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<document id="FD82656AA60D65DBC3037424F752700D" ID-DOI="10.5281/zenodo.7887652" ID-ISSN="1942-1354" ID-Zenodo-Dep="7887652" ID-ZooBank="F008A040-C404-46DA-B879-9543868A6D21O" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683049894525" checkinUser="felipe" docAuthor="Ziani, Stefano &amp; Keith, Denis" docDate="2023" docId="039487F9D415FFF3FF60257943F0B2BB" docLanguage="en" docName="InsectaMundi.2023.0974.14..pdf" docOrigin="Insecta Mundi 2023 (974)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Onthophagus aeneopiceus d'Orbigny 1902" docType="treatment" docVersion="3" lastPageNumber="3" masterDocId="FFADFF81D417FFF7FFA0230A466DB344" masterDocTitle="Third contribution to the knowledge of Arabian Onthophagini (Coleoptera: Scarabaeidae). Onthophagus aeneopiceus dOrbigny, 1902, a new record for Yemen and for the Palaearctic ecozone" masterLastPageNumber="4" masterPageNumber="1" pageNumber="1" updateTime="1699495793530" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="190DB3253BEB4815C92DF26975CEB7E7">Third contribution to the knowledge of Arabian Onthophagini (Coleoptera: Scarabaeidae). Onthophagus aeneopiceus dOrbigny, 1902, a new record for Yemen and for the Palaearctic ecozone</mods:title>
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<treatment id="039487F9D415FFF3FF60257943F0B2BB" LSID="urn:lsid:plazi:treatment:039487F9D415FFF3FF60257943F0B2BB" httpUri="http://treatment.plazi.org/id/039487F9D415FFF3FF60257943F0B2BB" lastPageId="4" lastPageNumber="3" pageId="2" pageNumber="1">
<subSubSection id="C3276564D415FFF5FF60257944B4B5D4" box="[192,729,1651,1680]" pageId="2" pageNumber="1" type="nomenclature">
<paragraph id="8B8236EFD415FFF5FF60257944B4B5D4" blockId="2.[192,729,1651,1713]" box="[192,729,1651,1680]" pageId="2" pageNumber="1">
<heading id="D0CA8183D415FFF5FF60257944B4B5D4" box="[192,729,1651,1680]" fontSize="12" level="2" pageId="2" pageNumber="1" reason="2">
<taxonomicName id="4C3D4D6CD415FFF5FF60257944B4B5D4" ID-CoL="7XM9B" authority="d'Orbigny, 1902" authorityName="d'Orbigny" authorityYear="1902" box="[192,729,1651,1680]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="aeneopiceus">
<emphasis id="B949EAFDD415FFF5FF60257944B4B5D4" bold="true" box="[192,729,1651,1680]" pageId="2" pageNumber="1">
<emphasis id="B949EAFDD415FFF5FF602579446AB5D4" bold="true" box="[192,519,1651,1680]" italics="true" pageId="2" pageNumber="1">Onthophagus aeneopiceus</emphasis>
dOrbigny, 1902
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3276564D415FFF5FF60259D4740B5F5" box="[192,301,1687,1713]" pageId="2" pageNumber="1" type="description">
<paragraph id="8B8236EFD415FFF5FF60259D4740B5F5" blockId="2.[192,729,1651,1713]" box="[192,301,1687,1713]" pageId="2" pageNumber="1">
(
<figureCitation id="13062A6AD415FFF5FF68259D4748B5F5" box="[200,293,1687,1713]" captionStart="Figures 14" captionStartId="3.[192,279,1251,1277]" captionTargetBox="[282,1298,252,1200]" captionTargetId="figure-440@3.[271,1361,233,1222]" captionTargetPageId="3" captionText="Figures 14. Onthophagus aeneopiceus dOrbigny, 1902 [Yemen]. 1) Male dorsal habitus. 2) Parameres, lateral view.3) Parameres, dorsal view. 4) Endophallus. Photos by A. Degiovanni, edited by G. Fiumi." figureDoi="http://doi.org/10.5281/zenodo.7887654" httpUri="https://zenodo.org/record/7887654/files/figure.png" pageId="2" pageNumber="1">Fig. 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3276564D415FFF5FF4025B5440CB450" pageId="2" pageNumber="1" type="reference_group">
<paragraph id="8B8236EFD415FFF5FF4025B5440CB450" blockId="2.[224,1440,1727,1812]" pageId="2" pageNumber="1">
<treatmentCitationGroup id="AB2D11C1D415FFF5FF4025B5440CB450" pageId="2" pageNumber="1">
<taxonomicName id="4C3D4D6CD415FFF5FF4025B544D3B593" ID-CoL="7XM9B" authority="d'Orbigny 1902: 156" authorityName="d'Orbigny" authorityPageNumber="156" authorityYear="1902" box="[224,702,1727,1751]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="aeneopiceus">
<emphasis id="B949EAFDD415FFF5FF4025B5478EB593" box="[224,483,1727,1751]" italics="true" pageId="2" pageNumber="1">Onthophagus aeneopiceus</emphasis>
dOrbigny 1902: 156
</taxonomicName>
; dOrbigny 1905: 488; dOrbigny 1913: 376; dOrbigny, 1922: 609;
<treatmentCitation id="0A9C10FED415FFF5FACD25CA444EB5B1" author="Boucomont A &amp; Gillet JJE" page="151" pageId="2" pageNumber="1" year="1927">
<bibRefCitation id="EFAC4B1ED415FFF5FACD25CA444EB5B1" author="Boucomont A &amp; Gillet JJE" pageId="2" pageNumber="1" pagination="103 - 263" refId="ref2174" refString="Boucomont A, Gillet JJE. 1927. Fam. Scarabaeidae. Subfam. Coprinae II. p. 103 - 263. In: Gillet JJE, Boucomont A (eds.). Scarabaeidae: Coprinae, Termitotroginae. Partes 38 et 90, Vol. 19 (3). In: Schenkling S (ed.). Coleopterorum Catalogus. W. Junk; Berlin. 265 p." type="book chapter" year="1927">Boucomont and Gillet 1927: 151</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9C10FED415FFF5FD8E25D4449BB5B2" author="Balthasar V." box="[558,758,1758,1782]" page="263" pageId="2" pageNumber="1" year="1963">
<bibRefCitation id="EFAC4B1ED415FFF5FD8E25D4449BB5B2" author="Balthasar V." box="[558,758,1758,1782]" pageId="2" pageNumber="1" pagination="1 - 628" refId="ref2013" refString="Balthasar V. 1963. Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 2. Coprinae. Tschechoslowakischen Akademie der Wissenschaften; Prag 2: 1 - 628." type="journal article" year="1963">Balthasar 1963: 263</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9C10FED415FFF5FCA125D445D0B5B1" author="Balthasar V." box="[769,957,1757,1782]" page="65" pageId="2" pageNumber="1" year="1969">
<bibRefCitation id="EFAC4B1ED415FFF5FCA125D445D0B5B1" author="Balthasar V." box="[769,957,1757,1782]" pageId="2" pageNumber="1" pagination="59 - 73" refId="ref2050" refString="Balthasar V. 1969. Contributions a la connaissance de la faune entomologique de la Cote-d'Ivoire (J. Decelle, 1961 - 1964). XXXII. Coleoptera Scarabaeidae et Aphodiidae. Annales du Musee Royal de l'Afrique Centrale / Sciences Zoologiques 175: 59 - 73." type="journal article" year="1969">Balthasar 1969: 65</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9C10FED415FFF5FC6825D442EFB5B1" author="Ferreira MC" box="[968,1154,1757,1782]" page="557" pageId="2" pageNumber="1" year="1972">
<bibRefCitation id="EFAC4B1ED415FFF5FC6825D442EFB5B1" author="Ferreira MC" box="[968,1154,1757,1782]" pageId="2" pageNumber="1" pagination="1 - 1088" refId="ref2248" refString="Ferreira MC. 1972. Os escarabideos de Africa (Sul do Saara). I. Revista de Entomologia de Mocambique 11 (1968 - 1969): 1 - 1088." type="journal article" year="1972">Ferreira 1972: 557</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9C10FED415FFF5FB2D25D44324B5B2" author="Balthasar V." box="[1165,1353,1758,1782]" page="22" pageId="2" pageNumber="1" year="1972">
<bibRefCitation id="EFAC4B1ED415FFF5FB2D25D44324B5B2" author="Balthasar V." box="[1165,1353,1758,1782]" pageId="2" pageNumber="1" pagination="18 - 32" refId="ref2099" refString="Balthasar V. 1972. Coleoptera aus Nordostafrika. Scarabaeoidea: Scarabaeidae und Aphodiidae. 135. Beitrag zur Kenntnis der Scarabaeoidea. Notulae Entomologicae 52: 18 - 32." type="journal article" year="1972">Balthasar 1972: 22</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9C10FED415FFF5FAF425D4470EB450" author="Zunino M." page="153" pageId="2" pageNumber="1" year="1975">
<bibRefCitation id="EFAC4B1ED415FFF5FAF425D4470EB450" author="Zunino M." pageId="2" pageNumber="1" pagination="151 - 194" refId="ref2709" refString="Zunino M. 1975. Revisione delle specie paleartiche del sottogenere Onthophagus (sensu strictu) Latr. (Coleoptera, Scarabaeoidea). I tipi di H. d'Orbigny, A. Raffray e A. Boucomont nel Museum National d'Histoire Naturelle di Parigi. Bollettino del Museo di Zoologia dell'Universita di Torino 7: 151 - 194." type="journal article" year="1975">Zunino 1975: 153</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A9C10FED415FFF5FECD25F64430B450" author="Barbero E &amp; Borghesio L &amp; Dellacasa G &amp; Dellacasa M." box="[365,605,1788,1812]" page="243" pageId="2" pageNumber="1" year="1998">
<bibRefCitation id="EFAC4B1ED415FFF5FECD25F64430B450" author="Barbero E &amp; Borghesio L &amp; Dellacasa G &amp; Dellacasa M." box="[365,605,1788,1812]" pageId="2" pageNumber="1" pagination="233 - 254" refId="ref2130" refString="Barbero E, Borghesio L, Dellacasa G, Dellacasa M. 1998. Contribution to the knowledge of Scarabaeidae and Aphodiidae from southern Ethiopia (Coleoptera Scarabaeoidea). Bollettino della Societa Entomologica Italiana 130 (3): 233 - 254." type="journal article" year="1998">Barbero et al. 1998: 243</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3276564D415FFF4FF60242E444AB6F5" lastPageId="3" lastPageNumber="2" pageId="2" pageNumber="1" type="materials_examined">
<paragraph id="8B8236EFD415FFF5FF60242E4429B47A" blockId="2.[192,580,1828,1854]" box="[192,580,1828,1854]" pageId="2" pageNumber="1">
<emphasis id="B949EAFDD415FFF5FF60242E4736B47A" bold="true" box="[192,347,1828,1854]" pageId="2" pageNumber="1">
<typeStatus id="5486884DD415FFF5FF60242E4694B47A" box="[192,249,1828,1854]" pageId="2" pageNumber="1">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32A767FD415FFF5FECB242E47A3B47A" box="[363,462,1828,1854]" name="Eritrea" pageId="2" pageNumber="1">Erythrée</collectingCountry>
” [
<collectingCountry id="F32A767FD415FFF5FE44242E4457B47A" box="[484,570,1828,1854]" name="Eritrea" pageId="2" pageNumber="1">Eritrea</collectingCountry>
].
</paragraph>
<paragraph id="8B8236EFD415FFF4FF602444444AB6F5" blockId="2.[192,1299,1869,1896]" lastBlockId="3.[192,1440,1364,1457]" lastPageId="3" lastPageNumber="2" pageId="2" pageNumber="1">
<emphasis id="B949EAFDD415FFF5FF6024444704B423" bold="true" box="[192,361,1869,1896]" pageId="2" pageNumber="1">Type material.</emphasis>
<materialsCitation id="3B553CB2D415FFF4FECF2447444AB6F5" collectionCode="MNHN" collectorName="Coll. H. d'Orbigny" country="Eritrea" lastPageId="3" lastPageNumber="2" location="Erythree" pageId="2" pageNumber="1" specimenCount="2" typeStatus="holotype">
<typeStatus id="5486884DD415FFF5FECF244747BAB423" box="[367,471,1869,1895]" pageId="2" pageNumber="1" type="holotype">Holotype</typeStatus>
male, fixed by original designation, at
<collectionCode id="ED2CAE2AD415FFF5FC262444458DB42C" box="[902,992,1870,1896]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="1">MNHN</collectionCode>
, examined through photos.
<emphasis id="B949EAFDD414FFF4FF60265E4702B62A" bold="true" box="[192,367,1364,1390]" pageId="3" pageNumber="2">Type labelling.</emphasis>
1
<superScript id="7C489BA7D414FFF4FE25265E47FDB626" attach="right" box="[389,400,1364,1378]" fontSize="6" pageId="3" pageNumber="2">st</superScript>
, light yellow, handwritten in black: “
<collectingCountry id="F32A767FD414FFF4FC95265E45F4B62A" box="[821,921,1364,1390]" name="Eritrea" pageId="3" pageNumber="2">Erythrée</collectingCountry>
”; 2
<superScript id="7C489BA7D414FFF4FC1E265E45A2B626" attach="none" box="[958,975,1364,1378]" fontSize="6" pageId="3" pageNumber="2">nd</superScript>
, light blue, printed in black: “MUSEUM PARIS /
<collectorName id="26C85339D414FFF4FE80267C4782B6D4" box="[288,495,1398,1424]" pageId="3" pageNumber="2">Coll. H. dOrbigny</collectorName>
/ 1915”; 3
<superScript id="7C489BA7D414FFF4FDC2267F441DB6C7" attach="none" box="[610,624,1397,1411]" fontSize="6" pageId="3" pageNumber="2">rd</superScript>
, white, handwritten and printed in black: “Vidit /
<bibRefCitation id="EFAC4B1ED414FFF4FB04267C435AB6D4" author="Zunino M." box="[1188,1335,1397,1424]" pageId="3" pageNumber="2" pagination="151 - 194" refId="ref2709" refString="Zunino M. 1975. Revisione delle specie paleartiche del sottogenere Onthophagus (sensu strictu) Latr. (Coleoptera, Scarabaeoidea). I tipi di H. d'Orbigny, A. Raffray e A. Boucomont nel Museum National d'Histoire Naturelle di Parigi. Bollettino del Museo di Zoologia dell'Universita di Torino 7: 151 - 194." type="journal article" year="1975">Zunino 1975</bibRefCitation>
”; 4
<superScript id="7C489BA7D414FFF4FAFA267F4305B6C7" attach="right" box="[1370,1384,1397,1411]" fontSize="6" pageId="3" pageNumber="2">th</superScript>
, red, printed in black: “
<typeStatus id="5486884DD414FFF4FE28269D4477B6F5" box="[392,538,1431,1457]" pageId="3" pageNumber="2" type="holotype">HOLOTYPE</typeStatus>
”.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF426667D414FFF4FF6027E942B2B65B" ID-DOI="http://doi.org/10.5281/zenodo.7887654" ID-Zenodo-Dep="7887654" httpUri="https://zenodo.org/record/7887654/files/figure.png" pageId="3" pageNumber="2" startId="3.[192,279,1251,1277]" targetBox="[282,1298,252,1200]" targetPageId="3" targetType="figure">
<paragraph id="8B8236EFD414FFF4FF6027E942B2B65B" blockId="3.[192,1440,1251,1312]" pageId="3" pageNumber="2">
<emphasis id="B949EAFDD414FFF4FF6027E9473CB7B9" bold="true" box="[192,337,1251,1277]" pageId="3" pageNumber="2">Figures 14.</emphasis>
<taxonomicName id="4C3D4D6CD414FFF4FEF827E9455EB7BA" authority="d'Orbigny, 1902" authorityName="d'Orbigny" authorityYear="1902" box="[344,819,1251,1278]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="aeneopiceus">
<emphasis id="B949EAFDD414FFF4FEF827E9441AB7B9" box="[344,631,1251,1277]" italics="true" pageId="3" pageNumber="2">Onthophagus aeneopiceus</emphasis>
dOrbigny, 1902
</taxonomicName>
[Yemen].
<emphasis id="B949EAFDD414FFF4FC0A27E945AFB7B9" bold="true" box="[938,962,1251,1277]" pageId="3" pageNumber="2">1)</emphasis>
Male dorsal habitus.
<emphasis id="B949EAFDD414FFF4FB1827E942BCB7B9" bold="true" box="[1208,1233,1251,1277]" pageId="3" pageNumber="2">2)</emphasis>
Parameres, lateral view.
<emphasis id="B949EAFDD414FFF4FF5E260F477AB65B" bold="true" box="[254,279,1285,1311]" pageId="3" pageNumber="2">3)</emphasis>
Parameres, dorsal view.
<emphasis id="B949EAFDD414FFF4FD89260F442FB65B" bold="true" box="[553,578,1285,1311]" pageId="3" pageNumber="2">4)</emphasis>
Endophallus. Photos by A. Degiovanni, edited by G. Fiumi.
</paragraph>
</caption>
<subSubSection id="C3276564D414FFF4FF6026CB43F3B69F" box="[192,1438,1473,1499]" pageId="3" pageNumber="2" type="distribution">
<paragraph id="8B8236EFD414FFF4FF6026CB43F3B69F" blockId="3.[192,1438,1473,1499]" box="[192,1438,1473,1499]" pageId="3" pageNumber="2">
<emphasis id="B949EAFDD414FFF4FF6026CB4736B69F" bold="true" box="[192,347,1473,1499]" pageId="3" pageNumber="2">Distribution.</emphasis>
<collectingCountry id="F32A767FD414FFF4FEC126CB47C3B69F" box="[353,430,1473,1499]" name="Eritrea" pageId="3" pageNumber="2">Eritrea</collectingCountry>
(dOrbigny 1902).
<collectingCountry id="F32A767FD414FFF4FD2026CB44A5B69F" box="[640,712,1473,1499]" name="Sudan" pageId="3" pageNumber="2">Sudan</collectingCountry>
,
<collectingCountry id="F32A767FD414FFF4FD7326CB4559B69F" box="[723,820,1473,1499]" name="Ethiopia" pageId="3" pageNumber="2">Ethiopia</collectingCountry>
,
<collectingCountry id="F32A767FD414FFF4FC9E26CB45E9B69F" box="[830,900,1473,1499]" name="Kenya" pageId="3" pageNumber="2">Kenya</collectingCountry>
(dOrbigny 1913).
<collectingCountry id="F32A767FD414FFF4FBF726CB42B4B69F" box="[1111,1241,1473,1499]" name="Ivory Coast" pageId="3" pageNumber="2">Ivory Coast</collectingCountry>
(
<bibRefCitation id="EFAC4B1ED414FFF4FB4726CB43FFB69F" author="Balthasar V." box="[1255,1426,1473,1499]" pageId="3" pageNumber="2" pagination="59 - 73" refId="ref2050" refString="Balthasar V. 1969. Contributions a la connaissance de la faune entomologique de la Cote-d'Ivoire (J. Decelle, 1961 - 1964). XXXII. Coleoptera Scarabaeidae et Aphodiidae. Annales du Musee Royal de l'Afrique Centrale / Sciences Zoologiques 175: 59 - 73." type="journal article" year="1969">Balthasar 1969</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3276564D414FFF4FF6026E043CCB562" pageId="3" pageNumber="2" type="materials_examined">
<paragraph id="8B8236EFD414FFF4FF6026E043CCB562" blockId="3.[192,1441,1514,1575]" pageId="3" pageNumber="2">
<emphasis id="B949EAFDD414FFF4FF6026E047CEB540" bold="true" box="[192,419,1514,1540]" pageId="3" pageNumber="2">Material examined.</emphasis>
<materialsCitation id="3B553CB2D414FFF4FE0A26E1434FB541" box="[426,1314,1514,1541]" collectingDate="1976-03-30" collectionCode="DKCC" collectorName="R. Naviaux" country="Yemen" location="Jabal Hadish" pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="Ibb">
<emphasis id="B949EAFDD414FFF4FE0A26E1447FB541" bold="true" box="[426,530,1515,1541]" pageId="3" pageNumber="2">
<collectingCountry id="F32A767FD414FFF4FE0A26E14460B541" box="[426,525,1515,1541]" name="Yemen" pageId="3" pageNumber="2">YEMEN</collectingCountry>
:
</emphasis>
<collectingRegion id="49F9F80DD414FFF4FDB826E04453B540" box="[536,574,1514,1540]" country="Yemen" name="Ibb" pageId="3" pageNumber="2">Ibb</collectingRegion>
gov., W of
<location id="8EE26034D414FFF4FD1C26E04520B540" LSID="urn:lsid:plazi:treatment:039487F9D415FFF3FF60257943F0B2BB:8EE26034D414FFF4FD1C26E04520B540" box="[700,845,1514,1540]" country="Yemen" name="Jabal Hadish" pageId="3" pageNumber="2" stateProvince="Ibb">Jabal Hadish</location>
,
<date id="FF83102FD414FFF4FCF726E045A1B540" box="[855,972,1514,1540]" pageId="3" pageNumber="2" value="1976-03-30">
<collectingDate id="EFC7E9C7D414FFF4FCF726E045A1B540" box="[855,972,1514,1540]" pageId="3" pageNumber="2" value="1976-03-30">30.iii.1976</collectingDate>
</date>
,
<collectorName id="26C85339D414FFF4FC7826E1423EB540" box="[984,1107,1514,1541]" pageId="3" pageNumber="2">R. Naviaux</collectorName>
legit
<specimenCount id="9D3BFD66D414FFF4FB3126E142D7B540" box="[1169,1210,1515,1541]" pageId="3" pageNumber="2" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED2CAE2AD414FFF4FB6826E14377B541" box="[1224,1306,1515,1541]" pageId="3" pageNumber="2">DKCC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B553CB2D414FFF4FA8D26E043F0B562" collectingDate="2005-10-26" collectingDateMax="2005-10-27" collectingDateMin="2005-10-26" collectionCode="NMPC" collectorName="D. Kral" country="Yemen" elevation="1600" latitude="13.874445" location="Jumai - Wadi Maytam" longLatPrecision="21" longitude="44.295834" pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="Ibb">
<collectingRegion id="49F9F80DD414FFF4FA8D26E0433EB540" box="[1325,1363,1514,1540]" country="Yemen" name="Ibb" pageId="3" pageNumber="2">Ibb</collectingRegion>
gov.,
<locationDeviation id="9A302FCDD414FFF4FA3326E0473DB562" location="Najd" pageId="3" pageNumber="2">
<quantity id="4CC59B0AD414FFF4FA3326E04688B562" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="2" unit="km" value="5.0">5 km</quantity>
SW Najd
</locationDeviation>
al
<location id="8EE26034D414FFF4FECF25064408B563" LSID="urn:lsid:plazi:treatment:039487F9D415FFF3FF60257943F0B2BB:8EE26034D414FFF4FECF25064408B563" box="[367,613,1548,1575]" country="Yemen" latitude="13.874445" longLatPrecision="21" longitude="44.295834" name="Jumai - Wadi Maytam" pageId="3" pageNumber="2" stateProvince="Ibb">Jumai - Wadi Maytam</location>
<quantity id="4CC59B0AD414FFF4FDCB250644ADB563" box="[619,704,1548,1575]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="2" unit="m" value="1600.0">
<elevation id="0010D1DCD414FFF4FDCB250644ADB563" box="[619,704,1548,1575]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="2" unit="m" value="1600.0">1600 m</elevation>
</quantity>
,
<geoCoordinate id="EE095028D414FFF4FD6A25064526B563" box="[714,843,1548,1575]" degrees="13" direction="north" minutes="52" orientation="latitude" pageId="3" pageNumber="2" precision="15" seconds="28" value="13.874445">13°5228″N</geoCoordinate>
<geoCoordinate id="EE095028D414FFF4FCF1250645B8B563" box="[849,981,1548,1575]" degrees="44" direction="east" minutes="17" orientation="longitude" pageId="3" pageNumber="2" precision="15" seconds="45" value="44.295834">44°1745″ E</geoCoordinate>
,
<date id="FF83102FD414FFF4FC7F2506421CB563" box="[991,1137,1548,1575]" pageId="3" pageNumber="2" value="2005-10-26" valueMax="2005-10-27" valueMin="2005-10-26">
<collectingDate id="EFC7E9C7D414FFF4FC7F2506421CB563" box="[991,1137,1548,1575]" pageId="3" pageNumber="2" value="2005-10-26" valueMax="2005-10-27" valueMin="2005-10-26">26-27.x.2005</collectingDate>
</date>
,
<collectorName id="26C85339D414FFF4FBDB250742A1B562" box="[1147,1228,1548,1575]" pageId="3" pageNumber="2">D. Král</collectorName>
legit
<specimenCount id="9D3BFD66D414FFF4FAA72506435DB562" box="[1287,1328,1548,1574]" pageId="3" pageNumber="2" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED2CAE2AD414FFF4FA9E250643F8B562" box="[1342,1429,1548,1574]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="2">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276564D414FFF3FF60253C43F0B2BB" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="discussion">
<paragraph id="8B8236EFD414FFF4FF60253C4350B535" blockId="3.[192,1361,1590,1650]" pageId="3" pageNumber="2">
<emphasis id="B949EAFDD414FFF4FF60253C4743B514" bold="true" box="[192,302,1590,1616]" pageId="3" pageNumber="2">Remarks.</emphasis>
New record for
<collectingCountry id="F32A767FD414FFF4FE44253C445DB514" box="[484,560,1590,1616]" name="Yemen" pageId="3" pageNumber="2">Yemen</collectingCountry>
and new record for the Palaearctic ecozone (
<emphasis id="B949EAFDD414FFF4FB86253C420FB50B" box="[1062,1122,1590,1615]" italics="true" pageId="3" pageNumber="2">sensu</emphasis>
<bibRefCitation id="EFAC4B1ED414FFF4FBC8253C4328B514" author="Lobl I &amp; Lobl D." box="[1128,1349,1590,1616]" pageId="3" pageNumber="2" refId="ref2281" refString="Lobl I, Lobl D. 2016. Catalogue of Palaearctic Coleoptera. Volume 3. Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. Brill; Leiden, Boston. 983 p. https: // doi. org / 10.1163 / 9789004309142" type="book" year="2016">Löbl and Löbl 2016</bibRefCitation>
). The species can be inserted in the key to the Arabian
<taxonomicName id="4C3D4D6CD414FFF4FCF2255D42ADB535" authority="(Ziani et al. 2019)" baseAuthorityName="Ziani" baseAuthorityYear="2019" box="[850,1216,1623,1650]" class="Insecta" family="Scarabaeidae" genus="Onthophagini" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
Onthophagini (
<bibRefCitation id="EFAC4B1ED414FFF4FBA0255D42D4B535" author="Ziani S &amp; Abdel-Dayem MS &amp; Aldhafer HM &amp; Barbero E." box="[1024,1209,1623,1650]" pageId="3" pageNumber="2" pagination="1 - 36" refId="ref2656" refString="Ziani S, Abdel-Dayem MS, Aldhafer HM, Barbero E. 2019. An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae). Zootaxa 4658 (1): 1 - 36. https: // doi. org / 10.11646 / zootaxa. 4658.1.1" type="journal article" year="2019">Ziani et al. 2019</bibRefCitation>
)
</taxonomicName>
as follows:
</paragraph>
<paragraph id="8B8236EFD414FFF4FF60258B43F0B420" blockId="3.[192,1442,1665,1927]" pageId="3" pageNumber="2">
26. Pronotal punctation simple or ocellate on disc; quite glossy; major males with clypeus produced to a blunt point, which is strongly reflexed, and with vertex of the head produced in a pair of unconnected horns, very broad and directed backward at the base, with a small sharp tooth on the outer edge of each, beyond which they curve upward and are tapered, bisinuate, and sharp at the tip; female margin of clypeus is truncate at middle; blackish brown, elytra with basal and apical yellowish red spots which are usually confluent. Length 5.5 to 7.0 mm....................................................................................
<taxonomicName id="4C3D4D6CD414FFF4FCD9244043F0B420" authority="Gillet, 1925" authorityName="Gillet" authorityYear="1925" box="[889,1437,1866,1893]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="falsus" subGenus="Onthophagus">
<emphasis id="B949EAFDD414FFF4FCD9244043F0B420" bold="true" box="[889,1437,1866,1893]" pageId="3" pageNumber="2">
<emphasis id="B949EAFDD414FFF4FCD924404279B421" bold="true" box="[889,1044,1866,1893]" italics="true" pageId="3" pageNumber="2">Onthophagus</emphasis>
(
<emphasis id="B949EAFDD414FFF4FB83244042D0B421" bold="true" box="[1059,1213,1866,1893]" italics="true" pageId="3" pageNumber="2">Onthophagus</emphasis>
)
<emphasis id="B949EAFDD414FFF4FB6E2440437DB421" bold="true" box="[1230,1296,1866,1893]" italics="true" pageId="3" pageNumber="2">falsus</emphasis>
Gillet, 1925
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8B8236EFD414FFF4FF60246743F0B4C1" blockId="3.[192,1442,1665,1927]" box="[192,1437,1900,1927]" pageId="3" pageNumber="2">
— Pronotal punctation raspish on disc..........................................................
<emphasis id="B949EAFDD414FFF4FADB246643FAB4C2" bold="true" box="[1403,1431,1900,1926]" pageId="3" pageNumber="2">26</emphasis>
</paragraph>
<paragraph id="8B8236EFD413FFF3FF6023DC43F0B211" blockId="4.[192,1440,214,512]" pageId="4" pageNumber="3">
26 Pygidial largest punctures close on disc, separated by less than one diameter; pronotum dull; head not sinuate near obviously produced genae; clypeofrontal carina with a very long backwards horn, in major males, sometimes extending beyond pronotal hind margin; pronotum without obvious gibbosities; blackish brown. Length 4.5 to 6.0 mm..............
<taxonomicName id="4C3D4D6CD413FFF3FC13223143F0B211" authority="d'Orbigny, 1902" authorityName="d'Orbigny" authorityYear="1902" box="[947,1437,315,342]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="aeneopiceus">
<emphasis id="B949EAFDD413FFF3FC13223143F0B211" bold="true" box="[947,1437,315,342]" pageId="4" pageNumber="3">
<emphasis id="B949EAFDD413FFF3FC13223142B0B211" bold="true" box="[947,1245,315,342]" italics="true" pageId="4" pageNumber="3">Onthophagus aeneopiceus</emphasis>
dOrbigny, 1902
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8B8236EFD413FFF3FF60225443F0B2BB" blockId="4.[192,1440,214,512]" pageId="4" pageNumber="3">
— Pygidial largest punctures spaced on disc, separated by one diameter, or more; pronotum glossy; head barely sinuate near obviously produced genae; clypeofrontal carina with a short backwards horn, in major males, as long as the occipital carina, or a little more, pronotum with an anteromedian bilobate gibbosity; blackish brown, elytra sometimes with apical dark red band. Length 5.0 to 6.0 mm..............................................
<taxonomicName id="4C3D4D6CD413FFF3FC0722EF43F0B2BB" authority="d'Orbigny, 1902" authorityName="d'Orbigny" authorityYear="1902" box="[935,1437,485,512]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="trapezicornis">
<emphasis id="B949EAFDD413FFF3FC0722EF43F0B2BB" bold="true" box="[935,1437,485,512]" pageId="4" pageNumber="3">
<emphasis id="B949EAFDD413FFF3FC0722EF42B3B2BB" bold="true" box="[935,1246,485,512]" italics="true" pageId="4" pageNumber="3">Onthophagus trapezicornis</emphasis>
dOrbigny, 1902
</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>