1545 lines
240 KiB
XML
1545 lines
240 KiB
XML
<document id="5B5928555613205248EAD7B1C3BE5390" ID-DOI="10.11646/zootaxa.5123.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6399827" ID-ZooBank="036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1648673534065" checkinUser="plazi" docAuthor="Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald" docDate="2022" docId="039487D1FF89FFBCFFBAF8886C2FFC2D" docLanguage="en" docName="zootaxa.5123.1.1.pdf" docOrigin="Zootaxa 5123 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5123.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Clavatulidae Gray 1853" docType="treatment" docVersion="4" lastPageNumber="21" masterDocId="FFADFFA9FF82FFA8FF2DFFAB6872FFC0" masterDocTitle="The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera" masterLastPageNumber="172" masterPageNumber="1" pageNumber="12" updateTime="1709132455857" updateUser="karina">
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<mods:titleInfo id="D02090199B97F1CCB1268894A7BCE79A">
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<mods:title id="939483ABD051656D76E8FA8D140F7948">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<mods:namePart id="5CA97EBFB7B8E24DFDB0759A067ED108">Harzhauser, Mathias</mods:namePart>
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<mods:nameIdentifier id="5DA2A0103FBED5B3E6399CBBA4FF1D88" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:roleTerm id="7A937745E2C9812443EB284D796503D9">Author</mods:roleTerm>
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<mods:namePart id="6ADFF78BAFB3D08AE8685A77B73B7C0E">Landau, Bernard</mods:namePart>
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<mods:nameIdentifier id="565A6DA069D4A262BC0B958D10FE6965" type="ORCID">0000-0002-7768-8494</mods:nameIdentifier>
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<mods:nameIdentifier id="2A0C7A3149F24305873219CE59C94A94" type="email">bernardmlandau@gmail.com</mods:nameIdentifier>
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<mods:namePart id="9EE221099F5E8CF0C006EEF3415F82AF">Janssen, Ronald</mods:namePart>
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<mods:nameIdentifier id="54E974651CBC305B4B8A589739AF9888" type="ORCID">0000-0002-4471-6655</mods:nameIdentifier>
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<mods:nameIdentifier id="64A14471DDC9E67B405C41BF36892FC0" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:typeOfResource id="4673241EACB0FB000CBA873C62907947">text</mods:typeOfResource>
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<mods:title id="397FFDCD40EDA4DDC40F9A854F2D5C39">Zootaxa</mods:title>
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<mods:part id="CE5CB5DC896561B0CADD223217941783">
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<mods:date id="B80355C081824E72D213315E1178D219">2022</mods:date>
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<mods:number id="5F2384AAD1ACE4954B3092A707E12737">2022-03-30</mods:number>
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<mods:detail id="0EF7F00E68D0F17F28963B7EF5E7B96F" type="volume">
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<mods:number id="8CE4D73465C0AAF0F0C9BB0549F0C07E">5123</mods:number>
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<mods:url id="76BFF24328184376951D82E0A29A54BE">http://dx.doi.org/10.11646/zootaxa.5123.1.1</mods:url>
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<mods:classification id="F5927BD7310621045431A73B426DB568">journal article</mods:classification>
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<treatment id="039487D1FF89FFBCFFBAF8886C2FFC2D" LSID="urn:lsid:plazi:treatment:039487D1FF89FFBCFFBAF8886C2FFC2D" httpUri="http://treatment.plazi.org/id/039487D1FF89FFBCFFBAF8886C2FFC2D" lastPageId="20" lastPageNumber="21" pageId="11" pageNumber="12">
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<subSubSection id="C327654CFF89FFA3FFBAF8886A55F8FD" box="[151,551,1827,1853]" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="8B8236C7FF89FFA3FFBAF8886A55F8FD" blockId="11.[151,551,1827,1853]" box="[151,551,1827,1853]" pageId="11" pageNumber="12">
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<heading id="D0CA81ABFF89FFA3FFBAF8886A55F8FD" bold="true" box="[151,551,1827,1853]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="3">
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<emphasis id="B949EAD5FF89FFA3FFBAF8886A55F8FD" bold="true" box="[151,551,1827,1853]" pageId="11" pageNumber="12">
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Family
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<taxonomicName id="4C3D4D44FF89FFA3FFDAF8886A55F8FD" ID-CoL="7NHYP" authority="Gray, 1853" authorityName="Gray" authorityYear="1853" box="[247,551,1827,1853]" class="Gastropoda" family="Clavatulidae" kingdom="Animalia" order="Neogastropoda" pageId="11" pageNumber="12" phylum="Mollusca" rank="family">
|
||
Clavatulidae
|
||
<bibRefCitation id="EFAC4B36FF89FFA3FEB3F8886A55F8FD" author="Gray, J. E." box="[414,551,1827,1853]" pageId="11" pageNumber="12" pagination="124 - 133" refId="ref133609" refString="Gray, J. E. (1853) On the division of Ctenobranchous Gasteropodous Mollusca into larger groups and families. Annals and Magazine of Natural History, Series 2, 11, 124 - 133. [https: // www. biodiversitylibrary. org / item / 48966 # page / 154 / mode / 1 up] https: // doi. org / 10.1080 / 03745485609496511" type="journal article" year="1853">Gray, 1853</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
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</paragraph>
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</subSubSection>
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<subSubSection id="C327654CFF89FFBBFFBAF8C0694FF931" lastPageId="19" lastPageNumber="20" pageId="11" pageNumber="12" type="description">
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<paragraph id="8B8236C7FF89FFA3FFBAF8C06BC5F831" blockId="11.[151,1436,1899,2034]" pageId="11" pageNumber="12">
|
||
Although all extant clavatulid species are superficially similar, which has led all authors until now to divide them into two genera (
|
||
<taxonomicName id="4C3D4D44FF89FFA3FE77F8246A09F86A" authority="Lamarck, 1801" authorityName="Lamarck" authorityYear="1801" box="[346,635,1935,1962]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF89FFA3FE77F82469BAF869" box="[346,456,1935,1961]" italics="true" pageId="11" pageNumber="12">Clavatula</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF89FFA3FEFDF8246A09F86A" author="Lamarck, J. B. P. A. de" box="[464,635,1935,1962]" pageId="11" pageNumber="12" refId="ref137273" refString="Lamarck, J. B. P. A. de (1801) Systeme des animaux sans vertebres, ou tableau general des classes, des ordres et des genres de ces animaux; Presentant leurs caracteres essentiels et leur distribution, d'apres la consideration de leurs rapports naturels et de leur organisation, et suivant l'arrangement etabli dans les galeries du Museum d'Histoire Naturelle, parmi leurs depouilles conservees; Precede du discours d'ouverture du Cours de Zoologie, donne dans le Museum National d'Histoire Naturelle l'an 8 de la Republique. Published by the author and Deterville, Paris, viii + 432 pp. [http: // biodiversitylibrary. org / page / 14117719] https: // doi. org / 10.5962 / bhl. title. 116650" type="book" year="1801">Lamarck, 1801</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3D4D44FF89FFA3FD99F83B6B9FF86A" authority="Schumacher, 1817" authorityName="Schumacher" authorityYear="1817" box="[692,1005,1935,1962]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF89FFA3FD99F83B6B62F869" box="[692,784,1936,1961]" italics="true" pageId="11" pageNumber="12">Perrona</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF89FFA3FC34F8246B9FF86A" author="Schumacher, H. C. F." box="[793,1005,1935,1962]" pageId="11" pageNumber="12" refId="ref143804" refString="Schumacher, H. C. F. (1817) Essais d'un Nuveau Systeme des Habitations des Vers Testaces. Schultz, Copenhagen, 287 pp. [https: // www. biodiversitylibrary. org / item / 81329 # page / 7 / mode / 1 up]" type="book" year="1817">Schumacher, 1817</bibRefCitation>
|
||
</taxonomicName>
|
||
), on closer examination there are also characters that are shared by some groups of clavatulids and not others. In order to clarify the descriptions given in the systematic section, some discussion on clavatulid shells is required.
|
||
</paragraph>
|
||
<caption id="DF42664FFF8EFFA5FFBAF9F86BADFD42" ID-DOI="http://doi.org/10.5281/zenodo.6399835" ID-Zenodo-Dep="6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" startId="12.[151,250,1619,1643]" targetBox="[151,1436,181,1595]" targetPageId="12">
|
||
<paragraph id="8B8236C7FF8EFFA4FFBAF9F86DEEF837" blockId="12.[151,1437,1619,2040]" pageId="12" pageNumber="13">
|
||
<emphasis id="B949EAD5FF8EFFA4FFBAF9F86967F9AB" bold="true" box="[151,277,1619,1643]" pageId="12" pageNumber="13">FIGURE 3.</emphasis>
|
||
Early teleoconch whorls of extant and fossil
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FDDFF9F86B00F9AB" authorityName="Gray" authorityYear="1853" box="[754,882,1619,1643]" class="Gastropoda" family="Clavatulidae" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="family">Clavatulidae</taxonomicName>
|
||
species.
|
||
<emphasis id="B949EAD5FF8EFFA4FCFEF9F86B9EF9AB" bold="true" box="[979,1004,1619,1643]" pageId="12" pageNumber="13">A.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FCD9F9F86D4FF9AC" authority="(Roding, 1798)" baseAuthorityName="Roding" baseAuthorityYear="1798" box="[1012,1341,1619,1644]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="regia">
|
||
<emphasis id="B949EAD5FF8EFFA4FCD9F9F86CE4F9AB" box="[1012,1174,1619,1643]" italics="true" pageId="12" pageNumber="13">Clavatula regia</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8EFFA4FB88F9F86D44F9AC" author="Roding, P. F." box="[1189,1334,1619,1644]" pageId="12" pageNumber="13" refId="ref142481" refString="Roding, P. F. (1798). Museum Boltenianum sive Catalogus cimeliorum e tribus regnis naturae quae olim collegerat Joa. Fried Bolten, M. D. p. d. per XL. annos proto physicus Hamburgensis. Pars secunda continens Conchylia sive Testacea univalvia, bivalvia & multivalvia. Trapp, Hamburg, viii + 199 pp. [https: // www. biodiversitylibrary. org / page / 16230659]" type="book" year="1798">Röding, 1798</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW 1683 6/28, West Africa.
|
||
<emphasis id="B949EAD5FF8EFFA4FEB8F9D369DEF950" bold="true" box="[405,428,1656,1680]" pageId="12" pageNumber="13">B.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FE99F9DC6B32F94F" authority="(Reeve, 1845)" baseAuthorityName="Reeve" baseAuthorityYear="1845" box="[436,832,1655,1680]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="rubrifasciata">
|
||
<emphasis id="B949EAD5FF8EFFA4FE99F9DC6AD6F94F" box="[436,676,1655,1679]" italics="true" pageId="12" pageNumber="13">Clavatula rubrifasciata</emphasis>
|
||
(Reeve, 1845)
|
||
</taxonomicName>
|
||
, NHMW 10203, Gorée (Senegal).
|
||
<emphasis id="B949EAD5FF8EFFA4FB84F9DC6CB0F94F" bold="true" box="[1193,1218,1655,1679]" pageId="12" pageNumber="13">C.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FBFFF9DC6D44F94F" authorityName="Lamarck" authorityYear="1801" box="[1234,1334,1655,1679]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8EFFA4FBFFF9DC6D44F94F" box="[1234,1334,1655,1679]" italics="true" pageId="12" pageNumber="13">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B949EAD5FF8EFFA4FA68F9DC6DEEF94F" box="[1349,1436,1655,1679]" italics="true" pageId="12" pageNumber="13">diadema</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8EFFA4FFB3F930695BF974" author="Kiener, L. C." box="[158,297,1691,1716]" pageId="12" pageNumber="13" refId="ref136494" refString="Kiener, L. C. (1839) Species general et iconographie des coquilles vivantes. Vol. 5. Famille des Canaliferes. Premiere partie. Pleurotome (Pleurotoma), Lamarck. Rousseau & J. B. Bailliere, Paris, 16 pp., 27 pls. [http: // www. biodiversitylibrary. org / item / 19762]" type="book" year="1839">Kiener, 1839</bibRefCitation>
|
||
), NHMW 1884/0010/0023, Gorée (Senegal).
|
||
<emphasis id="B949EAD5FF8EFFA4FC2AF9376B52F974" bold="true" box="[775,800,1692,1716]" pageId="12" pageNumber="13">D.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FC1FF9306BE4F973" authorityName="Lamarck" authorityYear="1801" box="[818,918,1691,1715]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8EFFA4FC1FF9306BE4F973" box="[818,918,1691,1715]" italics="true" pageId="12" pageNumber="13">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FC88F9376CE3F974" authority="(Reeve, 1843)" baseAuthorityName="Reeve" baseAuthorityYear="1843" box="[933,1169,1692,1716]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="mystica">
|
||
<emphasis id="B949EAD5FF8EFFA4FC88F9376B80F973" box="[933,1010,1692,1715]" italics="true" pageId="12" pageNumber="13">mystica</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8EFFA4FB2EF9376CFBF974" author="Reeve, L. A." box="[1027,1161,1692,1716]" pageId="12" pageNumber="13" refId="ref142338" refString="Reeve, L. A. (1843 & 1845) Monograph of the genus Pleurotoma. Conchologia Iconica, or, illustrations of the shells of molluscous animals. Vol. 1. Reeve & Co., London, 40 pls., unpaginated text. [https: // www. biodiversitylibrary. org / item / 41735 # page / 33 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 8129" type="url" year="1843">Reeve, 1843</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal).
|
||
<emphasis id="B949EAD5FF8EFFA4FEC4F96B6A72F918" bold="true" box="[489,512,1728,1752]" pageId="12" pageNumber="13">E.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FD3FF9146A04F917" authorityName="Lamarck" authorityYear="1801" box="[530,630,1727,1751]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8EFFA4FD3FF9146A04F917" box="[530,630,1727,1751]" italics="true" pageId="12" pageNumber="13">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B949EAD5FF8EFFA4FDAAF9146B71F917" box="[647,771,1727,1751]" italics="true" pageId="12" pageNumber="13">bimarginata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8EFFA4FC39F9146BC7F918" author="Lamarck, J. - B. M. de" box="[788,949,1727,1752]" pageId="12" pageNumber="13" refId="ref137485" refString="Lamarck, J. - B. M. de. (1822) Histoire naturelle des animaux sans vertebres. Tome Septieme. Published by the Author, Paris, 711 pp. [http: // www. biodiversitylibrary. org / item / 47433]" type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
), MNHN-IM-2013-43369, Dakar (Senegal).
|
||
<emphasis id="B949EAD5FF8EFFA4FAA4F96B6DEEF918" bold="true" box="[1417,1436,1728,1752]" pageId="12" pageNumber="13">F.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FF8DF9486976F93B" authorityName="Lamarck" authorityYear="1801" box="[160,260,1763,1787]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8EFFA4FF8DF9486976F93B" box="[160,260,1763,1787]" italics="true" pageId="12" pageNumber="13">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FE3FF9486A57F93C" authority="(Kiener, 1839)" baseAuthorityName="Kiener" baseAuthorityYear="1839" box="[274,549,1763,1788]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="pyramidata">
|
||
<emphasis id="B949EAD5FF8EFFA4FE3FF94869F5F93B" box="[274,391,1763,1787]" italics="true" pageId="12" pageNumber="13">pyramidata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8EFFA4FEB8F9486A6CF93C" author="Kiener, L. C." box="[405,542,1763,1788]" pageId="12" pageNumber="13" refId="ref136494" refString="Kiener, L. C. (1839) Species general et iconographie des coquilles vivantes. Vol. 5. Famille des Canaliferes. Premiere partie. Pleurotome (Pleurotoma), Lamarck. Rousseau & J. B. Bailliere, Paris, 16 pp., 27 pls. [http: // www. biodiversitylibrary. org / item / 19762]" type="book" year="1839">Kiener, 1839</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo).
|
||
<emphasis id="B949EAD5FF8EFFA4FA3AF9486D43F93B" bold="true" box="[1303,1329,1763,1787]" pageId="12" pageNumber="13">G.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FA15F94868A0F8DF" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="sorini" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF8EFFA4FA15F94868A0F8DF" italics="true" pageId="12" pageNumber="13">Clavatula sorini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8EFFA4FFF0F8A36943F8DF" box="[221,305,1800,1823]" pageId="12" pageNumber="13" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania).
|
||
<emphasis id="B949EAD5FF8EFFA4FCC4F8A36C70F8E0" bold="true" box="[1001,1026,1800,1824]" pageId="12" pageNumber="13">H.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FB3BF8AC6C08F8DF" authorityName="Lamarck" authorityYear="1801" box="[1046,1146,1799,1823]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8EFFA4FB3BF8AC6C08F8DF" box="[1046,1146,1799,1823]" italics="true" pageId="12" pageNumber="13">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FBA1F8A36DEAF8DF" authority="(Defrance, 1826)" baseAuthorityName="Defrance" baseAuthorityYear="1826" box="[1164,1432,1799,1823]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="romana">
|
||
<emphasis id="B949EAD5FF8EFFA4FBA1F8A36CA8F8DF" box="[1164,1242,1800,1823]" italics="true" pageId="12" pageNumber="13">romana</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8EFFA4FBC1F8AC6DE3F8DF" author="Defrance, M." box="[1260,1425,1799,1823]" pageId="12" pageNumber="13" pagination="388 - 396" refId="ref131223" refString="Defrance, M. (1826) Pleurotome. dans lequel on traite meithodiquement des diffeirens e ˆ tres de la nature, consideireis soit en eux-me ˆ mes, d'apreIs l'eitat actuel de nos connoissances, soit relativement aI l'utilitei qu'en peuvent retirer la meidecine, l'agriculture, le commerce et les artes. Vol. 41. Levrault, Le Normant, Paris, Strasbourg, 388 - 396. [https: // www. biodiversitylibrary. org / item / 81578 # page / 398 / mode / 1 up]" type="book chapter" year="1826">Defrance, 1826</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW 1999/0098/0004, Modra-Kráľová (Slovakia).
|
||
<emphasis id="B949EAD5FF8EFFA4FDFFF8876A90F884" bold="true" box="[722,738,1836,1860]" pageId="12" pageNumber="13">I.</emphasis>
|
||
‘
|
||
<emphasis id="B949EAD5FF8EFFA4FDDBF8806B12F883" box="[758,864,1835,1859]" italics="true" pageId="12" pageNumber="13">Clavatula’</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FC47F8876CF1F884" authority="(Brocchi, 1814)" baseAuthorityName="Brocchi" baseAuthorityYear="1814" box="[874,1155,1835,1860]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="interrupta">
|
||
<emphasis id="B949EAD5FF8EFFA4FC47F8876BA3F883" box="[874,977,1836,1859]" italics="true" pageId="12" pageNumber="13">interrupta</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8EFFA4FCCEF8806C0EF884" author="Brocchi, G." box="[995,1148,1835,1860]" pageId="12" pageNumber="13" refId="ref130110" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW 1861/0033/0420, Buituri (Romania).
|
||
<emphasis id="B949EAD5FF8EFFA4FE43F8FB69F3F8A8" bold="true" box="[366,385,1872,1896]" pageId="12" pageNumber="13">J.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FEB5F8E4698EF8A7" authorityName="Lamarck" authorityYear="1801" box="[408,508,1871,1895]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8EFFA4FEB5F8E4698EF8A7" box="[408,508,1871,1895]" italics="true" pageId="12" pageNumber="13">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FD3FF8E46BF3F8A8" authority="Kovacs & Vician, 2021" authorityName="Kovacs & Vician" authorityYear="2021" box="[530,897,1871,1896]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="hirmetzli">
|
||
<emphasis id="B949EAD5FF8EFFA4FD3FF8E46A1EF8A7" box="[530,620,1871,1895]" italics="true" pageId="12" pageNumber="13">hirmetzli</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF8EFFA4FD57F8E46BF3F8A8" author="Kovacs, Z. & Vician, Z." box="[634,897,1871,1896]" pageId="12" pageNumber="13" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
, NHMW 1868/0001/0069, Baden-Sooss (Austria).
|
||
<emphasis id="B949EAD5FF8EFFA4FFBAF8DF68C3F84C" bold="true" box="[151,177,1908,1932]" pageId="12" pageNumber="13">K.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FFEEF8D86955F84B" authorityName="Lamarck" authorityYear="1801" box="[195,295,1907,1931]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8EFFA4FFEEF8D86955F84B" box="[195,295,1907,1931]" italics="true" pageId="12" pageNumber="13">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B949EAD5FF8EFFA4FE17F8DF6906F84B" box="[314,372,1908,1931]" italics="true" pageId="12" pageNumber="13">irisae</emphasis>
|
||
nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria).
|
||
<emphasis id="B949EAD5FF8EFFA4FB07F8DF6C32F84C" bold="true" box="[1066,1088,1908,1932]" pageId="12" pageNumber="13">L.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FB7EF8D86CC5F84B" authorityName="Lamarck" authorityYear="1801" box="[1107,1207,1907,1931]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8EFFA4FB7EF8D86CC5F84B" box="[1107,1207,1907,1931]" italics="true" pageId="12" pageNumber="13">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FBE4F8D86940F870" authority="(Hoernes & Auinger, 1891)" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="sophiae">
|
||
<emphasis id="B949EAD5FF8EFFA4FBE4F8D86D6AF84B" box="[1225,1304,1907,1931]" italics="true" pageId="12" pageNumber="13">sophiae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8EFFA4FA04F8DF6959F870" author="Hoernes, R. & Auinger, M." pageId="12" pageNumber="13" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria).
|
||
<emphasis id="B949EAD5FF8EFFA4FC53F8336BEEF870" bold="true" box="[894,924,1944,1968]" pageId="12" pageNumber="13">M.</emphasis>
|
||
<emphasis id="B949EAD5FF8EFFA4FC8CF83C6C7FF86F" box="[929,1037,1943,1967]" italics="true" pageId="12" pageNumber="13">
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FC88F83C6C7FF86F" authorityName="Lamarck" authorityYear="1801" box="[933,1037,1943,1967]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">Clavatula</taxonomicName>
|
||
</emphasis>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FB36F8336DEAF870" authority="(Hoernes & Auinger, 1891)" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[1051,1432,1943,1968]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="veronicae">
|
||
<emphasis id="B949EAD5FF8EFFA4FB36F8336C0CF86F" box="[1051,1150,1944,1967]" italics="true" pageId="12" pageNumber="13">veronicae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8EFFA4FBA7F8336DE3F870" author="Hoernes, R. & Auinger, M." box="[1162,1425,1943,1968]" pageId="12" pageNumber="13" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania).
|
||
<emphasis id="B949EAD5FF8EFFA4FDE1F8176A96F814" bold="true" box="[716,740,1980,2004]" pageId="12" pageNumber="13">N.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FDDEF8106B25F813" authorityName="Lamarck" authorityYear="1801" box="[755,855,1979,2003]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8EFFA4FDDEF8106B25F813" box="[755,855,1979,2003]" italics="true" pageId="12" pageNumber="13">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FC4BF8106CD4F814" authority="Kovacs & Vician, 2021" authorityName="Kovacs & Vician" authorityYear="2021" box="[870,1190,1979,2004]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="santhai">
|
||
<emphasis id="B949EAD5FF8EFFA4FC4BF8106BC3F813" box="[870,945,1979,2003]" italics="true" pageId="12" pageNumber="13">santhai</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF8EFFA4FC9AF8106CD4F814" author="Kovacs, Z. & Vician, Z." box="[951,1190,1979,2004]" pageId="12" pageNumber="13" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).
|
||
<emphasis id="B949EAD5FF8EFFA4FEEEF87469AFF837" bold="true" box="[451,477,2015,2039]" pageId="12" pageNumber="13">O.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FEC6F8746A3DF837" authorityName="Lamarck" authorityYear="1801" box="[491,591,2015,2039]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8EFFA4FEC6F8746A3DF837" box="[491,591,2015,2039]" italics="true" pageId="12" pageNumber="13">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8EFFA4FD70F8746BBFF838" authority="(Hoernes & Auinger, 1891)" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[605,973,2015,2040]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="camillae">
|
||
<emphasis id="B949EAD5FF8EFFA4FD70F8746AC6F837" box="[605,692,2015,2039]" italics="true" pageId="12" pageNumber="13">camillae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8EFFA4FDEDF84B6BB4F838" author="Hoernes, R. & Auinger, M." box="[704,966,2015,2040]" pageId="12" pageNumber="13" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, lectotype, NHMW 1949/0005/0042, Lăpugiu
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF8FFFA5FFBAFF3C6BADFD42" blockId="13.[151,1437,150,643]" pageId="13" pageNumber="14">
|
||
de Sus (Romania).
|
||
<emphasis id="B949EAD5FF8FFFA5FE74FF3C691EFF6F" bold="true" box="[345,364,151,175]" pageId="13" pageNumber="14">P.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FE5EFF3C6BF2FF6F" authority="(Hoernes & Auinger, 1891)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[371,896,151,175]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="amaliae">
|
||
<emphasis id="B949EAD5FF8FFFA5FE5EFF3C6A11FF6F" box="[371,611,151,175]" italics="true" pageId="13" pageNumber="14">
|
||
<docIdISSN id="1331E051FF8FFFA5FE5EFF3C6A79FF6F" box="[371,523,151,175]" pageId="13" pageNumber="14">Megaclavatula</docIdISSN>
|
||
amaliae
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8FFFA5FD42FF3C6B0BFF6F" author="Hoernes, R. & Auinger, M." box="[623,889,151,175]" pageId="13" pageNumber="14" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8FFFA5FCA7FF3C6B98FF6F" box="[906,1002,151,175]" pageId="13" pageNumber="14" type="lectotype">lectotype</typeStatus>
|
||
, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania).
|
||
<emphasis id="B949EAD5FF8FFFA5FE3EFF11695FFF12" bold="true" box="[275,301,186,210]" pageId="13" pageNumber="14">Q.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FE1BFF106A58FF12" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[310,554,187,211]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="grunerti" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF8FFFA5FE1BFF106A58FF12" box="[310,554,187,211]" italics="true" pageId="13" pageNumber="14">Megaclavatula grunerti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8FFFA5FD1EFF106AF4FF12" box="[563,646,187,210]" pageId="13" pageNumber="14" rank="species">nov. sp.</taxonomicNameLabel>
|
||
,
|
||
<typeStatus id="54868865FF8FFFA5FDBEFF106A9FFF13" box="[659,749,187,211]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
|
||
, NHMW 2021/0127/0003, Enzesfeld (Austria).
|
||
<emphasis id="B949EAD5FF8FFFA5FBCEFF106C89FF13" bold="true" box="[1251,1275,187,211]" pageId="13" pageNumber="14">R.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FA29FF1069CBFF37" authority="(Eichwald, 1830)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Eichwald" baseAuthorityYear="1830" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="laevigata">
|
||
<emphasis id="B949EAD5FF8FFFA5FA29FF106887FF37" italics="true" pageId="13" pageNumber="14">Megaclavatula laevigata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8FFFA5FE24FF7469C0FF37" author="Eichwald, E." box="[265,434,223,247]" pageId="13" pageNumber="14" refId="ref131673" refString="Eichwald, E. (1830) Naturhistorische Skizze von Lithauen, Volhynien und Podolien in Geognostisch-Mineralogischer, Botanischer und Zoologischer Hinsicht. Wilna, Voss, 255 pp. [https: // reader. digitale-sammlungen. de / de / fs 1 / object / display / bsb 10691419 _ 00005. html]" type="book" year="1830">Eichwald, 1830</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW 1853/0038/0035a, Korytnica (Poland).
|
||
<emphasis id="B949EAD5FF8FFFA5FCE5FF756BAEFF36" bold="true" box="[968,988,222,246]" pageId="13" pageNumber="14">S.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FCC4FF746CB9FF37" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[1001,1227,223,247]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="pilleri" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF8FFFA5FCC4FF746CB9FF37" box="[1001,1227,223,247]" italics="true" pageId="13" pageNumber="14">Megaclavatula pilleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8FFFA5FBFAFF746D5FFF36" box="[1239,1325,223,246]" pageId="13" pageNumber="14" rank="species">nov. sp.</taxonomicNameLabel>
|
||
,
|
||
<typeStatus id="54868865FF8FFFA5FA10FF746DEAFF37" box="[1341,1432,223,247]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
, NHMW 1949/0005/0051a, Baden-Sooss (Austria).
|
||
<emphasis id="B949EAD5FF8FFFA5FD89FEA86AC8FEDB" bold="true" box="[676,698,259,283]" pageId="13" pageNumber="14">T.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FDEFFEA86C24FEDB" authority="(Pusch, 1837)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Pusch" baseAuthorityYear="1837" box="[706,1110,259,283]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="polonica">
|
||
<emphasis id="B949EAD5FF8FFFA5FDEFFEA86BCEFEDB" box="[706,956,259,283]" italics="true" pageId="13" pageNumber="14">Megaclavatula polonica</emphasis>
|
||
(Pusch, 1837)
|
||
</taxonomicName>
|
||
, NHMW 1862/0001/0324, Bad Vöslau (Austria).
|
||
<emphasis id="B949EAD5FF8FFFA5FE64FE8C6913FEFF" bold="true" box="[329,353,295,319]" pageId="13" pageNumber="14">U.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FE4AFE8C6B0FFEFF" authority="(Hoernes & Auinger, 1891)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[359,893,295,319]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="angelae">
|
||
<emphasis id="B949EAD5FF8FFFA5FE4AFE8C6A11FEFF" box="[359,611,295,319]" italics="true" pageId="13" pageNumber="14">Granulatocincta angelae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8FFFA5FD42FE8C6B07FEFF" author="Hoernes, R. & Auinger, M." box="[623,885,295,319]" pageId="13" pageNumber="14" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8FFFA5FCABFE8C6C63FEFF" box="[902,1041,295,319]" pageId="13" pageNumber="14" type="paralectotype">paralectotype</typeStatus>
|
||
, NHMW 1949/0005/0099, Möllersdorf (Austria).
|
||
<emphasis id="B949EAD5FF8FFFA5FE2EFEE0696AFEA3" bold="true" box="[259,280,331,355]" pageId="13" pageNumber="14">V.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FE0FFEE06A07FEA3" authority="granulatocincta" box="[290,629,331,355]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="granulatocincta">
|
||
<emphasis id="B949EAD5FF8FFFA5FE0FFEE06A07FEA3" box="[290,629,331,355]" italics="true" pageId="13" pageNumber="14">Granulatocincta granulatocincta</emphasis>
|
||
</taxonomicName>
|
||
(Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria).
|
||
<emphasis id="B949EAD5FF8FFFA5FFBAFEC468C6FE47" bold="true" box="[151,180,367,391]" pageId="13" pageNumber="14">W.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FF96FEC46AACFE47" authority="(Hoernes & Auinger, 1891)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[187,734,367,391]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="nataliae">
|
||
<emphasis id="B949EAD5FF8FFFA5FF96FEC469CFFE47" box="[187,445,367,391]" italics="true" pageId="13" pageNumber="14">Granulatocincta nataliae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8FFFA5FEE6FEC46AA4FE47" author="Hoernes, R. & Auinger, M." box="[459,726,367,391]" pageId="13" pageNumber="14" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW 1855/0035/0021,
|
||
<typeStatus id="54868865FF8FFFA5FCDBFEC46C23FE47" box="[1014,1105,367,391]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
,
|
||
<collectingRegion id="49F9F825FF8FFFA5FB71FEC46CC4FE47" box="[1116,1206,367,391]" country="Serbia" name="Beograd" pageId="13" pageNumber="14">Belgrade</collectingRegion>
|
||
/Rakovica (Serbia).
|
||
<emphasis id="B949EAD5FF8FFFA5FAA9FEC46DEEFE47" bold="true" box="[1412,1436,367,391]" pageId="13" pageNumber="14">X.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FFBAFE386A21FE6A" authority="(Hornes, 1854)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hornes" baseAuthorityYear="1854" box="[151,595,402,427]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="schreibersi">
|
||
<emphasis id="B949EAD5FF8FFFA5FFBAFE3869C6FE6B" box="[151,436,403,427]" italics="true" pageId="13" pageNumber="14">Granulatocincta schreibersi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8FFFA5FEEDFE386A3EFE6A" author="Hornes, M." box="[448,588,402,427]" pageId="13" pageNumber="14" pagination="297 - 382" refId="ref135553" refString="Hornes, M. (1854) Die fossilen Mollusken des Tertiar-Beckens von Wien. Abhandlungen der K. K. Geologischen Reichsanstalt, 3, 297 - 382, pls. XXXIII - XL (1854). [https: // opac. geologie. ac. at / ais 312 / dokumente / Abh- 3 - GRA-gesamt. pdf]" type="journal article" year="1854">Hörnes, 1854</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8FFFA5FD70FE386A9AFE6B" box="[605,744,403,427]" pageId="13" pageNumber="14" type="paralectotype">paralectotype</typeStatus>
|
||
, NHMW1997z0178/1133, Bad Vöslau (Austria).
|
||
<emphasis id="B949EAD5FF8FFFA5FBF4FE386C9DFE6B" bold="true" box="[1241,1263,403,427]" pageId="13" pageNumber="14">Y.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FBD8FE3868A4FE0F" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="callim" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF8FFFA5FBD8FE3868A4FE0F" italics="true" pageId="13" pageNumber="14">Granulatocincta callim</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8FFFA5FFF6FE1C6958FE0E" box="[219,298,439,462]" pageId="13" pageNumber="14" rank="species">nov. sp.</taxonomicNameLabel>
|
||
,
|
||
<typeStatus id="54868865FF8FFFA5FE1EFE1C69FFFE0F" box="[307,397,439,463]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
|
||
, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria).
|
||
<emphasis id="B949EAD5FF8FFFA5FC81FE1C6BB1FE0F" bold="true" box="[940,963,439,463]" pageId="13" pageNumber="14">Z.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FCE5FE1C6DEBFE0E" authority="(Svagrovsky, 1958)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Svagrovsky" baseAuthorityYear="1958" box="[968,1433,438,465]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="contorta">
|
||
<emphasis id="B949EAD5FF8FFFA5FCE5FE1C6CB9FE0E" box="[968,1227,439,463]" italics="true" pageId="13" pageNumber="14">Granulatocincta contorta</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8FFFA5FBFAFE136DE3FE0E" author="Svagrovsky, J." box="[1239,1425,438,465]" pageId="13" pageNumber="14" pagination="5 - 56" refId="ref145176" refString="Svagrovsky, J. (1958) Miocenne Pleurotomidae zapadokarpatskych panvi (Miozane Pleurotomidae der westkarpatischen Becken). Acta Geologica et Geographica Universitatis Comenianae, Geologica, 1, 5 - 56." type="journal article" year="1958">Švagrovský, 1958</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8FFFA5FFBAFE706880FE33" box="[151,242,475,499]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
, SNMB Z 3076, Kuzmice (Slovakia),
|
||
<emphasis id="B949EAD5FF8FFFA5FD55FE716AECFE32" bold="true" box="[632,670,474,498]" pageId="13" pageNumber="14">Aa.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FD89FE706BB7FE33" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[676,965,475,499]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="theoderichi" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF8FFFA5FD89FE706BB7FE33" box="[676,965,475,499]" italics="true" pageId="13" pageNumber="14">Granulatocincta theoderichi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8FFFA5FCE1FE706C69FE32" box="[972,1051,475,498]" pageId="13" pageNumber="14" rank="species">nov. sp.</taxonomicNameLabel>
|
||
,
|
||
<typeStatus id="54868865FF8FFFA5FB0BFE706CF2FE33" box="[1062,1152,475,499]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
|
||
, NHMW 2016/0026/0320b, Ritzing (Austria).
|
||
<emphasis id="B949EAD5FF8FFFA5FE63FE556907FDD6" bold="true" box="[334,373,510,534]" pageId="13" pageNumber="14">Ab.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FE57FE546ABFFDD6" authority="(Hornes, 1854)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hornes" baseAuthorityYear="1854" box="[378,717,510,535]" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="doderleini">
|
||
<emphasis id="B949EAD5FF8FFFA5FE57FE546A5FFDD7" box="[378,557,511,535]" italics="true" pageId="13" pageNumber="14">Olegia doderleini</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8FFFA5FD14FE546AB4FDD6" author="Hornes, M." box="[569,710,510,535]" pageId="13" pageNumber="14" pagination="297 - 382" refId="ref135553" refString="Hornes, M. (1854) Die fossilen Mollusken des Tertiar-Beckens von Wien. Abhandlungen der K. K. Geologischen Reichsanstalt, 3, 297 - 382, pls. XXXIII - XL (1854). [https: // opac. geologie. ac. at / ais 312 / dokumente / Abh- 3 - GRA-gesamt. pdf]" type="journal article" year="1854">Hörnes, 1854</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8FFFA5FDFAFE546B10FDD7" box="[727,866,511,535]" pageId="13" pageNumber="14" type="paralectotype">paralectotype</typeStatus>
|
||
, NHMW 1858/0015/0147, Wiesen (Austria).
|
||
<emphasis id="B949EAD5FF8FFFA5FA03FE556D20FDD6" bold="true" box="[1326,1362,510,534]" pageId="13" pageNumber="14">Ac.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FA7AFE54689BFDFB" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mandici" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF8FFFA5FA7AFE54689BFDFB" italics="true" pageId="13" pageNumber="14">Olegia mandici</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8FFFA5FFC3FD88694FFDFA" box="[238,317,547,570]" pageId="13" pageNumber="14" rank="species">nov. sp.</taxonomicNameLabel>
|
||
,
|
||
<typeStatus id="54868865FF8FFFA5FE6AFD8869D3FDFB" box="[327,417,547,571]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
|
||
, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia).
|
||
<emphasis id="B949EAD5FF8FFFA5FBEEFD896C95FDFA" bold="true" box="[1219,1255,546,570]" pageId="13" pageNumber="14">Ae.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FBC1FD8869CFFD9F" authority="(Hoernes & Auinger, 1891)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dorotheae">
|
||
<emphasis id="B949EAD5FF8FFFA5FBC1FD886DEEFDFB" box="[1260,1436,547,571]" italics="true" pageId="13" pageNumber="14">Olegia dorotheae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8FFFA5FFB3FDEC69C7FD9F" author="Hoernes, R. & Auinger, M." box="[158,437,583,607]" pageId="13" pageNumber="14" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8FFFA5FEE6FDEC6A24FD9F" box="[459,598,583,607]" pageId="13" pageNumber="14" type="paralectotype">paralectotype</typeStatus>
|
||
, NHMW 1949/0005/0040, Kleinebersdorf (
|
||
<collectingCountry id="F32A7657FF8FFFA5FB0EFDEC6C01FD9F" box="[1059,1139,583,607]" name="Austria" pageId="13" pageNumber="14">Austria</collectingCountry>
|
||
). Scale bar is
|
||
<quantity id="4CC59B22FF8FFFA5FA3CFDED6D3DFD9E" box="[1297,1359,582,606]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="mm" value="5.0">5 mm</quantity>
|
||
for all specimens except for
|
||
<emphasis id="B949EAD5FF8FFFA5FE5EFDC06B45FD43" box="[371,823,619,643]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FE5EFDC06A0BFD42" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Svagrovsky" baseAuthorityYear="1958" box="[371,633,619,643]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="contorta">Granulatocincta contorta</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FDA9FDC06B45FD43" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hornes" baseAuthorityYear="1854" box="[644,823,619,643]" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="doderleini">Olegia doderleini</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FC45FDC06BABFD43" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[872,985,619,643]" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mandici">
|
||
<emphasis id="B949EAD5FF8FFFA5FC45FDC06BABFD43" box="[872,985,619,643]" italics="true" pageId="13" pageNumber="14">O. mandici</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8236C7FF8FFFA5FFEAFD7E6BFAFC2B" blockId="13.[151,1437,725,2011]" pageId="13" pageNumber="14">
|
||
The protoconch of almost all known clavatulid species is paucispiral and almost always unsculptured (exceptions are
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FF92FD5269B9FCD3" authority="Gofas, 1990" authorityName="Gofas" authorityYear="1990" box="[191,459,761,787]" class="Gastropoda" family="Clavatulidae" genus="Scaevatula" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8FFFA5FF92FD526948FCD3" box="[191,314,761,787]" italics="true" pageId="13" pageNumber="14">Scaevatula</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF8FFFA5FE6DFD5269B9FCD3" author="Gofas, S." box="[320,459,761,787]" pageId="13" pageNumber="14" pagination="11 - 22" refId="ref133198" refString="Gofas, S. (1990) Scaevatula n. gen., a sinistral clavatuline Turrid from West Africa (Gastropoda: Turridae). Archiv fur Molluskenkunde, 120, 11 - 22. https: // doi. org / 10.1127 / arch. moll / 120 / 1990 / 11" type="journal article" year="1990">Gofas, 1990</bibRefCitation>
|
||
</taxonomicName>
|
||
, which has a ribbed protoconch and
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FC71FD526C60FCD3" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[860,1042,761,787]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="aueri" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF8FFFA5FC71FD526C60FCD3" box="[860,1042,761,787]" italics="true" pageId="13" pageNumber="14">Tomellana aueri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8FFFA5FB35FD526C1CFCD3" box="[1048,1134,761,787]" pageId="13" pageNumber="14" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, which has a protoconch of 2.5 whorls). This paucispiral
|
||
<typeStatus id="54868865FF8FFFA5FEFAFCB66A75FCF7" box="[471,519,797,823]" pageId="13" pageNumber="14">type</typeStatus>
|
||
of protoconch is suggestive of direct development, or at least a short planktotrophic phase, which tends to lead to stratigraphically short lived and geographically restricted species (
|
||
<bibRefCitation id="EFAC4B36FF8FFFA5FBFAFCEA68A1FCBF" author="Jablonski, D. & Lutz, R. A." pageId="13" pageNumber="14" pagination="323 - 377" refId="ref135715" refString="Jablonski, D. & Lutz, R. A. (1980) Molluscan Larval Shell Morphology, Ecological and Paleontological Applications. In: Rhoads, D. C. & Lutz, R. A. (Eds.), Skeletal Growth of Aquatic Organisms, Biological Records of Environmental Change. Plenum Press, New York and London, pp. 323 - 377. https: // doi. org / 10.1007 / 978 - 1 - 4899 - 4995 - 0 _ 10" type="book chapter" year="1980">Jablonski & Lutz 1980</bibRefCitation>
|
||
). This is certainly the pattern seen in both the fossil and present-day record and reaches its zenith in the Miocene Paratethys. Unfortunately, the protoconch is rarely preserved in adult specimens in the Paratethys record, however, based on the experience of other clavatulid populations (e.g., Pliocene Atlantic and Mediterranean), it is of limited taxonomic use in distinguishing species (own data B.L.).
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF8FFFA5FFEAFC5E6B31FAEF" blockId="13.[151,1437,725,2011]" pageId="13" pageNumber="14">
|
||
The spire in all species is relatively raised and conical, although the outline may be made irregular or somewhat gradate by a strongly developed raised subsutural collar and made coronate by spines or tubercles developed either on the adapical part of the collar or mid-collar. In many species sculpture changes markedly with ontogeny on about the fourth to sixth whorls. Early teleoconch whorls are often divided into two beaded cords separated by a narrow groove or concave area (bipartite), or bear three beaded portions, a subsutural cord, mid-cord and suprasutural cord (tripartite) (
|
||
<figureCitation id="13062A42FF8FFFA5FE37FB0269F0FB03" box="[282,386,1193,1219]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="12.[151,250,1619,1643]" captionStartId-1="14.[151,250,1619,1643]" captionTargetBox-0="[151,1436,181,1595]" captionTargetBox-1="[151,1436,181,1595]" captionTargetId-0="figure-26@12.[151,1436,181,1595]" captionTargetId-1="figure-26@14.[151,1436,181,1595]" captionTargetPageId-0="12" captionTargetPageId-1="14" captionText-0="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." captionText-1="FIGURE 4A. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), NHMW ex1949/0005/0110b, Buituri (Romania). B. ‘Clavatula’ mariae (Hoernes & Auinger, 1891), lectotype, NHMW 1849/0013/0009, Mold (Austria). C. ‘Clavatula’ olgae (Hoernes & Auinger, 1891), lectotype, NHMW 1862/0001/0552, Buituri (Romania). D. ‘Clavatula’ orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). E. ‘Clavatula’ juliae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0043c, Lăpugiu de Sus (Romania). F. ‘Clavatula’ apolloniae (Hoernes & Auinger, 1891), NHMW 1868/0001/0474a, Lăpugiu de Sus (Romania). G. Neoperrona harzhauseri (Kovács & Vicián, 2021), NHMW 2019/0151/019a, Letkés (Hungary). H. Neoperrona zoltanorum nov. sp., holotype, NHMW 2019/0151/0108, Letkés (Hungary). I. Scaevatula sidoniae (Hoernes & Auinger, 1891), NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). J. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0003, Várpalota (Hungary). K. Pusionella hofmanni nov. sp., NHMW 1867/0019/0136, holotype, Coşteiu de Sus (Romania). L. Perrona barbarae (Hoernes & Auinger, 1891) NHMW 1949/0005/0103a, Rückersdorf (Austria). M. Perrona descendens (Hilber, 1879), NHMW 1846/0049/0017, St. Florian (Austria). N. Perrona emmae (Hoernes & Auinger 1891), NHMW 2021/0127/0005, Enzesfeld (Austria), NHMW 1970/1396/1449, Baden (Austria). O. Perrona ernae (Šuklje, 1929), NHMW 2021/0156/0001, Weinsteig (Austria). P. Perrona floriana (Hilber, 1879), holotype, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria). Q. Perrona ilonae nov. nom., M.61.4367, Szentkút, Mátraverebély (Hungary). R. Perrona lydiae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0112, Gainfarn (Austria). S. Perrona oliviae (Hoernes & Auinger 1891), paralectotype, NHMW 1873/0026/0027, Lăpugiu de Sus (Romania). T. Perrona rosaliae (Hoernes & Auinger 1891), lectotype, NHMW 1999z0098/0002, Vienna/Pötzleinsdorf (Austria). U. Perrona sabinae (Hoernes & Auinger 1891), paralectotype, NHMW 2021/0154/0001, Lăpugiu de Sus (Romania). V. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309, Pöls (Austria). W. Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). X. Perrona vindobonensis (Quenstedt, 1884), paralectotype, NHMW 1997z0178/1467, Gainfarn (Austria), Y. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). Z. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). Aa. Tomellana aueri nov. sp., NHMW 2013/0078/0436, Baden (Austria). Ab. Striopusionella szekelyhidiae (Kovács & Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). Ac. Scaevatula sidoniae (Hoernes & Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). Scale bar is 5 mm for all specimens except for Perrona barbarae and protoconch of Scaevatula sidoniae." figureDoi-0="http://doi.org/10.5281/zenodo.6399835" figureDoi-1="http://doi.org/10.5281/zenodo.6399837" httpUri-0="https://zenodo.org/record/6399835/files/figure.png" httpUri-1="https://zenodo.org/record/6399837/files/figure.png" pageId="13" pageNumber="14">Figs 3–4</figureCitation>
|
||
). In some species the early teleoconch sculpture is neither bipartite nor tripartite but consists of very narrow adsutural cords and a wide concave mid-portion (e.g.,
|
||
<emphasis id="B949EAD5FF8FFFA5FC80FB666CD2FB27" box="[941,1184,1229,1255]" italics="true" pageId="13" pageNumber="14">‘Clavatula’ jarzynkae</emphasis>
|
||
nov. nom.), or a welldeveloped, but smooth, subsutural cord and a suprasutural row of beads separated by a concave portion, as is the case with many of the species placed in
|
||
<emphasis id="B949EAD5FF8FFFA5FD74FABE6A9CFAEF" box="[601,750,1301,1327]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FD74FABE6AB5FAEF" authorityName="Lamarck" authorityYear="1801" box="[601,711,1301,1327]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus" sensu="lato">Clavatula</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8FFFA5FDE3FABE6A9CFAEF" box="[718,750,1301,1327]" pageId="13" pageNumber="14" sensu="lato">s.l.</taxonomicNameLabel>
|
||
</emphasis>
|
||
herein.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF8FFFA5FFEAFA926A53FA23" blockId="13.[151,1437,725,2011]" pageId="13" pageNumber="14">
|
||
On later teleoconch whorls the subsutural cord swells to form the subsutural collar, the mid-portion beads fade and become smooth or finely spirally striate, and the beads on the suprasutural cord swell and become tubercles. In some species the subsutural cord is not visible, obscured by the succeeding whorl. On the last whorl it becomes clear that the mid-portion on the spire whorls represents the subsutural ramp and the suprasutural row of tubercles coincide with the shoulder (
|
||
<figureCitation id="13062A42FF8FFFA5FEE2FA626A66FA23" box="[463,532,1481,1507]" captionStart="FIGURE 2" captionStartId="7.[151,250,1264,1288]" captionTargetBox="[151,1436,597,1241]" captionTargetId="figure-187@7.[151,1436,597,1241]" captionTargetPageId="7" captionText="FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width." figureDoi="http://doi.org/10.5281/zenodo.6399831" httpUri="https://zenodo.org/record/6399831/files/figure.png" pageId="13" pageNumber="14">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF8FFFA5FFEAFA466900F97B" blockId="13.[151,1437,725,2011]" pageId="13" pageNumber="14">
|
||
In all
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FE20FA46690AF9C7" authorityName="J. Fleming" authorityYear="1822" box="[269,376,1517,1543]" class="Gastropoda" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="12" phylum="Mollusca" rank="superFamily" superFamily="Conoidea">Conoidea</taxonomicName>
|
||
the anal sinus is an important character, and often useful for separation at both generic and species level. In clavatulids the sinus is well-developed and clearly preserved in all species (
|
||
<figureCitation id="13062A42FF8FFFA5FBFBF9BA6D6CF9EC" box="[1238,1310,1553,1580]" captionStart="FIGURE 5" captionStartId="15.[151,250,1999,2023]" captionTargetBox="[151,1436,631,1976]" captionTargetId="figure-214@15.[151,1436,631,1976]" captionTargetPageId="15" captionText="FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae." figureDoi="http://doi.org/10.5281/zenodo.6399839" httpUri="https://zenodo.org/record/6399839/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
|
||
). On early teleoconch whorls the sinus can be represented by comma-shaped axial riblets but these structures are strong growth lines but do not represent true ribs. Ribs are uncommon in clavatulids, and when present are poorly defined and formed by axially aligned rows of tubercles (i.e., ‘
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FDF7F9D66B3AF957" authorityName="Lamarck" authorityYear="1801" box="[730,840,1661,1687]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8FFFA5FDF7F9D66B3AF957" box="[730,840,1661,1687]" italics="true" pageId="13" pageNumber="14">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FC75F9D66BA2F957" authorityName="Hoernes & Auinger" authorityYear="1891" box="[856,976,1661,1687]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="apolloniae">
|
||
<emphasis id="B949EAD5FF8FFFA5FC75F9D66BA2F957" box="[856,976,1661,1687]" italics="true" pageId="13" pageNumber="14">apolloniae</emphasis>
|
||
</taxonomicName>
|
||
), and are only present on the last whorl below the shoulder.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF8FFFA5FFEAF96E6D13F81B" blockId="13.[151,1437,725,2011]" pageId="13" pageNumber="14">
|
||
Spiral elements on the spire whorls are composed of sub- and suprasutural cords on the early whorls, which later become the subsutural collar and suprasutural row of tubercles (as discussed above). Many species have secondary spiral cords that cover the entire surface including the collar and suprasutural tubercles. Spiral sculpture on the last whorl is more variable. The shoulder cord is tubercular (occasionally spinous) in
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FB8BF89A6D1FF88B" box="[1190,1389,1841,1867]" pageId="13" pageNumber="14">
|
||
<emphasis id="B949EAD5FF8FFFA5FB8BF89A6D66F88B" box="[1190,1300,1841,1867]" italics="true" pageId="13" pageNumber="14">Clavatula</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
and usually smooth or rarely weakly tubercular in
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FD8FF8FD6B2CF8AF" box="[674,862,1877,1903]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B949EAD5FF8FFFA5FD8FF8FD6A8CF8AF" box="[674,766,1878,1903]" italics="true" pageId="13" pageNumber="14">Perrona</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
Usually, a peribasal and perifasciolar cord is more prominent, and less often a mid-whorl cord, all of which are more or less tubercular. Rarely a species may have five primary cords below the shoulder (i.e., ‘
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FD71F8366AB8F877" authorityName="Lamarck" authorityYear="1801" box="[604,714,1949,1975]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8FFFA5FD71F8366AB8F877" box="[604,714,1949,1975]" italics="true" pageId="13" pageNumber="14">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8FFFA5FDF4F8366B23F877" authorityName="Hoernes & Auinger" authorityYear="1891" box="[729,849,1949,1975]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="apolloniae">
|
||
<emphasis id="B949EAD5FF8FFFA5FDF4F8366B23F877" box="[729,849,1949,1975]" italics="true" pageId="13" pageNumber="14">apolloniae</emphasis>
|
||
</taxonomicName>
|
||
). Secondary and tertiary spiral sculpture tends to be rather irregular in clavatulids and is often not only present in the interspaces, but overrides the primary cords.
|
||
</paragraph>
|
||
<caption id="DF42664FFF8CFFA7FFBAF9F86C46FD9F" ID-DOI="http://doi.org/10.5281/zenodo.6399837" ID-Zenodo-Dep="6399837" httpUri="https://zenodo.org/record/6399837/files/figure.png" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" startId="14.[151,250,1619,1643]" targetBox="[151,1436,181,1595]" targetPageId="14">
|
||
<paragraph id="8B8236C7FF8CFFA6FFBAF9F86DEEF838" blockId="14.[151,1437,1619,2042]" pageId="14" pageNumber="15">
|
||
<emphasis id="B949EAD5FF8CFFA6FFBAF9F8695AF9AB" bold="true" box="[151,296,1619,1643]" pageId="14" pageNumber="15">FIGURE 4 A.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FE18F9F869EBF9AB" authorityName="Lamarck" authorityYear="1801" box="[309,409,1619,1643]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8CFFA6FE18F9F869EBF9AB" box="[309,409,1619,1643]" italics="true" pageId="14" pageNumber="15">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FE8AF9FF6B63F9AC" authority="(Hoernes & Auinger, 1891)" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[423,785,1619,1644]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="justinae">
|
||
<emphasis id="B949EAD5FF8CFFA6FE8AF9FF6985F9AB" box="[423,503,1620,1643]" italics="true" pageId="14" pageNumber="15">justinae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8CFFA6FD2EF9FF6B7BF9AC" author="Hoernes, R. & Auinger, M." box="[515,777,1619,1644]" pageId="14" pageNumber="15" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW ex1949/0005/0110b, Buituri (Romania).
|
||
<emphasis id="B949EAD5FF8CFFA6FA26F9FF6D50F9AC" bold="true" box="[1291,1314,1620,1644]" pageId="14" pageNumber="15">B.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FA1DF9F86DE6F9AB" authorityName="Lamarck" authorityYear="1801" box="[1328,1428,1619,1643]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8CFFA6FA1DF9F86DE6F9AB" box="[1328,1428,1619,1643]" italics="true" pageId="14" pageNumber="15">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FFBAF9D36A7BF950" authority="(Hoernes & Auinger, 1891)" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[151,521,1655,1680]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="mariae">
|
||
<emphasis id="B949EAD5FF8CFFA6FFBAF9D368ADF94F" box="[151,223,1656,1679]" italics="true" pageId="14" pageNumber="15">mariae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8CFFA6FFC2F9D36A70F950" author="Hoernes, R. & Auinger, M." box="[239,514,1655,1680]" pageId="14" pageNumber="15" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, lectotype, NHMW 1849/0013/0009, Mold (Austria).
|
||
<emphasis id="B949EAD5FF8CFFA6FB6DF9DC6C2BF94F" bold="true" box="[1088,1113,1655,1679]" pageId="14" pageNumber="15">C.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FB46F9DC6CBDF94F" authorityName="Lamarck" authorityYear="1801" box="[1131,1231,1655,1679]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8CFFA6FB46F9DC6CBDF94F" box="[1131,1231,1655,1679]" italics="true" pageId="14" pageNumber="15">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FBCCF9DC6946F974" authority="(Hoernes & Auinger, 1891)" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="olgae">
|
||
<emphasis id="B949EAD5FF8CFFA6FBCCF9DC6D6BF94F" box="[1249,1305,1655,1679]" italics="true" pageId="14" pageNumber="15">olgae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8CFFA6FA04F9D3695FF974" author="Hoernes, R. & Auinger, M." pageId="14" pageNumber="15" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, lectotype, NHMW 1862/0001/0552, Buituri (Romania).
|
||
<emphasis id="B949EAD5FF8CFFA6FC52F9376BE5F974" bold="true" box="[895,919,1692,1716]" pageId="14" pageNumber="15">D.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FC8AF9306C79F973" authorityName="Lamarck" authorityYear="1801" box="[935,1035,1691,1715]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8CFFA6FC8AF9306C79F973" box="[935,1035,1691,1715]" italics="true" pageId="14" pageNumber="15">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FB36F9376D40F973" authority="Baldi, 1960" authorityName="Baldi" authorityYear="1960" box="[1051,1330,1691,1715]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="orientoromana">
|
||
<emphasis id="B949EAD5FF8CFFA6FB36F9376CC0F973" box="[1051,1202,1692,1715]" italics="true" pageId="14" pageNumber="15">orientoromana</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF8CFFA6FB94F9306D40F973" author="Baldi, T." box="[1209,1330,1691,1715]" pageId="14" pageNumber="15" pagination="51 - 99" refId="ref128619" refString="Baldi, T. (1960) Tortonische Molluskenfauna von " Badener Tegelfazies' ans Szokolya, Nordungarn. Annales Historico- Naturales Musei Nationalis Hungarici, pars Mineralogica et Palaeontologica, 52, 51 - 99. [http: // publication. nhmus. hu / pdf / annHNHM / Annals _ HNHM _ 1960 _ Vol _ 52 _ 51. pdf]" type="journal article" year="1960">Báldi, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype, M.60.145, Szokolya (Hungary).
|
||
<emphasis id="B949EAD5FF8CFFA6FEC4F96B6A72F918" bold="true" box="[489,512,1728,1752]" pageId="14" pageNumber="15">E.</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FD3FF9146A04F917" authorityName="Lamarck" authorityYear="1801" box="[530,630,1727,1751]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF8CFFA6FD3FF9146A04F917" box="[530,630,1727,1751]" italics="true" pageId="14" pageNumber="15">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FDAAF9146B96F918" authority="(Hoernes & Auinger 1891)" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[647,996,1727,1752]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="juliae">
|
||
<emphasis id="B949EAD5FF8CFFA6FDAAF9146AB3F917" box="[647,705,1727,1751]" italics="true" pageId="14" pageNumber="15">juliae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8CFFA6FDFCF96B6BAFF918" author="Hoernes, R. & Auinger, M." box="[721,989,1727,1752]" pageId="14" pageNumber="15" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, paralectotype, NHMW 1949/0005/0043c, Lăpugiu de Sus (Romania).
|
||
<emphasis id="B949EAD5FF8CFFA6FEE2F94F6990F93C" bold="true" box="[463,482,1764,1788]" pageId="14" pageNumber="15">F.</emphasis>
|
||
<emphasis id="B949EAD5FF8CFFA6FEC2F9486A13F93B" box="[495,609,1763,1787]" italics="true" pageId="14" pageNumber="15">‘Clavatula’</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FD43F9486C61F93C" authority="(Hoernes & Auinger, 1891)" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[622,1043,1763,1788]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="apolloniae">
|
||
<emphasis id="B949EAD5FF8CFFA6FD43F9486AA8F93B" box="[622,730,1763,1787]" italics="true" pageId="14" pageNumber="15">apolloniae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8CFFA6FDC3F94F6C79F93C" author="Hoernes, R. & Auinger, M." box="[750,1035,1763,1788]" pageId="14" pageNumber="15" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW 1868/0001/0474a, Lăpugiu de Sus (Romania).
|
||
<emphasis id="B949EAD5FF8CFFA6FE48F8AC690DF8DF" bold="true" box="[357,383,1799,1823]" pageId="14" pageNumber="15">G.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FEA4F8A36BE8F8E0" authority="(Kovacs & Vician, 2021)" baseAuthorityName="Kovacs & Vician" baseAuthorityYear="2021" box="[393,922,1799,1824]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="harzhauseri">
|
||
<emphasis id="B949EAD5FF8CFFA6FEA4F8A36AF7F8DF" box="[393,645,1799,1823]" italics="true" pageId="14" pageNumber="15">Neoperrona harzhauseri</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8CFFA6FDBBF8AC6BE1F8E0" author="Kovacs, Z. & Vician, Z." box="[662,915,1799,1824]" pageId="14" pageNumber="15" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián, 2021</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW 2019/0151/019a, Letkés (Hungary).
|
||
<emphasis id="B949EAD5FF8CFFA6FAAFF8A36DEEF8E0" bold="true" box="[1410,1436,1800,1824]" pageId="14" pageNumber="15">H.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FFBAF88769FFF883" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[151,397,1835,1859]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="zoltanorum" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF8CFFA6FFBAF88769FFF883" box="[151,397,1835,1859]" italics="true" pageId="14" pageNumber="15">Neoperrona zoltanorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8CFFA6FEBBF887699AF883" box="[406,488,1836,1859]" pageId="14" pageNumber="15" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, holotype, NHMW 2019/0151/0108, Letkés (Hungary).
|
||
<emphasis id="B949EAD5FF8CFFA6FB1EF8876C31F884" bold="true" box="[1075,1091,1836,1860]" pageId="14" pageNumber="15">I.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FB61F8806941F8A8" authority="(Hoernes & Auinger, 1891)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" class="Gastropoda" family="Clavatulidae" genus="Scaevatula" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="sidoniae">
|
||
<emphasis id="B949EAD5FF8CFFA6FB61F8806D68F883" box="[1100,1306,1835,1859]" italics="true" pageId="14" pageNumber="15">Scaevatula sidoniae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8CFFA6FA07F887695EF8A8" author="Hoernes, R. & Auinger, M." pageId="14" pageNumber="15" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania).
|
||
<emphasis id="B949EAD5FF8CFFA6FCE1F8FB6BADF8A8" bold="true" box="[972,991,1872,1896]" pageId="14" pageNumber="15">J.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FCC8F8E46D4DF8A7" authority="Strausz, 1954" authorityName="Strausz" authorityYear="1954" box="[997,1343,1871,1896]" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="palatina">
|
||
<emphasis id="B949EAD5FF8CFFA6FCC8F8E46CDEF8A7" box="[997,1196,1871,1895]" italics="true" pageId="14" pageNumber="15">Pusionella palatina</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF8CFFA6FB9EF8FB6D4DF8A7" author="Strausz, L." box="[1203,1343,1871,1896]" pageId="14" pageNumber="15" pagination="1 - 150" refId="ref144830" refString="Strausz, L. (1954) Varpalotai felso-mediteean csigak. Geologica Hungarica, 25, 1 - 150. [http: // epa. oszk. hu / 02900 / 02989 / 00022 / pdf / EPA 02989 _ geologica _ hungarica _ ser _ paleo _ 25 _ 1954 _ 085 - 125. pdf]" type="journal article" year="1954">Strausz, 1954</bibRefCitation>
|
||
</taxonomicName>
|
||
, NHMW 2021/0159/0003, Várpalota (Hungary).
|
||
<emphasis id="B949EAD5FF8CFFA6FD02F8DF6A3BF84C" bold="true" box="[559,585,1908,1932]" pageId="14" pageNumber="15">K.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FD7CF8D86B54F84B" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[593,806,1907,1931]" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="hofmanni" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF8CFFA6FD7CF8D86B54F84B" box="[593,806,1907,1931]" italics="true" pageId="14" pageNumber="15">Pusionella hofmanni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8CFFA6FC03F8DF6B0DF84B" box="[814,895,1908,1931]" pageId="14" pageNumber="15" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, NHMW 1867/0019/0136, holotype, Coşteiu de Sus (Romania).
|
||
<emphasis id="B949EAD5FF8CFFA6FE3FF833695BF870" bold="true" box="[274,297,1944,1968]" pageId="14" pageNumber="15">L.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FE1FF8336BD7F86F" authority="(Hoernes & Auinger, 1891) NHMW 1949" authorityName="NHMW" authorityYear="1949" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[306,933,1943,1968]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="barbarae">
|
||
<emphasis id="B949EAD5FF8CFFA6FE1FF8336999F86F" box="[306,491,1943,1967]" italics="true" pageId="14" pageNumber="15">Perrona barbarae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8CFFA6FED6F8336B7AF870" author="Hoernes, R. & Auinger, M." box="[507,776,1943,1968]" pageId="14" pageNumber="15" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
) NHMW 1949
|
||
</taxonomicName>
|
||
/0005/0103a, Rückersdorf (Austria).
|
||
<emphasis id="B949EAD5FF8CFFA6FA0FF8336D33F870" bold="true" box="[1314,1345,1944,1968]" pageId="14" pageNumber="15">M.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FA64F83369DBF814" authority="(Hilber, 1879)" baseAuthorityName="Hilber" baseAuthorityYear="1879" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="descendens">
|
||
<emphasis id="B949EAD5FF8CFFA6FA64F8336978F813" italics="true" pageId="14" pageNumber="15">Perrona descendens</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8CFFA6FE36F81069D0F814" author="Hilber, V." box="[283,418,1979,2004]" pageId="14" pageNumber="15" pagination="416 - 464" refId="ref134915" refString="Hilber, V. (1879) Neue Conchylien aus den mittelsteierischen Mediterranschichten. Sitzungsberichte der kaiserlichen Akademie der Wissenschaften, der Mathematisch-Naturwissenschaftlichen Classe, 79, 416 - 464. [https: // www. zobodat. at / pdf / SBAWW _ 79 _ 0416 - 0464. pdf]" type="journal article" year="1879">Hilber, 1879</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW 1846/0049/0017, St. Florian (Austria).
|
||
<emphasis id="B949EAD5FF8CFFA6FC81F8176BB6F814" bold="true" box="[940,964,1980,2004]" pageId="14" pageNumber="15">N.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FCE3F8176DEAF814" authority="(Hoernes & Auinger 1891)" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[974,1432,1979,2004]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="emmae">
|
||
<emphasis id="B949EAD5FF8CFFA6FCE3F8176C01F813" box="[974,1139,1980,2003]" italics="true" pageId="14" pageNumber="15">Perrona emmae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8CFFA6FBAEF8176DE3F814" author="Hoernes, R. & Auinger, M." box="[1155,1425,1979,2004]" pageId="14" pageNumber="15" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, NHMW 2021/0127/0005, Enzesfeld (Austria), NHMW 1970/1396/1449, Baden (Austria).
|
||
<emphasis id="B949EAD5FF8CFFA6FB6FF8746C2EF837" bold="true" box="[1090,1116,2015,2039]" pageId="14" pageNumber="15">O.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8CFFA6FB49F84B6DEAF838" authority="(Suklje, 1929)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Suklje" baseAuthorityYear="1929" box="[1124,1432,2016,2042]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="ernae">
|
||
<emphasis id="B949EAD5FF8CFFA6FB49F84B6C88F837" box="[1124,1274,2016,2039]" italics="true" pageId="14" pageNumber="15">Perrona ernae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8CFFA6FA27F84A6DE3F838" author="Suklje, F." box="[1290,1425,2016,2042]" pageId="14" pageNumber="15" pagination="1 - 52" refId="ref145139" refString="Suklje, F. (1929) Mediteranska fauna zapresic Brijega u Samoborskoj Gori. Vijesti Geoloskoga zavoda, 3, 1 - 52. https: // doi. org / 10.3996 / nafa. 52.0001" type="journal article" year="1929">Šuklje, 1929</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF8DFFA7FFBAFF3C6C46FD9F" blockId="15.[151,1437,150,607]" pageId="15" pageNumber="16">
|
||
NHMW 2021/0156/0001, Weinsteig (Austria).
|
||
<emphasis id="B949EAD5FF8DFFA7FDB9FF3C6AD5FF6F" bold="true" box="[660,679,151,175]" pageId="15" pageNumber="16">P.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FD98FF3C6C62FF6F" authority="(Hilber, 1879)" baseAuthorityName="Hilber" baseAuthorityYear="1879" box="[693,1040,151,175]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="floriana">
|
||
<emphasis id="B949EAD5FF8DFFA7FD98FF3C6B1BFF6F" box="[693,873,151,175]" italics="true" pageId="15" pageNumber="16">Perrona floriana</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8DFFA7FC50FF3C6C7BFF6F" author="Hilber, V." box="[893,1033,151,175]" pageId="15" pageNumber="16" pagination="416 - 464" refId="ref134915" refString="Hilber, V. (1879) Neue Conchylien aus den mittelsteierischen Mediterranschichten. Sitzungsberichte der kaiserlichen Akademie der Wissenschaften, der Mathematisch-Naturwissenschaftlichen Classe, 79, 416 - 464. [https: // www. zobodat. at / pdf / SBAWW _ 79 _ 0416 - 0464. pdf]" type="journal article" year="1879">Hilber, 1879</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8DFFA7FB0FFF3C6C0FFF6F" box="[1058,1149,151,175]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria).
|
||
<emphasis id="B949EAD5FF8DFFA7FEC7FF116A76FF12" bold="true" box="[490,516,186,210]" pageId="15" pageNumber="16">Q.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FD20FF106ADBFF13" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[525,681,187,211]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="ilonae" status="nom. nov.">
|
||
<emphasis id="B949EAD5FF8DFFA7FD20FF106ADBFF13" box="[525,681,187,211]" italics="true" pageId="15" pageNumber="16">Perrona ilonae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8DFFA7FD9EFF106B6FFF12" box="[691,797,187,210]" pageId="15" pageNumber="16">nov. nom.</taxonomicNameLabel>
|
||
, M.61.4367, Szentkút, Mátraverebély (Hungary).
|
||
<emphasis id="B949EAD5FF8DFFA7FA0AFF106D4DFF13" bold="true" box="[1319,1343,187,211]" pageId="15" pageNumber="16">R.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FA64FF106A72FF37" authority="(Hoernes & Auinger, 1891)" baseAuthorityName="R. Hoernes & Auinger" baseAuthorityYear="1891" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="lydiae">
|
||
<emphasis id="B949EAD5FF8DFFA7FA64FF1068A7FF37" italics="true" pageId="15" pageNumber="16">Perrona lydiae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8DFFA7FFC8FF74698AFF37" author="Hoernes, R. & Auinger, M." box="[229,504,223,247]" pageId="15" pageNumber="16" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8DFFA7FD20FF746A1FFF37" box="[525,621,223,247]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
|
||
, NHMW 1949/0005/0112, Gainfarn (Austria).
|
||
<emphasis id="B949EAD5FF8DFFA7FB74FF756C1FFF36" bold="true" box="[1113,1133,222,246]" pageId="15" pageNumber="16">S.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FB5AFF74695DFEDB" authority="(Hoernes & Auinger 1891)" baseAuthorityName="R. Hoernes & Auinger" baseAuthorityYear="1891" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="oliviae">
|
||
<emphasis id="B949EAD5FF8DFFA7FB5AFF746D6BFF37" box="[1143,1305,223,247]" italics="true" pageId="15" pageNumber="16">Perrona oliviae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8DFFA7FA04FF746955FEDB" author="Hoernes, R. & Auinger, M." pageId="15" pageNumber="16" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8DFFA7FE17FEA869B7FEDB" box="[314,453,259,283]" pageId="15" pageNumber="16" type="paralectotype">paralectotype</typeStatus>
|
||
, NHMW 1873/0026/0027, Lăpugiu de Sus (Romania).
|
||
<emphasis id="B949EAD5FF8DFFA7FCD1FEA86C60FEDB" bold="true" box="[1020,1042,259,283]" pageId="15" pageNumber="16">T.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FB35FEA868A7FEFF" authority="(Hoernes & Auinger 1891)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="rosaliae">
|
||
<emphasis id="B949EAD5FF8DFFA7FB35FEA86CB7FEDB" box="[1048,1221,259,283]" italics="true" pageId="15" pageNumber="16">Perrona rosaliae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8DFFA7FBFEFEA868BFFEFF" author="Hoernes, R. & Auinger, M." pageId="15" pageNumber="16" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8DFFA7FFCCFE8C6933FEFF" box="[225,321,295,319]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
|
||
, NHMW 1999z0098/0002,
|
||
<collectingRegion id="49F9F825FF8DFFA7FD4CFE8C6ADBFEFF" box="[609,681,295,319]" country="Austria" name="Wien" pageId="15" pageNumber="16">Vienna</collectingRegion>
|
||
/Pötzleinsdorf (Austria).
|
||
<emphasis id="B949EAD5FF8DFFA7FC87FE8C6BB0FEFF" bold="true" box="[938,962,295,319]" pageId="15" pageNumber="16">U.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FCE6FE8C6DEAFEFF" authority="(Hoernes & Auinger 1891)" baseAuthorityName="R. Hoernes & Auinger" baseAuthorityYear="1891" box="[971,1432,295,319]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="sabinae">
|
||
<emphasis id="B949EAD5FF8DFFA7FCE6FE8C6C04FEFF" box="[971,1142,295,319]" italics="true" pageId="15" pageNumber="16">Perrona sabinae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8DFFA7FBABFE8C6DE3FEFF" author="Hoernes, R. & Auinger, M." box="[1158,1425,295,319]" pageId="15" pageNumber="16" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8DFFA7FFBAFEE06950FEA3" box="[151,290,331,355]" pageId="15" pageNumber="16" type="paralectotype">paralectotype</typeStatus>
|
||
, NHMW 2021/0154/0001, Lăpugiu de Sus (Romania).
|
||
<emphasis id="B949EAD5FF8DFFA7FC4AFEE06B0EFEA3" bold="true" box="[871,892,331,355]" pageId="15" pageNumber="16">V.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FCA8FEE06CA2FEA3" authority="(Hilber, 1879)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hilber" baseAuthorityYear="1879" box="[901,1232,331,355]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="styriaca">
|
||
<emphasis id="B949EAD5FF8DFFA7FCA8FEE06C41FEA2" box="[901,1075,331,354]" italics="true" pageId="15" pageNumber="16">Perrona styriaca</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8DFFA7FB6EFEE06CBBFEA3" author="Hilber, V." box="[1091,1225,331,355]" pageId="15" pageNumber="16" pagination="416 - 464" refId="ref134915" refString="Hilber, V. (1879) Neue Conchylien aus den mittelsteierischen Mediterranschichten. Sitzungsberichte der kaiserlichen Akademie der Wissenschaften, der Mathematisch-Naturwissenschaftlichen Classe, 79, 416 - 464. [https: // www. zobodat. at / pdf / SBAWW _ 79 _ 0416 - 0464. pdf]" type="journal article" year="1879">Hilber, 1879</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8DFFA7FBF3FEE06D4CFEA3" box="[1246,1342,331,355]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
|
||
, NHMW 1846/0037/0309, Pöls (Austria).
|
||
<emphasis id="B949EAD5FF8DFFA7FEC2FEC46A7EFE47" bold="true" box="[495,524,367,391]" pageId="15" pageNumber="16">W.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FD3BFEC46B9FFE47" authority="(Hoernes & Auinger, 1891)" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[534,1005,367,391]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="ursulae">
|
||
<emphasis id="B949EAD5FF8DFFA7FD3BFEC46AB2FE47" box="[534,704,367,391]" italics="true" pageId="15" pageNumber="16">Perrona ursulae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8DFFA7FDFCFEC46B97FE47" author="Hoernes, R. & Auinger, M." box="[721,997,367,391]" pageId="15" pageNumber="16" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8DFFA7FCD6FEC46CF4FE47" box="[1019,1158,367,391]" pageId="15" pageNumber="16" type="paralectotype">paralectotype</typeStatus>
|
||
, NHMW 1846/0001/1294, Buituri (Romania).
|
||
<emphasis id="B949EAD5FF8DFFA7FE72FE38690AFE6B" bold="true" box="[351,376,403,427]" pageId="15" pageNumber="16">X.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FE53FE386B5BFE6B" authority="(Quenstedt, 1884)" baseAuthorityName="Quenstedt" baseAuthorityYear="1884" box="[382,809,403,427]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="vindobonensis">
|
||
<emphasis id="B949EAD5FF8DFFA7FE53FE386A1BFE6B" box="[382,617,403,427]" italics="true" pageId="15" pageNumber="16">Perrona vindobonensis</emphasis>
|
||
(Quenstedt, 1884)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8DFFA7FC19FE386BCDFE6B" box="[820,959,403,427]" pageId="15" pageNumber="16" type="paralectotype">paralectotype</typeStatus>
|
||
, NHMW 1997z0178/1467, Gainfarn (Austria),
|
||
<emphasis id="B949EAD5FF8DFFA7FFBAFE1C68DFFE0F" bold="true" box="[151,173,439,463]" pageId="15" pageNumber="16">Y.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FF9EFE1C69E9FE0F" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[179,411,439,463]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="wanzenboecki" status="nom. nov.">
|
||
<emphasis id="B949EAD5FF8DFFA7FF9EFE1C69E9FE0F" box="[179,411,439,463]" italics="true" pageId="15" pageNumber="16">Perrona wanzenboecki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8DFFA7FE8CFE1C6A7AFE0E" box="[417,520,439,462]" pageId="15" pageNumber="16">nov. nom.</taxonomicNameLabel>
|
||
, NHMW 2021/0158/0001,
|
||
<typeStatus id="54868865FF8DFFA7FC31FE1C6B1DFE0F" box="[796,879,439,463]" pageId="15" pageNumber="16" type="neotype">neotype</typeStatus>
|
||
, St. Veit an der Triesting (Austria).
|
||
<emphasis id="B949EAD5FF8DFFA7FBF9FE1C6C99FE0F" bold="true" box="[1236,1259,439,463]" pageId="15" pageNumber="16">Z.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FBDCFE1C6DEEFE0F" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[1265,1436,439,463]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="dulaii" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF8DFFA7FBDCFE1C6DEEFE0F" box="[1265,1436,439,463]" italics="true" pageId="15" pageNumber="16">Tomellana dulaii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8DFFA7FFBAFE70689BFE32" box="[151,233,475,498]" pageId="15" pageNumber="16" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, NHMW 2021/0148/0002,
|
||
<typeStatus id="54868865FF8DFFA7FD28FE706A12FE33" box="[517,608,475,499]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
, Nemeşeşti (Romania).
|
||
<emphasis id="B949EAD5FF8DFFA7FC74FE716B0DFE32" bold="true" box="[857,895,474,498]" pageId="15" pageNumber="16">Aa.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FCAAFE706C5CFE32" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[903,1070,475,499]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="aueri" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF8DFFA7FCAAFE706C5CFE32" box="[903,1070,475,499]" italics="true" pageId="15" pageNumber="16">Tomellana aueri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF8DFFA7FB1AFE706CFAFE32" box="[1079,1160,475,498]" pageId="15" pageNumber="16" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, NHMW 2013/0078/0436, Baden (Austria).
|
||
<emphasis id="B949EAD5FF8DFFA7FE6AFE55691CFDD6" bold="true" box="[327,366,510,534]" pageId="15" pageNumber="16">Ab.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FE59FE546BD1FDD7" authority="(Kovacs & Vician, 2021)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Kovacs & Vician" baseAuthorityYear="2021" box="[372,931,511,535]" class="Gastropoda" family="Clavatulidae" genus="Striopusionella" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="szekelyhidiae">
|
||
<emphasis id="B949EAD5FF8DFFA7FE59FE546AEEFDD7" box="[372,668,511,535]" italics="true" pageId="15" pageNumber="16">Striopusionella szekelyhidiae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8DFFA7FD84FE546BEEFDD7" author="Kovacs, Z. & Vician, Z." box="[681,924,511,535]" pageId="15" pageNumber="16" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián, 2021</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="54868865FF8DFFA7FC83FE546C7BFDD7" box="[942,1033,511,535]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
, PAL 2021.19.1, Letkés (Hungary).
|
||
<emphasis id="B949EAD5FF8DFFA7FA55FE556DEFFDD6" bold="true" box="[1400,1437,510,534]" pageId="15" pageNumber="16">Ac.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FFBAFD886AF2FDFB" authority="(Hoernes & Auinger, 1891)" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[151,640,547,571]" class="Gastropoda" family="Clavatulidae" genus="Scaevatula" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="sidoniae">
|
||
<emphasis id="B949EAD5FF8DFFA7FFBAFD886910FDFB" box="[151,354,547,571]" italics="true" pageId="15" pageNumber="16">Scaevatula sidoniae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF8DFFA7FE42FD886A0AFDFB" author="Hoernes, R. & Auinger, M." box="[367,632,547,571]" pageId="15" pageNumber="16" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. NHMW 1855/0043/0045,
|
||
<typeStatus id="54868865FF8DFFA7FCB9FD886B9DFDFB" box="[916,1007,547,571]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
, Lăpugiu de Sus (Romania). Scale bar is
|
||
<quantity id="4CC59B22FF8DFFA7FABDFD8968CCFD9E" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="mm" value="5.0">5 mm</quantity>
|
||
for all specimens except for
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FEC8FDEC6AEEFD9F" authorityName="NHMW" authorityYear="1949" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[485,668,583,607]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="barbarae">
|
||
<emphasis id="B949EAD5FF8DFFA7FEC8FDEC6AEEFD9F" box="[485,668,583,607]" italics="true" pageId="15" pageNumber="16">Perrona barbarae</emphasis>
|
||
</taxonomicName>
|
||
and protoconch of
|
||
<emphasis id="B949EAD5FF8DFFA7FC4FFDEC6C46FD9F" box="[866,1076,583,607]" italics="true" pageId="15" pageNumber="16">
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FC4FFDEC6C42FD9F" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[866,1072,583,607]" class="Gastropoda" family="Clavatulidae" genus="Scaevatula" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="sidoniae">Scaevatula sidoniae</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF42664FFF8DFFA7FFBAF8646B5FF827" ID-DOI="http://doi.org/10.5281/zenodo.6399839" ID-Zenodo-Dep="6399839" box="[151,813,1999,2023]" httpUri="https://zenodo.org/record/6399839/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1999,2023]" targetBox="[151,1436,631,1976]" targetPageId="15">
|
||
<paragraph id="8B8236C7FF8DFFA7FFBAF8646B5FF827" blockId="15.[151,813,1999,2023]" box="[151,813,1999,2023]" pageId="15" pageNumber="16">
|
||
<emphasis id="B949EAD5FF8DFFA7FFBAF8646961F827" bold="true" box="[151,275,1999,2023]" pageId="15" pageNumber="16">FIGURE 5.</emphasis>
|
||
Anal sinus morphology of Paratethyan
|
||
<taxonomicName id="4C3D4D44FF8DFFA7FD8BF8646B5BF827" authorityName="Gray" authorityYear="1853" box="[678,809,1999,2023]" class="Gastropoda" family="Clavatulidae" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="family">Clavatulidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8236C7FF92FFB8FFEAFF3C699DFE81" blockId="16.[151,1436,151,321]" pageId="16" pageNumber="17">
|
||
Clavatulids often have rows of beads, tubercles, and spines. In this work beads are defined as small round, spirally-aligned structures in apposition to each other. Tubercles are larger, not in contact with each other and can be rounded, subtrigonal or subquadrate in outline and low or pointed, if sharp here called spinous. Spines are also present in some species, which is when the pointed structure, usually on the subsutural collar, is open on the abapical side (e.g.,
|
||
<taxonomicName id="4C3D4D44FF92FFB8FE26FE8C69ACFE81" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[267,478,295,321]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="eleonorae">
|
||
<emphasis id="B949EAD5FF92FFB8FE26FE8C69ACFE81" box="[267,478,295,321]" italics="true" pageId="16" pageNumber="17">Perrona eleonorae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF42664FFF92FFB8FFBAF83969BEF80F" ID-DOI="http://doi.org/10.5281/zenodo.6399841" ID-Zenodo-Dep="6399841" httpUri="https://zenodo.org/record/6399841/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1938,1962]" targetBox="[151,1436,345,1915]" targetPageId="16">
|
||
<paragraph id="8B8236C7FF92FFB8FFBAF83969BEF80F" blockId="16.[151,1436,1938,1999]" pageId="16" pageNumber="17">
|
||
<emphasis id="B949EAD5FF92FFB8FFBAF839696BF86A" bold="true" box="[151,281,1938,1962]" pageId="16" pageNumber="17">FIGURE 6.</emphasis>
|
||
Apertures of Paratethyan
|
||
<taxonomicName id="4C3D4D44FF92FFB8FD13F8396ACCF86A" authorityName="Gray" authorityYear="1853" box="[574,702,1938,1962]" class="Gastropoda" family="Clavatulidae" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="17" phylum="Mollusca" rank="family">Clavatulidae</taxonomicName>
|
||
(
|
||
<emphasis id="B949EAD5FF92FFB8FDFEF8386D0AF86A" box="[723,1400,1938,1962]" italics="true" pageId="16" pageNumber="17">
|
||
C. =
|
||
<taxonomicName id="4C3D4D44FF92FFB8FC38F8396B09F86A" authorityName="Lamarck" authorityYear="1801" box="[789,891,1938,1962]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">Clavatula</taxonomicName>
|
||
, ‘C.’ = ‘Clavatula’, G. =
|
||
<taxonomicName id="4C3D4D44FF92FFB8FB85F8396D20F86A" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[1192,1362,1938,1962]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">Granulatocincta</taxonomicName>
|
||
, M
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B949EAD5FF92FFB8FAA6F8386944F80E" italics="true" pageId="16" pageNumber="17">= Megaclavaltula</emphasis>
|
||
,
|
||
<emphasis id="B949EAD5FF92FFB8FE6EF81C69CCF80E" box="[323,446,1974,1998]" italics="true" pageId="16" pageNumber="17">
|
||
O. =
|
||
<taxonomicName id="4C3D4D44FF92FFB8FE54F81D69CCF80E" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[377,446,1974,1998]" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Olegia</taxonomicName>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8236C7FF93FFB9FFEAFF3C6BD6FE81" blockId="17.[151,1436,151,322]" pageId="17" pageNumber="18">
|
||
The aperture is pyriform to ovate in clavatulids, the outer lip does not possess a varix, and is usually smooth within (
|
||
<figureCitation id="13062A42FF93FFB9FFC6FF10693DFF16" box="[235,335,187,214]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="16.[151,250,1938,1962]" captionStartId-1="17.[151,250,1938,1962]" captionTargetBox-0="[151,1436,345,1915]" captionTargetBox-1="[151,1436,345,1915]" captionTargetId-0="figure-106@16.[151,1436,345,1915]" captionTargetId-1="figure-109@17.[151,1436,345,1915]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURE 6. Apertures of Paratethyan Clavatulidae (C. = Clavatula, ‘C.’ = ‘Clavatula’, G. = Granulatocincta, M. = Megaclavaltula, O. = Olegia)." captionText-1="FIGURE 7. Apertures of Paratethyan Clavatulidae (O. = Olegia, ‘C.’ = ‘Clavatula’, N. = Neoperrona, P. = Perrona, Pu. = Pusionella, Sc. = Scaevatula, S. = Striopusionella, T. = Tomellana)." figureDoi-0="http://doi.org/10.5281/zenodo.6399841" figureDoi-1="http://doi.org/10.5281/zenodo.6399845" httpUri-0="https://zenodo.org/record/6399841/files/figure.png" httpUri-1="https://zenodo.org/record/6399845/files/figure.png" pageId="17" pageNumber="18">Figs 6–7</figureCitation>
|
||
). Species with lirae within the outer lip do occur (e.g.,
|
||
<taxonomicName id="4C3D4D44FF93FFB9FCB3FF106D7DFF15" authority="granulatocincta" box="[926,1295,187,213]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="granulatocincta">
|
||
<emphasis id="B949EAD5FF93FFB9FCB3FF106D7DFF15" box="[926,1295,187,213]" italics="true" pageId="17" pageNumber="18">Granulatocincta granulatocincta</emphasis>
|
||
</taxonomicName>
|
||
, ‘
|
||
<taxonomicName id="4C3D4D44FF93FFB9FA09FF106DE0FF15" authorityName="Lamarck" authorityYear="1801" box="[1316,1426,187,213]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF93FFB9FA09FF106DE0FF15" box="[1316,1426,187,213]" italics="true" pageId="17" pageNumber="18">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B949EAD5FF93FFB9FFBAFF7468A5FF38" box="[151,215,223,248]" italics="true" pageId="17" pageNumber="18">irisae</emphasis>
|
||
, ‘
|
||
<taxonomicName id="4C3D4D44FF93FFB9FFDDFF74692CFF39" authorityName="Lamarck" authorityYear="1801" box="[240,350,223,249]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF93FFB9FFDDFF74692CFF39" box="[240,350,223,249]" italics="true" pageId="17" pageNumber="18">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF93FFB9FE5CFF7469BAFF39" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[369,456,223,249]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="sophiae">
|
||
<emphasis id="B949EAD5FF93FFB9FE5CFF7469BAFF39" box="[369,456,223,249]" italics="true" pageId="17" pageNumber="18">sophiae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B949EAD5FF93FFB9FEF5FF746A30FF39" box="[472,578,223,249]" italics="true" pageId="17" pageNumber="18">inter alia</emphasis>
|
||
), and if present are considered a species-specific character in most species (‘
|
||
<taxonomicName id="4C3D4D44FF93FFB9FF87FEA8696AFEDD" authorityName="Lamarck" authorityYear="1801" box="[170,280,259,285]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF93FFB9FF87FEA8696AFEDD" box="[170,280,259,285]" italics="true" pageId="17" pageNumber="18">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF93FFB9FE07FEA869F2FEDC" baseAuthorityName="Defrance" baseAuthorityYear="1826" box="[298,384,259,284]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="romana">
|
||
<emphasis id="B949EAD5FF93FFB9FE07FEA869F2FEDC" box="[298,384,259,284]" italics="true" pageId="17" pageNumber="18">romana</emphasis>
|
||
</taxonomicName>
|
||
is an exception). The siphonal canal is open in all species and usually weakly to moderately notched (exceptions exist). The length of the canal is species specific.
|
||
</paragraph>
|
||
<caption id="DF42664FFF93FFB9FFBAF8396B3BF80F" ID-DOI="http://doi.org/10.5281/zenodo.6399845" ID-Zenodo-Dep="6399845" httpUri="https://zenodo.org/record/6399845/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1938,1962]" targetBox="[151,1436,345,1915]" targetPageId="17">
|
||
<paragraph id="8B8236C7FF93FFB9FFBAF8396B3BF80F" blockId="17.[151,1436,1938,1999]" pageId="17" pageNumber="18">
|
||
<emphasis id="B949EAD5FF93FFB9FFBAF8396967F86B" bold="true" box="[151,277,1938,1963]" pageId="17" pageNumber="18">FIGURE 7.</emphasis>
|
||
Apertures of Paratethyan
|
||
<taxonomicName id="4C3D4D44FF93FFB9FD08F8396AD7F86A" authorityName="Gray" authorityYear="1853" box="[549,677,1938,1962]" class="Gastropoda" family="Clavatulidae" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="family">Clavatulidae</taxonomicName>
|
||
(
|
||
<emphasis id="B949EAD5FF93FFB9FD9BF8386B47F86A" box="[694,821,1938,1962]" italics="true" pageId="17" pageNumber="18">
|
||
O. =
|
||
<taxonomicName id="4C3D4D44FF93FFB9FDDDF8396B47F86A" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[752,821,1938,1962]" class="Insecta" family="Phylloxeridae" genus="Olegia" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Olegia</taxonomicName>
|
||
</emphasis>
|
||
, ‘
|
||
<emphasis id="B949EAD5FF93FFB9FC66F8386B2EF86A" box="[843,860,1939,1962]" italics="true" pageId="17" pageNumber="18">C</emphasis>
|
||
.’ = ‘
|
||
<taxonomicName id="4C3D4D44FF93FFB9FCBDF8396B86F86A" authorityName="Lamarck" authorityYear="1801" box="[912,1012,1938,1962]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF93FFB9FCBDF8396B86F86A" box="[912,1012,1938,1962]" italics="true" pageId="17" pageNumber="18">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’,
|
||
<emphasis id="B949EAD5FF93FFB9FB26F8386CCEF86A" box="[1035,1212,1939,1962]" italics="true" pageId="17" pageNumber="18">
|
||
N. =
|
||
<taxonomicName id="4C3D4D44FF93FFB9FB6EF8386CCEF86A" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[1091,1212,1939,1962]" class="Gastropoda" family="Clavatulidae" genus="Neoperrona" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">Neoperrona</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B949EAD5FF93FFB9FBE6F8386B49F80E" italics="true" pageId="17" pageNumber="18">
|
||
P. =
|
||
<taxonomicName id="4C3D4D44FF93FFB9FBD2F8386DF6F86A" authority=", Pu." authorityName="Pu." box="[1279,1412,1939,1962]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">Perrona, Pu.</taxonomicName>
|
||
=
|
||
<taxonomicName id="4C3D4D44FF93FFB9FFBAF81D695DF80E" authority=", Sc." authorityName="Sc." box="[151,303,1974,1998]" class="Gastropoda" family="Clavatulidae" genus="Pusionella" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">Pusionella, Sc.</taxonomicName>
|
||
=
|
||
<taxonomicName id="4C3D4D44FF93FFB9FE61F81D69CCF80E" authorityName="Gofas" authorityYear="1990" box="[332,446,1974,1998]" class="Gastropoda" family="Clavatulidae" genus="Scaevatula" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">Scaevatula</taxonomicName>
|
||
, S. =
|
||
<taxonomicName id="4C3D4D44FF93FFB9FED4F81D6AE4F80E" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[505,662,1974,1998]" class="Gastropoda" family="Clavatulidae" genus="Striopusionella" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">Striopusionella</taxonomicName>
|
||
, T. =
|
||
<taxonomicName id="4C3D4D44FF93FFB9FDFCF81D6B49F80E" authorityName="Wenz" authorityYear="1943" box="[721,827,1974,1998]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">Tomellana</taxonomicName>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8236C7FF90FFBAFFEAFF3C6B63FEDD" blockId="18.[151,1436,151,321]" pageId="18" pageNumber="19">The columella is more or less deeply excavated in the upper third or upper half, more or less twisted at the fasciole, and smooth in all species. The columellar callus is weakly to moderately developed, at most expanded onto the medial side of the venter, but never greatly thickened. The parietal portion of the callus is never thicker than the abapical portion and never develops a callus pad or fold.</paragraph>
|
||
<paragraph id="8B8236C7FF90FFBAFFEAFE8C6C15FE81" blockId="18.[151,1436,151,321]" box="[199,1127,295,321]" pageId="18" pageNumber="19">No color pattern is present in the Paratethyan specimens, nor is it enhanced by UV.</paragraph>
|
||
<paragraph id="8B8236C7FF90FFBAFFBAFE3869F6FE6C" blockId="18.[151,388,402,429]" box="[151,388,402,429]" pageId="18" pageNumber="19">
|
||
<emphasis id="B949EAD5FF90FFBAFFBAFE3869F6FE6C" bold="true" box="[151,388,402,429]" pageId="18" pageNumber="19">
|
||
What is
|
||
<taxonomicName id="4C3D4D44FF90FFBAFFD2FE396906FE6D" authorityName="Lamarck" authorityYear="1801" box="[255,372,402,429]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFFD2FE396906FE6D" bold="true" box="[255,372,402,429]" italics="true" pageId="18" pageNumber="19">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
?
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF90FFBAFFBAFE706A35FD45" blockId="18.[151,1437,475,1977]" pageId="18" pageNumber="19">
|
||
MolluscaBase (Eds.) (2021a) lists 37 species of
|
||
<taxonomicName id="4C3D4D44FF90FFBAFDEFFE706B42FE35" authorityName="Lamarck" authorityYear="1801" box="[706,816,475,501]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFDEFFE706B42FE35" box="[706,816,475,501]" italics="true" pageId="18" pageNumber="19">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
, including also
|
||
<taxonomicName id="4C3D4D44FF90FFBAFCC4FE7068A0FDD9" authority="Nolf & Verstraeten, 2006" authorityName="Nolf & Verstraeten" authorityYear="2006" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="xanteni">
|
||
<emphasis id="B949EAD5FF90FFBAFCC4FE706CC1FE34" box="[1001,1203,475,501]" italics="true" pageId="18" pageNumber="19">Clavatula xanteni</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFB91FE7068A0FDD9" author="Nolf, F. & Verstraeten, J." pageId="18" pageNumber="19" pagination="15 - 29" refId="ref140333" refString="Nolf, F. & Verstraeten, J. (2006) Recognition of two new Clavatula species (Mollusca: Gastropoda: Conoidea: Turridae) in a complex group from Gabon and north Angola. Neptunea, 5, 15 - 29. [https: // neptuneaorg. files. wordpress. com / 2019 / 02 / np 5 - 3 - tekst. pdf]" type="journal article" year="2006">Nolf & Verstraeten, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
, which was placed in
|
||
<taxonomicName id="4C3D4D44FF90FFBAFEE5FE546A56FDD8" box="[456,548,511,536]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFEE5FE546A56FDD8" box="[456,548,511,536]" italics="true" pageId="18" pageNumber="19">Perrona</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFD63FE546B4EFDD9" author="Kantor, Y. I. & Horro, J. & Rolan, E. & Puillandre, N." box="[590,828,511,537]" pageId="18" pageNumber="19" pagination="275 - 284" refId="ref136224" refString="Kantor, Y. I., Horro, J., Rolan, E. & Puillandre, N. (2018 a) Paraclavatula (Gastropoda: Conoidea: Clavatulidae), a new genus with a distinctive radula type from West Africa. Journal of Molluscan Studies, 84, 275 - 284. https: // doi. org / 10.1093 / mollus / eyy 012" type="journal article" year="2018">
|
||
Kantor
|
||
<emphasis id="B949EAD5FF90FFBAFD8FFE546AA6FDD9" box="[674,724,511,537]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
|
||
. (2018a)
|
||
</bibRefCitation>
|
||
.
|
||
<taxonomicName id="4C3D4D44FF90FFBAFC6BFE546C95FDD9" authority="(Kiener, 1839)" baseAuthorityName="Kiener" baseAuthorityYear="1839" box="[838,1255,511,537]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="pyramidata">
|
||
<emphasis id="B949EAD5FF90FFBAFC6BFE546C4FFDD9" box="[838,1085,511,537]" italics="true" pageId="18" pageNumber="19">Clavatula pyramidata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFB66FE546CADFDD9" author="Kiener, L. C." box="[1099,1247,511,537]" pageId="18" pageNumber="19" refId="ref136494" refString="Kiener, L. C. (1839) Species general et iconographie des coquilles vivantes. Vol. 5. Famille des Canaliferes. Premiere partie. Pleurotome (Pleurotoma), Lamarck. Rousseau & J. B. Bailliere, Paris, 16 pp., 27 pls. [http: // www. biodiversitylibrary. org / item / 19762]" type="book" year="1839">Kiener, 1839</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
is missing from that list, although
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFE70FD886A36FDFD" author="Kantor, Y. I. & Horro, J. & Rolan, E. & Puillandre, N." box="[349,580,547,573]" pageId="18" pageNumber="19" pagination="275 - 284" refId="ref136224" refString="Kantor, Y. I., Horro, J., Rolan, E. & Puillandre, N. (2018 a) Paraclavatula (Gastropoda: Conoidea: Clavatulidae), a new genus with a distinctive radula type from West Africa. Journal of Molluscan Studies, 84, 275 - 284. https: // doi. org / 10.1093 / mollus / eyy 012" type="journal article" year="2018">
|
||
Kantor
|
||
<emphasis id="B949EAD5FF90FFBAFE9CFD886993FDFD" box="[433,481,547,573]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
|
||
. (2018a)
|
||
</bibRefCitation>
|
||
placed that species in
|
||
<taxonomicName id="4C3D4D44FF90FFBAFC17FD886BDAFDFD" authorityName="Lamarck" authorityYear="1801" box="[826,936,547,573]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFC17FD886BDAFDFD" box="[826,936,547,573]" italics="true" pageId="18" pageNumber="19">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
based on molecular data.
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFBEAFD886D1CFDFD" author="Kilburn, R. N." box="[1223,1390,546,573]" pageId="18" pageNumber="19" pagination="417 - 470" refId="ref136560" refString="Kilburn, R. N. (1985) Turridae (Mollusca: Gastropoda) of southern Africa and Mozambique. Part 2. Subfamily Clavatulinae. Annals of the Natal Museum, 26, 417 - 470. [https: // journals. co. za / doi / abs / 10.10520 / AJA 03040798 _ 469]" type="journal article" year="1985">Kilburn (1985)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFFBAFDEC69BCFDA1" author="Boyer, F. & Hernandez, J. M." box="[151,462,583,609]" pageId="18" pageNumber="19" pagination="77 - 84" refId="ref130008" refString="Boyer, F. & Hernandez, J. M. (2004) Variability and distribution of Clavatula mystica (Reeve, 1843). Iberus, 22 (1), 77 - 84. [https: // zenodo. org / record / 4516361]" type="journal article" year="2004">Boyer & Hernández (2004)</bibRefCitation>
|
||
stated that
|
||
<taxonomicName id="4C3D4D44FF90FFBAFD7EFDEC6AB3FDA1" authorityName="Lamarck" authorityYear="1801" box="[595,705,583,609]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFD7EFDEC6AB3FDA1" box="[595,705,583,609]" italics="true" pageId="18" pageNumber="19">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
is a poorly characterized genus, lacking well-defined diagnostic features, but did not provide solutions.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF90FFBAFFEAFD246D70FC09" blockId="18.[151,1437,475,1977]" pageId="18" pageNumber="19">
|
||
New unpublished molecular data confirmed
|
||
<emphasis id="B949EAD5FF90FFBAFDE9FD246BD7FD69" box="[708,933,655,681]" italics="true" pageId="18" pageNumber="19">
|
||
<taxonomicName id="4C3D4D44FF90FFBAFDE9FD246B51FD68" box="[708,803,655,680]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">Perrona</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D4D44FF90FFBAFC1DFD246BD7FD69" authorityName="Wenz" authorityYear="1943" box="[816,933,655,681]" class="Gastropoda" family="Clavatulidae" genus="Tomellana" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">Tomellana</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C3D4D44FF90FFBAFCCDFD246C29FD69" authorityName="Gofas" authorityYear="1990" box="[992,1115,655,681]" class="Gastropoda" family="Clavatulidae" genus="Scaevatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFCCDFD246C29FD69" box="[992,1115,655,681]" italics="true" pageId="18" pageNumber="19">Scaevatula</emphasis>
|
||
</taxonomicName>
|
||
as genus-level taxa, but are in conflict with
|
||
<taxonomicName id="4C3D4D44FF90FFBAFE7DFD1869CCFD0D" authorityName="Lamarck" authorityYear="1801" box="[336,446,691,717]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFE7DFD1869CCFD0D" box="[336,446,691,717]" italics="true" pageId="18" pageNumber="19">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
as currently perceived (Nicolas Puillandre, pers. comm.,
|
||
<date id="FF831007FF90FFBAFB7EFD186C80FD0D" box="[1107,1266,691,717]" pageId="18" pageNumber="19" value="2021-10">October 2021</date>
|
||
). These results suggest that
|
||
<taxonomicName id="4C3D4D44FF90FFBAFE0AFD7C69E7FD31" authorityName="Lamarck" authorityYear="1801" box="[295,405,727,753]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFE0AFD7C69E7FD31" box="[295,405,727,753]" italics="true" pageId="18" pageNumber="19">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
is not monophyletic, which is also suggested by the morphologies of the early teleoconch whorls. We observe peculiar morphologies in certain species groups (
|
||
<figureCitation id="13062A42FF90FFBAFCBFFD506C7DFCD5" box="[914,1039,763,789]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="18" pageNumber="19">Figs 3A–E</figureCitation>
|
||
), which we discuss herein as clades.
|
||
<taxonomicName id="4C3D4D44FF90FFBAFFBAFCB46A4AFCF9" authority="(Lamarck, 1822)" authorityName="" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[151,568,799,825]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="muricata">
|
||
<emphasis id="B949EAD5FF90FFBAFFBAFCB46901FCF8" box="[151,371,799,825]" italics="true" pageId="18" pageNumber="19">Clavatula muricata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFEAFFCB46A42FCF9" author="Lamarck, J. - B. M. de" box="[386,560,799,825]" pageId="18" pageNumber="19" refId="ref137485" refString="Lamarck, J. - B. M. de. (1822) Histoire naturelle des animaux sans vertebres. Tome Septieme. Published by the Author, Paris, 711 pp. [http: // www. biodiversitylibrary. org / item / 47433]" type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(probably a synonym of
|
||
<taxonomicName id="4C3D4D44FF90FFBAFC7AFCB46BC1FCF8" baseAuthorityName="Roding" baseAuthorityYear="1798" box="[855,947,799,825]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="regia">
|
||
<emphasis id="B949EAD5FF90FFBAFC7AFCB46BC1FCF8" box="[855,947,799,825]" italics="true" pageId="18" pageNumber="19">C. regia</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="4C3D4D44FF90FFBAFCD8FCB46D77FCF9" authority="(Roding, 1798)" baseAuthorityName="Roding" baseAuthorityYear="1798" box="[1013,1285,799,825]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="regia">
|
||
<emphasis id="B949EAD5FF90FFBAFCD8FCB46C23FCF8" box="[1013,1105,799,825]" italics="true" pageId="18" pageNumber="19">C. regia</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFB4DFCB46C8FFCF9" author="Roding, P. F." box="[1120,1277,799,825]" pageId="18" pageNumber="19" refId="ref142481" refString="Roding, P. F. (1798). Museum Boltenianum sive Catalogus cimeliorum e tribus regnis naturae quae olim collegerat Joa. Fried Bolten, M. D. p. d. per XL. annos proto physicus Hamburgensis. Pars secunda continens Conchylia sive Testacea univalvia, bivalvia & multivalvia. Trapp, Hamburg, viii + 199 pp. [https: // www. biodiversitylibrary. org / page / 16230659]" type="book" year="1798">Röding, 1798</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13062A42FF90FFBAFA38FCB46DE6FCF9" box="[1301,1428,799,825]" captionStart="FIGURE 8" captionStartId="21.[151,250,1582,1606]" captionTargetBox="[151,1436,181,1558]" captionTargetId="figure-18@21.[151,1436,181,1558]" captionTargetPageId="21" captionText="FIGURE 8. Extant Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa; B. Clavatula regia (Röding, 1798), MNHN-IM-2009-24970, Ile de Principe (São Tomé et Principe). C. Clavatula regia (Röding, 1798) (labelled as Clavatula muricata (Lamarck, 1822)), MNHN-IM-2009-24979, Baie de Pointe-Noire (Republique du Congo). D. Clavatula regia (Röding, 1798), Gonçalo Rosa, Atoll Specimen Shells (Senegal). E. Clavatula regia (Röding, 1798) (NHMD- 230855), paralectotype, Western Africa. F. Clavatula regia (Röding, 1798), NHMD-916083, lectotype, Western Africa. G. Clavatula regia (Röding, 1798), NHMD-230855, paralectotype, Western Africa; labels of the type material from the Natural History Museum of Denmark. H. Clavatula regia (Röding, 1798), NHMD-90802, paralectotype, Guinea. Specimens B and C have been used for molecular analysis by Nicolas Puillandre (unpublished data)." figureDoi="http://doi.org/10.5281/zenodo.6399847" httpUri="https://zenodo.org/record/6399847/files/figure.png" pageId="18" pageNumber="19">Figs 8A–H</figureCitation>
|
||
) have a bipartite sculpture with a weak, smooth subsutural cord and a prominent suprasutural cord of large beads (overrun by weak spiral cords) (
|
||
<figureCitation id="13062A42FF90FFBAFD27FCCC6A15FC41" box="[522,615,871,897]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="18" pageNumber="19">Fig. 3A</figureCitation>
|
||
). The same early teleoconch sculpture is developed by
|
||
<taxonomicName id="4C3D4D44FF90FFBAFBCAFCCC6947FC65" authority="(Reeve, 1845)" baseAuthorityName="Reeve" baseAuthorityYear="1845" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="rubrifasciata">
|
||
<emphasis id="B949EAD5FF90FFBAFBCAFCCC6DEEFC41" box="[1255,1436,871,897]" italics="true" pageId="18" pageNumber="19">C. rubrifasciata</emphasis>
|
||
(Reeve, 1845)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13062A42FF90FFBAFE6EFC2069ECFC65" box="[323,414,907,933]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="18" pageNumber="19">Figs 3B</figureCitation>
|
||
,
|
||
<figureCitation id="13062A42FF90FFBAFE85FC20699FFC65" box="[424,493,907,933]" captionStart="FIGURE 9" captionStartId="22.[151,250,1808,1832]" captionTargetBox="[151,1436,777,1785]" captionTargetId="figure-260@22.[151,1436,777,1785]" captionTargetPageId="22" captionText="FIGURE 9. Extant Clavatulidae species. A. Clavatula rubrifasciata (Reeve, 1845), Gonçalo Rosa, Atoll Specimen Shells (Senegal). B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), Gonçalo Rosa, Atoll Specimen Shells (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ bimarginata (Lamarck, 1822), NHMW 2042, Gorée (Senegal).G. ‘Clavatula’ diadema (Kiener,1839), NHMW 1884/0010/0023, Gorée (Senegal). Specimens C and E have been used for molecular analysis by Nicolas Puillandre (unpublished data)." figureDoi="http://doi.org/10.5281/zenodo.6399849" httpUri="https://zenodo.org/record/6399849/files/figure.png" pageId="18" pageNumber="19">9A–B</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C3D4D44FF90FFBAFD04FC206B37FC65" authority="(Recluz, 1851)" baseAuthorityName="Recluz" baseAuthorityYear="1851" box="[553,837,906,933]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="lelieuri">
|
||
<emphasis id="B949EAD5FF90FFBAFD04FC206AE8FC65" box="[553,666,907,933]" italics="true" pageId="18" pageNumber="19">C. lelieuri</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFD85FC206B4FFC65" author="Recluz, C." box="[680,829,906,933]" pageId="18" pageNumber="19" pagination="194 - 216" refId="ref142286" refString="Recluz, C. (1851) Description de quelques coquilles nouvelles. Journal de Conchyliologie, 2, 194 - 216, pls. 5 - 6. [https: // www. biodiversitylibrary. org / item / 53852 # page / 200 / mode / 1 up]" type="journal article" year="1851">Récluz, 1851</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(see Nolf 2008: pl. 8, figs 42–49, pl. 9). The
|
||
<emphasis id="B949EAD5FF90FFBAFA1AFC206899FC08" italics="true" pageId="18" pageNumber="19">
|
||
<taxonomicName id="4C3D4D44FF90FFBAFA1AFC206DEEFC64" authorityName="" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[1335,1436,907,932]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="muricata">muricata</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C3D4D44FF90FFBAFF9CFC046899FC08" baseAuthorityName="Roding" baseAuthorityYear="1798" box="[177,235,943,968]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="regia">regia</taxonomicName>
|
||
</emphasis>
|
||
clade contains the
|
||
<typeStatus id="54868865FF90FFBAFEEFFC046980FC09" box="[450,498,943,969]" pageId="18" pageNumber="19">type</typeStatus>
|
||
species and therefore, these species represent
|
||
<emphasis id="B949EAD5FF90FFBAFCD4FC046C8EFC08" box="[1017,1276,943,969]" italics="true" pageId="18" pageNumber="19">
|
||
<taxonomicName id="4C3D4D44FF90FFBAFCD4FC046C15FC09" authorityName="Lamarck" authorityYear="1801" box="[1017,1127,943,969]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus" sensu="stricto">Clavatula</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF90FFBAFB42FC046C8EFC08" box="[1135,1276,943,968]" pageId="18" pageNumber="19" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF90FFBAFFEAFC786A8AFB05" blockId="18.[151,1437,475,1977]" pageId="18" pageNumber="19">
|
||
A close relation of ‘
|
||
<taxonomicName id="4C3D4D44FF90FFBAFE88FC786A61FC2D" authorityName="Lamarck" authorityYear="1801" box="[421,531,979,1005]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFE88FC786A61FC2D" box="[421,531,979,1005]" italics="true" pageId="18" pageNumber="19">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B949EAD5FF90FFBAFD0CFC786AF1FC2D" box="[545,643,979,1005]" italics="true" pageId="18" pageNumber="19">caerulea</emphasis>
|
||
(Weinkauff in
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFC06FC786C25FC2D" author="Weinkauff, H. C. & Kobelt, W." box="[811,1111,978,1005]" pageId="18" pageNumber="19" refId="ref146384" refString="Weinkauff, H. C. & Kobelt, W. (1875 - 1887) Die Familie Pleurotomidae. Systematisches Conchylien-Cabinet von Martini und Chemnitz. Vol. 4. Bauer & Raspe, Nurnberg, 248 pp., pls. A + 1 - 42. [https: // www. biodiversitylibrary. org / item / 217078 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 124643" type="book" year="1875">Weinkauff & Kobelt, 1875</bibRefCitation>
|
||
) and ‘
|
||
<taxonomicName id="4C3D4D44FF90FFBAFBB2FC786D7FFC2D" authorityName="Lamarck" authorityYear="1801" box="[1183,1293,979,1005]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFBB2FC786D7FFC2D" box="[1183,1293,979,1005]" italics="true" pageId="18" pageNumber="19">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF90FFBAFA36FC78694BFBD1" authority="(Kiener, 1839)" baseAuthorityName="Kiener" baseAuthorityYear="1839" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="pyramidata">
|
||
<emphasis id="B949EAD5FF90FFBAFA36FC786DEEFC2D" box="[1307,1436,979,1005]" italics="true" pageId="18" pageNumber="19">pyramidata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFFB2FC5C6943FBD1" author="Kiener, L. C." box="[159,305,1015,1041]" pageId="18" pageNumber="19" refId="ref136494" refString="Kiener, L. C. (1839) Species general et iconographie des coquilles vivantes. Vol. 5. Famille des Canaliferes. Premiere partie. Pleurotome (Pleurotoma), Lamarck. Rousseau & J. B. Bailliere, Paris, 16 pp., 27 pls. [http: // www. biodiversitylibrary. org / item / 19762]" type="book" year="1839">Kiener, 1839</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13062A42FF90FFBAFE6AFC5C69EFFBD1" box="[327,413,1015,1041]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="18" pageNumber="19">Figs 3F</figureCitation>
|
||
) with
|
||
<taxonomicName id="4C3D4D44FF90FFBAFECCFC5C6A3DFBD1" authorityName="Lamarck" authorityYear="1801" box="[481,591,1015,1041]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFECCFC5C6A3DFBD1" box="[481,591,1015,1041]" italics="true" pageId="18" pageNumber="19">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
is suggested by the early teleoconch sculpture with a prominent suprasutural cord of large beads. Both species, however, differ from typical
|
||
<taxonomicName id="4C3D4D44FF90FFBAFC4EFBB06BA3FBF5" authorityName="Lamarck" authorityYear="1801" box="[867,977,1051,1077]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFC4EFBB06BA3FBF5" box="[867,977,1051,1077]" italics="true" pageId="18" pageNumber="19">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
in their slender fusiform shell with high conical spire, the comparatively short last whorl, and the wider and shorter siphonal canal (
|
||
<figureCitation id="13062A42FF90FFBAFBB4FB946D56FB99" box="[1177,1316,1087,1113]" captionStart="FIGURE 10" captionStartId="23.[151,250,1355,1379]" captionTargetBox="[151,1436,181,1331]" captionTargetId="figure-18@23.[151,1436,181,1331]" captionTargetPageId="23" captionText="FIGURE 10. Extant Clavatulidae species. A. ‘Clavatula’ caerulea (Weinkauff in Weinkauff & Kobelt, 1875), MNHN-IM- 2009-19153, Libreville (Gabon). B. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). C. ‘Clavatula’ tripartita (Weinkauff, 1876), MNHN-IM-2009-14826, Cap Sainte Marie (Madagascar). D. ‘Clavatula’ tripartita (Weinkauff, 1876), MNHN-IM-2009-14985, Cap Sainte Marie (Madagascar). E. ‘Clavatula’ taxea (Röding, 1798), MNHN-IM-2009-14901, Fort-Dauphin (Madagascar). F. Pusionella aculeiformis (Lamarck, 1816), NHMW3051, Senegal. G. Pusionella buccinata (Lamarck, 1822); NHMW3052. Specimens A–E have been used for molecular analysis by Nicolas Puillandre (unpublished data)." figureDoi="http://doi.org/10.5281/zenodo.6399853" httpUri="https://zenodo.org/record/6399853/files/figure.png" pageId="18" pageNumber="19">Figs 10A, B</figureCitation>
|
||
) (see
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFA44FB9469FBFBBD" author="Nolf, F. & Verstraeten, J." pageId="18" pageNumber="19" pagination="17 - 29" refId="ref140403" refString="Nolf, F. & Verstraeten, J. (2008) Two new turrid species from Congo-Brazzaville: Clavatula hattenbergeri and Clavatula congoensis (Mollusca: Gastropoda: Clavatulidae). Neptunea, 7 / 1, 17 - 29. [https: // neptuneaorg. files. wordpress. com / 2019 / 02 / np 7 - 1 - tekst. pdf]" type="journal article" year="2008">Nolf and Verstraeten 2008</bibRefCitation>
|
||
: pls 1–2 for ‘
|
||
<emphasis id="B949EAD5FF90FFBAFD0FFBC86A4EFBBD" box="[546,572,1123,1149]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
|
||
’
|
||
<emphasis id="B949EAD5FF90FFBAFD66FBC86ADFFBBD" box="[587,685,1123,1149]" italics="true" pageId="18" pageNumber="19">caerulea</emphasis>
|
||
). ‘
|
||
<taxonomicName id="4C3D4D44FF90FFBAFDE2FBC86B4FFBBD" authorityName="Lamarck" authorityYear="1801" box="[719,829,1123,1149]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFDE2FBC86B4FFBBD" box="[719,829,1123,1149]" italics="true" pageId="18" pageNumber="19">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF90FFBAFC62FBC86CFCFBBD" authority="von Maltzan, 1883" authorityName="von Maltzan" authorityYear="1883" box="[847,1166,1123,1149]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="martensi">
|
||
<emphasis id="B949EAD5FF90FFBAFC62FBC86BC3FBBC" box="[847,945,1123,1148]" italics="true" pageId="18" pageNumber="19">martensi</emphasis>
|
||
von
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFCC6FBC86CFCFBBD" author="Maltzan, H. F. von" box="[1003,1166,1123,1149]" pageId="18" pageNumber="19" pagination="115 - 135" refId="ref138138" refString="Maltzan, H. F. von (1883) Beitrage zur Kenntnis der senegambischen Pleurotomiden. Jahrbucher der Deutschen Malakozoologischen Gesellschaft, 10, 115 - 135. [https: // www. biodiversitylibrary. org / item / 86735 # page / 131 / mode / 1 up]" type="journal article" year="1883">Maltzan, 1883</bibRefCitation>
|
||
</taxonomicName>
|
||
and ‘
|
||
<emphasis id="B949EAD5FF90FFBAFBFDFBC86C9BFBBD" box="[1232,1257,1123,1149]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF90FFBAFBD4FBC869FAFB61" authority="Nolf and Verstraeten, 2008" authorityName="Nolf and Verstraeten" authorityYear="2008" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="matthiasi">
|
||
<emphasis id="B949EAD5FF90FFBAFBD4FBC86D13FBBD" box="[1273,1377,1123,1149]" italics="true" pageId="18" pageNumber="19">matthiasi</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFA44FBC869FAFB61" author="Nolf, F. & Verstraeten, J." pageId="18" pageNumber="19" pagination="17 - 29" refId="ref140403" refString="Nolf, F. & Verstraeten, J. (2008) Two new turrid species from Congo-Brazzaville: Clavatula hattenbergeri and Clavatula congoensis (Mollusca: Gastropoda: Clavatulidae). Neptunea, 7 / 1, 17 - 29. [https: // neptuneaorg. files. wordpress. com / 2019 / 02 / np 7 - 1 - tekst. pdf]" type="journal article" year="2008">Nolf and Verstraeten, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
belong also to this group, based on conchological features (see
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFB67FB2C6D00FB61" author="Nolf, F. & Verstraeten, J." box="[1098,1394,1159,1185]" pageId="18" pageNumber="19" pagination="17 - 29" refId="ref140403" refString="Nolf, F. & Verstraeten, J. (2008) Two new turrid species from Congo-Brazzaville: Clavatula hattenbergeri and Clavatula congoensis (Mollusca: Gastropoda: Clavatulidae). Neptunea, 7 / 1, 17 - 29. [https: // neptuneaorg. files. wordpress. com / 2019 / 02 / np 7 - 1 - tekst. pdf]" type="journal article" year="2008">Nolf and Verstraeten 2008</bibRefCitation>
|
||
: pls 3–5). Herein, we treat these species as
|
||
<emphasis id="B949EAD5FF90FFBAFD67FB006ADEFB05" box="[586,684,1195,1221]" italics="true" pageId="18" pageNumber="19">caerulea</emphasis>
|
||
-clade.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF90FFBAFFEAFB646C93FAB9" blockId="18.[151,1437,475,1977]" pageId="18" pageNumber="19">
|
||
<emphasis id="B949EAD5FF90FFBAFFEAFB646937FB29" box="[199,325,1231,1257]" italics="true" pageId="18" pageNumber="19">‘Clavatula’</emphasis>
|
||
<taxonomicName id="4C3D4D44FF90FFBAFE67FB646A33FB29" authority="(Reeve, 1843)" baseAuthorityName="Reeve" baseAuthorityYear="1843" box="[330,577,1231,1257]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="mystica">
|
||
<emphasis id="B949EAD5FF90FFBAFE67FB6469D2FB28" box="[330,416,1231,1256]" italics="true" pageId="18" pageNumber="19">mystica</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFE81FB646A4BFB29" author="Reeve, L. A." box="[428,569,1231,1257]" pageId="18" pageNumber="19" refId="ref142338" refString="Reeve, L. A. (1843 & 1845) Monograph of the genus Pleurotoma. Conchologia Iconica, or, illustrations of the shells of molluscous animals. Vol. 1. Reeve & Co., London, 40 pls., unpaginated text. [https: // www. biodiversitylibrary. org / item / 41735 # page / 33 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 8129" type="url" year="1843">Reeve, 1843</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
differs from these species by its tripartite sculpture on the first four teleoconch whorls (
|
||
<figureCitation id="13062A42FF90FFBAFFDEFB586923FACD" box="[243,337,1267,1293]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="18" pageNumber="19">Figs 3D</figureCitation>
|
||
,
|
||
<figureCitation id="13062A42FF90FFBAFE70FB5869D7FACD" box="[349,421,1267,1293]" captionStart="FIGURE 9" captionStartId="22.[151,250,1808,1832]" captionTargetBox="[151,1436,777,1785]" captionTargetId="figure-260@22.[151,1436,777,1785]" captionTargetPageId="22" captionText="FIGURE 9. Extant Clavatulidae species. A. Clavatula rubrifasciata (Reeve, 1845), Gonçalo Rosa, Atoll Specimen Shells (Senegal). B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), Gonçalo Rosa, Atoll Specimen Shells (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ bimarginata (Lamarck, 1822), NHMW 2042, Gorée (Senegal).G. ‘Clavatula’ diadema (Kiener,1839), NHMW 1884/0010/0023, Gorée (Senegal). Specimens C and E have been used for molecular analysis by Nicolas Puillandre (unpublished data)." figureDoi="http://doi.org/10.5281/zenodo.6399849" httpUri="https://zenodo.org/record/6399849/files/figure.png" pageId="18" pageNumber="19">9C–D</figureCitation>
|
||
). It starts with weakly opisthocline axial ribs, which grade into three spiral rows of beads, with the suprasutural spiral row being most prominent. The whorl profile is almost flat-sided.A comparable sculpture is developed by ‘
|
||
<emphasis id="B949EAD5FF90FFBAFE7AFA906994FA95" box="[343,486,1339,1365]" italics="true" pageId="18" pageNumber="19">C.’ filograna</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFEC3FA906AF8FA95" author="Odhner, N. H." box="[494,650,1339,1365]" pageId="18" pageNumber="19" pagination="1 - 39" refId="ref140676" refString="Odhner, N. H. (1923) Contribution to the marine molluscan faunas of South and West Africa. Goteborgs Kungliga Vetenskapsoch Vitterhets-Samhalles Handlingar, 26 (7), 1 - 39." type="journal article" year="1923">Odhner, 1923</bibRefCitation>
|
||
, which has a tripartite sculpture on early teleoconch whorls consisting of three beaded spiral cords of similar strength. These species are grouped herein as
|
||
<taxonomicName id="4C3D4D44FF90FFBAFB6AFAF46CEFFAB8" baseAuthorityName="Reeve" baseAuthorityYear="1843" box="[1095,1181,1375,1400]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="mystica">
|
||
<emphasis id="B949EAD5FF90FFBAFB6AFAF46CEFFAB8" box="[1095,1181,1375,1400]" italics="true" pageId="18" pageNumber="19">mystica</emphasis>
|
||
</taxonomicName>
|
||
-clade
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF90FFBAFFEAFA286D1FF9C9" blockId="18.[151,1437,475,1977]" pageId="18" pageNumber="19">
|
||
‘
|
||
<emphasis id="B949EAD5FF90FFBAFFFCFA2869A9FA5D" box="[209,475,1411,1437]" italics="true" pageId="18" pageNumber="19">Clavatula’ bimarginata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFEDDFA286AD1FA5D" author="Lamarck, J. - B. M. de" box="[496,675,1411,1437]" pageId="18" pageNumber="19" refId="ref137485" refString="Lamarck, J. - B. M. de. (1822) Histoire naturelle des animaux sans vertebres. Tome Septieme. Published by the Author, Paris, 711 pp. [http: // www. biodiversitylibrary. org / item / 47433]" type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
) has a bipartite sculpture with a granulose subsutural cord and suprasutural spiral cord of equal strength with indistinct, bifid beads (
|
||
<figureCitation id="13062A42FF90FFBAFCE7FA0C6C68FA01" box="[970,1050,1447,1473]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="18" pageNumber="19">Fig 3F</figureCitation>
|
||
,
|
||
<figureCitation id="13062A42FF90FFBAFB04FA0C6C19FA01" box="[1065,1131,1447,1473]" captionStart="FIGURE 9" captionStartId="22.[151,250,1808,1832]" captionTargetBox="[151,1436,777,1785]" captionTargetId="figure-260@22.[151,1436,777,1785]" captionTargetPageId="22" captionText="FIGURE 9. Extant Clavatulidae species. A. Clavatula rubrifasciata (Reeve, 1845), Gonçalo Rosa, Atoll Specimen Shells (Senegal). B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), Gonçalo Rosa, Atoll Specimen Shells (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ bimarginata (Lamarck, 1822), NHMW 2042, Gorée (Senegal).G. ‘Clavatula’ diadema (Kiener,1839), NHMW 1884/0010/0023, Gorée (Senegal). Specimens C and E have been used for molecular analysis by Nicolas Puillandre (unpublished data)." figureDoi="http://doi.org/10.5281/zenodo.6399849" httpUri="https://zenodo.org/record/6399849/files/figure.png" pageId="18" pageNumber="19">9E–F</figureCitation>
|
||
). The moderately concave interspace bears prominent, comma-shaped axial riblets formed by growth lines. Spiral threads form a somewhat cancellate pattern especially in the abapical half of the whorls. This species is placed in the
|
||
<emphasis id="B949EAD5FF90FFBAFBB5FA446D53F9C9" box="[1176,1313,1519,1545]" italics="true" pageId="18" pageNumber="19">bimarginata</emphasis>
|
||
-clade.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF90FFBAFFEAF9B86BEBF879" blockId="18.[151,1437,475,1977]" pageId="18" pageNumber="19">
|
||
<emphasis id="B949EAD5FF90FFBAFFEAF9B869C4F9EC" box="[199,438,1555,1581]" italics="true" pageId="18" pageNumber="19">‘Clavatula’ tripartita</emphasis>
|
||
(Weinkauff, 1876) (
|
||
<figureCitation id="13062A42FF90FFBAFD8CF9B86B5CF9ED" box="[673,814,1555,1581]" captionStart="FIGURE 10" captionStartId="23.[151,250,1355,1379]" captionTargetBox="[151,1436,181,1331]" captionTargetId="figure-18@23.[151,1436,181,1331]" captionTargetPageId="23" captionText="FIGURE 10. Extant Clavatulidae species. A. ‘Clavatula’ caerulea (Weinkauff in Weinkauff & Kobelt, 1875), MNHN-IM- 2009-19153, Libreville (Gabon). B. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). C. ‘Clavatula’ tripartita (Weinkauff, 1876), MNHN-IM-2009-14826, Cap Sainte Marie (Madagascar). D. ‘Clavatula’ tripartita (Weinkauff, 1876), MNHN-IM-2009-14985, Cap Sainte Marie (Madagascar). E. ‘Clavatula’ taxea (Röding, 1798), MNHN-IM-2009-14901, Fort-Dauphin (Madagascar). F. Pusionella aculeiformis (Lamarck, 1816), NHMW3051, Senegal. G. Pusionella buccinata (Lamarck, 1822); NHMW3052. Specimens A–E have been used for molecular analysis by Nicolas Puillandre (unpublished data)." figureDoi="http://doi.org/10.5281/zenodo.6399853" httpUri="https://zenodo.org/record/6399853/files/figure.png" pageId="18" pageNumber="19">Figs 10C–D</figureCitation>
|
||
) and ‘
|
||
<emphasis id="B949EAD5FF90FFBAFC56F9B86BFCF9ED" box="[891,910,1555,1581]" italics="true" pageId="18" pageNumber="19">C</emphasis>
|
||
.’
|
||
<emphasis id="B949EAD5FF90FFBAFC8BF9B86B91F9EC" box="[934,995,1555,1580]" italics="true" pageId="18" pageNumber="19">taxea</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFCD9F9B86CE3F9ED" author="Roding, P. F." box="[1012,1169,1555,1581]" pageId="18" pageNumber="19" refId="ref142481" refString="Roding, P. F. (1798). Museum Boltenianum sive Catalogus cimeliorum e tribus regnis naturae quae olim collegerat Joa. Fried Bolten, M. D. p. d. per XL. annos proto physicus Hamburgensis. Pars secunda continens Conchylia sive Testacea univalvia, bivalvia & multivalvia. Trapp, Hamburg, viii + 199 pp. [https: // www. biodiversitylibrary. org / page / 16230659]" type="book" year="1798">Röding, 1798</bibRefCitation>
|
||
) (
|
||
<figureCitation id="13062A42FF90FFBAFB81F9B86D63F9ED" box="[1196,1297,1555,1581]" captionStart="FIGURE 10" captionStartId="23.[151,250,1355,1379]" captionTargetBox="[151,1436,181,1331]" captionTargetId="figure-18@23.[151,1436,181,1331]" captionTargetPageId="23" captionText="FIGURE 10. Extant Clavatulidae species. A. ‘Clavatula’ caerulea (Weinkauff in Weinkauff & Kobelt, 1875), MNHN-IM- 2009-19153, Libreville (Gabon). B. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). C. ‘Clavatula’ tripartita (Weinkauff, 1876), MNHN-IM-2009-14826, Cap Sainte Marie (Madagascar). D. ‘Clavatula’ tripartita (Weinkauff, 1876), MNHN-IM-2009-14985, Cap Sainte Marie (Madagascar). E. ‘Clavatula’ taxea (Röding, 1798), MNHN-IM-2009-14901, Fort-Dauphin (Madagascar). F. Pusionella aculeiformis (Lamarck, 1816), NHMW3051, Senegal. G. Pusionella buccinata (Lamarck, 1822); NHMW3052. Specimens A–E have been used for molecular analysis by Nicolas Puillandre (unpublished data)." figureDoi="http://doi.org/10.5281/zenodo.6399853" httpUri="https://zenodo.org/record/6399853/files/figure.png" pageId="18" pageNumber="19">Fig. 10E</figureCitation>
|
||
) are neither bipartite nor tripartite, but develop close-set spiral cords on flat-sided early teleoconch whorls. This morphology differs greatly from other
|
||
<taxonomicName id="4C3D4D44FF90FFBAFE9BF9F06A36F9B5" authorityName="Gray" authorityYear="1853" box="[438,580,1627,1653]" class="Gastropoda" family="Clavatulidae" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="family">Clavatulidae</taxonomicName>
|
||
discussed herein and are placed herein in the
|
||
<emphasis id="B949EAD5FF90FFBAFB13F9F06C09F9B4" box="[1086,1147,1627,1652]" italics="true" pageId="18" pageNumber="19">taxea</emphasis>
|
||
-clade. A separation of the ‘
|
||
<emphasis id="B949EAD5FF90FFBAFF8CF9D468C8F959" box="[161,186,1663,1689]" italics="true" pageId="18" pageNumber="19">C.</emphasis>
|
||
’
|
||
<emphasis id="B949EAD5FF90FFBAFFE4F9D4697DF958" box="[201,271,1663,1688]" italics="true" pageId="18" pageNumber="19">taxea-</emphasis>
|
||
clade from
|
||
<taxonomicName id="4C3D4D44FF90FFBAFEA6F9D4698BF959" authorityName="Lamarck" authorityYear="1801" box="[395,505,1663,1689]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF90FFBAFEA6F9D4698BF959" box="[395,505,1663,1689]" italics="true" pageId="18" pageNumber="19">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
is also suggested based on the comparatively slender, smooth and high spired shells, and the weak subsutural collar, which lacks spines. Consequently, already
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFCCEF9086CB5F97D" author="Kilburn, R. N." box="[995,1223,1698,1725]" pageId="18" pageNumber="19" pagination="417 - 470" refId="ref136560" refString="Kilburn, R. N. (1985) Turridae (Mollusca: Gastropoda) of southern Africa and Mozambique. Part 2. Subfamily Clavatulinae. Annals of the Natal Museum, 26, 417 - 470. [https: // journals. co. za / doi / abs / 10.10520 / AJA 03040798 _ 469]" type="journal article" year="1985">Kilburn (1985: 424)</bibRefCitation>
|
||
doubted that these South African species were congeneric with the western African
|
||
<taxonomicName id="4C3D4D44FF90FFBAFC45F96C6BA4F921" authorityName="Lamarck" authorityYear="1801" box="[872,982,1735,1761]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus" sensu="stricto">
|
||
<emphasis id="B949EAD5FF90FFBAFC45F96C6BA4F921" box="[872,982,1735,1761]" italics="true" pageId="18" pageNumber="19">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF90FFBAFCF0F96C6C73F921" box="[989,1025,1735,1761]" pageId="18" pageNumber="19" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
In addition, the anal sinus allows for a separation of ‘
|
||
<emphasis id="B949EAD5FF90FFBAFE64F940692EF8C5" box="[329,348,1771,1797]" italics="true" pageId="18" pageNumber="19">C</emphasis>
|
||
.’
|
||
<taxonomicName id="4C3D4D44FF90FFBAFE42F94069B7F8C4" baseAuthorityName="Reeve" baseAuthorityYear="1843" box="[367,453,1771,1796]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="mystica">
|
||
<emphasis id="B949EAD5FF90FFBAFE42F94069B7F8C4" box="[367,453,1771,1796]" italics="true" pageId="18" pageNumber="19">mystica</emphasis>
|
||
</taxonomicName>
|
||
, which has a moderately deep, widely V-shaped anal sinus, from
|
||
<taxonomicName id="4C3D4D44FF90FFBAFBB8F9406D1CF8C4" authorityName="" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[1173,1390,1771,1797]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="muricata">
|
||
<emphasis id="B949EAD5FF90FFBAFBB8F9406D1CF8C4" box="[1173,1390,1771,1797]" italics="true" pageId="18" pageNumber="19">Clavatula muricata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3D4D44FF90FFBAFFBAF8A46938F8E9" baseAuthorityName="Reeve" baseAuthorityYear="1845" box="[151,330,1807,1833]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="rubrifasciata">
|
||
<emphasis id="B949EAD5FF90FFBAFFBAF8A46938F8E9" box="[151,330,1807,1833]" italics="true" pageId="18" pageNumber="19">C. rubrifasciata</emphasis>
|
||
</taxonomicName>
|
||
, which have a deep, U-shaped anal sinus (
|
||
<figureCitation id="13062A42FF90FFBAFC05F8A46B1EF8E9" box="[808,876,1806,1833]" captionStart="FIGURE 5" captionStartId="15.[151,250,1999,2023]" captionTargetBox="[151,1436,631,1976]" captionTargetId="figure-214@15.[151,1436,631,1976]" captionTargetPageId="15" captionText="FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae." figureDoi="http://doi.org/10.5281/zenodo.6399839" httpUri="https://zenodo.org/record/6399839/files/figure.png" pageId="18" pageNumber="19">Fig. 5</figureCitation>
|
||
). An additional
|
||
<typeStatus id="54868865FF90FFBAFB0DF8A46C22F8E9" box="[1056,1104,1807,1833]" pageId="18" pageNumber="19">type</typeStatus>
|
||
of early teleoconch sculpture is represented by the elongate biconic, prominently spinose ‘
|
||
<emphasis id="B949EAD5FF90FFBAFC71F8986B1DF88D" box="[860,879,1843,1869]" italics="true" pageId="18" pageNumber="19">C</emphasis>
|
||
.’
|
||
<emphasis id="B949EAD5FF90FFBAFCA5F8986B9AF88D" box="[904,1000,1843,1869]" italics="true" pageId="18" pageNumber="19">diadema</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF90FFBAFCD7F8986CE3F88D" author="Kiener, L. C." box="[1018,1169,1843,1869]" pageId="18" pageNumber="19" refId="ref136494" refString="Kiener, L. C. (1839) Species general et iconographie des coquilles vivantes. Vol. 5. Famille des Canaliferes. Premiere partie. Pleurotome (Pleurotoma), Lamarck. Rousseau & J. B. Bailliere, Paris, 16 pp., 27 pls. [http: // www. biodiversitylibrary. org / item / 19762]" type="book" year="1839">Kiener, 1839</bibRefCitation>
|
||
) (
|
||
<figureCitation id="13062A42FF90FFBAFB81F8986D79F88D" box="[1196,1291,1843,1869]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="18" pageNumber="19">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation id="13062A42FF90FFBAFA36F8986D32F88D" box="[1307,1344,1843,1869]" captionStart="FIGURE 9" captionStartId="22.[151,250,1808,1832]" captionTargetBox="[151,1436,777,1785]" captionTargetId="figure-260@22.[151,1436,777,1785]" captionTargetPageId="22" captionText="FIGURE 9. Extant Clavatulidae species. A. Clavatula rubrifasciata (Reeve, 1845), Gonçalo Rosa, Atoll Specimen Shells (Senegal). B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), Gonçalo Rosa, Atoll Specimen Shells (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ bimarginata (Lamarck, 1822), NHMW 2042, Gorée (Senegal).G. ‘Clavatula’ diadema (Kiener,1839), NHMW 1884/0010/0023, Gorée (Senegal). Specimens C and E have been used for molecular analysis by Nicolas Puillandre (unpublished data)." figureDoi="http://doi.org/10.5281/zenodo.6399849" httpUri="https://zenodo.org/record/6399849/files/figure.png" pageId="18" pageNumber="19">9G</figureCitation>
|
||
), which has weakly opisthocline axial ribs on the first teleoconch whorl and a weak subsutural spiral cord on the second teleoconch whorl. The beaded suprasutural spiral cord evolves from the abapical tips of the axial ribs on subsequent whorls. In addition, this species is characterized by very long spines.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF91FFBBFFBAFF3C6BAAFF71" blockId="19.[151,1010,151,393]" box="[151,984,151,177]" pageId="19" pageNumber="20">
|
||
Thus, within the species, formerly referred to as
|
||
<taxonomicName id="4C3D4D44FF91FFBBFD9AFF3C6B57FF71" authorityName="Lamarck" authorityYear="1801" box="[695,805,151,177]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF91FFBBFD9AFF3C6B57FF71" box="[695,805,151,177]" italics="true" pageId="19" pageNumber="20">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
we distinguish:
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF91FFBBFFEAFF106B80FF15" blockId="19.[151,1010,151,393]" box="[199,1010,187,213]" pageId="19" pageNumber="20">
|
||
<emphasis id="B949EAD5FF91FFBBFFEAFF106917FF15" box="[199,357,187,213]" italics="true" pageId="19" pageNumber="20">
|
||
1.
|
||
<taxonomicName id="4C3D4D44FF91FFBBFFDAFF106917FF15" authorityName="Lamarck" authorityYear="1801" box="[247,357,187,213]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">Clavatula</taxonomicName>
|
||
</emphasis>
|
||
(‘
|
||
<emphasis id="B949EAD5FF91FFBBFE52FF1069E0FF15" box="[383,402,187,213]" italics="true" pageId="19" pageNumber="20">C</emphasis>
|
||
.’
|
||
<emphasis id="B949EAD5FF91FFBBFE84FF106998FF14" box="[425,490,187,212]" italics="true" pageId="19" pageNumber="20">
|
||
<taxonomicName id="4C3D4D44FF91FFBBFE84FF106994FF14" baseAuthorityName="Roding" baseAuthorityYear="1798" box="[425,486,187,212]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="regia">regia</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
‘
|
||
<emphasis id="B949EAD5FF91FFBBFED6FF106A7CFF15" box="[507,526,187,213]" italics="true" pageId="19" pageNumber="20">C</emphasis>
|
||
.’
|
||
<emphasis id="B949EAD5FF91FFBBFD0EFF106AFCFF14" box="[547,654,187,212]" italics="true" pageId="19" pageNumber="20">
|
||
<taxonomicName id="4C3D4D44FF91FFBBFD0EFF106AF9FF14" authorityName="" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[547,651,187,212]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="muricata">muricata</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
‘
|
||
<emphasis id="B949EAD5FF91FFBBFDB2FF106AC0FF15" box="[671,690,187,213]" italics="true" pageId="19" pageNumber="20">C</emphasis>
|
||
.’
|
||
<emphasis id="B949EAD5FF91FFBBFDEAFF106B12FF15" box="[711,864,187,213]" italics="true" pageId="19" pageNumber="20">
|
||
<taxonomicName id="4C3D4D44FF91FFBBFDEAFF106B2EFF15" baseAuthorityName="Reeve" baseAuthorityYear="1845" box="[711,860,187,213]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="rubrifasciata">rubrifasciata</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
‘
|
||
<emphasis id="B949EAD5FF91FFBBFC5DFF106BF1FF15" box="[880,899,187,213]" italics="true" pageId="19" pageNumber="20">C</emphasis>
|
||
.’
|
||
<taxonomicName id="4C3D4D44FF91FFBBFCB5FF106B9BFF15" baseAuthorityName="Recluz" baseAuthorityYear="1851" box="[920,1001,187,213]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="lelieuri">
|
||
<emphasis id="B949EAD5FF91FFBBFCB5FF106B9BFF15" box="[920,1001,187,213]" italics="true" pageId="19" pageNumber="20">lelieuri</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF91FFBBFFEAFF746B79FF39" blockId="19.[151,1010,151,393]" box="[199,779,223,249]" pageId="19" pageNumber="20">
|
||
<emphasis id="B949EAD5FF91FFBBFFEAFF74692BFF39" box="[199,345,223,249]" italics="true" pageId="19" pageNumber="20">2. caerulea</emphasis>
|
||
-clade (‘
|
||
<emphasis id="B949EAD5FF91FFBBFE9AFF7469B8FF39" box="[439,458,223,249]" italics="true" pageId="19" pageNumber="20">C</emphasis>
|
||
.’
|
||
<emphasis id="B949EAD5FF91FFBBFECCFF746B70FF39" box="[481,770,223,249]" italics="true" pageId="19" pageNumber="20">
|
||
caerulea, ‘C.’
|
||
<taxonomicName id="4C3D4D44FF91FFBBFDACFF746B70FF39" baseAuthorityName="Kiener" baseAuthorityYear="1839" box="[641,770,223,249]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="pyramidata">pyramidata</taxonomicName>
|
||
</emphasis>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF91FFBBFFEAFEA86A41FEDD" blockId="19.[151,1010,151,393]" box="[199,563,259,285]" pageId="19" pageNumber="20">
|
||
<emphasis id="B949EAD5FF91FFBBFFEAFEA8693FFEDC" box="[199,333,259,285]" italics="true" pageId="19" pageNumber="20">
|
||
3.
|
||
<taxonomicName id="4C3D4D44FF91FFBBFFDAFEA8693FFEDC" baseAuthorityName="Reeve" baseAuthorityYear="1843" box="[247,333,259,284]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="mystica">mystica</taxonomicName>
|
||
</emphasis>
|
||
-clade (‘
|
||
<emphasis id="B949EAD5FF91FFBBFE86FEA869CCFEDD" box="[427,446,259,285]" italics="true" pageId="19" pageNumber="20">C</emphasis>
|
||
.’
|
||
<taxonomicName id="4C3D4D44FF91FFBBFEF8FEA86A59FEDC" baseAuthorityName="Reeve" baseAuthorityYear="1843" box="[469,555,259,284]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="mystica">
|
||
<emphasis id="B949EAD5FF91FFBBFEF8FEA86A59FEDC" box="[469,555,259,284]" italics="true" pageId="19" pageNumber="20">mystica</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF91FFBBFFEAFE8C6AEBFE81" blockId="19.[151,1010,151,393]" box="[199,665,295,321]" pageId="19" pageNumber="20">
|
||
<emphasis id="B949EAD5FF91FFBBFFEAFE8C69F2FE81" box="[199,384,295,321]" italics="true" pageId="19" pageNumber="20">4. bimarginata</emphasis>
|
||
-clade (‘
|
||
<emphasis id="B949EAD5FF91FFBBFEF2FE8C6980FE81" box="[479,498,295,321]" italics="true" pageId="19" pageNumber="20">C</emphasis>
|
||
.’
|
||
<emphasis id="B949EAD5FF91FFBBFD2AFE8C6AE2FE81" box="[519,656,295,321]" italics="true" pageId="19" pageNumber="20">bimarginata</emphasis>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF91FFBBFFEAFEE06AD6FEA5" blockId="19.[151,1010,151,393]" box="[199,676,331,357]" pageId="19" pageNumber="20">
|
||
<emphasis id="B949EAD5FF91FFBBFFEAFEE06946FEA4" box="[199,308,331,357]" italics="true" pageId="19" pageNumber="20">5. taxea</emphasis>
|
||
-clade (‘
|
||
<emphasis id="B949EAD5FF91FFBBFEBFFEE069D7FEA5" box="[402,421,331,357]" italics="true" pageId="19" pageNumber="20">C</emphasis>
|
||
.’
|
||
<emphasis id="B949EAD5FF91FFBBFE97FEE06AEEFEA4" box="[442,668,331,357]" italics="true" pageId="19" pageNumber="20">taxea, ‘C.’ tripartita</emphasis>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF91FFBBFFEAFEC46A35FE49" blockId="19.[151,1010,151,393]" box="[199,583,367,393]" pageId="19" pageNumber="20">
|
||
<emphasis id="B949EAD5FF91FFBBFFEAFEC46925FE49" box="[199,343,367,393]" italics="true" pageId="19" pageNumber="20">6. diadema</emphasis>
|
||
-clade (‘
|
||
<emphasis id="B949EAD5FF91FFBBFE9BFEC469BBFE49" box="[438,457,367,393]" italics="true" pageId="19" pageNumber="20">C</emphasis>
|
||
.’
|
||
<emphasis id="B949EAD5FF91FFBBFEF3FEC46A4CFE49" box="[478,574,367,393]" italics="true" pageId="19" pageNumber="20">diadema</emphasis>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF91FFBBFFEAFA1C6B2EF945" blockId="19.[151,1437,1463,2029]" pageId="19" pageNumber="20">
|
||
As we focus on fossil species from the Neogene of Europe, we refrain from introducing genus names for these clades. Nevertheless, these data allow a refined systematic treatment of Paratethyan species, based on the hypothesis that early teleoconch sculpture is a useful trait to delimit genera within
|
||
<taxonomicName id="4C3D4D44FF91FFBBFC9BFA546C35F9D9" ID-CoL="7NHYP" authorityName="Gray" authorityYear="1853" box="[950,1095,1535,1561]" class="Gastropoda" family="Clavatulidae" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="family">Clavatulidae</taxonomicName>
|
||
. Only three of the Paratethyan
|
||
<taxonomicName id="4C3D4D44FF91FFBBFFBAF9886957F9FD" ID-CoL="7NHYP" authorityName="Gray" authorityYear="1853" box="[151,293,1571,1597]" class="Gastropoda" family="Clavatulidae" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="family">Clavatulidae</taxonomicName>
|
||
can be placed in one of these clades:
|
||
<taxonomicName id="4C3D4D44FF91FFBBFDEDF9886B07F9FC" ID-CoL="8YLQB" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[704,885,1571,1597]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="sorini" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF91FFBBFDEDF9886B07F9FC" box="[704,885,1571,1597]" italics="true" pageId="19" pageNumber="20">Clavatula sorini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF91FFBBFC56F9886BA2F9FD" box="[891,976,1571,1597]" pageId="19" pageNumber="20" rank="species">nov. sp.</taxonomicNameLabel>
|
||
is placed in
|
||
<taxonomicName id="4C3D4D44FF91FFBBFB75F9886CB4F9FD" ID-CoL="7NZRZ" box="[1112,1222,1571,1597]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus" sensu="stricto">
|
||
<emphasis id="B949EAD5FF91FFBBFB75F9886CB4F9FD" box="[1112,1222,1571,1597]" italics="true" pageId="19" pageNumber="20">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF91FFBBFBE1F9886C83F9FD" box="[1228,1265,1571,1597]" pageId="19" pageNumber="20" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
, ‘
|
||
<emphasis id="B949EAD5FF91FFBBFA2BF9886D52F9FD" box="[1286,1312,1571,1597]" italics="true" pageId="19" pageNumber="20">C.</emphasis>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF91FFBBFA02F98869C8F9A1" authority="Hoernes & Auinger, 1891" authorityName="Hoernes & Auinger" authorityYear="1891" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="veronicae">
|
||
<emphasis id="B949EAD5FF91FFBBFA02F9886DEEF9FC" box="[1327,1436,1571,1596]" italics="true" pageId="19" pageNumber="20">veronicae</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF91FFBBFFBAF9EC69C8F9A1" author="Hoernes, R. & Auinger, M." box="[151,442,1607,1633]" pageId="19" pageNumber="20" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
</taxonomicName>
|
||
is placed in the
|
||
<emphasis id="B949EAD5FF91FFBBFD59F9EC6A8FF9A1" box="[628,765,1607,1633]" italics="true" pageId="19" pageNumber="20">bimarginata</emphasis>
|
||
-clade and the ‘
|
||
<taxonomicName id="4C3D4D44FF91FFBBFC80F9EC6C69F9A1" ID-CoL="7NZRZ" box="[941,1051,1607,1633]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF91FFBBFC80F9EC6C69F9A1" box="[941,1051,1607,1633]" italics="true" pageId="19" pageNumber="20">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF91FFBBFB06F9EC6CEFF9A0" baseAuthorityName="Brocchi" baseAuthorityYear="1814" box="[1067,1181,1607,1632]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="interrupta">
|
||
<emphasis id="B949EAD5FF91FFBBFB06F9EC6CEFF9A0" box="[1067,1181,1607,1632]" italics="true" pageId="19" pageNumber="20">interrupta</emphasis>
|
||
</taxonomicName>
|
||
group and ‘
|
||
<taxonomicName id="4C3D4D44FF91FFBBFA0AF9EC6DE7F9A1" ID-CoL="7NZRZ" box="[1319,1429,1607,1633]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF91FFBBFA0AF9EC6DE7F9A1" box="[1319,1429,1607,1633]" italics="true" pageId="19" pageNumber="20">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF91FFBBFFBAF9C069C6F946" ID-CoL="8YLQ9" authority="(Defrance, 1826)" baseAuthorityName="Defrance" baseAuthorityYear="1826" box="[151,436,1643,1670]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="romana">
|
||
<emphasis id="B949EAD5FF91FFBBFFBAF9C0689FF944" box="[151,237,1643,1668]" italics="true" pageId="19" pageNumber="20">romana</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF91FFBBFFD1F9C069D9F946" author="Defrance, M." box="[252,427,1643,1670]" pageId="19" pageNumber="20" pagination="388 - 396" refId="ref131223" refString="Defrance, M. (1826) Pleurotome. dans lequel on traite meithodiquement des diffeirens e ˆ tres de la nature, consideireis soit en eux-me ˆ mes, d'apreIs l'eitat actuel de nos connoissances, soit relativement aI l'utilitei qu'en peuvent retirer la meidecine, l'agriculture, le commerce et les artes. Vol. 41. Levrault, Le Normant, Paris, Strasbourg, 388 - 396. [https: // www. biodiversitylibrary. org / item / 81578 # page / 398 / mode / 1 up]" type="book chapter" year="1826">Defrance, 1826</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
might be placed in the
|
||
<taxonomicName id="4C3D4D44FF91FFBBFD97F9C06B62F944" ID-CoL="VV4J" baseAuthorityName="Reeve" baseAuthorityYear="1843" box="[698,784,1643,1668]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="mystica">
|
||
<emphasis id="B949EAD5FF91FFBBFD97F9C06B62F944" box="[698,784,1643,1668]" italics="true" pageId="19" pageNumber="20">mystica</emphasis>
|
||
</taxonomicName>
|
||
-clade.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF91FFBBFFEAF924694FF931" blockId="19.[151,1437,1463,2029]" pageId="19" pageNumber="20">
|
||
Consequently, most of the remaining Paratethyan species should be placed into new genera, which we propose in the following. Nevertheless, some species remain, which do not fit in our scheme. These are listed provisionally as ‘
|
||
<taxonomicName id="4C3D4D44FF91FFBBFF92F97C695FF931" ID-CoL="7NZRZ" box="[191,301,1751,1777]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF91FFBBFF92F97C695FF931" box="[191,301,1751,1777]" italics="true" pageId="19" pageNumber="20">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFF91FFBBFFEAF951693EF82D" pageId="19" pageNumber="20" type="distribution">
|
||
<paragraph id="8B8236C7FF91FFBBFFEAF951693EF82D" blockId="19.[151,1437,1463,2029]" pageId="19" pageNumber="20">
|
||
<emphasis id="B949EAD5FF91FFBBFFEAF9516A40F8D4" bold="true" box="[199,562,1786,1813]" pageId="19" pageNumber="20">Distribution and stratigraphy.</emphasis>
|
||
The oldest records from the Eocene of
|
||
<collectingCountry id="F32A7657FF91FFBBFCDAF9506C58F8D5" box="[1015,1066,1787,1813]" name="Italy" pageId="19" pageNumber="20">Italy</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32A7657FF91FFBBFB4EF9506CC0F8D5" box="[1123,1202,1787,1813]" name="France" pageId="19" pageNumber="20">France</collectingCountry>
|
||
, placed in
|
||
<taxonomicName id="4C3D4D44FF91FFBBFA03F9506DEEF8D5" box="[1326,1436,1787,1813]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF91FFBBFA03F9506DEEF8D5" box="[1326,1436,1787,1813]" italics="true" pageId="19" pageNumber="20">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFAC4B36FF91FFBBFF94F8B469A3F8F9" author="Vinassa de Regny, P. E." box="[185,465,1823,1849]" pageId="19" pageNumber="20" pagination="103" refId="ref146196" refString="Vinassa de Regny, P. E. (1898) Synopsis dei molluschi terziari delle Alpi Venete. Palaeontographia Italica, 3, 103 (145) - 158 (200). [https: // www. biodiversitylibrary. org / item / 106874 # page / 155 / mode / 1 up]" type="journal article" year="1898">Vinassa de Regny (1898)</bibRefCitation>
|
||
and Boussac, (1911) represent other
|
||
<taxonomicName id="4C3D4D44FF91FFBBFC40F8B46BAAF8F9" authorityName="J. Fleming" authorityYear="1822" box="[877,984,1823,1849]" class="Gastropoda" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="12" phylum="Mollusca" rank="superFamily" superFamily="Conoidea">Conoidea</taxonomicName>
|
||
families, such as
|
||
<taxonomicName id="4C3D4D44FF91FFBBFBB0F8B46D0DF8F9" authorityName="J. P. E. Morrison" authorityYear="1966" box="[1181,1407,1823,1849]" class="Gastropoda" family="Pseudomelatomidae" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="family">Pseudomelatomidae</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C3D4D44FF91FFBBFFBAF8E8696CF89D" authorityName="Bellardi" authorityYear="1875" box="[151,286,1859,1885]" class="Gastropoda" family="Borsoniidae" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="family">Borsoniidae</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFAC4B36FF91FFBBFE76F8E86A1AF89D" author="Vinassa de Regny, P. E." box="[347,616,1859,1885]" pageId="19" pageNumber="20" pagination="103" refId="ref146196" refString="Vinassa de Regny, P. E. (1898) Synopsis dei molluschi terziari delle Alpi Venete. Palaeontographia Italica, 3, 103 (145) - 158 (200). [https: // www. biodiversitylibrary. org / item / 106874 # page / 155 / mode / 1 up]" type="journal article" year="1898">Vinassa de Regny 1898</bibRefCitation>
|
||
, pl. 7 (20), figs 40–45).
|
||
<taxonomicName id="4C3D4D44FF91FFBBFC51F8E86DF0F89D" authority="Vinassa de Regny, 1898" authorityName="Vinassa de Regny" authorityYear="1898" box="[892,1410,1859,1885]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="praegotica">
|
||
<emphasis id="B949EAD5FF91FFBBFC51F8E86C19F89C" box="[892,1131,1859,1885]" italics="true" pageId="19" pageNumber="20">Clavatula praegotica</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF91FFBBFB5EF8E86DF0F89D" author="Vinassa de Regny, P. E." box="[1139,1410,1859,1885]" pageId="19" pageNumber="20" pagination="103" refId="ref146196" refString="Vinassa de Regny, P. E. (1898) Synopsis dei molluschi terziari delle Alpi Venete. Palaeontographia Italica, 3, 103 (145) - 158 (200). [https: // www. biodiversitylibrary. org / item / 106874 # page / 155 / mode / 1 up]" type="journal article" year="1898">Vinassa de Regny, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
is reminiscent of
|
||
<taxonomicName id="4C3D4D44FF91FFBBFE12F8CC69E9F840" box="[319,411,1895,1920]" class="Gastropoda" family="Clavatulidae" genus="Perrona" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF91FFBBFE12F8CC69E9F840" box="[319,411,1895,1920]" italics="true" pageId="19" pageNumber="20">Perrona</emphasis>
|
||
</taxonomicName>
|
||
, but consists only of a fragment of the last whorl and lacks distinctive features (see
|
||
<bibRefCitation id="EFAC4B36FF91FFBBFA68F8CC6938F865" author="Vinassa de Regny, P. E." pageId="19" pageNumber="20" pagination="103" refId="ref146196" refString="Vinassa de Regny, P. E. (1898) Synopsis dei molluschi terziari delle Alpi Venete. Palaeontographia Italica, 3, 103 (145) - 158 (200). [https: // www. biodiversitylibrary. org / item / 106874 # page / 155 / mode / 1 up]" type="journal article" year="1898">Vinassa de Regny, 1898</bibRefCitation>
|
||
, pl. 7(20), figs 39a, b). Similarly, the numerous species from the Paleogene of
|
||
<collectingCountry id="F32A7657FF91FFBBFBEFF8206D3AF865" box="[1218,1352,1931,1957]" name="Kazakhstan" pageId="19" pageNumber="20">Kazakhstan</collectingCountry>
|
||
, placed in
|
||
<taxonomicName id="4C3D4D44FF91FFBBFF98F8046951F809" box="[181,291,1967,1993]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF91FFBBFF98F8046951F809" box="[181,291,1967,1993]" italics="true" pageId="19" pageNumber="20">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFAC4B36FF91FFBBFE63F8046A72F809" author="Amitrov, O. V." box="[334,512,1967,1993]" pageId="19" pageNumber="20" pagination="1 - 211" refId="ref128486" refString="Amitrov, O. V. (1973) Turrid pozdnego Eotzena - rannego Miotzena juga SSSR. Trudy Paleontologicheskogo Institutom, Akademij Nauk SSSR, 135, 1 - 211. [https: // www. geokniga. org / bookfiles / geokniga-amitrov- 1973. pdf]" type="journal article" year="1973">Amitrov (1973)</bibRefCitation>
|
||
are clearly not
|
||
<taxonomicName id="4C3D4D44FF91FFBBFD9FF8046B32F809" authorityName="Gray" authorityYear="1853" box="[690,832,1967,1993]" class="Gastropoda" family="Clavatulidae" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="20" phylum="Mollusca" rank="family">Clavatulidae</taxonomicName>
|
||
and belong to other conoidean families (see
|
||
<bibRefCitation id="EFAC4B36FF91FFBBFA10F80468A0F82D" author="Amitrov, O. V." pageId="19" pageNumber="20" pagination="1 - 211" refId="ref128486" refString="Amitrov, O. V. (1973) Turrid pozdnego Eotzena - rannego Miotzena juga SSSR. Trudy Paleontologicheskogo Institutom, Akademij Nauk SSSR, 135, 1 - 211. [https: // www. geokniga. org / bookfiles / geokniga-amitrov- 1973. pdf]" type="journal article" year="1973">Amitrov 1973</bibRefCitation>
|
||
, pls 9–11).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFF96FFBCFFEAFF3C6C2FFC2D" pageId="20" pageNumber="21" type="biology_ecology">
|
||
<paragraph id="8B8236C7FF96FFBCFFEAFF3C69B5FE12" blockId="20.[151,1437,151,1005]" pageId="20" pageNumber="21">
|
||
Species belonging to the ‘
|
||
<taxonomicName id="4C3D4D44FF96FFBCFED7FF3C6A1AFF71" box="[506,616,151,177]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFED7FF3C6A1AFF71" box="[506,616,151,177]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’ species group first appear during the Oligocene.
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFB95FF3C6DEEFF71" author="von Koenen, A." box="[1208,1436,151,177]" pageId="20" pageNumber="21" pagination="281 - 574" refId="ref146253" refString="von Koenen, A. (1890) Das Norddeutsche Unter-Oligocan und seine Molluskenfauna. Conidae. Abhandlungen zur Geologischen Specialkarte von Preussen und den Thuringischen Staaten, 10, 281 - 574, pls. 24 - 34. [https: // www. biodiversitylibrary. org / item / 110903 # page / 725 / mode / 1 up]" type="journal article" year="1890">Von Koenen (1890)</bibRefCitation>
|
||
described ‘
|
||
<taxonomicName id="4C3D4D44FF96FFBCFE38FF1069F1FF15" box="[277,387,187,213]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFE38FF1069F1FF15" box="[277,387,187,213]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’ species, such as ‘
|
||
<taxonomicName id="4C3D4D44FF96FFBCFD71FF106AB8FF15" box="[604,714,187,213]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFD71FF106AB8FF15" box="[604,714,187,213]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B949EAD5FF96FFBCFDF0FF106B5AFF15" box="[733,808,187,213]" italics="true" pageId="20" pageNumber="21">barthi,</emphasis>
|
||
‘
|
||
<emphasis id="B949EAD5FF96FFBCFC16FF106BC0FF15" box="[827,946,187,213]" italics="true" pageId="20" pageNumber="21">C.’ bifrons</emphasis>
|
||
and ‘
|
||
<emphasis id="B949EAD5FF96FFBCFCDAFF106C02FF14" box="[1015,1136,187,213]" italics="true" pageId="20" pageNumber="21">C.’ roeveri</emphasis>
|
||
, from the Rupelian of the North Sea. Several North Sea species such as ‘
|
||
<taxonomicName id="4C3D4D44FF96FFBCFD88FF746B61FF39" box="[677,787,223,249]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFD88FF746B61FF39" box="[677,787,223,249]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B949EAD5FF96FFBCFC08FF746BDDFF39" box="[805,943,223,249]" italics="true" pageId="20" pageNumber="21">struckmanni</emphasis>
|
||
and ‘
|
||
<emphasis id="B949EAD5FF96FFBCFCC2FF746CD9FF39" box="[1007,1195,223,249]" italics="true" pageId="20" pageNumber="21">C.’ millegranosa</emphasis>
|
||
, however, are clearly unrelated to
|
||
<taxonomicName id="4C3D4D44FF96FFBCFE0FFEA869E2FEDD" box="[290,400,259,285]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFE0FFEA869E2FEDD" box="[290,400,259,285]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
based on their morphologies (see
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFC21FEA86BA7FEDD" author="von Koenen, A." box="[780,981,259,285]" pageId="20" pageNumber="21" pagination="281 - 574" refId="ref146253" refString="von Koenen, A. (1890) Das Norddeutsche Unter-Oligocan und seine Molluskenfauna. Conidae. Abhandlungen zur Geologischen Specialkarte von Preussen und den Thuringischen Staaten, 10, 281 - 574, pls. 24 - 34. [https: // www. biodiversitylibrary. org / item / 110903 # page / 725 / mode / 1 up]" type="journal article" year="1890">von Koenen 1890</bibRefCitation>
|
||
, pl. 33, figs 8, 12). Similarly, ‘
|
||
<taxonomicName id="4C3D4D44FF96FFBCFA0AFEA86DE7FEDD" box="[1319,1429,259,285]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFA0AFEA86DE7FEDD" box="[1319,1429,259,285]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B949EAD5FF96FFBCFFBAFE8C6921FE80" box="[151,339,295,320]" italics="true" pageId="20" pageNumber="21">mogenstrupensis</emphasis>
|
||
Schnetler in
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFEC3FE8C6B79FE81" author="Schnetler, K. I. & Beyer, C." box="[494,779,295,321]" pageId="20" pageNumber="21" pagination="39 - 81" refId="ref143707" refString="Schnetler, K. I. & Beyer, C. (1990) A Late Oligocene (Chattian B) Molluscan Fauna from the Coastal Cliff at Mogenstrup, North of Skive, Jutland, Denmark. Contributions to Tertiary and Quaternary Geology, 27, 39 - 81." type="journal article" year="1990">Schnetler & Beyer, 1990</bibRefCitation>
|
||
, from the Chattian of
|
||
<collectingCountry id="F32A7657FF96FFBCFB26FE8C6C04FE81" box="[1035,1142,295,321]" name="Denmark" pageId="20" pageNumber="21">Denmark</collectingCountry>
|
||
, is unrelated to
|
||
<taxonomicName id="4C3D4D44FF96FFBCFA03FE8C6DEEFE81" box="[1326,1436,295,321]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFA03FE8C6DEEFE81" box="[1326,1436,295,321]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
based on its convex whorls and broad axial ribs (see
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFDC6FEE06B8DFEA5" author="Schnetler, K. I. & Beyer, C." box="[747,1023,331,357]" pageId="20" pageNumber="21" pagination="39 - 81" refId="ref143707" refString="Schnetler, K. I. & Beyer, C. (1990) A Late Oligocene (Chattian B) Molluscan Fauna from the Coastal Cliff at Mogenstrup, North of Skive, Jutland, Denmark. Contributions to Tertiary and Quaternary Geology, 27, 39 - 81." type="journal article" year="1990">Schnetler & Beyer 1990</bibRefCitation>
|
||
, pl. 3 fig. 5). ‘
|
||
<taxonomicName id="4C3D4D44FF96FFBCFB8CFEE06D7DFEA5" box="[1185,1295,331,357]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFB8CFEE06D7DFEA5" box="[1185,1295,331,357]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B949EAD5FF96FFBCFA0DFEE06DEEFEA4" box="[1312,1436,331,356]" italics="true" pageId="20" pageNumber="21">apenninica</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFFBAFEC4694EFE49" author="Bellardi, L" box="[151,316,367,393]" pageId="20" pageNumber="21" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi, 1877</bibRefCitation>
|
||
, from the Rupelian of
|
||
<collectingCountry id="F32A7657FF96FFBCFD12FEC46A01FE49" box="[575,627,367,393]" name="Italy" pageId="20" pageNumber="21">Italy</collectingCountry>
|
||
, seems to represent a
|
||
<taxonomicName id="4C3D4D44FF96FFBCFC43FEC46B8CFE49" authorityName="Gray" authorityYear="1853" box="[878,1022,367,393]" class="Gastropoda" family="Clavatulidae" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="family">Clavatulidae</taxonomicName>
|
||
, but does not fit in
|
||
<taxonomicName id="4C3D4D44FF96FFBCFBF2FEC46D3FFE49" box="[1247,1357,367,393]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFBF2FEC46D3FFE49" box="[1247,1357,367,393]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
due to its comparatively slender biconic shell with high last whorl and weak subsutural collar (see
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFB8AFE386887FE11" author="Ferrero Mortara, E. & Montefameglio, L. & Pavia, G. & Tampieri, R." pageId="20" pageNumber="21" pagination="1 - 327" refId="ref131938" refString="Ferrero Mortara, E., Montefameglio, L., Pavia, G. & Tampieri, R. (1981) Catalogo dei tipi e degli esemplari figurati della collezione Bellardi e Sacco. Parte I. Museo Regionale di Scienze Naturali, Catalogo, 6, 1 - 327. [https: // www. biodiversitylibrary. org / item / 269998 # page / 5 / mode / 1 up]" type="journal article" year="1981">
|
||
Ferrero Mortara
|
||
<emphasis id="B949EAD5FF96FFBCFA4EFE386DE7FE6D" box="[1379,1429,403,429]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
|
||
. 1981:77
|
||
</bibRefCitation>
|
||
, pl. 15, figs 1a–b).
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF96FFBCFFEAFE706D62FDA1" blockId="20.[151,1437,151,1005]" pageId="20" pageNumber="21">
|
||
During the Chattian, the ‘
|
||
<taxonomicName id="4C3D4D44FF96FFBCFEC6FE706A2BFE35" box="[491,601,475,501]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFEC6FE706A2BFE35" box="[491,601,475,501]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’ species group was widespread along European coasts, documented from the northeastern Atlantic [‘
|
||
<taxonomicName id="4C3D4D44FF96FFBCFEEAFE546A47FDD9" box="[455,565,511,537]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFEEAFE546A47FDD9" box="[455,565,511,537]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF96FFBCFD6AFE546BD1FDD9" authority="(Grateloup, 1832)" baseAuthorityName="Grateloup" baseAuthorityYear="1832" box="[583,931,510,537]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="concatenata">
|
||
<emphasis id="B949EAD5FF96FFBCFD6AFE546AA2FDD8" box="[583,720,511,536]" italics="true" pageId="20" pageNumber="21">concatenata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFDF2FE546BEEFDD9" author="Grateloup, J. P. S. de" box="[735,924,510,537]" pageId="20" pageNumber="21" pagination="314 - 344" refId="ref133310" refString="Grateloup, J. P. S. de (1832) Tableau des coquilles Fossiles qu'on rencontre dans les terrains calcaires tertiaries (faluns) des environs de Dax, dans le departement des Landes. Actes de la Societe Linneenne de Bordeaux, 5, 314 - 344. [https: // www. biodiversitylibrary. org / item / 102890 # page / 325 / mode / 1 up]" type="journal article" year="1832">Grateloup, 1832</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
] and the Central Paratethys Sea (‘
|
||
<taxonomicName id="4C3D4D44FF96FFBCFA0AFE546DE7FDD9" box="[1319,1429,511,537]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFA0AFE546DE7FDD9" box="[1319,1429,511,537]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B949EAD5FF96FFBCFFBAFD886899FDFD" box="[151,235,547,573]" italics="true" pageId="20" pageNumber="21">danuvii</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFFDCFD886A4EFDFD" author="Vician, Z. & Kovacs, Z. & Stein, G." box="[241,572,547,573]" pageId="20" pageNumber="21" pagination="149 - 162" refId="ref146137" refString="Vician, Z., Kovacs, Z. & Stein, G. (2019) Addition to the Egerian (late Oligocene - early Miocene) gastropod fauna of Esztergom (Hungary). Foldtani Kozlony, 149, 149 - 162. https: // doi. org / 10.23928 / foldt. kozl. 2019.149.2.149" type="journal article" year="2019">Vicián, Kovács & Stein, 2019</bibRefCitation>
|
||
) (
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFD7DFD886A9DFDFD" author="Lozouet, P." box="[592,751,547,573]" pageId="20" pageNumber="21" pagination="3 - 179" refId="ref137939" refString="Lozouet, P. (2017) Les Conoidea de l'Oligocene superieur (Chattien) du bassin de l'Adour (Sud-Ouest de la France). Cossmanniana, 19, 3 - 179." type="journal article" year="2017">Lozouet 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFDD4FD886BB3FDFD" author="Vician, Z. & Kovacs, Z. & Stein, G." box="[761,961,547,573]" pageId="20" pageNumber="21" pagination="149 - 162" refId="ref146137" refString="Vician, Z., Kovacs, Z. & Stein, G. (2019) Addition to the Egerian (late Oligocene - early Miocene) gastropod fauna of Esztergom (Hungary). Foldtani Kozlony, 149, 149 - 162. https: // doi. org / 10.23928 / foldt. kozl. 2019.149.2.149" type="journal article" year="2019">
|
||
Vicián
|
||
<emphasis id="B949EAD5FF96FFBCFC6AFD886B0CFDFD" box="[839,894,547,573]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
). The North Sea species
|
||
<taxonomicName id="4C3D4D44FF96FFBCFBE3FD886921FDA1" authority="R. Janssen, 1978" authorityName="R. Janssen" authorityYear="1978" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="chattica">
|
||
<emphasis id="B949EAD5FF96FFBCFBE3FD886DEEFDFD" box="[1230,1436,547,573]" italics="true" pageId="20" pageNumber="21">Clavatula chattica</emphasis>
|
||
R.
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFF95FDEC6921FDA1" author="Janssen, R." box="[184,339,583,609]" pageId="20" pageNumber="21" pagination="3 - 195" refId="ref135850" refString="Janssen, R. (1978) Die Scaphopoden und Gastropoden des Kasseler Meeressandes von Glimmerode (Niederhessen). Geologisches Jahrbuch A, 41, 3 - 195." type="journal article" year="1978">Janssen, 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C3D4D44FF96FFBCFECAFDEC6A27FDA1" box="[487,597,583,609]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFECAFDEC6A27FDA1" box="[487,597,583,609]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
in having convex whorls and axial ribs (see R.
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFB46FDEC6D71FDA1" author="Janssen, R." box="[1131,1283,583,609]" pageId="20" pageNumber="21" pagination="3 - 195" refId="ref135850" refString="Janssen, R. (1978) Die Scaphopoden und Gastropoden des Kasseler Meeressandes von Glimmerode (Niederhessen). Geologisches Jahrbuch A, 41, 3 - 195." type="journal article" year="1978">Janssen 1978</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF96FFBCFFEAFDC06C92FD31" blockId="20.[151,1437,151,1005]" pageId="20" pageNumber="21">
|
||
Species from the Miocene of the Western Atlantic, placed by
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFCAAFDC06C65FD45" author="Gabb, W. M." box="[903,1047,619,645]" pageId="20" pageNumber="21" pagination="270 - 274" refId="ref132742" refString="Gabb, W. M. (1873) Description of some new genera of Mollusca. Proceedings of the Academy of Natural Sciences of Philadelphia, 24 / 3, 270 - 274. [https: // www. biodiversitylibrary. org / item / 84756 # page / 298 / mode / 1 up]" type="journal article" year="1873">Gabb (1873)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFB7CFDC06C8DFD45" author="Gardner, J. A." box="[1105,1279,619,645]" pageId="20" pageNumber="21" pagination="493 - 656" refId="ref132892" refString="Gardner, J. A. (1947) The molluscan fauna of the Alum Bluff Group of Florida. Part VIII. Ctenobranchia (remainder), Aspidobranchia, and Scaphopoda. United States Geological Survey Professional Paper, 142 H, 493 - 656, pls. 52 - 62. [https: // pubs. usgs. gov / pp / 0142 f / report. pdf] https: // doi. org / 10.3133 / pp 142 H" type="journal article" year="1947">Gardner (1947)</bibRefCitation>
|
||
in
|
||
<taxonomicName id="4C3D4D44FF96FFBCFA0AFDC06DE7FD45" box="[1319,1429,619,645]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFA0AFDC06DE7FD45" box="[1319,1429,619,645]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
, are clearly unrelated with
|
||
<taxonomicName id="4C3D4D44FF96FFBCFE96FD246A3BFD69" authorityName="Gray" authorityYear="1853" box="[443,585,655,681]" class="Gastropoda" family="Clavatulidae" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="family">Clavatulidae</taxonomicName>
|
||
and represent rather members of the
|
||
<taxonomicName id="4C3D4D44FF96FFBCFCC7FD246CBEFD69" authorityName="J. P. E. Morrison" authorityYear="1966" box="[1002,1228,655,681]" class="Gastropoda" family="Pseudomelatomidae" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="family">Pseudomelatomidae</taxonomicName>
|
||
or other
|
||
<taxonomicName id="4C3D4D44FF96FFBCFA1CFD246DEEFD69" authorityName="J. Fleming" authorityYear="1822" box="[1329,1436,655,681]" class="Gastropoda" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="12" phylum="Mollusca" rank="superFamily" superFamily="Conoidea">Conoidea</taxonomicName>
|
||
families. Similarly, ‘
|
||
<taxonomicName id="4C3D4D44FF96FFBCFEABFD186986FD0D" box="[390,500,691,717]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFEABFD186986FD0D" box="[390,500,691,717]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’ species described from the Neogene of the Indo-West Pacific, by
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFA3DFD186DE5FD0E" author="Cox, L. R." box="[1296,1431,691,718]" pageId="20" pageNumber="21" pagination="1 - 69" refId="ref130729" refString="Cox, L. R. (1936) Fossil Mollusca from southern Persia (Iran) and Bahrein Island. Memoirs of the Geological Survey of India, Palaeontologia Indica, New Series, 22, 1 - 69." type="journal article" year="1936">Cox (1936)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFFBAFD7C693DFD31" author="Oostingh, C. H." box="[151,335,727,753]" pageId="20" pageNumber="21" pagination="17 - 46" refId="ref140756" refString="Oostingh, C. H. (1938) Die Mollusken des Pliocans von Sud-Bantam in Java. I. Abschnitt (Gastropoda I). De Ingenieur in Nederlandsch-Indie, IV. Mijnbouw en Geologie. " De Mijningenieur ", 5, 17 - 46." type="journal article" year="1938">Oostingh (1938)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFEABFD7C6A7DFD31" author="Beets, C." box="[390,527,727,753]" pageId="20" pageNumber="21" pagination="1 - 220" refId="ref129167" refString="Beets, C. (1941) Eine jungmiocane Mollusken-Fauna von der Halbinsei Mangkalihat, Ost-Borneo. Verhandelingen van het Geologisch-Mijnbouwkundig Genootschap voor Nederland en Kolonien, Geologische Serie, 13, 1 - 220." type="journal article" year="1941">Beets (1941</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFD37FD7C6A21FD31" author="Beets, C." box="[538,595,727,753]" pageId="20" pageNumber="21" pagination="255 - 328" refId="ref129203" refString="Beets, C. (1942) Beitrage zur Kenntniss der Angeblich Oberoligocanen Mollusken-Fauna der Insel Buton, Niederlandisch- Ostindien. Leidsche Geologische Mededeelingen, 13, 255 - 328. [https: // repository. naturalis. nl / pub / 505965]" type="journal article" year="1942">1942</bibRefCitation>
|
||
) represent other genera such as
|
||
<taxonomicName id="4C3D4D44FF96FFBCFC96FD7C6C4BFD31" authorityName="Makiyama" authorityYear="1940" box="[955,1081,727,753]" class="Gastropoda" family="Horaiclavidae" genus="Paradrillia" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFC96FD7C6C4BFD31" box="[955,1081,727,753]" italics="true" pageId="20" pageNumber="21">Paradrillia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3D4D44FF96FFBCFB5DFD7C6CA8FD31" authorityName="Schumacher" authorityYear="1817" box="[1136,1242,727,753]" class="Magnoliopsida" family="Namaceae" genus="Turricula" kingdom="Plantae" order="Boraginales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFB5DFD7C6CA8FD31" box="[1136,1242,727,753]" italics="true" pageId="20" pageNumber="21">Turricula</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF96FFBCFFEAFD506D07FC41" blockId="20.[151,1437,151,1005]" pageId="20" pageNumber="21">
|
||
Therefore, the ‘
|
||
<taxonomicName id="4C3D4D44FF96FFBCFE59FD506990FCD5" box="[372,482,763,789]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFE59FD506990FCD5" box="[372,482,763,789]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’ genus group is strictly Tethyan. It was widely distributed along the European Atlantic and Proto-Mediterranean coasts during the Oligocene. The group peaked in distribution and diversity during the Miocene, documented by numerous species from all European biogeographic regions except for the Eastern Paratethys (e.g.,
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFE7EFCCC699DFC41" author="Bellardi, L" box="[339,495,871,897]" pageId="20" pageNumber="21" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi 1877</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFED6FCCC6AE8FC41" author="Kautsky, F." box="[507,666,870,897]" pageId="20" pageNumber="21" pagination="1 - 255" refId="ref136369" refString="Kautsky, F. (1925) Das Miocan von Hemmoor und Basbek-Osten. Abhandlungen der Preussischen Geologischen Landesanstalt, Neue Folge, 97, 1 - 255." type="journal article" year="1925">Kautsky 1925</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFD8BFCCC6B45FC41" author="Glibert, M." box="[678,823,870,897]" pageId="20" pageNumber="21" pagination="1 - 75" refId="ref133027" refString="Glibert, M. (1954) Pleurotomes du Miocene de la Belgique et du bassin de la Loire. Memoires de l'Institut Royal des Sciences Naturelles de Belgique, 129, 1 - 75, 7 pls." type="journal article" year="1954">Glibert 1954</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFC6EFCCC6BBCFC41" author="Peyrot, A." box="[835,974,871,897]" pageId="20" pageNumber="21" pagination="5 - 288" refId="ref141284" refString="Peyrot, A. (1931 & 1933) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 83, 5 - 116 (1931), 84, 5 - 288 (1933). [https: // www. biodiversitylibrary. org / item / 185325 # page / 11 / mode / 1 up]" type="journal article" year="1931">Peyrot 1931</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFCF7FCCC6CDDFC41" author="Landau, B. M. & Harzhauser, M. & Yslamodlu, Y. & Marques da Silva, C." box="[986,1199,871,897]" pageId="20" pageNumber="21" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. & Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
|
||
Landau
|
||
<emphasis id="B949EAD5FF96FFBCFB18FCCC6C14FC41" box="[1077,1126,871,897]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAC4B36FF96FFBCFB97FCCC6C84FC41" author="Landau, B. M. & van Dingenen, F. & Ceulemans, L." box="[1210,1270,871,897]" pageId="20" pageNumber="21" pagination="3 - 107" refId="ref137618" refString="Landau, B. M., van Dingenen, F. & Ceulemans, L. (2020) The upper Miocene gastropods of northwestern France, 5. Conoidea. Cainozoic Research, 20, 3 - 107." type="journal article" year="2020">2020</bibRefCitation>
|
||
;
|
||
<emphasis id="B949EAD5FF96FFBCFA2DFCCC6D17FC40" box="[1280,1381,871,897]" italics="true" pageId="20" pageNumber="21">hoc opus</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF96FFBCFFEAFC206C2FFC2D" blockId="20.[151,1437,151,1005]" pageId="20" pageNumber="21">
|
||
After the late Pliocene cooling event, the group suffered a gradual range contraction, and today
|
||
<taxonomicName id="4C3D4D44FF96FFBCFBD3FC206D1EFC65" box="[1278,1388,907,933]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFBD3FC206D1EFC65" box="[1278,1388,907,933]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
and other ‘
|
||
<taxonomicName id="4C3D4D44FF96FFBCFFCCFC04693DFC09" box="[225,335,943,969]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="21" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF96FFBCFFCCFC04693DFC09" box="[225,335,943,969]" italics="true" pageId="20" pageNumber="21">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’-clades are restricted to tropical waters off West Africa, where they are represented by a relatively small number of genus groups and species compared to their older historical diversity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |