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<mods:title id="9253AA2CDF62FDD98928F2C0504BF53C">Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)</mods:title>
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<taxonomicName id="77C374D7FFBCFFA3FF06AD532C72FC9D" ID-CoL="85MCH" authority="Coutiere, 1903" authorityName="Coutiere" authorityYear="1903" box="[151,594,798,825]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="minikoensis">
<emphasis id="82B7D346FFBCFFA3FF06AD532C72FC9D" bold="true" box="[151,594,798,825]" pageId="12" pageNumber="191">
<emphasis id="82B7D346FFBCFFA3FF06AD532FB5FC9C" bold="true" box="[151,405,798,825]" italics="true" pageId="12" pageNumber="191">Athanas minikoensis</emphasis>
<bibRefCitation id="D45272A5FFBCFFA3FE0DAD532C72FC9D" author="Coutiere, H." box="[412,594,798,825]" pageId="12" pageNumber="191" pagination="72 - 90" refId="ref21591" refString="Coutiere, H. (1903) Note sur quelques Alpheidae des Maldives et Laquedives. Bulletin de la Societe Philomatique de Paris, Serie 9, 5, 72 - 90." type="journal article" year="1903">Coutière, 1903</bibRefCitation>
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(
<figureCitation id="28F813D1FFBCFFA3FF0EAD0E2EC3FCF8" box="[159,227,835,861]" captionStart="FIGURE 6" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[158,1430,181,1919]" captionTargetId="figure-19@13.[158,1430,181,1919]" captionTargetPageId="13" captionText="FIGURE 6. Athanas minikoensis Coutière, 1903, ovigerous female (cl 3.2 mm) from Masirah Island, Oman (FLMNH UF 71422); a, shrimp in life, lateral (right side); b, same, lateral (left side); c, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350789" httpUri="https://zenodo.org/record/10350789/files/figure.png" pageId="12" pageNumber="191">Fig. 6</figureCitation>
)
</paragraph>
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<subSubSection id="F8D95CDFFFBCFFA3FF06ADCA2DF5FBBF" pageId="12" pageNumber="191" type="reference_group">
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<treatmentCitationGroup id="90D3287AFFBCFFA3FF06ADCA2B7DFC3A" box="[151,1373,903,927]" pageId="12" pageNumber="191">
<treatmentCitation id="31622945FFBCFFA3FF06ADCA2C0BFC3A" author="Coutiere, H." box="[151,555,903,927]" page="76" pageId="12" pageNumber="191" year="1903">
<taxonomicName id="77C374D7FFBCFFA3FF06ADCA2C0BFC3A" ID-CoL="85MCH" authority="Coutiere 1903: 76" authorityName="Coutiere" authorityPageNumber="76" authorityYear="1903" box="[151,555,903,927]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="minikoensis">
<emphasis id="82B7D346FFBCFFA3FF06ADCA2F4BFC3A" box="[151,363,903,927]" italics="true" pageId="12" pageNumber="191">Athanas Minikoensis</emphasis>
<bibRefCitation id="D45272A5FFBCFFA3FEE0ADCA2C0BFC3A" author="Coutiere, H." box="[369,555,903,927]" pageId="12" pageNumber="191" pagination="72 - 90" refId="ref21591" refString="Coutiere, H. (1903) Note sur quelques Alpheidae des Maldives et Laquedives. Bulletin de la Societe Philomatique de Paris, Serie 9, 5, 72 - 90." type="journal article" year="1903">Coutière 1903: 76</bibRefCitation>
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, figs. 911;
<treatmentCitation id="31622945FFBCFFA3FD36ADCA2D4CFC3A" author="Coutiere, H." box="[679,876,903,927]" page="858" pageId="12" pageNumber="191" year="1905">
<bibRefCitation id="D45272A5FFBCFFA3FD36ADCA2D4CFC3A" author="Coutiere, H." box="[679,876,903,927]" pageId="12" pageNumber="191" pagination="852 - 921" refId="ref21624" refString="Coutiere, H. (1905) Les Alpheidae. In: Stanley Gardiner, J. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes, 2, pp. 852 - 921, pls. 70 - 87." type="journal article" year="1905">Coutière 1905: 858</bibRefCitation>
</treatmentCitation>
, fig. 130;
<treatmentCitation id="31622945FFBCFFA3FC42ADC52AB3FC3A" author="De Man, J. G." box="[979,1171,903,927]" page="149" pageId="12" pageNumber="191" year="1911">
<bibRefCitation id="D45272A5FFBCFFA3FC42ADC52AB3FC3A" author="De Man, J. G." box="[979,1171,903,927]" pageId="12" pageNumber="191" pagination="1 - 23" refId="ref22045" refString="De Man, J. G. (1911) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Siboga-Expeditie, 39 a 1 (2). E. J. Brill, Leiden, pp. 133 - 465, Supplement (1915), Explanations of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">De Man 1911: 149</bibRefCitation>
</treatmentCitation>
, (1915) pl. 2, fig. 5.
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</paragraph>
<paragraph id="B07C0F54FFBCFFA3FF06ADEB2F57FC5E" blockId="12.[151,1437,903,1050]" pageId="12" pageNumber="191">
<treatmentCitationGroup id="90D3287AFFBCFFA3FF06ADEB2F57FC5E" pageId="12" pageNumber="191">
<emphasis id="82B7D346FFBCFFA3FF06ADEB2F5EFC1B" box="[151,382,934,958]" italics="true" pageId="12" pageNumber="191">
<taxonomicName id="77C374D7FFBCFFA3FF06ADEB2F44FC1B" ID-CoL="85MCH" authorityName="Coutiere" authorityYear="1903" box="[151,356,934,958]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="minikoensis">Athanas minikoensis</taxonomicName>
</emphasis>
<treatmentCitation id="31622945FFBCFFA3FEEFADEB2C1FFC1B" author="Ramadan, M. M." box="[382,575,934,958]" page="13" pageId="12" pageNumber="191" year="1936">
<bibRefCitation id="D45272A5FFBCFFA3FEEFADEB2C1FFC1B" author="Ramadan, M. M." box="[382,575,934,958]" pageId="12" pageNumber="191" pagination="1 - 43" refId="ref23335" refString="Ramadan, M. M. (1936) Report on a collection of Stomatopoda and Decapoda from Ghardaga, Red Sea. Bulletin of the Faculty of Sciences, Egyptian University, 6, 1 - 43, pls. 1, 2." type="book chapter" year="1936">Ramadan 1936: 13</bibRefCitation>
</treatmentCitation>
; (?)
<treatmentCitation id="31622945FFBCFFA3FDFDADEB2D71FC1B" author="Fourmanoir, P." box="[620,849,934,958]" page="126" pageId="12" pageNumber="191" year="1958">
<bibRefCitation id="D45272A5FFBCFFA3FDFDADEB2D71FC1B" author="Fourmanoir, P." box="[620,849,934,958]" pageId="12" pageNumber="191" pagination="115 - 127" refId="ref22183" refString="Fourmanoir, P. (1958) Crevettes Alpheides de Nosy-Be. Naturaliste Malgache, 10, 115 - 127." type="journal article" year="1958">Fourmanoir 1958: 126</bibRefCitation>
</treatmentCitation>
, fig. 15; (?)
<treatmentCitation id="31622945FFBCFFA3FC5DADEB2AEEFC1B" box="[972,1230,934,958]" pageId="12" pageNumber="191">Macnae &amp; Kalk 1969: 43</treatmentCitation>
;
<treatmentCitation id="31622945FFBCFFA3FB49ADEB2BB8FC1B" author="Kensley, B. F." box="[1240,1432,934,958]" page="118" pageId="12" pageNumber="191" year="1970">
<bibRefCitation id="D45272A5FFBCFFA3FB49ADEB2BB8FC1B" author="Kensley, B. F." box="[1240,1432,934,958]" pageId="12" pageNumber="191" pagination="180 - 187" refId="ref22573" refString="Kensley, B. F. (1970) Some decapod Crustacea from northern South-West Africa, including a new species of Hippolyte. Cimbebasia, SWA Research, Series A, 1, 180 - 187." type="journal article" year="1970">Kensley 1970: 118</bibRefCitation>
</treatmentCitation>
, fig. 12;
<treatmentCitation id="31622945FFBCFFA3FE89AD892FEFFC79" box="[280,463,964,989]" pageId="12" pageNumber="191">
<bibRefCitation id="D45272A5FFBCFFA3FE89AD892FEFFC79" author="Kensley, B. F." box="[280,463,964,989]" pageId="12" pageNumber="191" refId="ref22611" refString="Kensley, B. F. (1972) Shrimps and prawns of southern Africa. South African Museum, Cape Town, 65 pp." type="book" year="1972">Kensley 1972: 54</bibRefCitation>
</treatmentCitation>
, fig. 25j;
<treatmentCitation id="31622945FFBCFFA3FDA0AD892CC7FC79" box="[561,743,964,989]" pageId="12" pageNumber="191">
<bibRefCitation id="D45272A5FFBCFFA3FDA0AD892CC7FC79" author="Kensley, B. F." box="[561,743,964,989]" pageId="12" pageNumber="191" pagination="1 - 64" refId="ref22637" refString="Kensley, B. F. (1981) On the zoogeography of southern African decapod Crustacea, with a distributional checklist of the species. Smithsonian Contributions to Zoology, 338, 1 - 64. https: // doi. org / 10.5479 / si. 00810282.338" type="journal article" year="1981">Kensley 1981: 26</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="31622945FFBCFFA3FD62AD882A2AFC78" box="[755,1034,965,989]" pageId="12" pageNumber="191">
<bibRefCitation id="D45272A5FFBCFFA3FD62AD882A2AFC78" author="Banner, D. M. &amp; Banner, A. H." box="[755,1034,965,989]" pageId="12" pageNumber="191" pagination="1 - 99" refId="ref20938" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981: 43</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="31622945FFBCFFA3FB87AD882B0DFC78" box="[1046,1325,965,989]" pageId="12" pageNumber="191">
<bibRefCitation id="D45272A5FFBCFFA3FB87AD882B0DFC78" author="Banner, A. H. &amp; Banner, D. M." box="[1046,1325,965,989]" pageId="12" pageNumber="191" pagination="1 - 164" refId="ref20768" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983: 81</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="31622945FFBCFFA3FAA9AD882F53FC5E" pageId="12" pageNumber="191">
<bibRefCitation id="D45272A5FFBCFFA3FAA9AD882BBDFC78" author="Banner, A. H. &amp; Banner, D. M." box="[1336,1437,965,989]" pageId="12" pageNumber="191" pagination="1 - 79" refId="ref20809" refString="Banner, A. H. &amp; Banner, D. M. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's &quot; The Decapoda of the Siboga Expedition, Part II: Family Alpheidae &quot; (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">Banner &amp;</bibRefCitation>
Banner 1985: 34
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.
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</paragraph>
<paragraph id="B07C0F54FFBCFFA3FF06AA4F2DF5FBBF" blockId="12.[151,1437,903,1050]" box="[151,981,1026,1050]" pageId="12" pageNumber="191">
<treatmentCitationGroup id="90D3287AFFBCFFA3FF06AA4F2DF5FBBF" box="[151,981,1026,1050]" pageId="12" pageNumber="191">
(?)
<taxonomicName id="77C374D7FFBCFFA3FF2BAA4F2F8BFBBF" ID-CoL="85MCH" authorityName="Coutiere" authorityYear="1903" box="[186,427,1026,1050]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="minikoensis">
<emphasis id="82B7D346FFBCFFA3FF2BAA4F2F2DFBBF" box="[186,269,1026,1050]" italics="true" pageId="12" pageNumber="191">Athanas</emphasis>
cf.
<emphasis id="82B7D346FFBCFFA3FEA2AA4F2F8BFBBF" box="[307,427,1026,1050]" italics="true" pageId="12" pageNumber="191">minikoensis</emphasis>
</taxonomicName>
<treatmentCitation id="31622945FFBCFFA3FE54AA4F2CA5FBBF" box="[453,645,1026,1050]" pageId="12" pageNumber="191">
<bibRefCitation id="D45272A5FFBCFFA3FE54AA4F2CA5FBBF" author="Barnard, K. H." box="[453,645,1026,1050]" pageId="12" pageNumber="191" pagination="361 - 392" refId="ref21045" refString="Barnard, K. H. (1947) Descriptions of new species of South African Decapoda Crustacea, with notes on synonymy and new records. Annals and Magazine of Natural History, Series 11, 13 (102), 361 - 392. https: // doi. org / 10.1080 / 00222934608654562" type="journal article" year="1947">Barnard 1947: 388</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="31622945FFBCFFA3FD01AA4F2D70FBBF" box="[656,848,1026,1050]" pageId="12" pageNumber="191">
<bibRefCitation id="D45272A5FFBCFFA3FD01AA4F2D70FBBF" author="Barnard, K. H." box="[656,848,1026,1050]" pageId="12" pageNumber="191" pagination="1 - 837" refId="ref21101" refString="Barnard, K. H. (1950) Descriptive catalogue of South African decapod Crustacea. South African Museum Annals, 38, 1 - 837." type="journal article" year="1950">Barnard 1950: 731</bibRefCitation>
</treatmentCitation>
, fig. 137ad.
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</paragraph>
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<emphasis id="82B7D346FFBCFFA3FF06AA052F5AFBC7" bold="true" box="[151,378,1096,1122]" pageId="12" pageNumber="191">Material examined</emphasis>
.
<materialsCitation id="00AB0509FFBCFFA3FE19AA052FAFFB0F" collectingDate="2022-11-20" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="2" latitude="20.264334" location="Masirah Island" longLatPrecision="1" longitude="58.62397" municipality="Ghab" pageId="12" pageNumber="191" specimenCode="UF 71422" specimenCount="1">
1 ovig. female (cl
<quantity id="773BA2B1FFBCFFA3FDC6AA052C91FBC7" box="[599,689,1096,1122]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="12" pageNumber="191" unit="mm" value="3.2">3.2 mm</quantity>
),
<collectionCode id="D6D29791FFBCFFA3FD55AA042D06FBC6" box="[708,806,1097,1123]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="12" pageNumber="191" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFBCFFA3FCBFAA042D82FBC7" box="[814,930,1096,1123]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="12" pageNumber="191" type="Museum">UF 71422</specimenCode>
,
<collectingCountry id="C8D44FC4FFBCFFA3FC3FAA052DD5FBC7" box="[942,1013,1096,1122]" name="Oman" pageId="12" pageNumber="191">Oman</collectingCountry>
,
<location id="B51C598FFFBCFFA3FB91AA052A8BFBC7" LSID="urn:lsid:plazi:treatment:386ABE42FFBCFFA3FF06AD532AE7F8F3:B51C598FFFBCFFA3FB91AA052A8BFBC7" box="[1024,1195,1096,1122]" country="Oman" latitude="20.264334" longLatPrecision="1" longitude="58.62397" municipality="Ghab" name="Masirah Island" pageId="12" pageNumber="191">Masirah Island</location>
,
<quantity id="773BA2B1FFBCFFA3FB26AA052AD0FBC7" box="[1207,1264,1096,1122]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="191" unit="km" value="2.0">2 km</quantity>
north of
<collectingMunicipality id="5018952EFFBCFFA3FAC8AA052BB8FBC7" box="[1369,1432,1096,1122]" pageId="12" pageNumber="191">Ghab</collectingMunicipality>
,
<geoCoordinate id="D5F76993FFBCFFA3FF06AA212F15FB22" box="[151,309,1132,1159]" degrees="20" direction="north" minutes="15" orientation="latitude" pageId="12" pageNumber="191" precision="1" seconds="51.6" value="20.264334">20°1551.6”N</geoCoordinate>
/
<geoCoordinate id="D5F76993FFBCFFA3FEDCAA212FCAFB23" box="[333,490,1132,1159]" degrees="58" direction="east" minutes="37" orientation="longitude" pageId="12" pageNumber="191" precision="1" seconds="26.3" value="58.62397">58°3726.3”E</geoCoordinate>
, coral reef flat, fragmenting of coral rubble, depth
<quantity id="773BA2B1FFBCFFA3FBBAAA212A4EFB22" box="[1067,1134,1132,1159]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="191" unit="m" value="1.5">
<elevation id="3BEEE867FFBCFFA3FBBAAA212A4EFB22" box="[1067,1134,1132,1159]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="191" unit="m" value="1.5">1.5 m</elevation>
</quantity>
, leg.
<collectorName id="1D366A82FFBCFFA3FB3BAA212B33FB23" box="[1194,1299,1132,1158]" pageId="12" pageNumber="191">A. Anker</collectorName>
,
<date id="C47D2994FFBCFFA3FA8EAA212BBCFB23" box="[1311,1436,1132,1158]" pageId="12" pageNumber="191" value="2022-11-20">
<collectingDate id="D439D07CFFBCFFA3FA8EAA212BBCFB23" box="[1311,1436,1132,1158]" pageId="12" pageNumber="191" value="2022-11-20">20.11.2022</collectingDate>
</date>
[fcn BOMAN-14832]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBCFFA3FF56AAF92D91FA6F" pageId="12" pageNumber="191" type="description">
<paragraph id="B07C0F54FFBCFFA3FF56AAF92ABFFAB3" blockId="12.[151,1437,1096,1879]" pageId="12" pageNumber="191">
<emphasis id="82B7D346FFBCFFA3FF56AAF92F71FB6B" bold="true" box="[199,337,1204,1230]" pageId="12" pageNumber="191">Description</emphasis>
. For description and illustrations of
<taxonomicName id="77C374D7FFBCFFA3FD6BAAF92DBFFB6B" authorityName="Coutiere" authorityYear="1903" box="[762,927,1204,1230]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="minikoensis">
<emphasis id="82B7D346FFBCFFA3FD6BAAF92DBFFB6B" box="[762,927,1204,1230]" italics="true" pageId="12" pageNumber="191">A. minikoensis</emphasis>
</taxonomicName>
see
<bibRefCitation id="D45272A5FFBCFFA3FC47AAF92AA4FB6B" author="Coutiere, H." box="[982,1156,1204,1230]" pageId="12" pageNumber="191" pagination="72 - 90" refId="ref21591" refString="Coutiere, H. (1903) Note sur quelques Alpheidae des Maldives et Laquedives. Bulletin de la Societe Philomatique de Paris, Serie 9, 5, 72 - 90." type="journal article" year="1903">Coutière (1903</bibRefCitation>
,
<bibRefCitation id="D45272A5FFBCFFA3FB03AAF92AEDFB6A" author="Coutiere, H." box="[1170,1229,1204,1231]" pageId="12" pageNumber="191" pagination="852 - 921" refId="ref21624" refString="Coutiere, H. (1905) Les Alpheidae. In: Stanley Gardiner, J. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes, 2, pp. 852 - 921, pls. 70 - 87." type="journal article" year="1905">1905</bibRefCitation>
),
<bibRefCitation id="D45272A5FFBCFFA3FB75AAF82BB7FB6B" author="De Man, J. G." box="[1252,1431,1204,1231]" pageId="12" pageNumber="191" pagination="1 - 23" refId="ref22045" refString="De Man, J. G. (1911) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Siboga-Expeditie, 39 a 1 (2). E. J. Brill, Leiden, pp. 133 - 465, Supplement (1915), Explanations of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">De Man (1911)</bibRefCitation>
,
<bibRefCitation id="D45272A5FFBCFFA3FF06AA952F1BFB56" author="Barnard, K. H." box="[151,315,1240,1267]" pageId="12" pageNumber="191" pagination="1 - 837" refId="ref21101" refString="Barnard, K. H. (1950) Descriptive catalogue of South African decapod Crustacea. South African Museum Annals, 38, 1 - 837." type="journal article" year="1950">Barnard (1950</bibRefCitation>
, as
<taxonomicName id="77C374D7FFBCFFA3FEF2AA952C07FB57" authorityName="Coutiere" authorityYear="1903" box="[355,551,1240,1266]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="minikoensis">
<emphasis id="82B7D346FFBCFFA3FEF2AA952F5BFB57" box="[355,379,1240,1266]" italics="true" pageId="12" pageNumber="191">A.</emphasis>
cf.
<emphasis id="82B7D346FFBCFFA3FE32AA952C07FB57" box="[419,551,1240,1266]" italics="true" pageId="12" pageNumber="191">minikoensis</emphasis>
</taxonomicName>
, with some reservation) and
<bibRefCitation id="D45272A5FFBCFFA3FCF6AA952A36FB57" author="Kensley, B. F." box="[871,1046,1240,1266]" pageId="12" pageNumber="191" pagination="180 - 187" refId="ref22573" refString="Kensley, B. F. (1970) Some decapod Crustacea from northern South-West Africa, including a new species of Hippolyte. Cimbebasia, SWA Research, Series A, 1, 180 - 187." type="journal article" year="1970">Kensley (1970)</bibRefCitation>
; colour pattern of the single Omani female specimen herein identified as
<taxonomicName id="77C374D7FFBCFFA3FDA8AAB12CFCFAB3" authorityName="Coutiere" authorityYear="1903" box="[569,732,1276,1302]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="minikoensis">
<emphasis id="82B7D346FFBCFFA3FDA8AAB12CFCFAB3" box="[569,732,1276,1302]" italics="true" pageId="12" pageNumber="191">A. minikoensis</emphasis>
</taxonomicName>
is described below and shown in
<figureCitation id="28F813D1FFBCFFA3FBC6AAB12ABBFAB3" box="[1111,1179,1276,1302]" captionStart="FIGURE 6" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[158,1430,181,1919]" captionTargetId="figure-19@13.[158,1430,181,1919]" captionTargetPageId="13" captionText="FIGURE 6. Athanas minikoensis Coutière, 1903, ovigerous female (cl 3.2 mm) from Masirah Island, Oman (FLMNH UF 71422); a, shrimp in life, lateral (right side); b, same, lateral (left side); c, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350789" httpUri="https://zenodo.org/record/10350789/files/figure.png" pageId="12" pageNumber="191">Fig. 6</figureCitation>
.
</paragraph>
<paragraph id="B07C0F54FFBCFFA3FF56AB6D2D91FA6F" blockId="12.[151,1437,1096,1879]" pageId="12" pageNumber="191">
<emphasis id="82B7D346FFBCFFA3FF56AB6D2F4BFA9F" bold="true" box="[199,363,1312,1338]" pageId="12" pageNumber="191">Colour in life</emphasis>
. Background largely translucent; mid-dorsal surface with broad longitudinal band formed by buff white chromatophores and expanding laterally near mid-length of carapace and each pleonite in trapezoid pattern; each anterolateral margin of carapace with red broad reddish band descending from each eye towards pterygostomial region; pleon with scattered red chromatophores, most disposed in chains; chelipeds hyaline white; remaining pereiopods, pleopods and tail fan largely colourless (
<figureCitation id="28F813D1FFBCFFA3FCCEABFD2D83FA6F" box="[863,931,1456,1482]" captionStart="FIGURE 6" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[158,1430,181,1919]" captionTargetId="figure-19@13.[158,1430,181,1919]" captionTargetPageId="13" captionText="FIGURE 6. Athanas minikoensis Coutière, 1903, ovigerous female (cl 3.2 mm) from Masirah Island, Oman (FLMNH UF 71422); a, shrimp in life, lateral (right side); b, same, lateral (left side); c, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350789" httpUri="https://zenodo.org/record/10350789/files/figure.png" pageId="12" pageNumber="191">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBCFFA3FF56AB992D8BF993" pageId="12" pageNumber="191" type="distribution">
<paragraph id="B07C0F54FFBCFFA3FF56AB992D8BF993" blockId="12.[151,1437,1096,1879]" pageId="12" pageNumber="191">
<emphasis id="82B7D346FFBCFFA3FF56AB992F79FA4B" bold="true" box="[199,345,1492,1518]" pageId="12" pageNumber="191">Distribution</emphasis>
. Indo-West Pacific from the Red Sea and
<collectingCountry id="C8D44FC4FFBCFFA3FCDFAB992DC0FA4B" box="[846,992,1492,1518]" name="South Africa" pageId="12" pageNumber="191">South Africa</collectingCountry>
(as
<taxonomicName id="77C374D7FFBCFFA3FB87AB992AC4FA4B" authorityName="Coutiere" authorityYear="1903" box="[1046,1252,1492,1518]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="minikoensis">
<emphasis id="82B7D346FFBCFFA3FB87AB992A0EFA4B" box="[1046,1070,1492,1518]" italics="true" pageId="12" pageNumber="191">A.</emphasis>
cf.
<emphasis id="82B7D346FFBCFFA3FBF1AB992AC4FA4B" box="[1120,1252,1492,1518]" italics="true" pageId="12" pageNumber="191">minikoensis</emphasis>
</taxonomicName>
) to
<collectingCountry id="C8D44FC4FFBCFFA3FA89AB982EC8F9B7" name="Papua New Guinea" pageId="12" pageNumber="191">Papua New Guinea</collectingCountry>
(
<bibRefCitation id="D45272A5FFBCFFA3FF67ABB52FB9F9B7" author="Coutiere, H." box="[246,409,1528,1554]" pageId="12" pageNumber="191" pagination="72 - 90" refId="ref21591" refString="Coutiere, H. (1903) Note sur quelques Alpheidae des Maldives et Laquedives. Bulletin de la Societe Philomatique de Paris, Serie 9, 5, 72 - 90." type="journal article" year="1903">Coutière 1903</bibRefCitation>
,
<bibRefCitation id="D45272A5FFBCFFA3FE32ABB52FFEF9B6" author="Coutiere, H." box="[419,478,1528,1555]" pageId="12" pageNumber="191" pagination="852 - 921" refId="ref21624" refString="Coutiere, H. (1905) Les Alpheidae. In: Stanley Gardiner, J. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes, 2, pp. 852 - 921, pls. 70 - 87." type="journal article" year="1905">1905</bibRefCitation>
;
<bibRefCitation id="D45272A5FFBCFFA3FE78ABB42CA4F9B7" author="De Man, J. G." box="[489,644,1528,1555]" pageId="12" pageNumber="191" pagination="1 - 23" refId="ref22045" refString="De Man, J. G. (1911) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Siboga-Expeditie, 39 a 1 (2). E. J. Brill, Leiden, pp. 133 - 465, Supplement (1915), Explanations of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">De Man 1911</bibRefCitation>
;
<bibRefCitation id="D45272A5FFBCFFA3FD1EABB52D1AF9B7" author="Ramadan, M. M." box="[655,826,1528,1554]" pageId="12" pageNumber="191" pagination="1 - 43" refId="ref23335" refString="Ramadan, M. M. (1936) Report on a collection of Stomatopoda and Decapoda from Ghardaga, Red Sea. Bulletin of the Faculty of Sciences, Egyptian University, 6, 1 - 43, pls. 1, 2." type="book chapter" year="1936">Ramadan 1936</bibRefCitation>
;
<bibRefCitation id="D45272A5FFBCFFA3FCD4ABB52DC0F9B6" author="Barnard, K. H." box="[837,992,1528,1555]" pageId="12" pageNumber="191" pagination="1 - 837" refId="ref21101" refString="Barnard, K. H. (1950) Descriptive catalogue of South African decapod Crustacea. South African Museum Annals, 38, 1 - 837." type="journal article" year="1950">Barnard 1950</bibRefCitation>
;
<bibRefCitation id="D45272A5FFBCFFA3FC7AABB52AA8F9B7" author="Kensley, B. F." box="[1003,1160,1528,1554]" pageId="12" pageNumber="191" pagination="180 - 187" refId="ref22573" refString="Kensley, B. F. (1970) Some decapod Crustacea from northern South-West Africa, including a new species of Hippolyte. Cimbebasia, SWA Research, Series A, 1, 180 - 187." type="journal article" year="1970">Kensley 1970</bibRefCitation>
,
<bibRefCitation id="D45272A5FFBCFFA3FB03ABB52AEDF9B7" author="Kensley, B. F." box="[1170,1229,1528,1554]" pageId="12" pageNumber="191" refId="ref22611" refString="Kensley, B. F. (1972) Shrimps and prawns of southern Africa. South African Museum, Cape Town, 65 pp." type="book" year="1972">1972</bibRefCitation>
;
<bibRefCitation id="D45272A5FFBCFFA3FB49ABB42EF1F993" author="Banner, D. M. &amp; Banner, A. H." pageId="12" pageNumber="191" pagination="1 - 99" refId="ref20938" refString="Banner, D. M. &amp; Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981</bibRefCitation>
,
<bibRefCitation id="D45272A5FFBCFFA3FF4CA8512F38F993" author="Banner, A. H. &amp; Banner, D. M." box="[221,280,1564,1590]" pageId="12" pageNumber="191" pagination="1 - 164" refId="ref20768" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">1983</bibRefCitation>
,
<bibRefCitation id="D45272A5FFBCFFA3FEB2A8512F7FF992" author="Banner, A. H. &amp; Banner, D. M." box="[291,351,1564,1591]" pageId="12" pageNumber="191" pagination="1 - 79" refId="ref20809" refString="Banner, A. H. &amp; Banner, D. M. (1985) The alpheid shrimp of Indonesia, based upon J. G. de Man's &quot; The Decapoda of the Siboga Expedition, Part II: Family Alpheidae &quot; (1911). Marine Research in Indonesia, 25, 1 - 79. https: // doi. org / 10.14203 / mri. v 25 i 0.431" type="journal article" year="1985">1985</bibRefCitation>
); apparently first record from
<collectingCountry id="C8D44FC4FFBCFFA3FD20A8512CD5F993" box="[689,757,1564,1590]" name="Oman" pageId="12" pageNumber="191">Oman</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBCFFA3FF56A80D2AF0F9DA" pageId="12" pageNumber="191" type="biology_ecology">
<paragraph id="B07C0F54FFBCFFA3FF56A80D2AF0F9DA" blockId="12.[151,1437,1096,1879]" pageId="12" pageNumber="191">
<emphasis id="82B7D346FFBCFFA3FF56A80D2F06F9FF" bold="true" box="[199,294,1600,1626]" pageId="12" pageNumber="191">Ecology</emphasis>
. Poorly known, however, appears to be bound to coral reef habitats (
<bibRefCitation id="D45272A5FFBCFFA3FBB5A80D2AE2F9FF" author="De Man, J. G." box="[1060,1218,1600,1627]" pageId="12" pageNumber="191" pagination="1 - 23" refId="ref22045" refString="De Man, J. G. (1911) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Siboga-Expeditie, 39 a 1 (2). E. J. Brill, Leiden, pp. 133 - 465, Supplement (1915), Explanations of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">De Man 1911</bibRefCitation>
). The single Omani specimen was collected by breaking pieces of coral rubble into a sieve at a depth of about
<quantity id="773BA2B1FFBCFFA3FB18A8292AECF9DA" box="[1161,1228,1636,1663]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="191" unit="m" value="1.5">1.5 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBCFFA3FF56A8C52AE7F8F3" pageId="12" pageNumber="191" type="discussion">
<paragraph id="B07C0F54FFBCFFA3FF56A8C52AE7F8F3" blockId="12.[151,1437,1096,1879]" pageId="12" pageNumber="191">
<emphasis id="82B7D346FFBCFFA3FF56A8C52F14F907" bold="true" box="[199,308,1672,1698]" pageId="12" pageNumber="191">Remarks</emphasis>
. The single ovigerous female from Masirah Island was identified as
<taxonomicName id="77C374D7FFBCFFA3FBC4A8C52ADBF907" authorityName="Coutiere" authorityYear="1903" box="[1109,1275,1672,1698]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="minikoensis">
<emphasis id="82B7D346FFBCFFA3FBC4A8C52ADBF907" box="[1109,1275,1672,1698]" italics="true" pageId="12" pageNumber="191">A. minikoensis</emphasis>
</taxonomicName>
based mainly on the configuration of the fronto-orbital region, especially the enlarged, protruding extra-orbital teeth, the much smaller infra-orbital teeth, the absence of supra-corneal teeth, as well as the asymmetrical chelipeds carried folded beneath the body (
<figureCitation id="28F813D1FFBCFFA3FEFDA8B92F93F8AB" box="[364,435,1780,1806]" captionStart="FIGURE 6" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[158,1430,181,1919]" captionTargetId="figure-19@13.[158,1430,181,1919]" captionTargetPageId="13" captionText="FIGURE 6. Athanas minikoensis Coutière, 1903, ovigerous female (cl 3.2 mm) from Masirah Island, Oman (FLMNH UF 71422); a, shrimp in life, lateral (right side); b, same, lateral (left side); c, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350789" httpUri="https://zenodo.org/record/10350789/files/figure.png" pageId="12" pageNumber="191">Fig. 6</figureCitation>
). Despite being widespread in the Indo-West Pacific,
<taxonomicName id="77C374D7FFBCFFA3FB8EA8B92AE4F8AB" authorityName="Coutiere" authorityYear="1903" box="[1055,1220,1780,1806]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="191" phylum="Arthropoda" rank="species" species="minikoensis">
<emphasis id="82B7D346FFBCFFA3FB8EA8B92AE4F8AB" box="[1055,1220,1780,1806]" italics="true" pageId="12" pageNumber="191">A. minikoensis</emphasis>
</taxonomicName>
is not well known morphologically and ecologically (
<bibRefCitation id="D45272A5FFBCFFA3FDB7A9552CE9F897" author="De Man, J. G." box="[550,713,1816,1843]" pageId="12" pageNumber="191" pagination="1 - 23" refId="ref22045" refString="De Man, J. G. (1911) The Decapoda of the Siboga-Expedition, II: Family Alpheidae. Siboga-Expeditie, 39 a 1 (2). E. J. Brill, Leiden, pp. 133 - 465, Supplement (1915), Explanations of plates of Alpheidae, pls. 1 - 23." type="book chapter" year="1911">De Man 1911</bibRefCitation>
;
<bibRefCitation id="D45272A5FFBCFFA3FD46A9552D56F896" author="Barnard, K. H." box="[727,886,1816,1843]" pageId="12" pageNumber="191" pagination="1 - 837" refId="ref21101" refString="Barnard, K. H. (1950) Descriptive catalogue of South African decapod Crustacea. South African Museum Annals, 38, 1 - 837." type="journal article" year="1950">Barnard 1950</bibRefCitation>
;
<bibRefCitation id="D45272A5FFBCFFA3FC15A9552A05F897" author="Kensley, B. F." box="[900,1061,1816,1842]" pageId="12" pageNumber="191" pagination="180 - 187" refId="ref22573" refString="Kensley, B. F. (1970) Some decapod Crustacea from northern South-West Africa, including a new species of Hippolyte. Cimbebasia, SWA Research, Series A, 1, 180 - 187." type="journal article" year="1970">Kensley 1970</bibRefCitation>
). More material, especially fully grown and complete males, are needed to confirm the present record of the species from
<collectingCountry id="C8D44FC4FFBCFFA3FBEBA9712AE2F8F3" box="[1146,1218,1852,1878]" name="Oman" pageId="12" pageNumber="191">Oman</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>