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<document id="CC9AED9FC8478DA95F7D0CCE7C0DEE32" ID-CLB-Dataset="158824" ID-DOI="10.5252/geodiversitas2022v44a27" ID-GBIF-Dataset="835cc227-e5c4-4453-a956-07a154f8fa5f" ID-ISSN="1638-9395" ID-Zenodo-Dep="7150329" ID-ZooBank="urn:lsid:zoobank.org:pub:E4896081-9312-4EA6-AE33-AAC44201748E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1664917095605" checkinUser="admin" docAuthor="Sandoval, José" docDate="2022" docId="0394878DFFBB7B231ACCFBF3FCE1DB6E" docLanguage="en" docName="Geodiversitas.44.27.801-851.pdf.imf" docOrigin="Geodiversitas 44 (27)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Witchellia nodatipinguis Buckman 1889" docType="treatment" docVersion="8" lastPageNumber="824" masterDocId="FFADFFF5FFA27B381A52FFC7FFE0D81E" masterDocTitle="Sonniniidae Ammonitina, Middle Jurassic from Southern Spain: taxonomic, biostratigraphical and palaeobiogeographical analysis" masterLastPageNumber="851" masterPageNumber="801" pageNumber="824" updateTime="1699387520007" updateUser="ExternalLinkService">
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<mods:title id="6FFB99DF731FD6AC01249310C58BD26E">Sonniniidae Ammonitina, Middle Jurassic from Southern Spain: taxonomic, biostratigraphical and palaeobiogeographical analysis</mods:title>
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<mods:namePart id="14F426E00FDCA871E0449D39C7F5BD79">Sandoval, José</mods:namePart>
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<mods:date id="89805D817ABD3628D5C1F67B4547464A">2022</mods:date>
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<treatment id="0394878DFFBB7B231ACCFBF3FCE1DB6E" ID-DOI="http://doi.org/10.5281/zenodo.7506337" ID-GBIF-Taxon="202146998" ID-Zenodo-Dep="7506337" LSID="urn:lsid:plazi:treatment:0394878DFFBB7B231ACCFBF3FCE1DB6E" httpUri="http://treatment.plazi.org/id/0394878DFFBB7B231ACCFBF3FCE1DB6E" lastPageId="27" lastPageNumber="824" pageId="25" pageNumber="824">
<subSubSection id="C3276510FFBB7B211ACCFBF3FDF5DC70" pageId="25" pageNumber="824" type="nomenclature">
<paragraph id="8B82369BFFBB7B211ACCFBF3FDF5DC70" blockId="25.[158,749,1076,1135]" pageId="25" pageNumber="824">
<taxonomicName id="4C3D4D18FFBB7B211ACCFBF3FD5ADC50" ID-CoL="884J" authority="(Buckman, 1923)" authorityName="Buckman" authorityYear="1889" baseAuthorityName="Buckman" baseAuthorityYear="1923" box="[158,698,1076,1103]" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="genus">
<emphasis id="B949EA89FFBB7B211ACCFBF3FEEFDC51" bold="true" box="[158,271,1076,1103]" italics="true" pageId="25" pageNumber="824">Witchellia</emphasis>
cf.
<emphasis id="B949EA89FFBB7B211B65FBF2FE14DC51" bold="true" box="[311,500,1076,1103]" italics="true" pageId="25" pageNumber="824">W. nodatipinguis</emphasis>
(Buckman, 1923)
</taxonomicName>
[M] (
<figureCitation id="13062A1EFFBB7B211B2CFB93FE34DC71" box="[382,468,1108,1135]" captionStart="FIG" captionStartId="19.[132,143,1963,1980]" captionTargetId="figure-26@20.[698,1267,768,1324]" captionTargetPageId="20" captionText="FIG. 9. — A, Sonninia (Sonninia) schlumbergeri Haug, 1893 [m], JAC4.39.10,Laeviuscula Zone,Sierra de Alta Coloma (JAC4 section); B, Sonninia (Euhoploceras) nannomorpha (Buckman, 1923), n. comb. ([m], JAC22.5.9, Ovale Zone, Sierra de Alta Coloma (JAC22 section); C, Sonninia (Papilliceras) arenata (Quenstedt, 1886) [M], A.9.9.2.2-1, Propinquans Zone?, Sierra de Alta Coloma; D, Sonninia (Papilliceras?) sp. [M], JAC22.6.13, Ovale Zone, Sierra de Alta Coloma (JAC22" figureDoi="http://doi.org/10.5281/zenodo.7150347" httpUri="https://zenodo.org/record/7150347/files/figure.png" pageId="25" pageNumber="824">Figs 9K</figureCitation>
;
<figureCitation id="13062A1EFFBB7B211B8DFB93FDEEDC70" box="[479,526,1108,1134]" captionStart="FIG" captionStartId="26.[132,143,1917,1934]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure-26@26.[698,1267,768,1324]" captionTargetPageId="26" captionText="FIG. 10. — A, Witchellia cf. W. nodatipinguis (Buckman, 1923) [M], JAC11.3.3, Ovale Zone,Sierra de Alta Coloma (JAC11 section);B-E, Witchellia connata (Buckman, 1927) [M]: B, C, JAC3.33.1, JAC3.33.2, Laeviuscula Zone, Trigonalis Subzone, Sierra de Alta Coloma (JAC3 section); D, E, JAC11.5.32, JAC11.R.19, Laeviuscula Zone, Trigonalis Subzone, Sierra de Alta Coloma (JAC11 section); F-K, Witchellia romanoides (Douvillé, 1885) [M]: F, J, JAC22.5.22, JAC22.5.8, Ovale Zone,Sierra de Alta Coloma (JAC22 section);G, JAC3.21.2, Laeviuscula Zone,Trigonalis Subzone, Sierra de Alta Coloma (JAC3 section);H, JFB.R.1, Laeviuscula Zone,Río Fardes area (JFB section);I, K, JAC11.R.20, JAC11.4.14,Laeviuscula Zone,Trigonalis Subzone,Sierra de Alta Coloma (JAC11 section).Scale bar:1 cm." figureDoi="http://doi.org/10.5281/zenodo.7150349" httpUri="https://zenodo.org/record/7150349/files/figure.png" pageId="25" pageNumber="824">10A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3276510FFBB7B211AD6FB4AFF15DD21" pageId="25" pageNumber="824" type="reference_group">
<paragraph id="8B82369BFFBB7B211AD6FB4AFF21DCDE" blockId="25.[132,775,1165,1216]" pageId="25" pageNumber="824">
cf.
<taxonomicName id="4C3D4D18FFBB7B211AF0FB4AFD8ADCBB" authority="Buckman, 1923" authorityName="Buckman" authorityYear="1923" box="[162,618,1165,1189]" class="Cephalopoda" family="Sonniniidae" genus="Stiphromorphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="nodatipinguis">
<emphasis id="B949EA89FFBB7B211AF0FB4AFE26DCBB" box="[162,454,1165,1189]" italics="true" pageId="25" pageNumber="824">Stiphromorphites nodatipinguis</emphasis>
Buckman, 1923
</taxonomicName>
: T.A. 4, pl. 398 (HT).
</paragraph>
<paragraph id="8B82369BFFBB7B211AD6FB1FFD07DCF1" blockId="25.[132,743,1239,1263]" box="[132,743,1239,1263]" pageId="25" pageNumber="824">
<treatmentCitationGroup id="AB2D11B5FFBB7B211AD6FB1FFD07DCF1" box="[132,743,1239,1263]" pageId="25" pageNumber="824">
<emphasis id="B949EA89FFBB7B211AD6FB1FFF3ADCF1" box="[132,218,1240,1263]" italics="true" pageId="25" pageNumber="824">Sonninia</emphasis>
cf.
<taxonomicName id="4C3D4D18FFBB7B211AACFB1FFE76DCF1" authorityName="Buckman, Horizon" authorityYear="1923" box="[254,406,1239,1263]" class="Cephalopoda" family="Sonniniidae" genus="Stiphromorphites" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="nodatipinguis">
<emphasis id="B949EA89FFBB7B211AACFB1FFEEADCF1" box="[254,266,1240,1263]" italics="true" pageId="25" pageNumber="824">S</emphasis>
.
<emphasis id="B949EA89FFBB7B211B45FB10FE76DCF1" box="[279,406,1239,1263]" italics="true" pageId="25" pageNumber="824">nodatipinguis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A9C108AFFBB7B211BFDFB10FDA9DCF1" author="IMLAY R. W." box="[431,585,1239,1263]" page="62" pageId="25" pageNumber="824" year="1973">
<bibRefCitation id="EFAC4B6AFFBB7B211BFDFB10FDA9DCF1" author="IMLAY R. W." box="[431,585,1239,1263]" pageId="25" pageNumber="824" pagination="1 - 100" refId="ref44018" refString="IMLAY R. W. 1973. - Middle Jurassic (Bajocian) Ammonites from Eastern Oregon. Professional Paper of the U. S. Geological Survey 756: 1 - 100. https: // doi. org / 10.3133 / pp 756" type="journal article" year="1973">Imlay 1973: 62</bibRefCitation>
</treatmentCitation>
, pl. 13, figs 1-4.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B82369BFFBB7B211AD6FACBFF15DD21" blockId="25.[132,774,1290,1343]" pageId="25" pageNumber="824">
cf.
<taxonomicName id="4C3D4D18FFBB7B211AF6FACCFE32DD3A" authorityName="Buckman, Horizon" authorityYear="1923" box="[164,466,1290,1317]" class="Cephalopoda" family="Sonniniidae" genus="Stiphromorphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="nodatipinguis" subGenus="Euhoploceras">
<emphasis id="B949EA89FFBB7B211AF6FACCFF58DD3B" box="[164,184,1291,1317]" italics="true" pageId="25" pageNumber="824">S.</emphasis>
(
<emphasis id="B949EA89FFBB7B211A98FACBFEA2DD3A" box="[202,322,1292,1316]" italics="true" pageId="25" pageNumber="824">Euhoploceras</emphasis>
)
<emphasis id="B949EA89FFBB7B211B00FACBFE32DD3A" box="[338,466,1292,1316]" italics="true" pageId="25" pageNumber="824">nodatipinguis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A9C108AFFBB7B211BBDFACBFCE1DD3A" box="[495,769,1292,1316]" pageId="25" pageNumber="824">
<bibRefCitation id="EFAC4B6AFFBB7B211BBDFACBFCE1DD3A" author="CHANDLER R. B. &amp; WHICHER J." box="[495,769,1292,1316]" pageId="25" pageNumber="824" refId="ref41103" refString="CHANDLER R. B. &amp; WHICHER J. 2015. - Fossils of Dorset. Inferior Oolite Ammonites Lower Bajocian. Commemorative Issue Celebrating the Work of John Callomon and the 200 th Year since the Production of William Smith's Geological Map of England and Wales. The Wessex Cephalopod. Burton upon Trent, Staffordshire, 76 p." type="book" year="2015">Chandler &amp; Whicher 2015</bibRefCitation>
</treatmentCitation>
: pl. 9, fig. 1.
</paragraph>
</subSubSection>
<subSubSection id="C3276510FFBB7B211AD6FA90FEE5DDBD" pageId="25" pageNumber="824" type="materials_examined">
<paragraph id="8B82369BFFBB7B211AD6FA90FEE5DDBD" blockId="25.[132,776,1366,1443]" pageId="25" pageNumber="824">
<collectionCode id="ED2CAE5EFFBB7B211AD6FA90FF0ADD70" box="[132,234,1367,1390]" pageId="25" pageNumber="824">MATERIAL</collectionCode>
EXAMINED. —
<materialsCitation id="3B553CC6FFBB7B211BDAFA91FD66DD70" ID-GBIF-Occurrence="3920964405" box="[392,646,1366,1390]" pageId="25" pageNumber="824">JAC11.3.3 to JAC11.3.8</materialsCitation>
,
<materialsCitation id="3B553CC6FFBB7B2118C1FA91FCE5DD70" ID-GBIF-Occurrence="3920964576" box="[659,773,1366,1390]" pageId="25" pageNumber="824">
JAC11.
<collectionCode id="ED2CAE5EFFBB7B2118B3FA91FD14DD70" box="[737,756,1366,1390]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="25" pageNumber="824">R</collectionCode>
.9
</materialsCitation>
,
<materialsCitation id="3B553CC6FFBB7B211AD6FAB6FEE9DD97" ID-GBIF-Occurrence="3920964537" box="[132,265,1393,1417]" pageId="25" pageNumber="824">
JAC11.
<collectionCode id="ED2CAE5EFFBB7B211A87FAB6FF09DD97" box="[213,233,1393,1417]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="25" pageNumber="824">R</collectionCode>
.43
</materialsCitation>
,
<materialsCitation id="3B553CC6FFBB7B211B45FAB6FE7CDD97" ID-GBIF-Occurrence="3920964400" box="[279,412,1393,1417]" pageId="25" pageNumber="824">
JAC11.
<collectionCode id="ED2CAE5EFFBB7B211B3AFAB6FE9CDD97" box="[360,380,1393,1417]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="25" pageNumber="824">R</collectionCode>
.44
</materialsCitation>
,
<materialsCitation id="3B553CC6FFBB7B211BF8FAB7FDCCDD96" ID-GBIF-Occurrence="3920964485" box="[426,556,1392,1416]" pageId="25" pageNumber="824">
JAC11.
<collectionCode id="ED2CAE5EFFBB7B211BABFAB7FDECDD96" box="[505,524,1392,1416]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="25" pageNumber="824">R</collectionCode>
.45
</materialsCitation>
and
<materialsCitation id="3B553CC6FFBB7B211834FAB6FEE1DDBD" ID-GBIF-Occurrence="3920964304" pageId="25" pageNumber="824">
JAC11.
<collectionCode id="ED2CAE5EFFBB7B2118E7FAB6FD28DD97" box="[693,712,1393,1417]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="25" pageNumber="824">R</collectionCode>
.48 to JAC11.
<collectionCode id="ED2CAE5EFFBB7B211A83FA4CFF04DDBD" box="[209,228,1419,1443]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="25" pageNumber="824">R</collectionCode>
.53
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276510FFBB7B211AD6FA7CFAF8DAAE" pageId="25" pageNumber="824" type="description">
<paragraph id="8B82369BFFBB7B211AD6FA7CFE2FDDCC" blockId="25.[132,463,1467,1491]" box="[132,463,1467,1491]" pageId="25" pageNumber="824">
MEASUREMENTS. — See
<tableCitation id="C6BF0320FFBB7B211B26FA7CFE2FDDCC" box="[372,463,1467,1491]" captionStart="TABLE" captionStartId="25.[450,460,219,236]" captionText="TABLE 11. — Measurements of Witchellia cf. W. nodatipinguis (Buckman, 1923) [M]." pageId="25" pageNumber="824">Table 11.</tableCitation>
</paragraph>
<paragraph id="8B82369BFFBB7B211AD6FA34FEF5DE13" blockId="25.[130,776,1523,2028]" box="[132,277,1523,1549]" pageId="25" pageNumber="824">DESCRIPTION</paragraph>
<paragraph id="8B82369BFFBB7B211AD6F9D5FAF8DAAE" blockId="25.[130,776,1523,2028]" lastBlockId="25.[812,1456,630,688]" pageId="25" pageNumber="824">Small to medium-sized shells, platycone with relatively involute coiling (U/D varying between 0.25 and 0.34). The whorl section is ovate (inner whorls) to almost rectangular, with a vertical umbilical wall, slightly convex flanks, and a tabulate venter with a low keel. The peristome is not present in the available specimens. The innermost whorls appear to have small tubercles, later periumbilical tuberculiform thickenings or primary ribs appear, from which two secondary ribs emerge, one more prominent than the other and, in addition, some weak intercalate ribs appear. On the outer whorls the thickenings progressively disappear, and distinct and less distinct radial ribs alternate, all weakening at the upper half of the flanks, where they gently bend forward and progressively erase, leaving a smooth ventrolateral shoulder near the keel. In some of the smaller specimens (maybe microconchs) the ribs are replaced by striae in the BC. The septal suture is not well preserved but appears to be quite simple.</paragraph>
</subSubSection>
<subSubSection id="C3276510FFBB7B21197FFD11FAEDDED3" pageId="25" pageNumber="824" type="discussion">
<paragraph id="8B82369BFFBB7B21197FFD11FC71DAEE" blockId="25.[811,1456,726,1741]" box="[813,913,726,752]" pageId="25" pageNumber="824">REMARKS</paragraph>
<paragraph id="8B82369BFFBB7B21197FFD32FB24DC31" blockId="25.[811,1456,726,1741]" pageId="25" pageNumber="824">
The specimens described resemble “
<taxonomicName id="4C3D4D18FFBB7B211EC0FD32FAD7DB11" authorityName="Buckman" authorityYear="1923" box="[1170,1335,757,783]" class="Cephalopoda" family="Sonniniidae" genus="Stiphromorphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="genus">
<emphasis id="B949EA89FFBB7B211EC0FD32FAD7DB11" box="[1170,1335,757,783]" italics="true" pageId="25" pageNumber="824">Stiphromorphites</emphasis>
</taxonomicName>
”
<taxonomicName id="4C3D4D18FFBB7B211F15FD32FBEBDB31" authority="Buckman, 1923" authorityName="Buckman, Horizon" authorityYear="1923" class="Cephalopoda" family="Sonniniidae" genus="Stiphromorphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="nodatipinguis">
<emphasis id="B949EA89FFBB7B211F15FD32FCB5DB31" italics="true" pageId="25" pageNumber="824">nodatipinguis</emphasis>
Buckman, 1923
</taxonomicName>
(Buckman 1923, T.A. 4: pl. 398), but the Subbetic specimens are more involute than the HT of “
<emphasis id="B949EA89FFBB7B21196AFC92FCB4DB71" box="[824,852,853,879]" italics="true" pageId="25" pageNumber="824">St.</emphasis>
”
<taxonomicName id="4C3D4D18FFBB7B211934FC92FC13DB71" authorityName="Buckman, Horizon" authorityYear="1923" box="[870,1011,853,879]" class="Cephalopoda" family="Sonniniidae" genus="Stiphromorphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="nodatipinguis">
<emphasis id="B949EA89FFBB7B211934FC92FC13DB71" box="[870,1011,853,879]" italics="true" pageId="25" pageNumber="824">nodatipinguis</emphasis>
</taxonomicName>
and have different ornamentation on the inner whorls, and the two species occupy different stratigraphic ranges. Thus while “
<emphasis id="B949EA89FFBB7B211E0FFC52FB99DBB1" box="[1117,1145,917,943]" italics="true" pageId="25" pageNumber="824">St.</emphasis>
”
<taxonomicName id="4C3D4D18FFBB7B211ED8FC52FAF7DBB1" authorityName="Buckman, Horizon" authorityYear="1923" box="[1162,1303,917,943]" class="Cephalopoda" family="Sonniniidae" genus="Stiphromorphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="nodatipinguis">
<emphasis id="B949EA89FFBB7B211ED8FC52FAF7DBB1" box="[1162,1303,917,943]" italics="true" pageId="25" pageNumber="824">nodatipinguis</emphasis>
</taxonomicName>
” constitutes a characteristic horizon (Bj-8a) in the Trigonalis Subzone of the Laeviuscula Zone (
<bibRefCitation id="EFAC4B6AFFBB7B2119A8FC13FB3ADBF0" author="CHANDLER R. B. &amp; CALLOMON J. H. &amp; KING A. &amp; JEFFREYS K. &amp; VARAH M. &amp; BENTLEY A." box="[1018,1242,980,1006]" pageId="25" pageNumber="824" pagination="345 - 375" refId="ref40976" refString="CHANDLER R. B., CALLOMON J. H., KING A., JEFFREYS K., VARAH M. &amp; BENTLEY A. 2006. - The stratigraphy of the Inferior Oolite at South Main Road Quarry, Dundry, Avon. Proceedings of the Geologists' Association 117: 345 - 375. https: // doi. org / 10.1016 / S 0016 - 7878 (06) 80043 - X" type="journal article" year="2006">
Chandler
<emphasis id="B949EA89FFBB7B211E35FC12FB7BDBF0" box="[1127,1179,980,1006]" italics="true" pageId="25" pageNumber="824">et al.</emphasis>
2006
</bibRefCitation>
;
<bibRefCitation id="EFAC4B6AFFBB7B211EBAFC13FA48DBF1" author="DIETZE V. &amp; STAPPENBECK G. &amp; WANNENMACHER N. &amp; SCHWEIGERT G." box="[1256,1448,980,1007]" pageId="25" pageNumber="824" pagination="141 - 165" refId="ref42017" refString="DIETZE V., STAPPENBECK G., WANNENMACHER N. &amp; SCHWEIGERT G. 2008. - Stratigraphie und Ammoniten-Faunenhorizonte im Grenzbereich Sauzei- / Humphriesianum-Zone (Unter-Bajocium, Mitteljura) der westlichen Schwabischen Alb (SW-Deutschland). Palaeodiversity 1: 141 - 165." type="journal article" year="2008">
Dietze
<emphasis id="B949EA89FFBB7B211F67FC12FA89DBF0" box="[1333,1385,980,1006]" italics="true" pageId="25" pageNumber="824">et al.</emphasis>
2008
</bibRefCitation>
,
<bibRefCitation id="EFAC4B6AFFBB7B21197FFC32FC85DC11" author="DIETZE V. &amp; KUTZ M. &amp; FRANZ M. &amp; BOSCH K." box="[813,869,1013,1039]" pageId="25" pageNumber="824" pagination="19 - 65" refId="ref42063" refString="DIETZE V., KUTZ M., FRANZ M. &amp; BOSCH K. 2009. - Stratigraphy of the Kahlenberg near Ringsheim (Upper Rhine Valley, SW Germany) with emphasis on the Laeviuscula and Sauzei zones (Lower Bajocian, Middle Jurassic). Palaeodiversity 2: 19 - 65." type="journal article" year="2009">2009</bibRefCitation>
,
<bibRefCitation id="EFAC4B6AFFBB7B211921FC32FC57DC11" author="DIETZE V. &amp; FRANZ M. &amp; DIETL G." box="[883,951,1013,1039]" pageId="25" pageNumber="824" pagination="23 - 30" refId="ref42118" refString="DIETZE V., FRANZ M. &amp; DIETL G. 2010 a. - Ammoniten aus dem Ubergangsbereich Ovale- / Laeviuscula-Zone (Unter-Bajocium) des Unteren Wedelsandsteins von Zillhausen (Westalb, Suddeutschland). Jahreshefte der Gesellschaft fur Naturkunde in Wurttemberg 166: 23 - 30." type="journal article" year="2010">2010a</bibRefCitation>
), the Subbetic forms are from the Ovale Zone or even the uppermost Discites Zone.
</paragraph>
<paragraph id="8B82369BFFBB7B211911FBF3FAEDDED3" blockId="25.[811,1456,726,1741]" pageId="25" pageNumber="824">
According to
<bibRefCitation id="EFAC4B6AFFBB7B21198CFBF3FB04DC51" author="DIETZE V. &amp; CALLOMON J. H. &amp; SCHWEIGERT G. &amp; CHANDLER R. B." box="[990,1252,1076,1103]" pageId="25" pageNumber="824" pagination="1 - 82" refId="ref41866" refString="DIETZE V., CALLOMON J. H., SCHWEIGERT G. &amp; CHANDLER R. B. 2005. - The ammonite fauna and biostratigraphy of the Lower Bajocian (Ovale and Laeviuscula zones) of E Swabia (S Germany). Stuttgarter Beitrage zur Naturkunde B 353: 1 - 82." type="journal article" year="2005">
Dietze
<emphasis id="B949EA89FFBB7B211E7EFBF2FB82DC50" box="[1068,1122,1076,1102]" italics="true" pageId="25" pageNumber="824">et al.</emphasis>
(2005: 25)
</bibRefCitation>
, this “species” is a small adult form synonymous with
<taxonomicName id="4C3D4D18FFBB7B211EE7FB93FC3CDC91" authority="(Waagen, 1867)" baseAuthorityName="Waagen" baseAuthorityYear="1867" class="Cephalopoda" family="Sonniniidae" genus="Stiphromorphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="adicra" subGenus="Euhoploceras">
<emphasis id="B949EA89FFBB7B211EE7FB93FB2ADC70" box="[1205,1226,1108,1134]" italics="true" pageId="25" pageNumber="824">S.</emphasis>
(
<emphasis id="B949EA89FFBB7B211E8BFB93FABDDC70" box="[1241,1373,1108,1134]" italics="true" pageId="25" pageNumber="824">Euhoploceras</emphasis>
)
<emphasis id="B949EA89FFBB7B211F3DFB93FA4FDC70" box="[1391,1455,1108,1134]" italics="true" pageId="25" pageNumber="824">adicra</emphasis>
(
<bibRefCitation id="EFAC4B6AFFBB7B211966FBB3FC33DC91" author="WAAGEN W." box="[820,979,1140,1167]" pageId="25" pageNumber="824" pagination="507 - 668" refId="ref47792" refString="WAAGEN W. 1867. - Uber die Zone des Ammonites Sowerbyi. Geognostische-Palaontologie Beitrage 1: 507 - 668." type="journal article" year="1867">Waagen, 1867</bibRefCitation>
)
</taxonomicName>
, but this could not be corroborated because actually
<taxonomicName id="4C3D4D18FFBB7B2119D4FB53FB9CDCB0" baseAuthorityName="Waagen" baseAuthorityYear="1867" box="[902,1148,1172,1198]" class="Cephalopoda" family="Sonniniidae" genus="Stiphromorphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="adicra" subGenus="Euhoploceras">
<emphasis id="B949EA89FFBB7B2119D4FB53FC7ADCB0" box="[902,922,1172,1198]" italics="true" pageId="25" pageNumber="824">S.</emphasis>
(
<emphasis id="B949EA89FFBB7B2119FBFB53FBCDDCB0" box="[937,1069,1172,1198]" italics="true" pageId="25" pageNumber="824">Euhoploceras</emphasis>
)
<emphasis id="B949EA89FFBB7B211E6EFB53FB9CDCB0" box="[1084,1148,1172,1198]" italics="true" pageId="25" pageNumber="824">adicra</emphasis>
</taxonomicName>
has an ovate rather than rectangular whorl section on outer whorls and a well developed tuberculate stage that is not present on
<emphasis id="B949EA89FFBB7B211EEEFB13FA8EDCF0" box="[1212,1390,1236,1262]" italics="true" pageId="25" pageNumber="824">
<taxonomicName id="4C3D4D18FFBB7B211EEEFB13FA8ADCF0" authorityName="Buckman, Horizon" authorityYear="1923" box="[1212,1386,1236,1262]" class="Cephalopoda" family="Sonniniidae" genus="Stiphromorphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="nodatipinguis">W. nodatipinguis</taxonomicName>
.
</emphasis>
Some strongly ribbed
<taxonomicName id="4C3D4D18FFBB7B211981FB34FB6EDD13" box="[979,1166,1267,1293]" pageId="25" pageNumber="824">
<emphasis id="B949EA89FFBB7B211981FB34FBDDDD13" box="[979,1085,1267,1293]" italics="true" pageId="25" pageNumber="824">Witchellia</emphasis>
species
</taxonomicName>
such as
<taxonomicName id="4C3D4D18FFBB7B211EB1FB33FC8CDD30" authority="(Branco, 1879)" baseAuthorityName="Branco" baseAuthorityYear="1879" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="sutneri">
<emphasis id="B949EA89FFBB7B211EB1FB33FAABDD13" box="[1251,1355,1268,1293]" italics="true" pageId="25" pageNumber="824">W. sutneri</emphasis>
(
<bibRefCitation id="EFAC4B6AFFBB7B211F08FB33FC84DD30" author="BRANCO W." pageId="25" pageNumber="824" pagination="1 - 160" refId="ref40218" refString="BRANCO W. 1879. - Der Untere Doggers Deutsch-Lothringens. Abhandlungen Geologische Spezialkarte Elsass-Lothringen 2 (2): 1 - 160. https: // doi. org / 10.3931 / e-rara- 18884" type="journal article" year="1879">Branco, 1879</bibRefCitation>
)
</taxonomicName>
(HT refigured in
<bibRefCitation id="EFAC4B6AFFBB7B211E67FAD4FAEADD30" author="SCHLEGELMILCH R." box="[1077,1290,1299,1326]" pageId="25" pageNumber="824" refId="ref47166" refString="SCHLEGELMILCH R. 1985. - Die Ammoniten der suddeutschen Doggers. Gustav Fisher, Stuttgart, 276 p. https: // doi. org / 10.1007 / 978 - 3 - 8274 - 3105 - 9" type="book" year="1985">Schlegelmilch 1985</bibRefCitation>
: pl. 19, fig. 4),
<taxonomicName id="4C3D4D18FFBB7B21197FFAF3FBA3DD50" authority="Buckman, 1925" authorityName="Buckman" authorityYear="1925" box="[813,1091,1331,1358]" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="glauca">
<emphasis id="B949EA89FFBB7B21197FFAF3FC72DD53" box="[813,914,1331,1357]" italics="true" pageId="25" pageNumber="824">W. glauca</emphasis>
Buckman, 1925
</taxonomicName>
(Buckman 1925, T.A. 6: pl. 594),
<taxonomicName id="4C3D4D18FFBB7B21197EFA93FB73DD73" authority="Buckman, 1926" authorityName="Buckman" authorityYear="1926" box="[812,1171,1363,1389]" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="platymorpha">
<emphasis id="B949EA89FFBB7B21197EFA93FC36DD73" box="[812,982,1363,1389]" italics="true" pageId="25" pageNumber="824">W. platymorpha</emphasis>
Buckman, 1926
</taxonomicName>
(Buckman 1926, T.A. 6: pl. 580A),
<taxonomicName id="4C3D4D18FFBB7B2119F0FAB3FB5EDD93" authority="Buckman, 1926" authorityName="Buckman" authorityYear="1926" box="[930,1214,1395,1421]" class="Cephalopoda" family="Sonniniidae" genus="Sonninia" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="falcata">
<emphasis id="B949EA89FFBB7B2119F0FAB3FBEADD93" box="[930,1034,1395,1421]" italics="true" pageId="25" pageNumber="824">W. falcata</emphasis>
Buckman, 1926
</taxonomicName>
(Buckman 1926, T.A. 6: pl. 688), and
<taxonomicName id="4C3D4D18FFBB7B2119B0FA53FAD6DDB3" authority="Buckman, 1926" authorityName="Buckman" authorityYear="1926" box="[994,1334,1427,1453]" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="actinophora">
<emphasis id="B949EA89FFBB7B2119B0FA53FB60DDB3" box="[994,1152,1427,1453]" italics="true" pageId="25" pageNumber="824">W. actinophora</emphasis>
Buckman, 1926
</taxonomicName>
(Buckman 1926, T.A. 6: pl. 689) also bear similarities with the Subbetic specimens in whorl section, coiling, and septal suture, but the four Buckman’ species have sigmoid rather than radial ribbing as well as a different stratigraphic position. The
<typeStatus id="54868839FFBB7B211F6AF9D4FA86DE33" box="[1336,1382,1555,1581]" pageId="25" pageNumber="824">type</typeStatus>
of
<taxonomicName id="4C3D4D18FFBB7B211FC3F9D4FC93DE52" baseAuthorityName="Branco" baseAuthorityYear="1879" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="sutneri">
<emphasis id="B949EA89FFBB7B211FC3F9D4FA48DE32" box="[1425,1448,1555,1580]" italics="true" pageId="25" pageNumber="824">W</emphasis>
.
<emphasis id="B949EA89FFBB7B21197FF9F4FC93DE52" box="[813,883,1587,1612]" italics="true" pageId="25" pageNumber="824">sutneri</emphasis>
</taxonomicName>
has well differentiated ventral sulci and a clear separation between the primary and secondary ribs. The relatively simple septal suture, the tabulate venter, and the absence of well-marked tubercles of the Subbetic specimens makes it closer to
<taxonomicName id="4C3D4D18FFBB7B2119DCF975FC18DED2" authorityName="Buckman" authorityYear="1889" box="[910,1016,1714,1740]" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="genus">
<emphasis id="B949EA89FFBB7B2119DCF975FC18DED2" box="[910,1016,1714,1740]" italics="true" pageId="25" pageNumber="824">Witchellia</emphasis>
</taxonomicName>
than to
<emphasis id="B949EA89FFBB7B211E07F975FB89DED2" box="[1109,1129,1714,1740]" italics="true" pageId="25" pageNumber="824">S.</emphasis>
(
<emphasis id="B949EA89FFBB7B211E2BF975FB1DDED2" box="[1145,1277,1714,1740]" italics="true" pageId="25" pageNumber="824">Euhoploceras</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C3276510FFBB7B23197FF935FCE1DB6E" lastPageId="27" lastPageNumber="826" pageId="25" pageNumber="824" type="distribution">
<paragraph id="8B82369BFFBB7B21197FF935FC28DF12" blockId="25.[813,1457,1778,2028]" box="[813,968,1778,1804]" pageId="25" pageNumber="824">DISTRIBUTION</paragraph>
<paragraph id="8B82369BFFBB7B23197FF8D5FCE1DB6E" blockId="25.[813,1457,1778,2028]" lastBlockId="27.[130,777,661,880]" lastPageId="27" lastPageNumber="826" pageId="25" pageNumber="824">
In the section at Redhole Lane, described byBuckman (1893),
<bibRefCitation id="EFAC4B6AFFBB7B21197FF8F5FC34DF52" author="PARSONS C. F." box="[813,980,1842,1868]" pageId="25" pageNumber="824" pagination="153 - 180" refId="ref45442" refString="PARSONS C. F. 1974. - The sauzei and ' so called' sowerbyi Zones of the Lower Bajocian. Newsletter on Stratigraphy 3: 153 - 180. https: // doi. org / 10.1127 / nos / 3 / 1974 / 153" type="journal article" year="1974">Parsons (1974)</bibRefCitation>
, and
<bibRefCitation id="EFAC4B6AFFBB7B211E44F8F6FB2FDF52" author="HUXTABLE J. G." box="[1046,1231,1841,1868]" pageId="25" pageNumber="824" pagination="101 - 106" refId="ref43843" refString="HUXTABLE J. G. 2000. - A review of the stratigraphy of the Inferior Oolite (Bajocian Stage, Middle Jurassic) near Clatcombe, Sherborne, with a description of temporary sections. Proceedings of the Dorset natural History and archaeological Society 121: 101 - 106." type="journal article" year="2000">Huxtable (2000)</bibRefCitation>
, the critical interval lies in the so-called Blue Bed, bed 3. There, the upper part, 3b in the section as described by Huxtable, constitutes the
<typeStatus id="54868839FFBB7B21197CF855FCBCDFB2" box="[814,860,1938,1964]" pageId="25" pageNumber="824">type</typeStatus>
horizon of ‘
<taxonomicName id="4C3D4D18FFBB7B2119B6F856FB73DFB5" authorityName="Buckman" authorityYear="1923" box="[996,1171,1937,1963]" class="Cephalopoda" family="Sonniniidae" genus="Stiphromorphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="genus">
<emphasis id="B949EA89FFBB7B2119B6F856FB73DFB5" box="[996,1171,1937,1963]" italics="true" pageId="25" pageNumber="824">Stiphromorphites</emphasis>
</taxonomicName>
’
<taxonomicName id="4C3D4D18FFBB7B211EF0F856FBCCDFD5" authority="Buckman, Horizon" authorityName="Buckman, Horizon" authorityYear="1923" class="Cephalopoda" family="Sonniniidae" genus="Stiphromorphites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="25" pageNumber="824" phylum="Mollusca" rank="species" species="nodatipinguis" subGenus="Euhoploceras">
(
<emphasis id="B949EA89FFBB7B211EF9F856FACFDFB5" box="[1195,1327,1937,1963]" italics="true" pageId="25" pageNumber="824">Euhoploceras</emphasis>
)
<emphasis id="B949EA89FFBB7B211F13F856FCB5DFD5" italics="true" pageId="25" pageNumber="824">nodatipinguis</emphasis>
Buckman, Horizon
</taxonomicName>
Bj-8a, Laeviuscula Zone, Trigonalis Subzone (see
<bibRefCitation id="EFAC4B6AFFBB7B211993F816FB52DFF2" author="DIETZE V. &amp; CALLOMON J. H. &amp; SCHWEIGERT G. &amp; CHANDLER R. B." box="[961,1202,2001,2028]" pageId="25" pageNumber="824" pagination="1 - 82" refId="ref41866" refString="DIETZE V., CALLOMON J. H., SCHWEIGERT G. &amp; CHANDLER R. B. 2005. - The ammonite fauna and biostratigraphy of the Lower Bajocian (Ovale and Laeviuscula zones) of E Swabia (S Germany). Stuttgarter Beitrage zur Naturkunde B 353: 1 - 82." type="journal article" year="2005">
Dietze
<emphasis id="B949EA89FFBB7B211E5CF815FBA3DFF5" box="[1038,1091,2001,2027]" italics="true" pageId="25" pageNumber="824">et al.</emphasis>
2005: 76
</bibRefCitation>
). The specimen figured by
<bibRefCitation id="EFAC4B6AFFB97B231AF6FD51FE39DAAE" author="CHANDLER R. B. &amp; WHICHER J." box="[164,473,661,688]" pageId="27" pageNumber="826" refId="ref41103" refString="CHANDLER R. B. &amp; WHICHER J. 2015. - Fossils of Dorset. Inferior Oolite Ammonites Lower Bajocian. Commemorative Issue Celebrating the Work of John Callomon and the 200 th Year since the Production of William Smith's Geological Map of England and Wales. The Wessex Cephalopod. Burton upon Trent, Staffordshire, 76 p." type="book" year="2015">Chandler &amp; Whicher (2015)</bibRefCitation>
comes from the same faunal horizon. The highly similar specimens, from
<collectingRegion id="49F9F879FFB97B23180BFD71FD4CDACE" box="[601,684,694,720]" country="United States of America" name="Oregon" pageId="27" pageNumber="826">Oregon</collectingRegion>
(
<collectingCountry id="F32A760BFFB97B2318EBFD72FF24DAEE" name="United States of America" pageId="27" pageNumber="826">United States</collectingCountry>
), described and figured by
<bibRefCitation id="EFAC4B6AFFB97B231BB5FD12FD95DAEE" author="IMLAY R. W." box="[487,629,725,752]" pageId="27" pageNumber="826" pagination="1 - 100" refId="ref44018" refString="IMLAY R. W. 1973. - Middle Jurassic (Bajocian) Ammonites from Eastern Oregon. Professional Paper of the U. S. Geological Survey 756: 1 - 100. https: // doi. org / 10.3133 / pp 756" type="journal article" year="1973">Imlay (1973)</bibRefCitation>
are also from the Trigonalis Subzone. All the Subbetic specimens appear to be stratigraphically older (lowermost part of the Trigonalis Subzone?, Ovale Zone or even the upper part of the Discites Zone) than any of the strongly ribbed species of
<taxonomicName id="4C3D4D18FFB97B2318C2FC92FD1ADB71" authorityName="Buckman" authorityYear="1889" box="[656,762,853,879]" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="27" pageNumber="826" phylum="Mollusca" rank="genus">
<emphasis id="B949EA89FFB97B2318C2FC92FD1ADB71" box="[656,762,853,879]" italics="true" pageId="27" pageNumber="826">Witchellia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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