197 lines
23 KiB
XML
197 lines
23 KiB
XML
<document id="C670776A05D94A5F4B8CAC35A753FC68" ID-DOI="10.1206/0003-0082(2006)3535[1:PATROT]2.0.CO;2" ID-ISSN="0003-0082" ID-Zenodo-Dep="5385333" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630301546832" checkinUser="carolina" docAuthor="STIASSNY, MELANIE L. J & SPARKS, JOHN S" docDate="2006" docId="039487E1F04EFFBBD781FA1B06EFF955" docLanguage="en" docName="N3535.pdf" docOrigin="American Museum Novitates 3535 (1)" docSource="http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0082(2006)3535%5B1%3APATROT%5D2.0.CO%3B2" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.3:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="3" docTitle="Katria STIASSNY & SPARKS 2006, new genus" docType="treatment" docUuid="994D87BC-800B-4491-BFAE-5E561D3689C3" docUuidSource="ZooBank" docVersion="5" lastPageNumber="52" masterDocId="FFADFF99F07DFF88D776FE63052CFF8C" masterDocTitle="Phylogeny and Taxonomic Revision of the Endemic Malagasy Genus Ptychochromis (Teleostei: Cichlidae), with the Description of Five New Species and a Diagnosis for Katria, New Genus" masterLastPageNumber="56" masterPageNumber="1" pageNumber="52" updateTime="1699220694966" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C479AC9640D56E4BF56F7C507A727F1F">Phylogeny and Taxonomic Revision of the Endemic Malagasy Genus Ptychochromis (Teleostei: Cichlidae), with the Description of Five New Species and a Diagnosis for Katria, New Genus</mods:title>
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<mods:namePart id="2B2FA8C81D483A61338FAA7C9DCB8A00">STIASSNY, MELANIE L. J</mods:namePart>
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<mods:namePart id="ADB93185D9C7719388341BD8850ACCCE">SPARKS, JOHN S</mods:namePart>
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<mods:title id="93C56C5BB892F1C77AE8D8069963539A">American Museum Novitates</mods:title>
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<mods:part id="343016D667B521FF7A4A7A0F79A8171F">
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<mods:date id="13BCB27C8C931E194ADD35D94DCED6FB">2006</mods:date>
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<mods:number id="E7C72651141EAABB2C7B02AA63C65AAC">2006-09-08</mods:number>
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<mods:identifier id="4A471A479D41EB42A3B76FAFBA144AAB" type="DOI">10.1206/0003-0082(2006)3535[1:PATROT]2.0.CO;2</mods:identifier>
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<treatment id="039487E1F04EFFBBD781FA1B06EFF955" ID-DOI="http://doi.org/10.5281/zenodo.5466935" ID-Zenodo-Dep="5466935" LSID="urn:lsid:zoobank.org:act:994D87BC-800B-4491-BFAE-5E561D3689C3" httpUri="http://treatment.plazi.org/id/039487E1F04EFFBBD781FA1B06EFF955" lastPageNumber="52" pageId="51" pageNumber="52">
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<subSubSection id="C327657CF04EFFBBD781FA1B04EBFB1C" box="[247,455,1144,1168]" pageId="51" pageNumber="52" type="nomenclature">
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<paragraph id="8B8236F7F04EFFBBD781FA1B04EBFB1C" blockId="51.[247,455,1144,1168]" box="[247,455,1144,1168]" pageId="51" pageNumber="52">
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<heading id="D0CA819BF04EFFBBD781FA1B04EBFB1C" box="[247,455,1144,1168]" centered="true" fontSize="36" level="2" pageId="51" pageNumber="52" reason="8">
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<taxonomicName id="4C3D4D74F04EFFBBD781FA1B0413FB1C" ID-CoL="57QT" authority="STIASSNY & SPARKS, 2006" authorityName="STIASSNY & SPARKS" authorityYear="2006" box="[247,319,1144,1168]" class="Actinopterygii" family="Cichlidae" genus="Katria" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="B949EAE5F04EFFBBD781FA1B0413FB1C" bold="true" box="[247,319,1144,1168]" italics="true" pageId="51" pageNumber="52">Katria</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27A579EF04EFFBBD639FA1B04EBFB1C" box="[335,455,1144,1168]" pageId="51" pageNumber="52" rank="genus">new genus</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C327657CF04EFFBBD704FAC80140FEAA" pageId="51" pageNumber="52" type="diagnosis">
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<paragraph id="8B8236F7F04EFFBBD704FAC80140FEAA" blockId="51.[88,614,1195,1754]" lastBlockId="51.[662,1188,211,1754]" pageId="51" pageNumber="52">
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DIAGNOSIS: A ptychochromin genus diagnosed by a unique pigmentation and coloration pattern consisting of two prominent black vertical bars on an iridescent golden background (plate 1H and 2F), the presence of reduced anterior gas bladder horns that do not approach or contact the exoccipital region of the neurocranium (
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<emphasis id="B949EAE5F04EFFBBD64DFB1804DFFA1F" box="[315,499,1403,1427]" italics="true" pageId="51" pageNumber="52">Character 17: 0</emphasis>
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, fig. 3A), a reduced number of gill rakers on the lower limb of the first gill arch (8–9 [mode 8] vs. 8–12 [mode 10] in other ptychochromins), an elevated vertebral count (29–30 [mode 30] vs. 26–28 [mode 27] in
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<taxonomicName id="4C3D4D74F04EFFBBD658F87304FEF9A4" box="[302,466,1552,1576]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromis" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
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<emphasis id="B949EAE5F04EFFBBD658F87304FEF9A4" box="[302,466,1552,1576]" italics="true" pageId="51" pageNumber="52">Ptychochromis</emphasis>
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</taxonomicName>
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), and a more slender caudal peduncle (length to depth ratio modally 1.4 vs.
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<quantity id="4CC59B12F04EFFBBD664F82804AFF9EF" box="[274,387,1611,1635]" metricMagnitude="-2" metricUnit="m" metricValue="2.2859999999999996" metricValueMax="3.048" metricValueMin="1.524" pageId="51" pageNumber="52" unit="in" value="0.8999999999999999" valueMax="1.2" valueMin="0.6">0.6–1.2 in</quantity>
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<taxonomicName id="4C3D4D74F04EFFBBD6FBF828071DF9EF" box="[397,561,1611,1635]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromis" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
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<emphasis id="B949EAE5F04EFFBBD6FBF828071DF9EF" box="[397,561,1611,1635]" italics="true" pageId="51" pageNumber="52">Ptychochromis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C3D4D74F04EFFBBD72EF80A040FF90D" authorityName="Kiener & Mauge" authorityYear="1966" box="[88,291,1641,1665]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromoides" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
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<emphasis id="B949EAE5F04EFFBBD72EF80A040FF90D" box="[88,291,1641,1665]" italics="true" pageId="51" pageNumber="52">Ptychochromoides</emphasis>
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</taxonomicName>
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).
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<taxonomicName id="4C3D4D74F04EFFBBD64BF80A04A9F90D" authorityName="STIASSNY & SPARKS" authorityYear="2006" box="[317,389,1641,1665]" class="Actinopterygii" family="Cichlidae" genus="Katria" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
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<emphasis id="B949EAE5F04EFFBBD64BF80A04A9F90D" box="[317,389,1641,1665]" italics="true" pageId="51" pageNumber="52">Katria</emphasis>
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</taxonomicName>
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is readily differentiated from
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<taxonomicName id="4C3D4D74F04EFFBBD797F8E40480F913" authorityName="Kiener & Mauge" authorityYear="1966" box="[225,428,1671,1695]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromoides" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
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<emphasis id="B949EAE5F04EFFBBD797F8E40480F913" box="[225,428,1671,1695]" italics="true" pageId="51" pageNumber="52">Ptychochromoides</emphasis>
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</taxonomicName>
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by the presence of two toothrows (fig. 7B) on Pb2 (vs. 1;
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<emphasis id="B949EAE5F04EFFBBD72EF8A10428F956" box="[88,260,1730,1754]" italics="true" pageId="51" pageNumber="52">Character 19:1</emphasis>
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), and from
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<taxonomicName id="4C3D4D74F04EFFBBD6EFF8A10711F956" box="[409,573,1730,1754]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromis" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
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<emphasis id="B949EAE5F04EFFBBD6EFF8A10711F956" box="[409,573,1730,1754]" italics="true" pageId="51" pageNumber="52">Ptychochromis</emphasis>
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</taxonomicName>
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by the absence (fig. 7B) of a ‘‘free’’ toothplate associated with Ep2 (vs. presence of ‘‘free’’ Ep2 toothplate;
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<emphasis id="B949EAE5F04EFFBBD424FF6C06D7FEAB" box="[850,1019,271,295]" italics="true" pageId="51" pageNumber="52">Character 10:1</emphasis>
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,
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<figureCitation id="13062A72F04EFFBBD37DFF6D014CFEAA" box="[1035,1120,270,294]" captionStart="Fig" captionStartId="12.[151,189,564,586]" captionTargetBox="[297,1056,211,530]" captionTargetId="figure-152@12.[297,1056,211,530]" captionTargetPageId="12" captionText="Fig. 7. Isolated upper pharyngeal jaw (ventral view) of (A) Ptychochromis onilahy (MNHN 1962-0201) and (B) Katria katria (AMNH 93701). Ptychochromis is diagnosed among cichlids by the presence of a ‘‘free’’ toothplate associated with, but not fused to, the second epibranchial bone (indicated by arrow)." figureDoi="http://doi.org/10.5281/zenodo.5385347" httpUri="https://zenodo.org/record/5385347/files/figure.png" pageId="51" pageNumber="52">Fig 7A</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C327657CF04EFFBBD5C6FF4E06DDFEEE" pageId="51" pageNumber="52" type="materials_examined">
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<paragraph id="8B8236F7F04EFFBBD5C6FF4E06DDFEEE" blockId="51.[662,1188,211,1754]" pageId="51" pageNumber="52">
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<typeStatus id="54868855F04EFFBBD5C6FF4E07C4FECF" box="[688,744,301,325]" pageId="51" pageNumber="52">TYPE</typeStatus>
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SPECIES:
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<taxonomicName id="4C3D4D74F04EFFBBD413FF4E066FFEEE" authority="katria (Reinthal and Stiassny, 1997)" authorityName="katria (Reinthal and Stiassny" authorityYear="1997" baseAuthorityName="Reinthal and Stiassny" baseAuthorityYear="1997" class="Actinopterygii" family="Cichlidae" genus="Katria" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="katria">
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<emphasis id="B949EAE5F04EFFBBD413FF4E06D6FEC9" box="[869,1018,301,325]" italics="true" pageId="51" pageNumber="52">Katria katria</emphasis>
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(
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<bibRefCitation id="EFAC4B06F04EFFBBD37AFF4E0617FEEE" author="Reinthal, P. N. & M. L. J. Stiassny" pageId="51" pageNumber="52" pagination="353 - 368" refId="ref30071" refString="Reinthal, P. N., and M. L. J. Stiassny. 1997. Revision of the Madagascan genus Ptychochromoides (Teleostei: Cichlidae), with description of a new species. Ichthyological Exploration of Freshwaters 7: 353 - 368." type="journal article" year="1997">Reinthal and Stiassny, 1997</bibRefCitation>
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)
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</taxonomicName>
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; by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="C327657CF04EFFBBD5C6FF0A06EFF955" pageId="51" pageNumber="52" type="discussion">
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<paragraph id="8B8236F7F04EFFBBD5C6FF0A06EFF955" blockId="51.[662,1188,211,1754]" pageId="51" pageNumber="52">
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REMARKS: At the time of its original description,
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<bibRefCitation id="EFAC4B06F04EFFBBD454FFE50144FE12" author="Reinthal, P. N. & M. L. J. Stiassny" box="[802,1128,390,414]" pageId="51" pageNumber="52" pagination="353 - 368" refId="ref30071" refString="Reinthal, P. N., and M. L. J. Stiassny. 1997. Revision of the Madagascan genus Ptychochromoides (Teleostei: Cichlidae), with description of a new species. Ichthyological Exploration of Freshwaters 7: 353 - 368." type="journal article" year="1997">Reinthal and Stiassny (1997)</bibRefCitation>
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were unable to provide unambiguous morphological support for their placement of
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<taxonomicName id="4C3D4D74F04EFFBBD36FFFA10170FE56" authorityName="katria (Reinthal and Stiassny" authorityYear="1997" box="[1049,1116,450,474]" class="Actinopterygii" family="Cichlidae" genus="Katria" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="katria">
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<emphasis id="B949EAE5F04EFFBBD36FFFA10170FE56" box="[1049,1116,450,474]" italics="true" pageId="51" pageNumber="52">katria</emphasis>
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</taxonomicName>
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in the genus
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<taxonomicName id="4C3D4D74F04EFFBBD597FF830680FE74" authorityName="Kiener & Mauge" authorityYear="1966" box="[737,940,480,504]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromoides" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
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<emphasis id="B949EAE5F04EFFBBD597FF830680FE74" box="[737,940,480,504]" italics="true" pageId="51" pageNumber="52">Ptychochromoides</emphasis>
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</taxonomicName>
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. They proposed that the presence of a pronounced nuchal hump in adult
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<taxonomicName id="4C3D4D74F04EFFBBD5AEFC7F0637FDB8" authorityName="katria (Reinthal and Stiassny" authorityYear="1997" box="[728,795,540,564]" class="Actinopterygii" family="Cichlidae" genus="Katria" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="katria">
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<emphasis id="B949EAE5F04EFFBBD5AEFC7F0637FDB8" box="[728,795,540,564]" italics="true" pageId="51" pageNumber="52">katria</emphasis>
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</taxonomicName>
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and in
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<taxonomicName id="4C3D4D74F04EFFBBD402FC7F0113FDB8" authorityName="Kiener & Mauge" authorityYear="1966" box="[884,1087,540,564]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromoides" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
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<emphasis id="B949EAE5F04EFFBBD402FC7F0113FDB8" box="[884,1087,540,564]" italics="true" pageId="51" pageNumber="52">Ptychochromoides</emphasis>
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</taxonomicName>
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, and the possession of an elevated vertebral count (relative to that of
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<taxonomicName id="4C3D4D74F04EFFBBD4DCFC340163FDE3" box="[938,1103,599,623]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromis" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
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<emphasis id="B949EAE5F04EFFBBD4DCFC340163FDE3" box="[938,1103,599,623]" italics="true" pageId="51" pageNumber="52">Ptychochromis</emphasis>
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</taxonomicName>
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) were suggestive of relationship, but cautioned that ‘‘further systematic analysis may necessitate an eventual reassignment and generic reorganization’’. The molecular analyses of Sparks (2004) and
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<bibRefCitation id="EFAC4B06F04EFFBBD46BFC8F016DFC88" author="Sparks, J. S. & W. L. Smith" box="[797,1089,748,772]" pageId="51" pageNumber="52" pagination="501 - 517" refId="ref30568" refString="Sparks, J. S., and W. L. Smith. 2004. Phylogeny and biogeography of cichlid fishes (Teleostei: Perciformes: Cichlidae). Cladistics 20: 501 - 517." type="journal article" year="2004">Sparks and Smith (2004)</bibRefCitation>
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provide strong support for the placement of
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<taxonomicName id="4C3D4D74F04EFFBBD33AFD6A064AFCB3" authorityName="Reinthal & Stiassny" authorityYear="1997" class="Actinopterygii" family="Cichlidae" genus="Ptychochromoides" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="katria">
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<emphasis id="B949EAE5F04EFFBBD33AFD6A064AFCB3" italics="true" pageId="51" pageNumber="52">Ptychochromoides katria</emphasis>
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</taxonomicName>
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as the sister taxon to a monophyletic
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<taxonomicName id="4C3D4D74F04EFFBBD441FD2606F7FCD1" box="[823,987,837,861]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromis" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
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<emphasis id="B949EAE5F04EFFBBD441FD2606F7FCD1" box="[823,987,837,861]" italics="true" pageId="51" pageNumber="52">Ptychochromis</emphasis>
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</taxonomicName>
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(e.g., in the study of
|
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<bibRefCitation id="EFAC4B06F04EFFBBD5C0FD0106F8FCF6" author="Sparks, J. S. & W. L. Smith" box="[694,980,866,890]" pageId="51" pageNumber="52" pagination="501 - 517" refId="ref30568" refString="Sparks, J. S., and W. L. Smith. 2004. Phylogeny and biogeography of cichlid fishes (Teleostei: Perciformes: Cichlidae). Cladistics 20: 501 - 517." type="journal article" year="2004">Sparks and Smith [2004]</bibRefCitation>
|
||
, which comprised fragments of two mitochondrial and two nuclear genes, this sister group relationship is robustly supported [Bremer support: 11; jackknife 100%]). The current study corroborates that placement and, consequently, we remove
|
||
<taxonomicName id="4C3D4D74F04EFFBBD5E0FA7607F5FBA1" authorityName="Reinthal & Stiassny" authorityYear="1997" box="[662,729,1045,1069]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromoides" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="katria">
|
||
<emphasis id="B949EAE5F04EFFBBD5E0FA7607F5FBA1" box="[662,729,1045,1069]" italics="true" pageId="51" pageNumber="52">katria</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C3D4D74F04EFFBBD453FA7606DCFBA1" authorityName="Kiener & Mauge" authorityYear="1966" box="[805,1008,1045,1069]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromoides" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B949EAE5F04EFFBBD453FA7606DCFBA1" box="[805,1008,1045,1069]" italics="true" pageId="51" pageNumber="52">Ptychochromoides</emphasis>
|
||
</taxonomicName>
|
||
and place it in
|
||
<taxonomicName id="4C3D4D74F04EFFBBD5E0FA5007F2FBC7" authority="STIASSNY & SPARKS, 2006" authorityName="STIASSNY & SPARKS" authorityYear="2006" box="[662,734,1075,1099]" class="Actinopterygii" family="Cichlidae" genus="Katria" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="B949EAE5F04EFFBBD5E0FA5007F2FBC7" box="[662,734,1075,1099]" italics="true" pageId="51" pageNumber="52">Katria</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A579EF04EFFBBD59DFA500646FBC7" box="[747,874,1075,1099]" pageId="51" pageNumber="52" rank="genus">new genus</taxonomicNameLabel>
|
||
. Uniquely among ptychochromins,
|
||
<taxonomicName id="4C3D4D74F04EFFBBD462FA320670FBE5" authorityName="STIASSNY & SPARKS" authorityYear="2006" box="[788,860,1105,1129]" class="Actinopterygii" family="Cichlidae" genus="Katria" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B949EAE5F04EFFBBD462FA320670FBE5" box="[788,860,1105,1129]" italics="true" pageId="51" pageNumber="52">Katria</emphasis>
|
||
</taxonomicName>
|
||
shares with
|
||
<taxonomicName id="4C3D4D74F04EFFBBD489FA32018FFBE5" box="[1023,1187,1105,1129]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromis" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B949EAE5F04EFFBBD489FA32018FFBE5" box="[1023,1187,1105,1129]" italics="true" pageId="51" pageNumber="52">Ptychochromis</emphasis>
|
||
</taxonomicName>
|
||
the following synapomorphic features: Cb4 bears a characteristically elongate process and deeply rounded indentation on its inner margin (fig. 4C;
|
||
<emphasis id="B949EAE5F04EFFBBD42AFAAB06D6FB6C" box="[860,1018,1224,1248]" italics="true" pageId="51" pageNumber="52">Character 6:1</emphasis>
|
||
); hypertrophy of the laterosensory canal system of the neurocranium, preopercle, mandible, and infraorbital series (
|
||
<emphasis id="B949EAE5F04EFFBBD479FB420680FAB5" box="[783,940,1313,1337]" italics="true" pageId="51" pageNumber="52">Character 7:1</emphasis>
|
||
, fig. 8E); two supraneural elements located anterior to the first neural spine (fig. 3A;
|
||
<emphasis id="B949EAE5F04EFFBBD4C6FB3E0179FAF9" box="[944,1109,1373,1397]" italics="true" pageId="51" pageNumber="52">Character 8:1</emphasis>
|
||
); and a well-developed, expansive and dorsally elevated lateral ethmoid process on the dorsal margin of the palatine head (fig. 6E;
|
||
<emphasis id="B949EAE5F04EFFBBD5E0FBB70615FA60" box="[662,825,1492,1516]" italics="true" pageId="51" pageNumber="52">Character 9:1</emphasis>
|
||
).
|
||
<taxonomicName id="4C3D4D74F04EFFBBD42FFBB7068DFA60" authorityName="STIASSNY & SPARKS" authorityYear="2006" box="[857,929,1492,1516]" class="Actinopterygii" family="Cichlidae" genus="Katria" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B949EAE5F04EFFBBD42FFBB7068DFA60" box="[857,929,1492,1516]" italics="true" pageId="51" pageNumber="52">Katria</emphasis>
|
||
</taxonomicName>
|
||
lacks the four apomorphic features identified herein as diagnostic of its sister taxon,
|
||
<taxonomicName id="4C3D4D74F04EFFBBD4CAF86C014CF9AB" box="[956,1120,1551,1575]" class="Actinopterygii" family="Cichlidae" genus="Ptychochromis" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="genus">
|
||
<emphasis id="B949EAE5F04EFFBBD4CAF86C014CF9AB" box="[956,1120,1551,1575]" italics="true" pageId="51" pageNumber="52">Ptychochromis</emphasis>
|
||
</taxonomicName>
|
||
(i.e., presence of a ‘‘free’’ toothplate associated with Ep2,
|
||
<emphasis id="B949EAE5F04EFFBBD5A0F82806AFF9EF" box="[726,899,1611,1635]" italics="true" pageId="51" pageNumber="52">Character 10:1</emphasis>
|
||
; IO2 excluded from the orbit by IO3,
|
||
<emphasis id="B949EAE5F04EFFBBD426F80B012DF90C" box="[848,1025,1640,1664]" italics="true" pageId="51" pageNumber="52">Character 11:1</emphasis>
|
||
; presence of a palatine groove,
|
||
<emphasis id="B949EAE5F04EFFBBD4FFF8E50117F912" box="[905,1083,1670,1694]" italics="true" pageId="51" pageNumber="52">Character 12:1</emphasis>
|
||
; ventral displacement of the palato-palatine ligament insertion,
|
||
<emphasis id="B949EAE5F04EFFBBD47FF8A1069FF956" box="[777,947,1730,1754]" italics="true" pageId="51" pageNumber="52">Character 13:1</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |