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<mods:title id="F3DE41F6E7DF1355A4D7258834BE7A18">Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)</mods:title>
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<mods:affiliation id="E41992EE4F2C2EDF4023E675DCD745C8">Universidade Federal de Pelotas (UFPEL), Departamento de Ecologia, Zoologia e Genética, Instituto de Biologia, Campus Universitário Capão do Leão, RS, 96010 - 610, Brazil &amp; Red Sea Research Center, King Abdullah University of Science and Technology (KAUST), Thuwal, Saudi Arabia</mods:affiliation>
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<taxonomicName id="77C374D7FFBEFFA1FF06AEDA2C2AFF14" ID-CoL="85MCR" authority="De Man, 1910" authorityName="De Man" authorityYear="1910" box="[151,522,151,178]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="193" phylum="Arthropoda" rank="species" species="parvus">
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<emphasis id="82B7D346FFBEFFA1FF06AEDA2F75FF14" bold="true" box="[151,341,151,178]" italics="true" pageId="14" pageNumber="193">Athanas parvus</emphasis>
<bibRefCitation id="D45272A5FFBEFFA1FECDAEDA2C2AFF14" author="De Man, J. G." box="[348,522,151,177]" pageId="14" pageNumber="193" pagination="287 - 319" refId="ref22004" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;, V. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2 (11), 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
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<paragraph id="B07C0F54FFBEFFA1FF06AEF62ECBFF70" blockId="14.[151,522,151,213]" box="[151,235,187,213]" pageId="14" pageNumber="193">
(
<figureCitation id="28F813D1FFBEFFA1FF0EAEF62EC3FF70" box="[159,227,187,213]" captionStart="FIGURE 7" captionStartId="14.[151,250,1948,1972]" captionTargetBox="[151,1436,673,1924]" captionTargetId="figure-201@14.[151,1436,673,1924]" captionTargetPageId="14" captionText="FIGURE 7. Athanas parvus De Man, 1910, male (cl 2.9 mm) from Masirah Island, Oman (FLMNH UF 71418); a, shrimp in life, lateral (left side); b, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350791" httpUri="https://zenodo.org/record/10350791/files/figure.png" pageId="14" pageNumber="193">Fig. 7</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="90D3287AFFBEFFA1FF06AF4D2BBCFEBD" box="[151,1436,255,280]" pageId="14" pageNumber="193">
<treatmentCitation id="31622945FFBEFFA1FF06AF4D2C25FEB2" author="De Man, J. G." box="[151,517,255,280]" page="315" pageId="14" pageNumber="193" year="1910">
<taxonomicName id="77C374D7FFBEFFA1FF06AF4D2C25FEB2" ID-CoL="85MCR" authority="De Man, 1910: 315" authorityName="De Man" authorityPageNumber="315" authorityYear="1910" box="[151,517,255,280]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="193" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="82B7D346FFBEFFA1FF06AF4D2F16FEB2" box="[151,310,256,280]" italics="true" pageId="14" pageNumber="193">Athanas parvus</emphasis>
<bibRefCitation id="D45272A5FFBEFFA1FEADAF4D2C25FEB2" author="De Man, J. G." box="[316,517,255,280]" pageId="14" pageNumber="193" pagination="287 - 319" refId="ref22004" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;, V. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2 (11), 287 - 319." type="journal article" year="1910">De Man, 1910: 315</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="31622945FFBEFFA1FD9EAF4D2CF0FEBD" author="De Man, J. G." box="[527,720,256,280]" page="148" pageId="14" pageNumber="193" year="1911">
<bibRefCitation id="D45272A5FFBEFFA1FD9EAF4D2CF0FEBD" author="De Man, J. G." box="[527,720,256,280]" pageId="14" pageNumber="193" pagination="1 - 23" refId="ref22045" refString="De Man, J. G. (1911) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Siboga-Expeditie, 39 a 1 (2). E. J. Brill, Leiden, pp. 133 - 465, Supplement (1915), Explanations of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">De Man 1911: 148</bibRefCitation>
</treatmentCitation>
, (1915) pl. 1, fig. 4;
<treatmentCitation id="31622945FFBEFFA1FC33AF4D2A56FEBD" author="Tattersall, W. M." box="[930,1142,256,280]" page="372" pageId="14" pageNumber="193" year="1921">
<bibRefCitation id="D45272A5FFBEFFA1FC33AF4D2A56FEBD" author="Tattersall, W. M." box="[930,1142,256,280]" pageId="14" pageNumber="193" pagination="345 - 398" refId="ref23551" refString="Tattersall, W. M. (1921) Report on the Stomatopoda and macrurous Decapoda collected by Mr. Cyril Crossland in the Sudanese Red Sea. Journal of Linnean Society of London, Zoology, 34, 345 - 398, pls. 27 + 28. https: // doi. org / 10.1111 / j. 1096 - 3642.1921. tb 01838. x" type="journal article" year="1921">Tattersall, 1921: 372</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="31622945FFBEFFA1FB11AF4D2B69FEB2" author="Ledoyer, M." box="[1152,1353,255,280]" page="126" pageId="14" pageNumber="193" year="1970">
<bibRefCitation id="D45272A5FFBEFFA1FB11AF4D2B69FEB2" author="Ledoyer, M." box="[1152,1353,255,280]" pageId="14" pageNumber="193" pagination="121 - 157" refId="ref22778" refString="Ledoyer, M. (1970) Etude systematique et remarques ecologiques sur les Caridea recueillis principalement dans les biotopes de substrat meuble. Regions de Tulear et de Nosy-Be. Annales de l'Universite de Madagascar (Sciences Naturelles et Mathematiques), 7, 121 - 157, pls. 1 - 25." type="journal article" year="1970">Ledoyer, 1970: 126</bibRefCitation>
</treatmentCitation>
, fig. 11;
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</paragraph>
<paragraph id="B07C0F54FFBEFFA1FF56AF532BBDFED1" blockId="14.[151,1437,255,433]" pageId="14" pageNumber="193">
<bibRefCitation id="D45272A5FFBEFFA1FF56AF532FC2FE93" author="Al-Kandari, M. &amp; Anker, A. &amp; Al-Hasheem, A. &amp; Al-Yassen, S. &amp; Sattari, Z. &amp; De Grave, S." box="[199,482,286,310]" pageId="14" pageNumber="193" pagination="251 - 280" refId="ref19182" refString="Al-Kandari, M., Anker, A., Al-Hasheem, A., Al-Yassen, S., Sattari, Z. &amp; De Grave, S. (2020) New records of decapod crustaceans from Kuwait (Malacostraca: Decapoda). Zootaxa, 4803 (2), 251 - 280. https: // doi. org / 10.11646 / zootaxa. 4803.2.2" type="journal article" year="2020">
Al-Kandari
<emphasis id="82B7D346FFBEFFA1FED0AF522F54FE93" box="[321,372,286,310]" italics="true" pageId="14" pageNumber="193">et al.</emphasis>
2020: 257
</bibRefCitation>
, fig. 4.
<taxonomicName id="77C374D7FFBEFFA1FF06AF702C27FEF0" authority="De Man 1910: 314" authorityName="De Man" authorityPageNumber="314" authorityYear="1910" box="[151,519,317,341]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="193" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="82B7D346FFBEFFA1FF06AF702F63FEF0" box="[151,323,317,341]" italics="true" pageId="14" pageNumber="193">Athanas Sibogae</emphasis>
<bibRefCitation id="D45272A5FFBEFFA1FED8AF702C27FEF0" author="De Man, J. G." box="[329,519,317,341]" pageId="14" pageNumber="193" pagination="287 - 319" refId="ref22004" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;, V. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2 (11), 287 - 319." type="journal article" year="1910">De Man 1910: 314</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="D45272A5FFBEFFA1FD8AAF702CFBFEF0" author="De Man, J. G." box="[539,731,317,341]" pageId="14" pageNumber="193" pagination="1 - 23" refId="ref22045" refString="De Man, J. G. (1911) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Siboga-Expeditie, 39 a 1 (2). E. J. Brill, Leiden, pp. 133 - 465, Supplement (1915), Explanations of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">De Man 1911: 151</bibRefCitation>
, (1915) pl. 2, fig. 6.
<emphasis id="82B7D346FFBEFFA1FF06AF112F77FED1" box="[151,343,348,372]" italics="true" pageId="14" pageNumber="193">
<taxonomicName id="77C374D7FFBEFFA1FF06AF112F1DFED1" authorityName="De Man" authorityYear="1910" box="[151,317,348,372]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="193" phylum="Arthropoda" rank="species" species="sibogae">Athanas sibogae</taxonomicName>
—
</emphasis>
<bibRefCitation id="D45272A5FFBEFFA1FEC6AF112C5DFED1" author="Banner, D. M. &amp; Banner, A. H." box="[343,637,348,372]" pageId="14" pageNumber="193" pagination="291 - 382" refId="ref20883" refString="Banner, D. M. &amp; Banner, A. H. (1973) The alpheid shrimp of Australia, part I: the lower genera. Records of the Australian Museum, 28, 291 - 382. https: // doi. org / 10.3853 / j. 0067 - 1975.28.1973.407" type="journal article" year="1973">Banner &amp; Banner 1973: 321</bibRefCitation>
, fig. 9;
<bibRefCitation id="D45272A5FFBEFFA1FD5FAF112DC8FED1" author="Banner, D. M. &amp; Banner, A. H." box="[718,1000,348,372]" pageId="14" pageNumber="193" pagination="1 - 99" refId="ref20938" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981: 43</bibRefCitation>
;
<bibRefCitation id="D45272A5FFBEFFA1FC65AF112ABAFED1" author="Titgen, R. H." box="[1012,1178,348,372]" pageId="14" pageNumber="193" refId="ref23619" refString="Titgen, R. H. (1982) The systematics and ecology of the decapods of Dubai, and their zoogeographic relationships to the Arabian Gulf and the western Indian Ocean. PhD thesis, Texas A &amp; M University, College Station, Texas, 269 pp." type="book" year="1982">Titgen 1982: 80</bibRefCitation>
;
<bibRefCitation id="D45272A5FFBEFFA1FB37AF112BBDFED1" author="Banner, A. H. &amp; Banner, D. M." box="[1190,1437,348,372]" pageId="14" pageNumber="193" pagination="1 - 164" refId="ref20768" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983:</bibRefCitation>
</paragraph>
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<paragraph id="B07C0F54FFBEFFA1FF56AF372B42FE14" blockId="14.[151,1437,255,433]" pageId="14" pageNumber="193">81. (Note: selective synonymy, with focus on important morphological accounts and records from the western Indian Ocean)</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBEFFA1FF06AF922C4BFDE4" pageId="14" pageNumber="193" type="materials_examined">
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<emphasis id="82B7D346FFBEFFA1FF06AF922F5FFE5C" bold="true" box="[151,383,479,505]" pageId="14" pageNumber="193">Material examined</emphasis>
.
<materialsCitation id="00AB0509FFBEFFA1FE03AF922C46FDE4" collectingDate="2022-11-20" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="2" latitude="20.264334" location="Masirah Island" longLatPrecision="1" longitude="58.62397" municipality="Ghab" pageId="14" pageNumber="193" specimenCode="UF 71418" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFBEFFA1FE03AF922FC3FE5C" box="[402,483,479,505]" count="1" pageId="14" pageNumber="193" type="male">1 male</specimenCount>
(cl
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),
<collectionCode id="D6D29791FFBEFFA1FD03AFAD2CD4FE5F" box="[658,756,480,506]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="14" pageNumber="193" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFBEFFA1FC91AFAD2D5AFE5C" box="[768,890,479,506]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="14" pageNumber="193" type="Museum">UF 71418</specimenCode>
,
<collectingCountry id="C8D44FC4FFBEFFA1FC1BAF922DF1FE5C" box="[906,977,479,505]" name="Oman" pageId="14" pageNumber="193">Oman</collectingCountry>
,
<location id="B51C598FFFBEFFA1FC73AF922AB1FE5C" LSID="urn:lsid:plazi:treatment:386ABE42FFBEFFA0FF06AEDA2D0BFD0C:B51C598FFFBEFFA1FC73AF922AB1FE5C" box="[994,1169,479,505]" country="Oman" latitude="20.264334" longLatPrecision="1" longitude="58.62397" municipality="Ghab" name="Masirah Island" pageId="14" pageNumber="193">Masirah Island</location>
,
<quantity id="773BA2B1FFBEFFA1FB33AF922AC1FE5C" box="[1186,1249,479,505]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="193" unit="km" value="2.0">2 km</quantity>
north of
<collectingMunicipality id="5018952EFFBEFFA1FAC8AF922BB8FE5C" box="[1369,1432,479,505]" pageId="14" pageNumber="193">Ghab</collectingMunicipality>
,
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/
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, coral reef flat, under coral rubble on sand, hand collecting, depth
<quantity id="773BA2B1FFBEFFA1FB75AC4E2B64FDBB" box="[1252,1348,515,542]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="14" pageNumber="193" unit="m" value="1.75" valueMax="2.0" valueMin="1.5">
<elevation id="3BEEE867FFBEFFA1FB75AC4E2B64FDBB" box="[1252,1348,515,542]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="14" pageNumber="193" unit="m" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 m</elevation>
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, leg.
<collectorName id="1D366A82FFBEFFA1FA10AC4E2EFFFDE4" pageId="14" pageNumber="193">A. Anker</collectorName>
,
<date id="C47D2994FFBEFFA1FF7BAC6A2F47FDE4" box="[234,359,551,577]" pageId="14" pageNumber="193" value="2022-11-20">
<collectingDate id="D439D07CFFBEFFA1FF7BAC6A2F47FDE4" box="[234,359,551,577]" pageId="14" pageNumber="193" value="2022-11-20">20.11.2022</collectingDate>
</date>
[fcn BOMAN-14839]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBEFFA0FF56AC062B72FEE4" lastPageId="15" lastPageNumber="194" pageId="14" pageNumber="193" type="description">
<paragraph id="B07C0F54FFBEFFA1FF56AC062D62FD2F" blockId="14.[151,1436,479,650]" pageId="14" pageNumber="193">
<emphasis id="82B7D346FFBEFFA1FF56AC062F71FDC0" bold="true" box="[199,337,587,613]" pageId="14" pageNumber="193">Description</emphasis>
. For description and illustrations of
<taxonomicName id="77C374D7FFBEFFA1FD69AC062D45FDC0" authorityName="De Man" authorityYear="1910" box="[760,869,587,613]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="193" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="82B7D346FFBEFFA1FD69AC062D45FDC0" box="[760,869,587,613]" italics="true" pageId="14" pageNumber="193">A. parvus</emphasis>
</taxonomicName>
see, for instance,
<bibRefCitation id="D45272A5FFBEFFA1FBA8AC012AC4FDC0" author="De Man, J. G." box="[1081,1252,587,614]" pageId="14" pageNumber="193" pagination="287 - 319" refId="ref22004" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;, V. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2 (11), 287 - 319." type="journal article" year="1910">De Man (1910</bibRefCitation>
,
<bibRefCitation id="D45272A5FFBEFFA1FB60AC062B0BFDC0" author="De Man, J. G." box="[1265,1323,587,613]" pageId="14" pageNumber="193" pagination="1 - 23" refId="ref22045" refString="De Man, J. G. (1911) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Siboga-Expeditie, 39 a 1 (2). E. J. Brill, Leiden, pp. 133 - 465, Supplement (1915), Explanations of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">1911</bibRefCitation>
),
<bibRefCitation id="D45272A5FFBEFFA1FAD3AC062F1EFD2C" author="Miya, Y. &amp; S. Miyake" pageId="14" pageNumber="193" pagination="129 - 162" refId="ref23069" refString="Miya, Y. &amp; S. Miyake. 1968 a. Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 1, 129 - 162." type="journal article" year="1968">Miya &amp; Miyake (1968)</bibRefCitation>
and
<bibRefCitation id="D45272A5FFBEFFA1FEE4AC3D2CB0FD2C" author="Banner, D. M. &amp; Banner, A. H." box="[373,656,623,650]" pageId="14" pageNumber="193" pagination="291 - 382" refId="ref20883" refString="Banner, D. M. &amp; Banner, A. H. (1973) The alpheid shrimp of Australia, part I: the lower genera. Records of the Australian Museum, 28, 291 - 382. https: // doi. org / 10.3853 / j. 0067 - 1975.28.1973.407" type="journal article" year="1973">Banner &amp; Banner (1973)</bibRefCitation>
; see also
<figureCitation id="28F813D1FFBEFFA1FD6BAC222D1EFD2F" box="[762,830,623,650]" captionStart="FIGURE 7" captionStartId="14.[151,250,1948,1972]" captionTargetBox="[151,1436,673,1924]" captionTargetId="figure-201@14.[151,1436,673,1924]" captionTargetPageId="14" captionText="FIGURE 7. Athanas parvus De Man, 1910, male (cl 2.9 mm) from Masirah Island, Oman (FLMNH UF 71418); a, shrimp in life, lateral (left side); b, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350791" httpUri="https://zenodo.org/record/10350791/files/figure.png" pageId="14" pageNumber="193">Fig. 7</figureCitation>
.
</paragraph>
<caption id="E4BC5FDCFFBEFFA1FF06A9D12CCFF87D" ID-DOI="http://doi.org/10.5281/zenodo.10350791" ID-Zenodo-Dep="10350791" httpUri="https://zenodo.org/record/10350791/files/figure.png" pageId="14" pageNumber="193" startId="14.[151,250,1948,1972]" targetBox="[151,1436,673,1924]" targetPageId="14" targetType="figure">
<paragraph id="B07C0F54FFBEFFA1FF06A9D12CCFF87D" blockId="14.[151,1437,1948,2008]" pageId="14" pageNumber="193">
<emphasis id="82B7D346FFBEFFA1FF06A9D12F34F810" bold="true" box="[151,276,1948,1973]" pageId="14" pageNumber="193">FIGURE 7.</emphasis>
<taxonomicName id="77C374D7FFBEFFA1FE8AA9D12C77F810" authority="De Man, 1910" authorityName="De Man" authorityYear="1910" box="[283,599,1948,1973]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="193" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="82B7D346FFBEFFA1FE8AA9D12F9AF811" box="[283,442,1948,1972]" italics="true" pageId="14" pageNumber="193">Athanas parvus</emphasis>
<bibRefCitation id="D45272A5FFBEFFA1FE50A9D02C77F810" author="De Man, J. G." box="[449,599,1949,1973]" pageId="14" pageNumber="193" pagination="287 - 319" refId="ref22004" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;, V. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2 (11), 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
</taxonomicName>
, male (cl 2.9 mm) from Masirah Island, Oman (FLMNH UF 71418); a, shrimp in life, lateral (left side); b, dorsal. Photographs by the author.
</paragraph>
</caption>
<paragraph id="B07C0F54FFBFFFA0FF56AEDB2B72FEE4" blockId="15.[151,1437,150,681]" pageId="15" pageNumber="194">
<emphasis id="82B7D346FFBFFFA0FF56AEDB2F41FF15" bold="true" box="[199,353,150,176]" pageId="15" pageNumber="194">Colour in life</emphasis>
. Body evenly speckled with bright red chromatophores, carapace and pleon with broad, somewhat nerve cord-shaped band composed of bright white and pinkish white chromatophores, extending along mid-dorsal surface from tip of the rostrum to posterior margin of sixth pleonite; antennular peduncles with large white patches; antenna mostly translucent with red spots; chelipeds hyaline with red spotting; remaining pereiopods mostly translucent with occasional red spotting; tail fan bright red (
<figureCitation id="28F813D1FFBFFFA0FCA4AF6A2D5AFEE4" box="[821,890,295,321]" captionStart="FIGURE 7" captionStartId="14.[151,250,1948,1972]" captionTargetBox="[151,1436,673,1924]" captionTargetId="figure-201@14.[151,1436,673,1924]" captionTargetPageId="14" captionText="FIGURE 7. Athanas parvus De Man, 1910, male (cl 2.9 mm) from Masirah Island, Oman (FLMNH UF 71418); a, shrimp in life, lateral (left side); b, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350791" httpUri="https://zenodo.org/record/10350791/files/figure.png" pageId="15" pageNumber="194">Fig. 7</figureCitation>
). See also
<bibRefCitation id="D45272A5FFBFFFA0FC60AF6A2ADCFEE4" author="Al-Kandari, M. &amp; Anker, A. &amp; Al-Hasheem, A. &amp; Al-Yassen, S. &amp; Sattari, Z. &amp; De Grave, S." box="[1009,1276,295,321]" pageId="15" pageNumber="194" pagination="251 - 280" refId="ref19182" refString="Al-Kandari, M., Anker, A., Al-Hasheem, A., Al-Yassen, S., Sattari, Z. &amp; De Grave, S. (2020) New records of decapod crustaceans from Kuwait (Malacostraca: Decapoda). Zootaxa, 4803 (2), 251 - 280. https: // doi. org / 10.11646 / zootaxa. 4803.2.2" type="journal article" year="2020">
Al-Kandari
<emphasis id="82B7D346FFBFFFA0FBE6AF6A2A90FEE4" box="[1143,1200,295,321]" italics="true" pageId="15" pageNumber="194">et al.</emphasis>
(2020
</bibRefCitation>
: fig. 4).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBFFFA0FF56AF072D9FFE08" pageId="15" pageNumber="194" type="distribution">
<paragraph id="B07C0F54FFBFFFA0FF56AF072D9FFE08" blockId="15.[151,1437,150,681]" pageId="15" pageNumber="194">
<emphasis id="82B7D346FFBFFFA0FF56AF072F79FEC1" bold="true" box="[199,345,330,356]" pageId="15" pageNumber="194">Distribution</emphasis>
. Indo-West Pacific from the Red Sea and
<collectingCountry id="C8D44FC4FFBFFFA0FCD3AF062DE9FEC0" box="[834,969,331,357]" name="Madagascar" pageId="15" pageNumber="194">Madagascar</collectingCountry>
to
<collectingCountry id="C8D44FC4FFBFFFA0FC60AF062A14FEC0" box="[1009,1076,331,357]" name="Japan" pageId="15" pageNumber="194">Japan</collectingCountry>
,
<collectingCountry id="C8D44FC4FFBFFFA0FBAEAF062A86FEC0" box="[1087,1190,331,357]" name="Australia" pageId="15" pageNumber="194">Australia</collectingCountry>
and
<collectingCountry id="C8D44FC4FFBFFFA0FB71AF062B0CFEC0" box="[1248,1324,331,357]" name="Samoa" pageId="15" pageNumber="194">Samoa</collectingCountry>
(
<bibRefCitation id="D45272A5FFBFFFA0FAAFAF062EF1FE2C" author="De Man, J. G." pageId="15" pageNumber="194" pagination="1 - 23" refId="ref22045" refString="De Man, J. G. (1911) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Siboga-Expeditie, 39 a 1 (2). E. J. Brill, Leiden, pp. 133 - 465, Supplement (1915), Explanations of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">De Man 1911</bibRefCitation>
;
<bibRefCitation id="D45272A5FFBFFFA0FF4CAF222FF2FE2C" author="Miya, Y. &amp; S. Miyake" box="[221,466,367,393]" pageId="15" pageNumber="194" pagination="129 - 162" refId="ref23069" refString="Miya, Y. &amp; S. Miyake. 1968 a. Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 1, 129 - 162." type="journal article" year="1968">Miya &amp; Miyake 1968</bibRefCitation>
;
<bibRefCitation id="D45272A5FFBFFFA0FE4CAF222CC3FE2C" author="Banner, D. M. &amp; Banner, A. H." box="[477,739,367,393]" pageId="15" pageNumber="194" pagination="291 - 382" refId="ref20883" refString="Banner, D. M. &amp; Banner, A. H. (1973) The alpheid shrimp of Australia, part I: the lower genera. Records of the Australian Museum, 28, 291 - 382. https: // doi. org / 10.3853 / j. 0067 - 1975.28.1973.407" type="journal article" year="1973">Banner &amp; Banner 1973</bibRefCitation>
,
<bibRefCitation id="D45272A5FFBFFFA0FD7FAF232D09FE2C" author="Banner, A. H. &amp; Banner, D. M." box="[750,809,366,393]" pageId="15" pageNumber="194" pagination="1 - 79" refId="ref20809" refString="Banner, A. H. &amp; Banner, D. M. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's &quot; The Decapoda of the Siboga Expedition, Part II: Family Alpheidae &quot; (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">1985</bibRefCitation>
;
<bibRefCitation id="D45272A5FFBFFFA0FCA5AF222D9CFE2C" author="Chace, F. A. Jr." box="[820,956,367,393]" pageId="15" pageNumber="194" pagination="1 - 99" refId="ref21274" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, i - v + 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace 1988</bibRefCitation>
; Anker 2001;
<bibRefCitation id="D45272A5FFBFFFA0FBC6AF222B4EFE2C" author="Anker, A. &amp; De Grave, S." box="[1111,1390,367,393]" pageId="15" pageNumber="194" pagination="343 - 454" refId="ref19934" refString="Anker, A. &amp; De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore (Crustacea, Decapoda). The Raffles Bulletin of Zoology, Supplement 34, 343 - 454." type="journal article" year="2016">Anker &amp; De Grave 2016</bibRefCitation>
;
<bibRefCitation id="D45272A5FFBFFFA0FAE6AF222F54FE08" author="Al-Kandari, M. &amp; Anker, A. &amp; Al-Hasheem, A. &amp; Al-Yassen, S. &amp; Sattari, Z. &amp; De Grave, S." pageId="15" pageNumber="194" pagination="251 - 280" refId="ref19182" refString="Al-Kandari, M., Anker, A., Al-Hasheem, A., Al-Yassen, S., Sattari, Z. &amp; De Grave, S. (2020) New records of decapod crustaceans from Kuwait (Malacostraca: Decapoda). Zootaxa, 4803 (2), 251 - 280. https: // doi. org / 10.11646 / zootaxa. 4803.2.2" type="journal article" year="2020">
Al-Kandari
<emphasis id="82B7D346FFBFFFA0FF69AFDE2F11FE08" box="[248,305,403,429]" italics="true" pageId="15" pageNumber="194">et al.</emphasis>
2020
</bibRefCitation>
); apparently first record from
<collectingCountry id="C8D44FC4FFBFFFA0FD54AFDE2D29FE08" box="[709,777,403,429]" name="Oman" pageId="15" pageNumber="194">Oman</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBFFFA0FF56AFFA2CE3FE50" pageId="15" pageNumber="194" type="biology_ecology">
<paragraph id="B07C0F54FFBFFFA0FF56AFFA2CE3FE50" blockId="15.[151,1437,150,681]" pageId="15" pageNumber="194">
<emphasis id="82B7D346FFBFFFA0FF56AFFA2F06FE74" bold="true" box="[199,294,439,465]" pageId="15" pageNumber="194">Ecology</emphasis>
. Commonly encountered under rocks, living and dead corals, or in coral crevices, from the lower intertidal to at least
<quantity id="773BA2B1FFBFFFA0FEE7AF962F8FFE50" box="[374,431,475,501]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="15" pageNumber="194" unit="m" value="70.0">70 m</quantity>
(
<bibRefCitation id="D45272A5FFBFFFA0FE2CAF962C96FE50" author="Miya, Y. &amp; S. Miyake" box="[445,694,475,501]" pageId="15" pageNumber="194" pagination="129 - 162" refId="ref23069" refString="Miya, Y. &amp; S. Miyake. 1968 a. Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 1, 129 - 162." type="journal article" year="1968">Miya &amp; Miyake 1968</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBFFFA0FF56AFB22D0BFD0C" pageId="15" pageNumber="194" type="discussion">
<paragraph id="B07C0F54FFBFFFA0FF56AFB22D0BFD0C" blockId="15.[151,1437,150,681]" pageId="15" pageNumber="194">
<emphasis id="82B7D346FFBFFFA0FF56AFB22F14FDBC" bold="true" box="[199,308,511,537]" pageId="15" pageNumber="194">Remarks</emphasis>
.
<taxonomicName id="77C374D7FFBFFFA0FED2AFB22FD4FDBD" authorityName="De Man" authorityYear="1910" box="[323,500,511,537]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="194" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="82B7D346FFBFFFA0FED2AFB22FD4FDBD" box="[323,500,511,537]" italics="true" pageId="15" pageNumber="194">Athanas parvus</emphasis>
</taxonomicName>
is one of the most common species of the genus, found throughout the Indo-West Pacific. The species can be identified using the combination of the colour pattern (
<figureCitation id="28F813D1FFBFFFA0FBFFAC6E2A96FD98" box="[1134,1206,547,573]" captionStart="FIGURE 7" captionStartId="14.[151,250,1948,1972]" captionTargetBox="[151,1436,673,1924]" captionTargetId="figure-201@14.[151,1436,673,1924]" captionTargetPageId="14" captionText="FIGURE 7. Athanas parvus De Man, 1910, male (cl 2.9 mm) from Masirah Island, Oman (FLMNH UF 71418); a, shrimp in life, lateral (left side); b, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350791" httpUri="https://zenodo.org/record/10350791/files/figure.png" pageId="15" pageNumber="194">Fig. 7</figureCitation>
) with the enlarged, ventrally folding chelipeds, and the presence of biunguiculate dactylus on the third to fifth pereiopods. However, no phylogeographic studies on
<taxonomicName id="77C374D7FFBFFFA0FE5EAC262C19FD21" authorityName="De Man" authorityYear="1910" box="[463,569,619,645]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="194" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="82B7D346FFBFFFA0FE5EAC262C19FD21" box="[463,569,619,645]" italics="true" pageId="15" pageNumber="194">A. parvus</emphasis>
</taxonomicName>
, including comparison of the populations from the western Indian Ocean with those from the western Pacific have been performed so far.
</paragraph>
</subSubSection>
</treatment>
</document>
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