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<document id="71610A90CCF595054E1C44871B653061" ID-CLB-Dataset="20492" ID-DOI="10.5169/seals-787049" ID-GBIF-Dataset="ce0756e2-5184-4b74-8472-ab69fed7951c" ID-ISSN="2624-9170" ID-Zenodo-Dep="6285683" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637935326602" checkinUser="carolina" docAuthor="Huber, Charles &amp; Schmidt, Joachim" docDate="2017" docId="03948799FFF4FFF2FF2BFC6FFE6CFCAE" docLanguage="en" docName="ContribNatHist.36.1-85.pdf.imf" docOrigin="Contributions to Natural History 36" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Nebria (Epinebriola) tuberculata Huber &amp; Schmidt, 2017, sp. nov." docType="treatment" docVersion="12" lastPageNumber="51" masterDocId="FFADFFE1FFDBFFC0FF9CFFE9FFB0FFA7" masterDocTitle="Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)" masterLastPageNumber="85" masterPageNumber="1" pageNumber="48" updateTime="1699470525451" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="80534A218BFD58AF67CE9EC71590FFDA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A47257DAB51D195B408A77482553B90D">Notes on the Nebria subgenus Epinebriola K. DANIEL, 1904 with the description of Barbonebriola subgen. nov. and 13 new species from the Himalaya-Tibet orogen (Coleoptera, Carabidae, Nebriini)</mods:title>
</mods:titleInfo>
<mods:name id="E60500E4378368913E9108279833ABCA" type="personal">
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<mods:roleTerm id="C602CE0F17E5DBCE41BCD8B9080B5266">Author</mods:roleTerm>
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<mods:namePart id="5551518930C3903BFC4C9CD975BCF9BD">Huber, Charles</mods:namePart>
</mods:name>
<mods:name id="CB46C56EE3BF53A01C90C31D7DFA0B1E" type="personal">
<mods:role id="F9C58CC8C9507EA00871BBB746008CC6">
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</mods:role>
<mods:namePart id="71B37BE2A777B47EF881F548724407F8">Schmidt, Joachim</mods:namePart>
</mods:name>
<mods:typeOfResource id="C2FEAA0AC7C0A809A6B3EB753718BE65">text</mods:typeOfResource>
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<mods:title id="A8C3335CD1F4C1961F0A723B49C6B4F7">Contributions to Natural History</mods:title>
</mods:titleInfo>
<mods:part id="4167B71D76B1516B17AD478FAEDDE921">
<mods:date id="E0828F88D647C155936A258A892EAD52">2017</mods:date>
<mods:detail id="877FA7D36347CF3784B44386820CC11E" type="pubDate">
<mods:number id="1C496C6DFC95DB65D03A23628681E709">2017-07-17</mods:number>
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<mods:detail id="77E2EC2AFE5A5856BB335E59363FAE0E" type="volume">
<mods:number id="1CC0C170D044F6932F2DE7AACAB8EF31">36</mods:number>
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<mods:start id="6627D7E95DAE7B237E70066D3783AB1A">1</mods:start>
<mods:end id="C8B4D56FF5B542729BD98C87A98827F6">85</mods:end>
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<mods:classification id="EB421B2970BFFE94F62401A1855AC08B">journal article</mods:classification>
<mods:identifier id="71474F82186DD24224D52AD279D6F909" type="CLB-Dataset">20492</mods:identifier>
<mods:identifier id="86E13C57FCDA994BC2BE5C04BB20D33F" type="DOI">10.5169/seals-787049</mods:identifier>
<mods:identifier id="43CAF67D3C85CFADC9F33ECF570E8F73" type="GBIF-Dataset">ce0756e2-5184-4b74-8472-ab69fed7951c</mods:identifier>
<mods:identifier id="351B472687054B3C2548560408444C8D" type="ISSN">2624-9170</mods:identifier>
<mods:identifier id="BDF014071F6CC137E57FAE5307F06DA6" type="Zenodo-Dep">6285683</mods:identifier>
</mods:mods>
<treatment id="03948799FFF4FFF2FF2BFC6FFE6CFCAE" ID-DOI="http://doi.org/10.5281/zenodo.6315668" ID-GBIF-Taxon="193820247" ID-Zenodo-Dep="6315668" LSID="urn:lsid:plazi:treatment:03948799FFF4FFF2FF2BFC6FFE6CFCAE" httpUri="http://treatment.plazi.org/id/03948799FFF4FFF2FF2BFC6FFE6CFCAE" lastPageId="50" lastPageNumber="51" pageId="47" pageNumber="48">
<subSubSection id="C3276504FFF4FFEFFF2BFC6FFD58FC39" box="[183,744,902,927]" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="8B82368FFFF4FFEFFF2BFC6FFD58FC39" blockId="47.[183,744,902,927]" box="[183,744,902,927]" pageId="47" pageNumber="48">
<heading id="D0CA81E3FFF4FFEFFF2BFC6FFD58FC39" box="[183,744,902,927]" centered="true" fontSize="12" level="2" pageId="47" pageNumber="48" reason="2">
<emphasis id="B949EA9DFFF4FFEFFF2BFC6FFD35FC38" bold="true" box="[183,645,902,927]" pageId="47" pageNumber="48">
<taxonomicName id="4C3D4D0CFFF4FFEFFF2BFC6FFD97FC38" ID-CoL="8MNFP" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[183,551,902,927]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov." subGenus="Epinebriola">
<emphasis id="B949EA9DFFF4FFEFFF2BFC6FFEB3FC38" bold="true" box="[183,259,902,927]" italics="true" pageId="47" pageNumber="48">Nebria</emphasis>
(
<emphasis id="B949EA9DFFF4FFEFFE8FFC6FFE25FC38" bold="true" box="[275,405,902,927]" italics="true" pageId="47" pageNumber="48">Epinebriola</emphasis>
)
<emphasis id="B949EA9DFFF4FFEFFE39FC6FFD97FC38" bold="true" box="[421,551,902,927]" italics="true" pageId="47" pageNumber="48">tuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFF4FFEFFDB1FC6FFD35FC38" box="[557,645,902,927]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13062A0AFFF4FFEFFD0DFC6FFD6EFC39" box="[657,734,902,926]" captionStart="Fig" captionStartId="48.[223,251,160,178]" captionTargetBox="[463,1106,153,1268]" captionTargetId="figure-104@48.[463,1106,153,1268]" captionTargetPageId="48" captionText="Fig. 24: Nebria (Epinebriola) tuberculata sp. nov. Holotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285786" httpUri="https://zenodo.org/record/6285786/files/figure.png" pageId="47" pageNumber="48">Fig. 24</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3276504FFF4FFEFFF2BFC38FDFAFB03" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="8B82368FFFF4FFEFFF2BFC38FE24FBA9" blockId="47.[183,1069,974,1636]" pageId="47" pageNumber="48">
<materialsCitation id="3B553CD2FFF4FFEFFF2BFC38FE24FBA9" ID-GBIF-Occurrence="3497686348" collectingDate="1998-08-05" collectionCode="cSCHM" collectorName="C. Berndt" country="Nepal" elevation="4600" location="Helambu" municipality="Ganja La" pageId="47" pageNumber="48" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="5486882DFFF4FFEFFF2BFC38FEAFFC4E" box="[183,287,977,1001]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
<specimenCount id="9D3BFD06FFF4FFEFFEB9FC27FE89FC4E" box="[293,313,974,1001]" pageId="47" pageNumber="48" type="female">♀</specimenCount>
:
<collectingCountry id="F32A761FFFF4FFEFFEDBFC38FE3EFC4E" box="[327,398,977,1001]" name="Nepal" pageId="47" pageNumber="48">Nepal</collectingCountry>
,
<location id="8EE26054FFF4FFEFFE0BFC38FDB1FC4E" LSID="urn:lsid:plazi:treatment:03948799FFF4FFF2FF2BFC6FFE6CFCAE:8EE26054FFF4FFEFFE0BFC38FDB1FC4E" box="[407,513,977,1001]" country="Nepal" municipality="Ganja La" name="Helambu" pageId="47" pageNumber="48">Helambu</location>
,
<location id="8EE26054FFF4FFEFFD96FC38FD7BFC4E" LSID="urn:lsid:plazi:treatment:03948799FFF4FFF2FF2BFC6FFE6CFCAE:8EE26054FFF4FFEFFD96FC38FD7BFC4E" box="[522,715,977,1001]" country="Nepal" municipality="Ganja La" name="S-slope Ganja La" pageId="47" pageNumber="48">S-slope Ganja La</location>
,
<quantity id="4CC59B6AFFF4FFEFFD48FC38FCCAFC4E" box="[724,890,977,1001]" metricMagnitude="3" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="47" pageNumber="48" unit="m" value="4600.0" valueMax="4800.0" valueMin="4400.0">
<elevation id="0010D1BCFFF4FFEFFD48FC38FCCAFC4E" box="[724,890,977,1001]" metricMagnitude="3" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="47" pageNumber="48" unit="m" value="4600.0" valueMax="4800.0" valueMin="4400.0">4400 –4800 m</elevation>
</quantity>
,
<date id="FF83104FFFF4FFEFFC18FC38FC45FC4E" box="[900,1013,977,1001]" pageId="47" pageNumber="48" value="1998-08-05">
<collectingDate id="EFC7E9A7FFF4FFEFFC18FC38FC45FC4E" box="[900,1013,977,1001]" pageId="47" pageNumber="48" value="1998-08-05">5. 8. 1998</collectingDate>
</date>
, leg.
<collectorName id="26C85359FFF4FFEFFF2BFC1FFE91FBA9" box="[183,289,1014,1038]" pageId="47" pageNumber="48">C. Berndt</collectorName>
(
<collectionCode id="ED2CAE4AFFF4FFEFFEB1FC1FFE3AFBA9" box="[301,394,1014,1038]" pageId="47" pageNumber="48">cSCHM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B82368FFFF4FFEFFF41FBF5FDFAFB03" blockId="47.[183,1069,974,1636]" pageId="47" pageNumber="48">
Body length
<quantity id="4CC59B6AFFF4FFEFFEEEFBF5FE75FB93" box="[370,453,1052,1076]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="47" pageNumber="48" unit="mm" value="12.0">12 mm</quantity>
. Colour dark brown, appendages of the head brown. Antennae brown. Legs brown, femur darkened with brown knees. Pronotum brownish lightened, slightly lighter than the black head and the elytra. 1
<superScript id="7C489BC7FFF4FFEFFC47FB8AFC5CFBD1" attach="left" box="[987,1004,1123,1142]" fontSize="8" pageId="47" pageNumber="48">st</superScript>
interval of the elytra brownish lightened.
</paragraph>
</subSubSection>
<subSubSection id="C3276504FFF4FFF1FF41FB58FB94FA23" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48" type="description">
<paragraph id="8B82368FFFF4FFEFFF41FB58FCD6FA53" blockId="47.[183,1069,974,1636]" pageId="47" pageNumber="48">
Head with two shallow oblique impressions between the eyes. Labrum with anterior margin straight, bearing six setae. Apical margin of clypeus straight. Vertex impunctate, transversally wrinkled. Eyes weakly prominent. Antennae long and slender extending to one third of the elytra. Antennal scape elongate, a little longer than the eye's diameter, basally narrowed, subcylindrical apically, with one dorsal seta. 2
<superScript id="7C489BC7FFF4FFEFFD17FA81FD12FADC" attach="left" box="[651,674,1384,1403]" fontSize="8" pageId="47" pageNumber="48">nd</superScript>
antennomere with one seta ventroapically. Maxillary stipes normal. Penultimate labial palpomere trisetose. Ligula spiny, with two lateral setae. Mentum with bifid medial tooth. Submentum with a row of 14 setae. Microreticulation of the head isodiametric.
</paragraph>
<paragraph id="8B82368FFFF4FFF1FF41F9E8FCF9FE79" blockId="47.[183,1069,974,1636]" lastBlockId="49.[184,1069,156,1598]" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48">
Pronotum faintly transverse, ratio width/length of the pronotum = 1.26. Lateral margin distinctly convex, narrowed basally, widely rounded to the anterior angles, convexly narrowed to the posterior angles, concavely sinuate just in front of the small posterior angle (
<figureCitation id="13062A0AFFEBFFF0FD32FAAFFCBEFAF9" box="[686,782,1350,1374]" captionStart="Fig" captionStartId="61.[184,213,1317,1335]" captionTargetBox="[185,1068,153,1276]" captionTargetId="figure-106@61.[551,733,151,716]" captionTargetPageId="61" captionText="Fig. 31: Left part of the pronotum of Epinebriola species. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) tuberculata sp. nov.; F: N. (E.) numburica sp. nov.; G: N. (E.) retingensis sp. nov.; H: N. (E.) delicata sp. nov.; I: N. (E.) incarinata sp. nov.; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6285816" httpUri="https://zenodo.org/record/6285816/files/figure.png" pageId="48" pageNumber="49">Fig. 31E</figureCitation>
). Lateral explanation narrow with a conspicuous groove also apically, broadened basally. Lateral margin basally blade-like, obliquely upturned. Posterior angles acute but very small, slightly protruding as a tubercle (
<figureCitation id="13062A0AFFEBFFF0FDCAFA5FFD13FA69" box="[598,675,1462,1486]" captionStart="Fig" captionStartId="50.[223,252,160,178]" captionTargetBox="[463,1106,154,605]" captionTargetId="figure-276@50.[463,1106,154,606]" captionTargetPageId="50" captionText="Fig. 25: Acutely pro- truding posterior angle (arrow) of the pronotum of Nebria (Epinebriola) tuberculata sp. nov. in dorso-lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285790" httpUri="https://zenodo.org/record/6285790/files/figure.png" pageId="48" pageNumber="49">Fig. 25</figureCitation>
). Anterior angles widely rounded, not protruding. Basal margin bisinuate, with a faint incision between the basal fovea and the small posterior angles. Pronotal disc weakly convex. Basal fovea deep, anterior and posterior transverse impression distinct; median line shallow, faintly transversally wrinkled. Lateral groove, basal fovea, anterior and posterior transverse sulci impunctate, only the area between the apical margin and the anterior transverse impression faintly punctate. Apical margination of the pronotum restricted to lateral one-fourths, merges with a faint ridge at the anterior angle; the ridge continues along the lateral groove to the lateral seta. Base of the pronotum not margined. Basolateral seta present. 1–2 midlateral setae in the apical half of the pronotum, inserted in distinct grooves at the inside of the slightly upturned margin. Microreticulation of the pronotum isodiametric, impunctate on disc. Proepisternum smooth. Prosternal process laterally margined, apex bulging and unmargined.
</paragraph>
<caption id="DF426607FFEBFFF0FF43FF49FECAFEB2" ID-DOI="http://doi.org/10.5281/zenodo.6285786" ID-Zenodo-Dep="6285786" httpUri="https://zenodo.org/record/6285786/files/figure.png" pageId="48" pageNumber="49" startId="48.[223,251,160,178]" targetBox="[463,1106,153,1268]" targetPageId="48">
<paragraph id="8B82368FFFEBFFF0FF43FF49FECAFEB2" blockId="48.[221,392,160,277]" pageId="48" pageNumber="49">
Fig. 24:
<taxonomicName id="4C3D4D0CFFEBFFF0FEBFFF49FEFEFF6D" authority="(Epinebriola)" authorityName="LATREILLE" authorityYear="1802" baseAuthorityName="Epinebriola" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
Nebria
<emphasis id="B949EA9DFFEBFFF0FF41FF51FEFEFF6D" bold="true" box="[221,334,184,202]" pageId="48" pageNumber="49">
(
<emphasis id="B949EA9DFFEBFFF0FF78FF51FEF7FF6D" bold="true" box="[228,327,184,202]" italics="true" pageId="48" pageNumber="49">Epinebriola</emphasis>
)
</emphasis>
</taxonomicName>
<emphasis id="B949EA9DFFEBFFF0FF41FF39FECAFEB2" bold="true" pageId="48" pageNumber="49">
<taxonomicName id="4C3D4D0CFFEBFFF0FF41FF39FE8FFF45" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[221,319,208,226]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="B949EA9DFFEBFFF0FF41FF39FE8FFF45" bold="true" box="[221,319,208,226]" italics="true" pageId="48" pageNumber="49">tuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFEBFFF0FED8FF39FE38FF45" box="[324,392,208,226]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
<typeStatus id="5486882DFFEBFFF0FF41FF01FE9DFF5D" box="[221,301,232,250]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
. Scale bar =
<quantity id="4CC59B6AFFEBFFF0FED9FEEAFEC6FEB2" box="[325,374,259,277]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="48" pageNumber="49" unit="mm" value="5.0">5 mm</quantity>
.
</emphasis>
</paragraph>
</caption>
<paragraph id="8B82368FFFEAFFF1FF41FE05FE27FCF3" blockId="49.[184,1069,156,1598]" pageId="49" pageNumber="50">Elytral silhouette convex, regularly ovoid-elongate, widest slightly behind the middle. Ratio length/width of the elytra = 1.60. 1.7 times wider than the pronotum. Elytral apex rounded, apical carina absent. Basal margination straight, smoothly merged with the lateral margination. Humeral carina faintly developed. Striae distinct on disc, coarsely and tightly punctate. Striae obliterate towards the apex. Intervals rather convex, more convex on disc. Interval 3 with 7–8 setae adjoining stria 3. Scutellar seta absent. Microreticulation oblong. Mesepisterna smooth. Metepisterna 1.8 times as long as wide, smooth. Metacoxa with 4–5 basal and 1–2 apical setae. Hindwings present as a strap-like vestige.</paragraph>
<paragraph id="8B82368FFFEAFFF1FF41FC88FEE2FC63" blockId="49.[184,1069,156,1598]" pageId="49" pageNumber="50">Sternum 3 with one seta, faintly transversally wrinkled. Sterna 4–6 each with 5–7 posterior paramedial setae. Anal sternum with three paramedial setae in the female.</paragraph>
<paragraph id="8B82368FFFEAFFF1FF41FC38FC68FBFE" blockId="49.[184,1069,156,1598]" pageId="49" pageNumber="50">Legs long and slender. All tarsi dorsally glabrous. Metatarsomere 4 ventroapically markedly projected. Meso- and metatarsomeres 2–4 ventroapically with long setae. All tarsomeres 5 ventrally with two rows of 3–4 short setae. Metatarsomere 5 long, as long or longer than the metatarsomeres 3+4.</paragraph>
<paragraph id="8B82368FFFEAFFF1FF41FB8FFD46FAB3" blockId="49.[184,1069,156,1598]" pageId="49" pageNumber="50">
Female genitalia: Gonocoxa (
<figureCitation id="13062A0AFFEAFFF1FDB2FB8FFD23FBD9" box="[558,659,1126,1150]" captionStart="Fig" captionStartId="20.[221,250,1430,1448]" captionTargetBox="[228,1106,153,1365]" captionTargetId="figure-16@20.[574,757,751,1260]" captionTargetPageId="20" captionText="Fig. 10: Right gonocoxae of Epinebriola species, in ventral view. A: Nebria (Epinebriola) triseriata sp. nov.; B: N. (E.) rupina sp. nov.; C: N. (E.) montisanimae sp. nov.; D: N. (E.) impunctata sp. nov.; E: N. (E.) tuberculata sp. nov.; F: N. (E.) numburica sp. nov.; G: N. (E.) numburica sp. nov. in lateral view with the distale gonocoxite 2 dorsally grooved (arrow); H: N. (E.) christinae HUBER &amp; SCHMIDT, 2007; I: N. (E.) pseudorestias sp. nov.; J: N. (E.) retingensis sp. nov.; Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6285726" httpUri="https://zenodo.org/record/6285726/files/figure.png" pageId="49" pageNumber="50">Fig. 10E</figureCitation>
): Gonocoxite 2 elongate-triangular, broad at base, half the length of gonocoxite 1. Apex dorsally arcuate, dorsally distinctly grooved. Ventral preapical insertion furrow elongate-oval, with two nematiforme setae. Gonocoxites ventrally unjointed and continuously sclerotized, dorsally separated by a membranous area.
</paragraph>
<paragraph id="8B82368FFFEAFFF1FF41FAC8FE3BFA9E" blockId="49.[184,1069,156,1598]" box="[221,395,1313,1337]" pageId="49" pageNumber="50">Male unknown.</paragraph>
<paragraph id="8B82368FFFEAFFF1FF41FAAFFB94FA23" blockId="49.[184,1069,156,1598]" pageId="49" pageNumber="50">Body ratios: hea.w/fro.w = 1.43; prm.w/hea.w = 1.34; prm.w/prp.w = 1.54; pra.w/prp.w = 1.07; ely.w/prm.w = 1.74; prm.w/prm.l = 1.27; ely.l/ely.w = 1.59.</paragraph>
</subSubSection>
<subSubSection id="C3276504FFEAFFF1FF41FA78FBB4FA69" pageId="49" pageNumber="50" type="etymology">
<paragraph id="8B82368FFFEAFFF1FF41FA78FBB4FA69" blockId="49.[184,1069,156,1598]" pageId="49" pageNumber="50">
Etymology: The name
<taxonomicName id="4C3D4D0CFFEAFFF1FE46FA78FDE9FA0E" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[474,601,1425,1449]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B949EA9DFFEAFFF1FE46FA78FDE9FA0E" box="[474,601,1425,1449]" italics="true" pageId="49" pageNumber="50">tuberculata</emphasis>
</taxonomicName>
(
<emphasis id="B949EA9DFFEAFFF1FDF6FA78FD59FA0E" box="[618,745,1425,1449]" italics="true" pageId="49" pageNumber="50">tuberculum</emphasis>
, latin = small protuberance) refers to the acutely protruding tip of the posterior angles of the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C3276504FFEAFFF1FF41FA35FBACF999" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="8B82368FFFEAFFF1FF41FA35FBACF999" blockId="49.[184,1069,156,1598]" pageId="49" pageNumber="50">
Diagnosis:
<taxonomicName id="4C3D4D0CFFEAFFF1FEFFFA35FDBCFA53" authorityName="C.Huber &amp; J.Schmidt" authorityYear="2017" box="[355,524,1500,1524]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="B949EA9DFFEAFFF1FEFFFA35FDBCFA53" box="[355,524,1500,1524]" italics="true" pageId="49" pageNumber="50">N. tuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E6FFEAFFF1FD8AFA35FDC4FA53" box="[534,628,1500,1524]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
is recognizable within the subgenus
<taxonomicName id="4C3D4D0CFFEAFFF1FF24F9E8FE8AF9BE" authorityName="K. DANIEL" authorityYear="1904" box="[184,314,1537,1561]" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="subGenus" subGenus="Epinebriola">
<emphasis id="B949EA9DFFEAFFF1FF24F9E8FE8AF9BE" box="[184,314,1537,1561]" italics="true" pageId="49" pageNumber="50">Epinebriola</emphasis>
</taxonomicName>
by its plurisetose metacoxa with 4–5 setae, its plurisetose elytral interval 3 with 7–8 setae and its tuberculous posterior angles of the pronotum.
</paragraph>
</subSubSection>
<caption id="DF426607FFE9FFF2FF43FF49FE3AFEF5" ID-DOI="http://doi.org/10.5281/zenodo.6285790" ID-Zenodo-Dep="6285790" httpUri="https://zenodo.org/record/6285790/files/figure.png" pageId="50" pageNumber="51" startId="50.[223,252,160,178]" targetBox="[463,1106,154,605]" targetPageId="50">
<paragraph id="8B82368FFFE9FFF2FF43FF49FE3AFEF5" blockId="50.[221,430,160,338]" pageId="50" pageNumber="51">
Fig. 25: Acutely
<emphasis id="B949EA9DFFE9FFF2FEF4FF49FE3AFEF5" bold="true" pageId="50" pageNumber="51">
protruding posterior angle (arrow) of the pronotum of
<taxonomicName id="4C3D4D0CFFE9FFF2FF6FFF19FE14FEA5" box="[243,420,240,258]" pageId="50" pageNumber="51" status="sp. nov.">
<emphasis id="B949EA9DFFE9FFF2FF6FFF19FE9DFEA5" bold="true" box="[243,301,240,258]" italics="true" pageId="50" pageNumber="51">Nebria</emphasis>
(
<emphasis id="B949EA9DFFE9FFF2FEA5FF19FE2DFEA5" bold="true" box="[313,413,240,258]" italics="true" pageId="50" pageNumber="51">Epinebriola</emphasis>
)
</taxonomicName>
<emphasis id="B949EA9DFFE9FFF2FF41FEE2FEF1FEBA" bold="true" box="[221,321,267,285]" italics="true" pageId="50" pageNumber="51">tuberculata</emphasis>
<taxonomicNameLabel id="A27A57E6FFE9FFF2FED9FEE2FE39FEBA" box="[325,393,267,285]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
in dorso-lateral view. Scale bar =
<quantity id="4CC59B6AFFE9FFF2FED9FEA9FE36FEF5" box="[325,390,320,338]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="50" pageNumber="51" unit="mm" value="0.5">0.5 mm</quantity>
.
</emphasis>
</paragraph>
</caption>
<subSubSection id="C3276504FFE9FFF2FE9FFD4FFB91FD43" pageId="50" pageNumber="51" type="materials_examined">
<paragraph id="8B82368FFFE9FFF2FE9FFD4FFB91FD43" blockId="50.[221,1112,678,777]" pageId="50" pageNumber="51">
Distribution (
<figureCitation id="13062A0AFFE9FFF2FE0BFD4FFE54FD19" box="[407,484,678,702]" captionStart="Fig" captionStartId="76.[221,250,160,178]" captionTargetId="figure-8@75.[184,1068,152,1645]" captionTargetPageId="75" captionText="Fig. 41: Locations with occurrences of Nebria (Epinebriola) species from the Central and Eastern Himalaya and from South Tibet." figureDoi="http://doi.org/10.5281/zenodo.6285892" httpUri="https://zenodo.org/record/6285892/files/figure.png" pageId="50" pageNumber="51">Fig. 41</figureCitation>
): Known only from the
<typeStatus id="5486882DFFE9FFF2FD72FD4FFC91FD19" box="[750,801,678,702]" pageId="50" pageNumber="51">type</typeStatus>
locality, the pass Ganja La (= Kanja La) in the Helambu mountain range, Northwestern Central
<collectingCountry id="F32A761FFFE9FFF2FC4BFD25FBADFD43" box="[983,1053,716,740]" name="Nepal" pageId="50" pageNumber="51">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276504FFE9FFF2FE9FFD18FE6CFCAE" box="[259,476,753,777]" pageId="50" pageNumber="51" type="biology_ecology">
<paragraph id="8B82368FFFE9FFF2FE9FFD18FE6CFCAE" blockId="50.[221,1112,678,777]" box="[259,476,753,777]" pageId="50" pageNumber="51">Habitat: Not noted.</paragraph>
</subSubSection>
</treatment>
</document>
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