347 lines
45 KiB
XML
347 lines
45 KiB
XML
<document id="F47A99A89684D3030199F1058CCD7BD8" ID-DOI="10.11646/zootaxa.5123.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6399827" ID-ZooBank="036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1648673534065" checkinUser="plazi" docAuthor="Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald" docDate="2022" docId="039487D1FFB9FF94FFBAFEE16B97FEDD" docLanguage="en" docName="zootaxa.5123.1.1.pdf" docOrigin="Zootaxa 5123 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5123.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Megaclavatula Harzhauser & Landau & Janssen 2022, nov. gen." docType="treatment" docVersion="1" lastPageNumber="61" masterDocId="FFADFFA9FF82FFA8FF2DFFAB6872FFC0" masterDocTitle="The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera" masterLastPageNumber="172" masterPageNumber="1" pageNumber="60" updateTime="1709123294374" updateUser="karina">
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<mods:titleInfo id="1626FC4FA3F890F89AA4E976C4E5BBBB">
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<mods:title id="F0493708443C528C5E184B12047F209E">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<mods:namePart id="2919B741970C018D0CE35829B11550BE">Harzhauser, Mathias</mods:namePart>
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<mods:nameIdentifier id="BD1CB7F9B4EABB277B1F2D71B2893113" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:roleTerm id="D54E06A8C1F75842D00E43E8B8978A43">Author</mods:roleTerm>
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<mods:namePart id="1C7C3CE9F8966DF69E33B53212DF60DC">Landau, Bernard</mods:namePart>
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<mods:nameIdentifier id="1796D01B9BF6F76DCCB3C69FABB55B54" type="ORCID">0000-0002-7768-8494</mods:nameIdentifier>
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<mods:nameIdentifier id="C094F4110CF6829229E5963CEEC53AE0" type="email">bernardmlandau@gmail.com</mods:nameIdentifier>
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<mods:namePart id="F7E2E761944C7D4FE3EC0A8CBE3AC72A">Janssen, Ronald</mods:namePart>
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<mods:nameIdentifier id="7FB263A66A4BC7483151ED10EF981ADD" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:title id="E342ABDE3DFF6072A08E8A9FFFFF96CC">Zootaxa</mods:title>
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<mods:date id="24D3F7F3FD87D97E6FDD5A87F8EDB037">2022</mods:date>
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<mods:number id="1182E36B92D42718E62867F7EADC700B">2022-03-30</mods:number>
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<mods:number id="D5BAAF7069588547837597D27CDA5A95">5123</mods:number>
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<treatment id="039487D1FFB9FF94FFBAFEE16B97FEDD" LSID="urn:lsid:plazi:treatment:039487D1FFB9FF94FFBAFEE16B97FEDD" httpUri="http://treatment.plazi.org/id/039487D1FFB9FF94FFBAFEE16B97FEDD" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
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<subSubSection id="C327654CFFB9FF93FFBAFEE16A65FEA5" box="[151,535,330,357]" pageId="59" pageNumber="60" type="distribution">
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<paragraph id="8B8236C7FFB9FF93FFBAFEE16A65FEA5" blockId="59.[151,535,330,357]" box="[151,535,330,357]" pageId="59" pageNumber="60">
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<emphasis id="B949EAD5FFB9FF93FFBAFEE16A65FEA5" bold="true" box="[151,535,330,357]" pageId="59" pageNumber="60">
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Genus
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<taxonomicName id="4C3D4D44FFB9FF93FFC3FEE169D3FEA5" ID-CoL="8YKXM" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[238,417,330,357]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="B949EAD5FFB9FF93FFC3FEE169D3FEA5" bold="true" box="[238,417,330,357]" italics="true" pageId="59" pageNumber="60">Megaclavatula</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A57AEFFB9FF93FE84FEE06A65FEA5" box="[425,535,331,357]" pageId="59" pageNumber="60" rank="genus">nov. gen.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C327654CFFB9FF93FFBAFE3868B5FE11" pageId="59" pageNumber="60" type="type_taxon">
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<paragraph id="8B8236C7FFB9FF93FFBAFE3868B5FE11" blockId="59.[151,1437,402,2013]" pageId="59" pageNumber="60">
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<materialsCitation id="3B553C9AFFB9FF93FFBAFE3868B1FE11" country="Austria" county="Central Paratethys Sea" location="Miocene" municipality="Langhian" pageId="59" pageNumber="60" specimenCount="1" typeStatus="holotype">
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<emphasis id="B949EAD5FFB9FF93FFBAFE386940FE6C" bold="true" box="[151,306,402,429]" pageId="59" pageNumber="60">
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<typeStatus id="54868865FFB9FF93FFBAFE3868A0FE6D" box="[151,210,403,429]" pageId="59" pageNumber="60">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C3D4D44FFB9FF93FE14FE386A36FE6C" ID-CoL="8YN4B" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[313,580,403,429]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="grunerti" status="sp. nov.">
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<emphasis id="B949EAD5FFB9FF93FE14FE386A36FE6C" box="[313,580,403,429]" italics="true" pageId="59" pageNumber="60">Megaclavatula grunerti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A57AEFFB9FF93FD66FE386AD1FE6D" box="[587,675,403,429]" pageId="59" pageNumber="60" rank="species">nov. sp.</taxonomicNameLabel>
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, middle
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<location id="8EE2601CFFB9FF93FC29FE386B1BFE6D" LSID="urn:lsid:plazi:treatment:039487D1FFB9FF94FFBAFEE16B97FEDD:8EE2601CFFB9FF93FC29FE386B1BFE6D" box="[772,873,403,429]" country="Austria" county="Central Paratethys Sea" municipality="Langhian" name="Miocene" pageId="59" pageNumber="60">Miocene</location>
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,
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<collectingMunicipality id="6BE6ACBDFFB9FF93FC59FE386BACFE6D" box="[884,990,403,429]" pageId="59" pageNumber="60">Langhian</collectingMunicipality>
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(Badenian),
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<collectingCountry id="F32A7657FFB9FF93FB43FE386CB1FE6D" box="[1134,1219,403,429]" name="Austria" pageId="59" pageNumber="60">Austria</collectingCountry>
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,
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<collectingCounty id="62E34E4BFFB9FF93FBE3FE3868B1FE11" pageId="59" pageNumber="60">Central Paratethys Sea</collectingCounty>
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C327654CFFB9FF93FFEAFE706BD4FE35" box="[199,934,475,502]" pageId="59" pageNumber="60" type="etymology">
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<paragraph id="8B8236C7FFB9FF93FFEAFE706BD4FE35" blockId="59.[151,1437,402,2013]" box="[199,934,475,502]" pageId="59" pageNumber="60">
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<emphasis id="B949EAD5FFB9FF93FFEAFE70693CFE35" bold="true" box="[199,334,475,501]" pageId="59" pageNumber="60">Etymology.</emphasis>
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Combination of
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<emphasis id="B949EAD5FFB9FF93FD21FE706A3EFE34" box="[524,588,475,500]" italics="true" pageId="59" pageNumber="60">Mega</emphasis>
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- (Greek: large) and
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<taxonomicName id="4C3D4D44FFB9FF93FC1DFE706BECFE35" box="[816,926,475,501]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="genus">
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<emphasis id="B949EAD5FFB9FF93FC1DFE706BECFE35" box="[816,926,475,501]" italics="true" pageId="59" pageNumber="60">Clavatula</emphasis>
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</taxonomicName>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C327654CFFB9FF93FFEAFE55699FFD45" pageId="59" pageNumber="60" type="diagnosis">
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||
<paragraph id="8B8236C7FFB9FF93FFEAFE55699FFD45" blockId="59.[151,1437,402,2013]" pageId="59" pageNumber="60">
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<emphasis id="B949EAD5FFB9FF93FFEAFE556930FDD8" bold="true" box="[199,322,510,536]" pageId="59" pageNumber="60">Diagnosis.</emphasis>
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Medium to large sized, moderately slender fusiform, with high conical spire characterized by tripartite sculpture on early teleoconch whorls with two prominent, beaded adsutural cords and weaker, beaded mid-whorl cord. Subsutural collar coronate with pointed tubercles or spines. Base with weak sculpture; siphonal canal moderately long to long.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C327654CFFB9FF93FFEAFD256B4AF97D" pageId="59" pageNumber="60" type="description">
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||
<paragraph id="8B8236C7FFB9FF93FFEAFD256A69FB61" blockId="59.[151,1437,402,2013]" pageId="59" pageNumber="60">
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||
<emphasis id="B949EAD5FFB9FF93FFEAFD256923FD68" bold="true" box="[199,337,654,680]" pageId="59" pageNumber="60">Description</emphasis>
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. Medium to large sized, moderately slender fusiform, with high conical spire. Protoconch not preserved. Early teleoconch whorls with prominent tripartite spiral sculpture comprising prominent, beaded subsutural cord. Slightly weaker, beaded cord in concave mid-whorl portion, and prominent beaded suprasutural cord. Beads in mid-whorl position and at the suprasutural cord form at ends of comma-shaped axial riblets. On later teleoconch whorls subsutural collar usually swollen, coronate, with small, pointed tubercles or trigonal spines (tubercles may be reduced in some species, e.g.,
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<taxonomicName id="4C3D4D44FFB9FF93FD91FCE86B67FC9C" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[700,789,835,860]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="evae">
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<emphasis id="B949EAD5FFB9FF93FD91FCE86B67FC9C" box="[700,789,835,860]" italics="true" pageId="59" pageNumber="60">M. evae</emphasis>
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</taxonomicName>
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). Mid-whorl cord weakens, suprasutural cord weakens to subobsolete, bearing weak, wide-spaced tubercles. Secondary spiral threads may be present on spire whorls. Suture narrowly incised, weakly undulating. Last whorl typically ~ 55–60% of total height, attaining up to 65% in species with lower spire (e.g.,
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<taxonomicName id="4C3D4D44FFB9FF93FEBAFC046A73FC09" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[407,513,943,969]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="pilleri" status="sp. nov.">
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<emphasis id="B949EAD5FFB9FF93FEBAFC046A73FC09" box="[407,513,943,969]" italics="true" pageId="59" pageNumber="60">M. pilleri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A57AEFFB9FF93FD24FC046A10FC09" box="[521,610,943,969]" pageId="59" pageNumber="60" rank="species">nov. sp.</taxonomicNameLabel>
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). Weakly concave subsutural ramp, rounded periphery. Weakly delimited shoulder. Base moderately constricted, sculpture weak. Siphonal fasciole of medium length, weakly delimited. Aperture moderately wide, pyriform. Outer lip not thickened, smooth within. Anal sinus wide, moderately deep to deep, asymmetrically V-shaped, with apex below subsutural collar in deepest part of subsutural concavity. Siphonal canal moderately long to long, wide, often twisted and recurved, shallowly notched at tip. Columella straight to weakly twisted, smooth. Columellar and parietal callus thickened, sharply delimited, forming broad callus rim, without forming pseudoumbilicus.
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</paragraph>
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<paragraph id="8B8236C7FFB9FF93FFEAFB016C28FA5D" blockId="59.[151,1437,402,2013]" pageId="59" pageNumber="60">
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<emphasis id="B949EAD5FFB9FF93FFEAFB016A0FFB05" bold="true" box="[199,637,1194,1221]" pageId="59" pageNumber="60">Stratigraphic and geographic range.</emphasis>
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Middle Miocene (Langhian to Serravallian) of the Central Paratethys. Additional occurrences in the Miocene of the northeastern Atlantic and the Proto-Mediterranean Sea will need verification.
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<taxonomicName id="4C3D4D44FFB9FF93FE0BFB586AB3FACD" authority="Sacco, 1890" authorityName="Sacco" authorityYear="1890" box="[294,705,1267,1293]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="taurofusulata">
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<emphasis id="B949EAD5FFB9FF93FE0BFB586A43FACD" box="[294,561,1267,1293]" italics="true" pageId="59" pageNumber="60">Clavatula taurofusulata</emphasis>
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<bibRefCitation id="EFAC4B36FFB9FF93FD15FB586AB3FACD" author="Sacco, F." box="[568,705,1267,1293]" pageId="59" pageNumber="60" pagination="189 - 340" refId="ref143023" refString="Sacco, F. (1890) Catalogo paleontologico del bacino terziaro del Piemonte. Bollettino della Societa Geologica Italiana, 9, 189 - 340. [https: // www. biodiversitylibrary. org / item / 210076 # page / 207 / mode / 1 up]" type="journal article" year="1890">Sacco, 1890</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C3D4D44FFB9FF93FDDAFB586CFBFACD" authority="Sacco, 1890" authorityName="Sacco" authorityYear="1890" box="[759,1161,1267,1293]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="subdepressa">
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<emphasis id="B949EAD5FFB9FF93FDDAFB586B87FACD" box="[759,1013,1267,1293]" italics="true" pageId="59" pageNumber="60">Clavatula subdepressa</emphasis>
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<bibRefCitation id="EFAC4B36FFB9FF93FCD0FB586CFBFACD" author="Sacco, F." box="[1021,1161,1267,1293]" pageId="59" pageNumber="60" pagination="189 - 340" refId="ref143023" refString="Sacco, F. (1890) Catalogo paleontologico del bacino terziaro del Piemonte. Bollettino della Societa Geologica Italiana, 9, 189 - 340. [https: // www. biodiversitylibrary. org / item / 210076 # page / 207 / mode / 1 up]" type="journal article" year="1890">Sacco, 1890</bibRefCitation>
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</taxonomicName>
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, from the Burdigalian of the Colli Torinesi (
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<collectingCountry id="F32A7657FFB9FF93FE4BFABC69ECFAF2" box="[358,414,1303,1330]" name="Italy" pageId="59" pageNumber="60">Italy</collectingCountry>
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), might be a
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<taxonomicName id="4C3D4D44FFB9FF93FD07FABC6AA0FAF1" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[554,722,1303,1329]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="genus">
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<emphasis id="B949EAD5FFB9FF93FD07FABC6AA0FAF1" box="[554,722,1303,1329]" italics="true" pageId="59" pageNumber="60">Megaclavatula</emphasis>
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</taxonomicName>
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, but the early teleoconch whorls are poorly preserved and do not allow a clear identification. Similarly,
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<taxonomicName id="4C3D4D44FFB9FF93FD63FA906BBAFA95" authority="Peyrot, 1931" authorityName="Peyrot" authorityYear="1931" box="[590,968,1339,1365]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="tortonica">
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<emphasis id="B949EAD5FFB9FF93FD63FA906B5EFA94" box="[590,812,1339,1365]" italics="true" pageId="59" pageNumber="60">Clavatula tortonica</emphasis>
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<bibRefCitation id="EFAC4B36FFB9FF93FC18FA906BBAFA95" author="Peyrot, A." box="[821,968,1339,1365]" pageId="59" pageNumber="60" pagination="5 - 288" refId="ref141284" refString="Peyrot, A. (1931 & 1933) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 83, 5 - 116 (1931), 84, 5 - 288 (1933). [https: // www. biodiversitylibrary. org / item / 185325 # page / 11 / mode / 1 up]" type="journal article" year="1931">Peyrot, 1931</bibRefCitation>
|
||
</taxonomicName>
|
||
, might represent
|
||
<taxonomicName id="4C3D4D44FFB9FF93FBA2FA906D45FA95" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[1167,1335,1339,1365]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFB9FF93FBA2FA906D45FA95" box="[1167,1335,1339,1365]" italics="true" pageId="59" pageNumber="60">Megaclavatula</emphasis>
|
||
</taxonomicName>
|
||
, judging from the illustrations in
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FE83FAF46A3CFAB9" author="Peyrot, A." box="[430,590,1375,1401]" pageId="59" pageNumber="60" pagination="5 - 288" refId="ref141284" refString="Peyrot, A. (1931 & 1933) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 83, 5 - 116 (1931), 84, 5 - 288 (1933). [https: // www. biodiversitylibrary. org / item / 185325 # page / 11 / mode / 1 up]" type="journal article" year="1931">Peyrot (1931)</bibRefCitation>
|
||
, but we have not seen specimens.
|
||
<taxonomicName id="4C3D4D44FFB9FF93FCF1FAF46DEAFAB9" authority="Lamarck, 1822" authorityName="Lamarck" authorityYear="1822" box="[988,1432,1375,1401]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="asperulata">
|
||
<emphasis id="B949EAD5FFB9FF93FCF1FAF46C93FAB9" box="[988,1249,1375,1401]" italics="true" pageId="59" pageNumber="60">Pleurotoma asperulata</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FBC7FAF46DEAFAB9" author="Lamarck, J. - B. M. de" box="[1258,1432,1375,1401]" pageId="59" pageNumber="60" refId="ref137485" refString="Lamarck, J. - B. M. de. (1822) Histoire naturelle des animaux sans vertebres. Tome Septieme. Published by the Author, Paris, 711 pp. [http: // www. biodiversitylibrary. org / item / 47433]" type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
</taxonomicName>
|
||
, however, can clearly be placed in
|
||
<taxonomicName id="4C3D4D44FFB9FF93FD39FA286ACEFA5D" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[532,700,1411,1437]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFB9FF93FD39FA286ACEFA5D" box="[532,700,1411,1437]" italics="true" pageId="59" pageNumber="60">Megaclavatula</emphasis>
|
||
</taxonomicName>
|
||
based on early teleoconch sculpture.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFB9FF93FFEAFA0C6955F991" blockId="59.[151,1437,402,2013]" pageId="59" pageNumber="60">
|
||
<emphasis id="B949EAD5FFB9FF93FFEAFA0C69E4FA00" bold="true" box="[199,406,1446,1473]" pageId="59" pageNumber="60">Included species.</emphasis>
|
||
Paratethys: Miocene:
|
||
<taxonomicName id="4C3D4D44FFB9FF93FDB2FA0C6BDCFA00" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[671,942,1447,1473]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="grunerti" status="sp. nov.">
|
||
<emphasis id="B949EAD5FFB9FF93FDB2FA0C6BDCFA00" box="[671,942,1447,1473]" italics="true" pageId="59" pageNumber="60">Megaclavatula grunerti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFFB9FF93FC97FA0C6C64FA01" box="[954,1046,1447,1473]" pageId="59" pageNumber="60" rank="species">nov. sp.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C3D4D44FFB9FF93FB0AFA0C69B0FA25" authority="Hoernes & Auinger, 1891" authorityName="Hoernes & Auinger" authorityYear="1891" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="amaliae" subGenus="Clavatula">
|
||
<emphasis id="B949EAD5FFB9FF93FB0AFA0C6CD8FA01" box="[1063,1194,1447,1473]" italics="true" pageId="59" pageNumber="60">Pleurotoma</emphasis>
|
||
(
|
||
<emphasis id="B949EAD5FFB9FF93FB92FA0C6D5FFA01" box="[1215,1325,1447,1473]" italics="true" pageId="59" pageNumber="60">Clavatula</emphasis>
|
||
)
|
||
<emphasis id="B949EAD5FFB9FF93FA6FFA0C6DEEFA01" box="[1346,1436,1447,1473]" italics="true" pageId="59" pageNumber="60">amaliae</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FFBAFA6069B0FA25" author="Hoernes, R. & Auinger, M." box="[151,450,1483,1509]" pageId="59" pageNumber="60" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FFB9FF93FEFCFA606C0DFA25" authority="Hoernes & Auinger, 1891" authorityName="Hoernes & Auinger" authorityYear="1891" box="[465,1151,1483,1509]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="antoniae" subGenus="Clavatula">
|
||
<emphasis id="B949EAD5FFB9FF93FEFCFA606A26FA25" box="[465,596,1483,1509]" italics="true" pageId="59" pageNumber="60">Pleurotoma</emphasis>
|
||
(
|
||
<emphasis id="B949EAD5FFB9FF93FD4AFA606AA7FA25" box="[615,725,1483,1509]" italics="true" pageId="59" pageNumber="60">Clavatula</emphasis>
|
||
)
|
||
<emphasis id="B949EAD5FFB9FF93FDC4FA606B39FA24" box="[745,843,1483,1508]" italics="true" pageId="59" pageNumber="60">antoniae</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FC78FA606C0DFA25" author="Hoernes, R. & Auinger, M." box="[853,1151,1483,1509]" pageId="59" pageNumber="60" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FFB9FF93FBA3FA60699DF9C9" authority="Hoernes & Auinger, 1891" authorityName="Hoernes & Auinger" authorityYear="1891" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="evae" subGenus="Clavatula">
|
||
<emphasis id="B949EAD5FFB9FF93FBA3FA606D63FA25" box="[1166,1297,1483,1509]" italics="true" pageId="59" pageNumber="60">Pleurotoma</emphasis>
|
||
(
|
||
<emphasis id="B949EAD5FFB9FF93FA09FA606DE0FA25" box="[1316,1426,1483,1509]" italics="true" pageId="59" pageNumber="60">Clavatula</emphasis>
|
||
)
|
||
<emphasis id="B949EAD5FFB9FF93FFBAFA4468B8F9C8" box="[151,202,1519,1544]" italics="true" pageId="59" pageNumber="60">evae</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FFFDFA44699DF9C9" author="Hoernes, R. & Auinger, M." box="[208,495,1519,1545]" pageId="59" pageNumber="60" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FFB9FF93FED4FA446BF6F9C9" authority="Baluk, 2003" authorityName="Baluk" authorityYear="2003" box="[505,900,1519,1545]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="kowalewskii">
|
||
<emphasis id="B949EAD5FFB9FF93FED4FA446A85F9C9" box="[505,759,1519,1545]" italics="true" pageId="59" pageNumber="60">Clavatula kowalewskii</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FDD1FA446BF6F9C9" author="Baluk, W." box="[764,900,1519,1545]" pageId="59" pageNumber="60" pagination="29 - 78" refId="ref128777" refString="Baluk, W. (2003) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part IV - Turridae. Acta Geologica Polonica, 53, 29 - 78. [https: // geojournals. pgi. gov. pl / agp / article / view / 9720 / 8257]" type="journal article" year="2003">Bałuk, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C3D4D44FFB9FF93FC85FA446C78F9C8" authorityName="Hoernes & Auinger" authorityYear="1891" box="[936,1034,1519,1544]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="antoniae">
|
||
<emphasis id="B949EAD5FFB9FF93FC85FA446C78F9C8" box="[936,1034,1519,1544]" italics="true" pageId="59" pageNumber="60">antoniae</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="4C3D4D44FFB9FF93FB0DFA4468A0F9ED" authority="Eichwald, 1830" authorityName="Eichwald" authorityYear="1830" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="laevigatum">
|
||
<emphasis id="B949EAD5FFB9FF93FB0DFA446D57F9C9" box="[1056,1317,1519,1545]" italics="true" pageId="59" pageNumber="60">Pleurotoma laevigatum</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FA06FA4468A0F9ED" author="Eichwald, E." pageId="59" pageNumber="60" refId="ref131673" refString="Eichwald, E. (1830) Naturhistorische Skizze von Lithauen, Volhynien und Podolien in Geognostisch-Mineralogischer, Botanischer und Zoologischer Hinsicht. Wilna, Voss, 255 pp. [https: // reader. digitale-sammlungen. de / de / fs 1 / object / display / bsb 10691419 _ 00005. html]" type="book" year="1830">Eichwald, 1830</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FFB9FF93FFF6F9B86AEEF9ED" authority="Schaffer, 1898" authorityName="Schaffer" authorityYear="1898" box="[219,668,1555,1581]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="neudorfensis">
|
||
<emphasis id="B949EAD5FFB9FF93FFF6F9B86981F9ED" box="[219,499,1555,1581]" italics="true" pageId="59" pageNumber="60">Pleurotoma neudorfensis</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FED5F9B86AEEF9ED" author="Schaffer, F." box="[504,668,1555,1581]" pageId="59" pageNumber="60" pagination="533 - 548" refId="ref143392" refString="Schaffer, F. (1898) Der marine Tegel von Theben - Neudorf in Ungarn. Jahrbuch der k. k. Geologischen Reichsanstalt, 47, 533 - 548. [https: // opac. geologie. ac. at / ais 312 / dokumente / JB 0473 _ 533 _ A. pdf]" type="journal article" year="1898">Schaffer, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FFB9FF93FD88F9B86BE4F9ED" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[677,918,1555,1581]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="pilleri" status="sp. nov.">
|
||
<emphasis id="B949EAD5FFB9FF93FD88F9B86BE4F9ED" box="[677,918,1555,1581]" italics="true" pageId="59" pageNumber="60">Megaclavatula pilleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFFB9FF93FCB1F9B86B80F9ED" box="[924,1010,1555,1581]" pageId="59" pageNumber="60" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C3D4D44FFB9FF93FB07F9B86951F991" authority="Pusch, 1837" authorityName="Pusch, 1937 Megaclavatula polonica (Pusch" authorityYear="1837" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="subSpecies" species="tuberculosa" subSpecies="polonica">
|
||
<emphasis id="B949EAD5FFB9FF93FB07F9B86DEEF9ED" box="[1066,1436,1555,1581]" italics="true" pageId="59" pageNumber="60">Pleurotoma tuberculosa polonica</emphasis>
|
||
Pusch, 1837
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFB9FF93FFEAF9F06B4AF97D" blockId="59.[151,1437,402,2013]" pageId="59" pageNumber="60">
|
||
Proto-Mediterranean Sea: Miocene:
|
||
<taxonomicName id="4C3D4D44FFB9FF93FD72F9F06B8EF9B5" authority="Sacco, 1890" authorityName="Sacco" authorityYear="1890" box="[607,1020,1627,1653]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="taurofusulata">
|
||
<emphasis id="B949EAD5FFB9FF93FD72F9F06B18F9B5" box="[607,874,1627,1653]" italics="true" pageId="59" pageNumber="60">Clavatula taurofusulata</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FC5CF9F06B8EF9B5" author="Sacco, F." box="[881,1020,1627,1653]" pageId="59" pageNumber="60" pagination="189 - 340" refId="ref143023" refString="Sacco, F. (1890) Catalogo paleontologico del bacino terziaro del Piemonte. Bollettino della Societa Geologica Italiana, 9, 189 - 340. [https: // www. biodiversitylibrary. org / item / 210076 # page / 207 / mode / 1 up]" type="journal article" year="1890">Sacco, 1890</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FFB9FF93FB2AF9F06DEAF9B5" authority="Sacco, 1890" authorityName="Sacco" authorityYear="1890" box="[1031,1432,1627,1653]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="subdepressa">
|
||
<emphasis id="B949EAD5FFB9FF93FB2AF9F06D77F9B5" box="[1031,1285,1627,1653]" italics="true" pageId="59" pageNumber="60">Clavatula subdepressa</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FA20F9F06DEAF9B5" author="Sacco, F." box="[1293,1432,1627,1653]" pageId="59" pageNumber="60" pagination="189 - 340" refId="ref143023" refString="Sacco, F. (1890) Catalogo paleontologico del bacino terziaro del Piemonte. Bollettino della Societa Geologica Italiana, 9, 189 - 340. [https: // www. biodiversitylibrary. org / item / 210076 # page / 207 / mode / 1 up]" type="journal article" year="1890">Sacco, 1890</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FFB9FF93FFBAF9D46989F959" authority="Toula, 1901" authorityName="Toula" authorityYear="1901" box="[151,507,1663,1689]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="ernesti">
|
||
<emphasis id="B949EAD5FFB9FF93FFBAF9D4691FF958" box="[151,365,1663,1689]" italics="true" pageId="59" pageNumber="60">Pleurotoma ernesti</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FE5EF9D46989F959" author="Toula, F." box="[371,507,1663,1689]" pageId="59" pageNumber="60" pagination="247 - 263" refId="ref145622" refString="Toula, F. (1901) Eine marine Neogenfauna aus Cilicien. Jahrbuch der k. k. Geologischen Reichsanstalt, 51, 247 - 263. [https: // opac. geologie. ac. at / ais 312 / dokumente / JB 0512 _ 247 _ A. pdf]" type="journal article" year="1901">
|
||
<collectingRegion id="49F9F825FFB9FF93FE5EF9D469C7F959" box="[371,437,1663,1689]" country="Russia" name="Tula" pageId="59" pageNumber="60">Toula</collectingRegion>
|
||
, 1901
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
. Northeastern Atlantic:
|
||
<taxonomicName id="4C3D4D44FFB9FF93FC2EF9D46CC4F959" authority="Lamarck, 1822" authorityName="Lamarck" authorityYear="1822" box="[771,1206,1663,1689]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="asperulata">
|
||
<emphasis id="B949EAD5FFB9FF93FC2EF9D46C76F959" box="[771,1028,1663,1689]" italics="true" pageId="59" pageNumber="60">Pleurotoma asperulata</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FB27F9D46CC4F959" author="Lamarck, J. - B. M. de" box="[1034,1206,1663,1689]" pageId="59" pageNumber="60" refId="ref137485" refString="Lamarck, J. - B. M. de. (1822) Histoire naturelle des animaux sans vertebres. Tome Septieme. Published by the Author, Paris, 711 pp. [http: // www. biodiversitylibrary. org / item / 47433]" type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FFB9FF93FBECF9D4695AF97D" authority="Peyrot, 1931" authorityName="Peyrot" authorityYear="1931" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="tortonica">
|
||
<emphasis id="B949EAD5FFB9FF93FBECF9D46DEFF958" box="[1217,1437,1663,1689]" italics="true" pageId="59" pageNumber="60">Clavatula tortonica</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FFBAF908695AF97D" author="Peyrot, A." box="[151,296,1699,1725]" pageId="59" pageNumber="60" pagination="5 - 288" refId="ref141284" refString="Peyrot, A. (1931 & 1933) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 83, 5 - 116 (1931), 84, 5 - 288 (1933). [https: // www. biodiversitylibrary. org / item / 185325 # page / 11 / mode / 1 up]" type="journal article" year="1931">Peyrot, 1931</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FFB9FF93FE19F9086B46F97D" authority="Peyrot, 1931" authorityName="Peyrot" authorityYear="1931" box="[308,820,1699,1725]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="subSpecies" species="saubrigiana" subSpecies="laurensii">
|
||
<emphasis id="B949EAD5FFB9FF93FE19F9086AE9F97D" box="[308,667,1699,1725]" italics="true" pageId="59" pageNumber="60">Clavatula saubrigiana laurensii</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FD8FF9086B46F97D" author="Peyrot, A." box="[674,820,1699,1725]" pageId="59" pageNumber="60" pagination="5 - 288" refId="ref141284" refString="Peyrot, A. (1931 & 1933) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 83, 5 - 116 (1931), 84, 5 - 288 (1933). [https: // www. biodiversitylibrary. org / item / 185325 # page / 11 / mode / 1 up]" type="journal article" year="1931">Peyrot, 1931</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFB9FF93FFEAF96D6934F8B1" pageId="59" pageNumber="60" type="discussion">
|
||
<paragraph id="8B8236C7FFB9FF93FFEAF96D6934F8B1" blockId="59.[151,1437,402,2013]" pageId="59" pageNumber="60">
|
||
<emphasis id="B949EAD5FFB9FF93FFEAF96D693EF920" bold="true" box="[199,332,1734,1760]" pageId="59" pageNumber="60">Discussion.</emphasis>
|
||
<taxonomicName id="4C3D4D44FFB9FF93FE7AF96C698DF921" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[343,511,1735,1761]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFB9FF93FE7AF96C698DF921" box="[343,511,1735,1761]" italics="true" pageId="59" pageNumber="60">Megaclavatula</emphasis>
|
||
</taxonomicName>
|
||
is reminiscent of
|
||
<taxonomicName id="4C3D4D44FFB9FF93FDF9F96C6B30F921" box="[724,834,1735,1761]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFB9FF93FDF9F96C6B30F921" box="[724,834,1735,1761]" italics="true" pageId="59" pageNumber="60">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
in general shape and the coronate spire, but differs distinctly in its tripartite early teleoconch sculpture and larger size. In addition,
|
||
<taxonomicName id="4C3D4D44FFB9FF93FB6CF9406D37F8C5" box="[1089,1349,1771,1797]" pageId="59" pageNumber="60">
|
||
<emphasis id="B949EAD5FFB9FF93FB6CF9406C9BF8C5" box="[1089,1257,1771,1797]" italics="true" pageId="59" pageNumber="60">Megaclavatula</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
have a more constricted base and longer siphonal canal resulting in a more slender outline. A further difference is the deep U-shaped anal sinus of
|
||
<taxonomicName id="4C3D4D44FFB9FF93FEC8F8986A21F88D" box="[485,595,1843,1869]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFB9FF93FEC8F8986A21F88D" box="[485,595,1843,1869]" italics="true" pageId="59" pageNumber="60">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
, which contrasts with the moderately deep, broad V-shaped anal sinus of
|
||
<taxonomicName id="4C3D4D44FFB9FF93FFBAF8FC694DF8B1" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[151,319,1879,1905]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFB9FF93FFBAF8FC694DF8B1" box="[151,319,1879,1905]" italics="true" pageId="59" pageNumber="60">Megaclavatula</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFFB9FF94FFEAF8D06B97FEDD" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="reference_group">
|
||
<paragraph id="8B8236C7FFB9FF94FFEAF8D06B0DFF3A" blockId="59.[151,1437,402,2013]" lastBlockId="60.[151,1436,150,285]" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
|
||
<taxonomicName id="4C3D4D44FFB9FF93FFEAF8D0691DF855" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[199,367,1915,1941]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFB9FF93FFEAF8D0691DF855" box="[199,367,1915,1941]" italics="true" pageId="59" pageNumber="60">Megaclavatula</emphasis>
|
||
</taxonomicName>
|
||
is mainly known so far from the Central Paratethys Sea and for most species from the Proto- Mediterraean Sea and the northeastern Atlantic, included in
|
||
<taxonomicName id="4C3D4D44FFB9FF93FC07F8346BA0F879" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[810,978,1951,1977]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="59" pageNumber="60" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFB9FF93FC07F8346BA0F879" box="[810,978,1951,1977]" italics="true" pageId="59" pageNumber="60">Megaclavatula</emphasis>
|
||
</taxonomicName>
|
||
above, we have not seen early teleoconch whorls. Some morphologically similar species from the Serravallian of
|
||
<collectingCountry id="F32A7657FFB9FF93FC91F8686C7FF81D" box="[956,1037,1987,2013]" name="Turkey" pageId="59" pageNumber="60">Turkey</collectingCountry>
|
||
, described by
|
||
<bibRefCitation id="EFAC4B36FFB9FF93FB82F8686DEAF81D" author="Landau, B. M. & Harzhauser, M. & Yslamodlu, Y. & Marques da Silva, C." box="[1199,1432,1987,2013]" pageId="59" pageNumber="60" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. & Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
|
||
Landau
|
||
<emphasis id="B949EAD5FFB9FF93FA26F8686D31F81D" box="[1291,1347,1987,2013]" italics="true" pageId="59" pageNumber="60">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
, display distinctly different early teleoconch morphology.
|
||
<taxonomicName id="4C3D4D44FFBEFF94FC31FF3C6C91FF71" authority="Erunal-Erentoz, 1958" authorityName="Erunal-Erentoz" authorityYear="1958" box="[796,1251,150,177]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="gracilis">
|
||
<emphasis id="B949EAD5FFBEFF94FC31FF3C6B9AFF71" box="[796,1000,151,177]" italics="true" pageId="60" pageNumber="61">Clavatula gracilis</emphasis>
|
||
<bibRefCitation id="EFAC4B36FFBEFF94FCC2FF3C6C91FF71" author="Erunal-Erentoz, L." box="[1007,1251,150,177]" pageId="60" pageNumber="61" pagination="1 - 232" refId="ref131888" refString="Erunal-Erentoz, L. (1958) Mollusques du Neogene des Bassins de Karaman, Adana et Hatay (Turquie). Premiere these, 1 ere partie. Publications de l'Institut d'Etude et du Recherches Minieres de Turquie, (C), 4, 1 - 232." type="journal article" year="1958">Erünal-Erentöz, 1958</bibRefCitation>
|
||
</taxonomicName>
|
||
, has two beaded adsutural spiral cords and
|
||
<taxonomicName id="4C3D4D44FFBEFF94FEE5FF106B9BFF15" authority="(Toula, 1901)" authorityName="Toula" authorityYear="1901" baseAuthorityName="Toula" baseAuthorityYear="1901" box="[456,1001,187,213]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="60" pageNumber="61" phylum="Mollusca" rank="species" species="francisci" subGenus="Clavatula">
|
||
<emphasis id="B949EAD5FFBEFF94FEE5FF106A39FF15" box="[456,587,187,213]" italics="true" pageId="60" pageNumber="61">Pleurotoma</emphasis>
|
||
(
|
||
<emphasis id="B949EAD5FFBEFF94FD72FF106ABFFF15" box="[607,717,187,213]" italics="true" pageId="60" pageNumber="61">Clavatula</emphasis>
|
||
)
|
||
<emphasis id="B949EAD5FFBEFF94FDCDFF106B30FF15" box="[736,834,187,213]" italics="true" pageId="60" pageNumber="61">francisci</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FFBEFF94FC79FF106B92FF15" author="Toula, F." box="[852,992,187,213]" pageId="60" pageNumber="61" pagination="247 - 263" refId="ref145622" refString="Toula, F. (1901) Eine marine Neogenfauna aus Cilicien. Jahrbuch der k. k. Geologischen Reichsanstalt, 51, 247 - 263. [https: // opac. geologie. ac. at / ais 312 / dokumente / JB 0512 _ 247 _ A. pdf]" type="journal article" year="1901">Toula, 1901</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, develops a
|
||
<taxonomicName id="4C3D4D44FFBEFF94FB56FF106CA5FF14" box="[1147,1239,187,212]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="60" pageNumber="61" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FFBEFF94FB56FF106CA5FF14" box="[1147,1239,187,212]" italics="true" pageId="60" pageNumber="61">Perrona</emphasis>
|
||
</taxonomicName>
|
||
-like sculpture of opisthocline axial riblets (see
|
||
<bibRefCitation id="EFAC4B36FFBEFF94FECBFF746AC8FF39" author="Landau, B. M. & Harzhauser, M. & Yslamodlu, Y. & Marques da Silva, C." box="[486,698,223,249]" pageId="60" pageNumber="61" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. & Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
|
||
Landau
|
||
<emphasis id="B949EAD5FFBEFF94FD6DFF746A0BFF39" box="[576,633,223,249]" italics="true" pageId="60" pageNumber="61">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
, pl. 70, figs 6–9).
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FFBEFF94FFEAFEA96B97FEDD" blockId="60.[151,1436,150,285]" box="[199,997,258,285]" pageId="60" pageNumber="61">
|
||
<emphasis id="B949EAD5FFBEFF94FFEAFEA969DAFEDC" bold="true" box="[199,424,258,284]" pageId="60" pageNumber="61">Paleoenvironment.</emphasis>
|
||
Coastal inner neritic to outer neritic environments.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |