal article" year="1919">Lessert, 1919</bibRefCitation>
</taxonomicName>
, epigyne, ventral view; (35, 36)
<taxonomicName authority="Niekerk &amp; Dippenaar-Schoeman, 2013" authorityName="Niekerk &amp; Dippenaar-Schoeman" authorityYear="2013" box="[453,655,1241,1263]" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="462" phylum="Arthropoda" rank="species" species="madagascar" status="sp. n.">
<emphasis box="[453,655,1241,1263]" italics="true" pageId="15" pageNumber="462">Heriaeus madagascar</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[662,715,1242,1263]" pageId="15" pageNumber="462" rank="species">sp. n.</taxonomicNameLabel>
: (35) male palp, ventral view; (36) male palp, lateral view.
</paragraph>
</caption>
<paragraph blockId="16.[144,1104,203,1434]" box="[144,332,373,399]" pageId="16" pageNumber="463">
<emphasis box="[144,202,373,399]" italics="true" pageId="16" pageNumber="463">Male</emphasis>
(
<typeStatus box="[216,321,373,399]" pageId="16" pageNumber="463">holotype</typeStatus>
).
</paragraph>
<paragraph blockId="16.[144,1104,203,1434]" pageId="16" pageNumber="463">
<emphasis box="[144,189,413,439]" italics="true" pageId="16" pageNumber="463">Size</emphasis>
: TL 3.18, CL 1.66, CW 1.74.
<emphasis box="[544,623,413,439]" italics="true" pageId="16" pageNumber="463">Colour</emphasis>
: Carapace yellow-brown, with two broad irregularly shaped longitudinal dark brown stripes; setae dark brown; clypeus brown; chelicerae with brown marks; eye region white between anterior eyes, brown between posterior eyes; eye tubercles white or greyish white. Abdomen yellow-brown, mottled with brown; some males with faint circular pattern similar to that of female; setae dark brown. Legs with white markings on coxae IIV; leg I (patella to tarsus) much darker than legs IIIV.
<emphasis box="[407,517,605,631]" italics="true" pageId="16" pageNumber="463">Carapace</emphasis>
:
<emphasis box="[533,742,606,631]" italics="true" pageId="16" pageNumber="463">Eye measurements</emphasis>
: AMEAME 0.16, ALEAME 0.09, PMEPME 0.12, PLEPME 0.21, AMEPME 0.22, MOQL AMEPME 0.27, MOQAW AMEAME 0.22, MOQPW PMEPME 0.20.
<emphasis box="[792,1001,670,695]" italics="true" pageId="16" pageNumber="463">Leg measurements</emphasis>
: leg I Fe 1.96, Pat 0.78, Tib 1.54, Mt 1.61, Ta 0.94, total 5.83; leg II Fe 1.67, Pat 0.69 Tib 1.34, Mt 1.25, Ta 0.68, total 5.63; leg III Fe 0.91, Pat 0.46, Tib 0.82, Mt 0.66, Ta 0.47, total 3.32; leg IV Fe 1.08, Pat 0.47, Tib 0.85, Mt 0.57, Ta 0.39, total 3.36.
<emphasis box="[902,1008,765,791]" italics="true" pageId="16" pageNumber="463">Abdomen</emphasis>
: Round, truncated anteriorly; dorsum with scattered long spiniform, acute setae.
<emphasis box="[924,976,798,823]" italics="true" pageId="16" pageNumber="463">Legs</emphasis>
: Very hairy with numerous long setae.
<emphasis box="[451,567,829,855]" italics="true" pageId="16" pageNumber="463">Male palp</emphasis>
: Embolus long, winding once around tegulum, tip slender and straight; VTA large and two-lobed; RTA with prolaterally curved distal tooth and basal lobe; cymbial apophysis present (
<figureCitation box="[695,825,893,919]" captionStart="Figs 3036" captionStartId="15.[181,220,1193,1215]" captionTargetBox="[189,1133,212,1162]" captionTargetId="figure-164@15.[181,1141,204,1174]" captionTargetPageId="15" captionText="Figs 3036.(3033) Heriaeus foordi sp. n.: (30) male palp, ventral view; (31) male palp, lateral view; (32) epigyne, ventral view; (33) epigyne, dorsal view; (34) Heriaeus latifrons Lessert, 1919, epigyne, ventral view; (35, 36) Heriaeus madagascar sp. n.: (35) male palp, ventral view; (36) male palp, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7918671" httpUri="https://zenodo.org/record/7918671/files/figure.png" pageId="16" pageNumber="463">Figs 30, 31</figureCitation>
).
</paragraph>
<paragraph blockId="16.[144,1104,203,1434]" pageId="16" pageNumber="463">
<emphasis box="[144,255,933,959]" italics="true" pageId="16" pageNumber="463">Juveniles.</emphasis>
Immature males can usually be recognized by brown dorsum, mottled with white, with dark brown circular markings forming a characteristic pattern.
</paragraph>
<paragraph blockId="16.[144,1104,203,1434]" pageId="16" pageNumber="463">
<materialsCitation collectingDate="2009-09-11" collectionCode="NCA" collectorName="S. Foord" country="South Africa" latitude="-26.75" location="Limpopo" longLatPrecision="744" longitude="24.42" pageId="16" pageNumber="463" specimenCode="NCA 2011" specimenCount="1" specimenCount-male="1" stateProvince="Limpopo" typeStatus="holotype">
<typeStatus box="[144,232,1003,1025]" pageId="16" pageNumber="463">Holotype</typeStatus>
:
<specimenCount box="[241,258,1004,1025]" pageId="16" pageNumber="463" type="male"></specimenCount>
<collectingCountry box="[264,432,1003,1024]" name="South Africa" pageId="16" pageNumber="463">SOUTH AFRICA</collectingCountry>
:
<emphasis box="[441,522,1004,1025]" italics="true" pageId="16" pageNumber="463">
<collectingRegion box="[441,522,1004,1025]" country="South Africa" name="Limpopo" pageId="16" pageNumber="463">Limpopo</collectingRegion>
</emphasis>
: Atherstone Nat. Res. (
<geoCoordinate box="[737,810,1003,1025]" degrees="26.75" direction="south" orientation="latitude" pageId="16" pageNumber="463" precision="555" value="-26.75">26.75°S</geoCoordinate>
<geoCoordinate box="[817,894,1003,1024]" degrees="24.42" direction="east" orientation="longitude" pageId="16" pageNumber="463" precision="555" value="24.42">24.42°E</geoCoordinate>
),
<date box="[911,1009,1003,1025]" pageId="16" pageNumber="463" value="2009-09-11">
<collectingDate box="[911,1009,1003,1025]" pageId="16" pageNumber="463" value="2009-09-11">11.ix.2009</collectingDate>
</date>
,
<collectorName box="[1019,1100,1003,1025]" pageId="16" pageNumber="463">S. Foord</collectorName>
, sweeping grass (
<specimenCode box="[293,393,1027,1048]" collectionCode="NCA" pageId="16" pageNumber="463">NCA 2011</specimenCode>
/2044).
</materialsCitation>
</paragraph>
<paragraph blockId="16.[144,1104,203,1434]" box="[144,655,1059,1081]" pageId="16" pageNumber="463">
<materialsCitation box="[144,655,1059,1081]" collectingDate="2009-09-11" collectionCode="NCA" collectorName="S. Foord" country="South Africa" latitude="-26.75" location="Limpopo" longLatPrecision="744" longitude="24.42" pageId="16" pageNumber="463" specimenCode="NCA 2011" specimenCount="1" specimenCount-female="1" stateProvince="Limpopo" typeStatus="allotype">
<typeStatus box="[144,227,1059,1081]" pageId="16" pageNumber="463">Allotype</typeStatus>
:
<specimenCount box="[235,263,1059,1080]" pageId="16" pageNumber="463" type="female">1♀</specimenCount>
same data as holotype (
<specimenCode box="[480,580,1059,1081]" collectionCode="NCA" pageId="16" pageNumber="463">NCA 2011</specimenCode>
/2045a).
</materialsCitation>
</paragraph>
<paragraph blockId="16.[144,1104,203,1434]" pageId="16" pageNumber="463">
<materialsCitation collectingDate="2003-11-26" collectionCode="NCA" collectorName="M. Hamer" country="South Africa" latitude="-27.63" location="Umkhuze Game Reserve" longLatPrecision="742" longitude="32.25" pageId="16" pageNumber="463" specimenCode="NCA 2004" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<typeStatus box="[144,236,1092,1113]" pageId="16" pageNumber="463">Paratypes</typeStatus>
:
<collectingCountry box="[242,407,1091,1112]" name="South Africa" pageId="16" pageNumber="463">SOUTH AFRICA</collectingCountry>
:
<emphasis box="[414,555,1091,1112]" italics="true" pageId="16" pageNumber="463">
<collectingRegion box="[414,555,1091,1112]" country="South Africa" name="KwaZulu-Natal" pageId="16" pageNumber="463">KwaZulu­Natal</collectingRegion>
</emphasis>
:
<specimenCount box="[565,594,1091,1112]" pageId="16" pageNumber="463" type="female">1♀</specimenCount>
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE5A93F21A5FB3BFD47FA7C:8EAE3599FFFBA93F2203FBDAFCD1FBBF" box="[598,821,1091,1113]" country="South Africa" latitude="-27.63" longLatPrecision="742" longitude="32.25" name="Umkhuze Game Reserve" pageId="16" pageNumber="463" stateProvince="KwaZulu-Natal">Umkhuze Game Reserve</location>
(
<geoCoordinate box="[832,905,1091,1113]" degrees="27.63" direction="south" orientation="latitude" pageId="16" pageNumber="463" precision="555" value="-27.63">27.63°S</geoCoordinate>
<geoCoordinate box="[910,987,1091,1113]" degrees="32.25" direction="east" orientation="longitude" pageId="16" pageNumber="463" precision="555" value="32.25">32.25°E</geoCoordinate>
),
<date box="[1001,1100,1091,1113]" pageId="16" pageNumber="463" value="2003-11-26">
<collectingDate box="[1001,1100,1091,1113]" pageId="16" pageNumber="463" value="2003-11-26">26.xi.2003</collectingDate>
</date>
,
<collectorName box="[144,241,1116,1137]" pageId="16" pageNumber="463">M. Hamer</collectorName>
, beating tree layer (
<specimenCode box="[428,533,1115,1136]" collectionCode="NCA" pageId="16" pageNumber="463">NCA 2004</specimenCode>
/884)
</materialsCitation>
;
<materialsCitation collectingDate="2003-02-01" collectionCode="NCA" collectorName="S. Lovell" country="South Africa" latitude="-27.63" location="Umkhuze Game Reserve" longLatPrecision="742" longitude="32.25" pageId="16" pageNumber="463" specimenCode="NCA 2004" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount box="[592,621,1115,1136]" pageId="16" pageNumber="463" type="female">1♀</specimenCount>
same locality,
<date box="[764,847,1115,1137]" pageId="16" pageNumber="463" value="2003-02-01">
<collectingDate box="[764,847,1115,1137]" pageId="16" pageNumber="463" value="2003-02-01">1.ii.2003</collectingDate>
</date>
,
<collectorName box="[858,946,1115,1137]" pageId="16" pageNumber="463">S. Lovell</collectorName>
,
<collectingMethod box="[958,1103,1115,1137]" pageId="16" pageNumber="463">yellow pan trap</collectingMethod>
in
<taxonomicName box="[171,351,1139,1161]" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="16" pageNumber="463" phylum="Tracheophyta" rank="species" species="sericea">
<emphasis box="[171,351,1139,1161]" italics="true" pageId="16" pageNumber="463">Terminalia sericea</emphasis>
</taxonomicName>
field (
<specimenCode box="[419,529,1139,1160]" collectionCode="NCA" pageId="16" pageNumber="463">NCA 2004</specimenCode>
/122)
</materialsCitation>
;
<materialsCitation collectingDate="2007-01-12" collectionCode="NCA" collectorName="C. Haddad" country="South Africa" latitude="-26.87" location="Crocodile" longLatPrecision="744" longitude="32.24" municipality="Ndumo Game Reserve" pageId="16" pageNumber="463" specimenCode="NCA 2007" specimenCount="1" specimenCount-female="1" stateProvince="KwaZulu-Natal" typeStatus="paratype">
<specimenCount box="[591,621,1139,1160]" pageId="16" pageNumber="463" type="female">1♀</specimenCount>
<collectingMunicipality box="[629,852,1139,1161]" pageId="16" pageNumber="463">Ndumo Game Reserve</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE5A93F21A5FB3BFD47FA7C:8EAE3599FFFBA93F2334FBEAFC5AFB6F" box="[865,958,1139,1161]" country="South Africa" latitude="-26.87" longLatPrecision="744" longitude="32.24" municipality="Ndumo Game Reserve" name="Crocodile" pageId="16" pageNumber="463" stateProvince="KwaZulu-Natal">Crocodile</location>
farm (
<geoCoordinate box="[1027,1104,1139,1161]" degrees="26.87" direction="south" orientation="latitude" pageId="16" pageNumber="463" precision="555" value="-26.87">26.87°S</geoCoordinate>
<geoCoordinate box="[144,221,1163,1184]" degrees="32.24" direction="east" orientation="longitude" pageId="16" pageNumber="463" precision="555" value="32.24">32.24°E</geoCoordinate>
),
<date box="[236,324,1163,1185]" pageId="16" pageNumber="463" value="2007-01-12">
<collectingDate box="[236,324,1163,1185]" pageId="16" pageNumber="463" value="2007-01-12">12.i.2007</collectingDate>
</date>
,
<collectorName box="[333,433,1163,1185]" pageId="16" pageNumber="463">C. Haddad</collectorName>
, rocky area (
<specimenCode box="[548,649,1163,1185]" collectionCode="NCA" pageId="16" pageNumber="463">NCA 2007</specimenCode>
/4526)
</materialsCitation>
.
<materialsCitation collectingDate="2009-09-11" collectionCode="NCA" collectorName="S. Foord" country="South Africa" latitude="-26.75" location="Limpopo" longLatPrecision="744" longitude="24.42" pageId="16" pageNumber="463" specimenCode="NCA 2011" specimenCount="4" specimenCount-male="4" stateProvince="Limpopo" typeStatus="paratype">
<emphasis box="[718,799,1164,1185]" italics="true" pageId="16" pageNumber="463">
<collectingRegion box="[718,799,1164,1185]" country="South Africa" name="Limpopo" pageId="16" pageNumber="463">Limpopo</collectingRegion>
</emphasis>
:
<specimenCount box="[810,838,1163,1184]" pageId="16" pageNumber="463" type="male">4♂</specimenCount>
same data as holotype (
<specimenCode collectionCode="NCA" pageId="16" pageNumber="463">NCA 2011</specimenCode>
/2045b)
</materialsCitation>
.
</paragraph>
<paragraph blockId="16.[144,1104,203,1434]" pageId="16" pageNumber="463">
Other material examined:
<materialsCitation collectingDate="2008-08-03" collectionCode="NCA" collectorName="X. Combrink" country="South Africa" latitude="-27.63" location="Umkhuze Game Reserve" longLatPrecision="742" longitude="32.25" pageId="16" pageNumber="463" specimenCode="NCA 2011" specimenCount="1" stateProvince="KwaZulu-Natal">
<collectingCountry box="[384,552,1219,1240]" name="South Africa" pageId="16" pageNumber="463">SOUTH AFRICA</collectingCountry>
:
<emphasis box="[562,703,1219,1240]" italics="true" pageId="16" pageNumber="463">
<collectingRegion box="[562,703,1219,1240]" country="South Africa" name="KwaZulu-Natal" pageId="16" pageNumber="463">KwaZulu­Natal</collectingRegion>
</emphasis>
:
<specimenCount box="[716,782,1219,1241]" pageId="16" pageNumber="463" type="generic">1 imm.</specimenCount>
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE5A93F21A5FB3BFD47FA7C:8EAE3599FFFBA93F2341FB5AFC1DFB3F" box="[788,1017,1219,1241]" country="South Africa" latitude="-27.63" longLatPrecision="742" longitude="32.25" name="Umkhuze Game Reserve" pageId="16" pageNumber="463" stateProvince="KwaZulu-Natal">Umkhuze Game Reserve</location>
(
<geoCoordinate box="[1030,1103,1219,1241]" degrees="27.63" direction="south" orientation="latitude" pageId="16" pageNumber="463" precision="555" value="-27.63">27.63°S</geoCoordinate>
<geoCoordinate box="[144,212,1243,1265]" degrees="32.25" direction="east" orientation="longitude" pageId="16" pageNumber="463" precision="555" value="32.25">32.25E</geoCoordinate>
),
<date box="[226,325,1243,1265]" pageId="16" pageNumber="463" value="2008-08-03">
<collectingDate box="[226,325,1243,1265]" pageId="16" pageNumber="463" value="2008-08-03">3.viii.2008</collectingDate>
</date>
,
<collectorName box="[333,453,1243,1265]" pageId="16" pageNumber="463">X. Combrink</collectorName>
,
<collectingMethod box="[461,563,1243,1265]" pageId="16" pageNumber="463">pitfall traps</collectingMethod>
(
<specimenCode box="[573,671,1243,1264]" collectionCode="NCA" pageId="16" pageNumber="463">NCA 2011</specimenCode>
/2048)
</materialsCitation>
.
<materialsCitation collectingDate="2007-06-16" collectionCode="NCA" collectorName="G. Ellis" country="South Africa" latitude="-25.38" location="Makhuthwanini" longLatPrecision="748" longitude="31.6" municipality="Kruger National Park" pageId="16" pageNumber="463" specimenCode="NCA 2008" specimenCount="3" specimenCount-male="3" stateProvince="Mpumalanga">
<emphasis box="[739,859,1243,1264]" italics="true" pageId="16" pageNumber="463">
<collectingRegion box="[739,859,1243,1264]" country="South Africa" name="Mpumalanga" pageId="16" pageNumber="463">Mpumalanga</collectingRegion>
</emphasis>
:
<specimenCount box="[870,953,1243,1265]" pageId="16" pageNumber="463" type="male">3 imm. ♂</specimenCount>
<collectingMunicipality pageId="16" pageNumber="463">Kruger National Park</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE5A93F21A5FB3BFD47FA7C:8EAE3599FFFBA93F2091FB6AFEB0FAEF" box="[196,340,1267,1289]" country="South Africa" latitude="-25.38" longLatPrecision="748" longitude="31.6" municipality="Kruger National Park" name="Makhuthwanini" pageId="16" pageNumber="463" stateProvince="Mpumalanga">Makhuthwanini</location>
(
<geoCoordinate box="[352,425,1267,1289]" degrees="25.38" direction="south" orientation="latitude" pageId="16" pageNumber="463" precision="555" value="-25.38">25.38°S</geoCoordinate>
<geoCoordinate box="[432,509,1267,1289]" degrees="31.60" direction="east" orientation="longitude" pageId="16" pageNumber="463" precision="555" value="31.6">31.60°E</geoCoordinate>
),
<date box="[524,623,1267,1289]" pageId="16" pageNumber="463" value="2007-06-16">
<collectingDate box="[524,623,1267,1289]" pageId="16" pageNumber="463" value="2007-06-16">16.vi.2007</collectingDate>
</date>
,
<collectingMethod box="[633,738,1267,1289]" pageId="16" pageNumber="463">pitfall traps</collectingMethod>
,
<collectorName box="[748,818,1267,1289]" pageId="16" pageNumber="463">G. Ellis</collectorName>
(
<specimenCode box="[829,930,1267,1289]" collectionCode="NCA" pageId="16" pageNumber="463">NCA 2008</specimenCode>
/68, 2008/69)
</materialsCitation>
.
</paragraph>
<paragraph blockId="16.[144,1104,203,1434]" box="[144,1068,1303,1330]" pageId="16" pageNumber="463">
Distribution:
<collectingCountry box="[294,437,1304,1330]" name="South Africa" pageId="16" pageNumber="463">South Africa</collectingCountry>
(
<collectingRegion box="[451,631,1304,1330]" country="South Africa" name="KwaZulu-Natal" pageId="16" pageNumber="463">KwaZulu­Natal</collectingRegion>
,
<collectingRegion box="[642,745,1304,1330]" country="South Africa" name="Limpopo" pageId="16" pageNumber="463">Limpopo</collectingRegion>
and
<collectingRegion box="[799,949,1304,1330]" country="South Africa" name="Mpumalanga" pageId="16" pageNumber="463">Mpumalanga</collectingRegion>
) (
<figureCitation box="[972,1056,1303,1330]" captionStart="Figs 5759" captionStartId="27.[347,386,1654,1676]" captionTargetBox="[181,1141,609,1633]" captionTargetId="figure-176@27.[132,1190,602,1636]" captionTargetPageId="27" captionText="Figs 5759. Distribution of Heriaeus spp. in the Afrotropical Region." figureDoi="http://doi.org/10.5281/zenodo.7918679" httpUri="https://zenodo.org/record/7918679/files/figure.png" pageId="16" pageNumber="463">Fig. 58</figureCitation>
).
</paragraph>
<paragraph blockId="16.[144,1104,203,1434]" pageId="16" pageNumber="463">Natural history: Collected from yellow pan traps and pitfall traps as well as sweeping of vegetation. Adult females collected during October, males during September and October, and juveniles during June and August.</paragraph>
</subSubSection>
</treatment>
</document>