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<document id="8FD2EEAFE5A80E5D51DA438F63D66FEE" ID-DOI="10.11646/zootaxa.5383.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10350767" ID-ZooBank="7E434B40-6346-4E6D-BC55-38EBAE24BD52" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1702283478140" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2023" docId="386ABE42FFB8FFA5FF06ADC72CC3FC48" docLanguage="en" docName="zootaxa.5383.2.5.pdf" docOrigin="Zootaxa 5383 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.2.5/52448" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athanas dentirostris Anker, Jeng &amp; Chan 2001" docType="treatment" docVersion="3" lastPageNumber="189" masterDocId="C453C63AFFB0FFAFFF91AE4D2E20FFA5" masterDocTitle="Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)" masterLastPageNumber="215" masterPageNumber="179" pageNumber="187" updateTime="1702658426235" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="C615E040841BCBA5526662DB15BE9322">Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)</mods:title>
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<mods:namePart id="517E42A400EF959FFC0D53DBC1D016A1">Anker, Arthur</mods:namePart>
<mods:affiliation id="03B4E7E366D3E9CD06BA0CA18CB5BA87">Universidade Federal de Pelotas (UFPEL), Departamento de Ecologia, Zoologia e Genética, Instituto de Biologia, Campus Universitário Capão do Leão, RS, 96010 - 610, Brazil &amp; Red Sea Research Center, King Abdullah University of Science and Technology (KAUST), Thuwal, Saudi Arabia</mods:affiliation>
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<subSubSection id="F8D95CDFFFB8FFA7FF06ADC72D23FC01" box="[151,771,906,933]" pageId="8" pageNumber="187" type="nomenclature">
<paragraph id="B07C0F54FFB8FFA7FF06ADC72D23FC01" blockId="8.[151,771,906,969]" box="[151,771,906,933]" pageId="8" pageNumber="187">
<heading id="EB34B838FFB8FFA7FF06ADC72D23FC01" bold="true" box="[151,771,906,933]" fontSize="11" level="1" pageId="8" pageNumber="187" reason="1">
<taxonomicName id="77C374D7FFB8FFA7FF06ADC72D23FC01" ID-CoL="85MBQ" authority="Anker, Jeng &amp; Chan, 2001" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[151,771,906,933]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFB8FFA7FF06ADC72D23FC01" bold="true" box="[151,771,906,933]" pageId="8" pageNumber="187">
<emphasis id="82B7D346FFB8FFA7FF06ADC72EDCFC00" bold="true" box="[151,252,906,933]" italics="true" pageId="8" pageNumber="187">Athanas</emphasis>
cf.
<emphasis id="82B7D346FFB8FFA7FEB8ADC72F92FC00" bold="true" box="[297,434,906,933]" italics="true" pageId="8" pageNumber="187">dentirostris</emphasis>
<bibRefCitation id="D45272A5FFB8FFA7FE29ADC72D23FC01" author="Anker, A. &amp; Jeng, M. S. &amp; Chan, T. Y." box="[440,771,906,933]" pageId="8" pageNumber="187" pagination="1049 - 1061" refId="ref20272" refString="Anker, A., Jeng, M. S. &amp; Chan, T. Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea), associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049 - 1061. https: // doi. org / 10.1163 / 20021975 - 99990197" type="journal article" year="2001">Anker, Jeng &amp; Chan, 2001</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB8FFA7FF06ADE22ECBFC6C" box="[151,235,943,969]" pageId="8" pageNumber="187" type="description">
<paragraph id="B07C0F54FFB8FFA7FF06ADE22ECBFC6C" blockId="8.[151,771,906,969]" box="[151,235,943,969]" pageId="8" pageNumber="187">
(
<figureCitation id="28F813D1FFB8FFA7FF0EADE22EC3FC6C" box="[159,227,943,969]" captionStart="FIGURE 4" captionStartId="9.[151,250,1533,1557]" captionTargetBox="[151,1435,181,1510]" captionTargetId="figure-19@9.[151,1435,181,1510]" captionTargetPageId="9" captionText="FIGURE 4. Athanas cf. dentirostris Anker, Jeng &amp; Chan, 2001, male (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71432); a, shrimp in life, lateral (right side); b, same, lateral (left side); c, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350781" httpUri="https://zenodo.org/record/10350781/files/figure.png" pageId="8" pageNumber="187">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB8FFA7FF06ADBE2CE7FBAE" box="[151,711,1011,1035]" pageId="8" pageNumber="187" type="reference_group">
<paragraph id="B07C0F54FFB8FFA7FF06ADBE2CE7FBAE" blockId="8.[151,711,1011,1035]" box="[151,711,1011,1035]" pageId="8" pageNumber="187">
<treatmentCitationGroup id="90D3287AFFB8FFA7FF06ADBE2CE7FBAE" box="[151,711,1011,1035]" pageId="8" pageNumber="187">
<treatmentCitation id="31622945FFB8FFA7FF06ADBE2C7DFBAE" author="Anker, A. &amp; Jeng, M. S. &amp; Chan, T. Y." box="[151,605,1011,1035]" page="1049" pageId="8" pageNumber="187" year="2001">
<taxonomicName id="77C374D7FFB8FFA7FF06ADBE2C7DFBAE" ID-CoL="85MBQ" authority="Anker et al. 2001: 1049" authorityName="Anker et al. 2001:" authorityPageNumber="1049" authorityYear="2001" box="[151,605,1011,1035]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFB8FFA7FF06ADBE2F45FBAE" box="[151,357,1011,1035]" italics="true" pageId="8" pageNumber="187">Athanas dentirostris</emphasis>
<bibRefCitation id="D45272A5FFB8FFA7FEF8ADBE2C7DFBAE" author="Anker, A. &amp; Jeng, M. S. &amp; Chan, T. Y." box="[361,605,1011,1035]" pageId="8" pageNumber="187" pagination="1049 - 1061" refId="ref20272" refString="Anker, A., Jeng, M. S. &amp; Chan, T. Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea), associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049 - 1061. https: // doi. org / 10.1163 / 20021975 - 99990197" type="journal article" year="2001">
Anker
<emphasis id="82B7D346FFB8FFA7FE3EADB92FC2FBAE" box="[431,482,1011,1035]" italics="true" pageId="8" pageNumber="187">et al.</emphasis>
2001: 1049
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 13.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB8FFA7FF06AA742F40F8A5" pageId="8" pageNumber="187" type="materials_examined">
<paragraph id="B07C0F54FFB8FFA7FF06AA742F40F8A5" blockId="8.[151,1437,1081,1900]" pageId="8" pageNumber="187">
<emphasis id="82B7D346FFB8FFA7FF06AA742F5EFBF6" bold="true" box="[151,382,1081,1107]" pageId="8" pageNumber="187">Material examined</emphasis>
.
<materialsCitation id="00AB0509FFB8FFA7FE03AA742ECAFB1A" collectingDate="2022-08-11" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="8" pageNumber="187" specimenCode="UF 71432" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFB8FFA7FE03AA742FC2FBF6" box="[402,482,1081,1107]" count="1" pageId="8" pageNumber="187" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFB8FFA7FD89AA742C57FBF6" box="[536,631,1081,1107]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="8" pageNumber="187" unit="mm" value="4.4">4.4 mm</quantity>
),
<collectionCode id="D6D29791FFB8FFA7FD1EAA772CD1FBF1" box="[655,753,1082,1108]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FD6FAA772D57FBF6" box="[766,887,1081,1108]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71432</specimenCode>
,
<collectingCountry id="C8D44FC4FFB8FFA7FC16AA742DEEFBF6" box="[903,974,1081,1107]" name="Oman" pageId="8" pageNumber="187">Oman</collectingCountry>
,
<location id="B51C598FFFB8FFA7FC4FAA742AADFBF6" LSID="urn:lsid:plazi:treatment:386ABE42FFB8FFA5FF06ADC72CC3FC48:B51C598FFFB8FFA7FC4FAA742AADFBF6" box="[990,1165,1081,1107]" country="Oman" latitude="20.587528" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" name="Masirah Island" pageId="8" pageNumber="187">Masirah Island</location>
, channel side,
<collectingMunicipality id="5018952EFFB8FFA7FAD0AA742BB8FBF6" box="[1345,1432,1081,1107]" pageId="8" pageNumber="187">Dafiyat</collectingMunicipality>
,
<geoCoordinate id="D5F76993FFB8FFA7FF06AA102F15FBDD" box="[151,309,1117,1144]" degrees="20" direction="north" minutes="35" orientation="latitude" pageId="8" pageNumber="187" precision="1" seconds="15.1" value="20.587528">20°3515.1”N</geoCoordinate>
/
<geoCoordinate id="D5F76993FFB8FFA7FEC5AA102FD1FBDD" box="[340,497,1117,1144]" degrees="58" direction="east" minutes="49" orientation="longitude" pageId="8" pageNumber="187" precision="1" seconds="36.9" value="58.826916">58°4936.9”E</geoCoordinate>
, intertidal and shallow subtidal sandflat with patches of coral rubble, rocks and seagrass, depth at low tide less than
<quantity id="773BA2B1FFB8FFA7FDD3AACC2CA7FB39" box="[578,647,1153,1180]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="187" unit="m" value="0.5">
<elevation id="3BEEE867FFB8FFA7FDD3AACC2CA7FB39" box="[578,647,1153,1180]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="187" unit="m" value="0.5">0.5 m</elevation>
</quantity>
, suction pump, in burrow, leg.
<collectorName id="1D366A82FFB8FFA7FC62AACF2A7FFB3E" box="[1011,1119,1153,1180]" pageId="8" pageNumber="187">A. Anker</collectorName>
,
<date id="C47D2994FFB8FFA7FBFCAACC2AC9FB3E" box="[1133,1257,1153,1179]" pageId="8" pageNumber="187" value="2022-08-11">
<collectingDate id="D439D07CFFB8FFA7FBFCAACC2AC9FB3E" box="[1133,1257,1153,1179]" pageId="8" pageNumber="187" value="2022-08-11">08.11.2022</collectingDate>
</date>
[fcn BOMAN-13044]
</materialsCitation>
;
<materialsCitation id="00AB0509FFB8FFA7FF67AAE82C2CFAAD" collectingDate="2022-08-11" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="8" pageNumber="187" specimenCode="UF 71419, UF 71434" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFB8FFA7FF67AAE82F62FB1A" box="[246,322,1189,1215]" count="1" pageId="8" pageNumber="187" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFB8FFA7FEFEAAE82FEAFB1A" box="[367,458,1189,1216]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="187" unit="mm" value="3.5">3.5 mm</quantity>
), 1 ovig. female (cl 4.0 mm),
<collectionCode id="D6D29791FFB8FFA7FC8DAAEB2D5EFB65" box="[796,894,1190,1216]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FC17AAEB2DDAFB1A" box="[902,1018,1189,1216]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71419</specimenCode>
, same collection data as for previous specimen [fcn BOMAN-13052]; 1 ovig. female (cl
<quantity id="773BA2B1FFB8FFA7FD5BAA842D02FB46" box="[714,802,1225,1251]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="187" unit="mm" value="3.6">3.6 mm</quantity>
),
<collectionCode id="D6D29791FFB8FFA7FCA2AA872DB5FB41" box="[819,917,1226,1252]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FC0BAA872A2BFB46" box="[922,1035,1225,1252]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71434</specimenCode>
, same collection data as for previous specimens [fcn BOMAN-13051]
</materialsCitation>
;
<materialsCitation id="00AB0509FFB8FFA7FD88AAA02A05FA89" collectingDate="2022-08-11" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="8" pageNumber="187" specimenCode="UF 71438" specimenCount="3" specimenCount-female="3">
<specimenCount id="A6C5C4DDFFB8FFA7FD88AAA02CA5FAA2" box="[537,645,1261,1287]" count="3" pageId="8" pageNumber="187" type="female">3 females</specimenCount>
(incl. 2 ovig., cl not measured, all missing major chelipeds),
<collectionCode id="D6D29791FFB8FFA7FAABAAA32BBCFAAD" box="[1338,1436,1262,1288]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FF06AB5F2F2BFA8E" box="[151,267,1297,1324]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71438</specimenCode>
, same collection data as for previous specimens [fcn BOMAN-13053]
</materialsCitation>
;
<materialsCitation id="00AB0509FFB8FFA7FBA0AB5C2FAFFA19" collectingDate="2022-11-13" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="8" pageNumber="187" specimenCode="UF 71433, UF 71415, UF 71416" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFB8FFA7FBA0AB5C2A5CFA8E" box="[1073,1148,1297,1323]" count="1" pageId="8" pageNumber="187" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFB8FFA7FB39AB5C2B20FA89" box="[1192,1280,1297,1324]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="187" unit="mm" value="3.6">3.6 mm</quantity>
, missing both chelipeds),
<collectionCode id="D6D29791FFB8FFA7FE88AB7B2F5BFAF5" box="[281,379,1334,1360]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FE13AB7B2FD7FAEA" box="[386,503,1333,1360]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71433</specimenCode>
, same collection locality as for previous specimens, leg.
<collectorName id="1D366A82FFB8FFA7FBE7AB7B2AFFFAEA" box="[1142,1247,1333,1360]" pageId="8" pageNumber="187">A. Anker</collectorName>
,
<date id="C47D2994FFB8FFA7FB7AAB782B47FAEA" box="[1259,1383,1333,1359]" pageId="8" pageNumber="187" value="2022-11-13">
<collectingDate id="D439D07CFFB8FFA7FB7AAB782B47FAEA" box="[1259,1383,1333,1359]" pageId="8" pageNumber="187" value="2022-11-13">13.11.2022</collectingDate>
</date>
[fcn BOMAN-13817]; 1 ovig. female (cl
<quantity id="773BA2B1FFB8FFA7FDA8AB142CB4FAD6" box="[569,660,1369,1395]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="8" pageNumber="187" unit="mm" value="4.7">4.7 mm</quantity>
),
<collectionCode id="D6D29791FFB8FFA7FD39AB172D2AFAD1" box="[680,778,1370,1396]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FC83AB172DA7FAD1" box="[786,903,1369,1396]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71415</specimenCode>
, same collection data as for previous specimen [fcn BOMAN-13826]; 1 ovig. female (cl
<quantity id="773BA2B1FFB8FFA7FDCAAB302C93FA32" box="[603,691,1405,1431]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="8" pageNumber="187" unit="mm" value="4.6">4.6 mm</quantity>
),
<collectionCode id="D6D29791FFB8FFA7FD55AB332D06FA3D" box="[708,806,1406,1432]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FCBAAB332DBDFA32" box="[811,925,1405,1432]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71416</specimenCode>
, same collection data as for previous specimen [fcn BOMAN-13825]
</materialsCitation>
;
<materialsCitation id="00AB0509FFB8FFA7FE0AABEC2CC2FA45" collectingDate="2022-11-13" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="8" pageNumber="187" specimenCode="UF 71436" specimenCount="4" specimenCount-female="4">
<specimenCount id="A6C5C4DDFFB8FFA7FE0AABEC2C27FA1E" box="[411,519,1441,1467]" count="4" pageId="8" pageNumber="187" type="female">4 females</specimenCount>
(incl. 3 ovig., all but one without chelipeds),
<collectionCode id="D6D29791FFB8FFA7FB97ABEF2A48FA19" box="[1030,1128,1442,1468]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FBFEABEF2AC2FA1E" box="[1135,1250,1441,1468]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71436</specimenCode>
, same collection data as for previous specimen [fcn BOMAN-13827]
</materialsCitation>
;
<materialsCitation id="00AB0509FFB8FFA7FD7FAB8B2C94F9A1" collectingDate="2022-11-13" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="8" pageNumber="187" specimenCode="UF 71430" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFB8FFA7FD7FAB8B2D18FA7A" box="[750,824,1477,1504]" count="1" pageId="8" pageNumber="187" type="male">1 male</specimenCount>
(cl 5.0 mm),
<collectionCode id="D6D29791FFB8FFA7FC40AB8B2A13FA45" box="[977,1075,1478,1504]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FBA8AB8B2A8DFA45" box="[1081,1197,1478,1504]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71430</specimenCode>
, same collection data as for previous specimen [fcn BOMAN-13824]
</materialsCitation>
;
<materialsCitation id="00AB0509FFB8FFA7FD51ABA72C83F982" collectingDate="2022-11-13" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="8" pageNumber="187" specimenCode="UF 71431" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFB8FFA7FD51ABA72D2DF9A6" box="[704,781,1513,1540]" count="1" pageId="8" pageNumber="187" type="male">1 male</specimenCount>
(cl 5.0 mm),
<collectionCode id="D6D29791FFB8FFA7FC3BABA72A2CF9A1" box="[938,1036,1514,1540]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FB85ABA72AA9F9A1" box="[1044,1161,1514,1540]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71431</specimenCode>
, same collection data as for previous specimens [fcn BOMAN-13837]
</materialsCitation>
;
<materialsCitation id="00AB0509FFB8FFA7FD20A8402BB8F9EE" collectingDate="2022-11-13" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="8" pageNumber="187" specimenCode="UF 71413" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFB8FFA7FD20A8402CDEF982" box="[689,766,1549,1575]" count="1" pageId="8" pageNumber="187" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFB8FFA7FCBFA8402DABF982" box="[814,907,1549,1575]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="8" pageNumber="187" unit="mm" value="4.7">4.7 mm</quantity>
), 2 ovig. females (cl
<quantity id="773BA2B1FFB8FFA7FB11A8402AFAF98D" box="[1152,1242,1549,1576]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="8" pageNumber="187" unit="mm" value="3.9">3.9 mm</quantity>
, 4.0 mm, former missing both chelipeds),
<collectionCode id="D6D29791FFB8FFA7FE3CA87F2C2FF9E9" box="[429,527,1586,1612]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FD84A87F2CA8F9EE" box="[533,648,1585,1612]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71413</specimenCode>
, same collection data as for previous specimens [fcn BOMAN-13814]
</materialsCitation>
;
<materialsCitation id="00AB0509FFB8FFA7FF06A8182BA2F9CA" box="[151,1410,1621,1648]" collectingDate="2022-11-13" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="8" pageNumber="187" specimenCode="UF 71421" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFB8FFA7FF06A8182EC3F9CA" box="[151,227,1621,1647]" count="1" pageId="8" pageNumber="187" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFB8FFA7FE80A8182F4DF9CA" box="[273,365,1621,1648]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="8" pageNumber="187" unit="mm" value="5.3">5.3 mm</quantity>
),
<collectionCode id="D6D29791FFB8FFA7FE10A81B2FC3F9D5" box="[385,483,1622,1648]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FE7AA81B2C40F9CA" box="[491,608,1621,1648]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71421</specimenCode>
, same collection data as for previous specimens [fcn BOMAN-13823]
</materialsCitation>
;
<materialsCitation id="00AB0509FFB8FFA7FA1FA8182FAFF91D" collectingDate="2022-11-15" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="8" pageNumber="187" specimenCode="UF 71439" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFB8FFA7FA1FA8182EEDF936" count="1" pageId="8" pageNumber="187" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFB8FFA7FF66A8342F70F936" box="[247,336,1657,1684]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="8" pageNumber="187" unit="mm" value="5.1">5.1 mm</quantity>
),
<collectionCode id="D6D29791FFB8FFA7FEF3A8372FE4F931" box="[354,452,1658,1684]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FE5BA8372C1DF936" box="[458,573,1657,1684]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71439</specimenCode>
, same collection locality as for previous specimens, leg.
<collectorName id="1D366A82FFB8FFA7FB3FA8372B35F936" box="[1198,1301,1657,1684]" pageId="8" pageNumber="187">A. Anker</collectorName>
,
<date id="C47D2994FFB8FFA7FA8EA8342BBCF931" box="[1311,1436,1657,1684]" pageId="8" pageNumber="187" value="2022-11-15">
<collectingDate id="D439D07CFFB8FFA7FA8EA8342BBCF931" box="[1311,1436,1657,1684]" pageId="8" pageNumber="187" value="2022-11-15">15.11.2022</collectingDate>
</date>
[fcn BOMAN-14056]
</materialsCitation>
;
<materialsCitation id="00AB0509FFB8FFA7FE0AA8D02F7BF97E" collectingDate="2022-11-15" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="8" pageNumber="187" specimenCode="UF 71435" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFB8FFA7FE0AA8D02FC6F912" box="[411,486,1693,1719]" count="1" pageId="8" pageNumber="187" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFB8FFA7FD83A8D02C4BF912" box="[530,619,1693,1719]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="187" unit="mm" value="4.8">4.8 mm</quantity>
),
<collectionCode id="D6D29791FFB8FFA7FDEFA8D32CC0F91D" box="[638,736,1694,1720]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FD76A8D32D7BF91D" box="[743,859,1693,1720]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71435</specimenCode>
, same collection data as for previous specimen [fcn BOMAN-14062]
</materialsCitation>
;
<materialsCitation id="00AB0509FFB8FFA7FEFBA88C2F7BF8A5" collectingDate="2022-11-15" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="8" pageNumber="187" specimenCode="UF 71420" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFB8FFA7FEFBA88C2F98F97E" box="[362,440,1729,1755]" count="1" pageId="8" pageNumber="187" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFB8FFA7FE7AA88C2C68F97E" box="[491,584,1729,1755]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="8" pageNumber="187" unit="mm" value="2.9">2.9 mm</quantity>
),
<collectionCode id="D6D29791FFB8FFA7FDCFA88F2CE0F979" box="[606,704,1730,1756]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="8" pageNumber="187" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB8FFA7FD5BA88F2D61F97E" box="[714,833,1729,1756]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="8" pageNumber="187" type="Museum">UF 71420</specimenCode>
, same collection data as for previous specimen [fcn BOMAN-14053]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB8FFA6FF56A9442E95F84F" lastPageId="9" lastPageNumber="188" pageId="8" pageNumber="187" type="description">
<paragraph id="B07C0F54FFB8FFA7FF56A9442DA5F8CE" blockId="8.[151,1437,1081,1900]" pageId="8" pageNumber="187">
<emphasis id="82B7D346FFB8FFA7FF56A9442F71F886" bold="true" box="[199,337,1801,1827]" pageId="8" pageNumber="187">Description</emphasis>
. For description and illustrations of
<taxonomicName id="77C374D7FFB8FFA7FD4EA9472D5CF886" authorityName="Anker et al. 2001:" authorityYear="1049" box="[735,892,1801,1828]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFB8FFA7FD4EA9472D5CF886" box="[735,892,1801,1828]" italics="true" pageId="8" pageNumber="187">A. dentirostris</emphasis>
</taxonomicName>
see
<bibRefCitation id="D45272A5FFB8FFA7FC38A9442A5CF886" author="Anker, A. &amp; Jeng, M. S. &amp; Chan, T. Y." box="[937,1148,1801,1827]" pageId="8" pageNumber="187" pagination="1049 - 1061" refId="ref20272" refString="Anker, A., Jeng, M. S. &amp; Chan, T. Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea), associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049 - 1061. https: // doi. org / 10.1163 / 20021975 - 99990197" type="journal article" year="2001">
Anker
<emphasis id="82B7D346FFB8FFA7FC65A9472A0AF886" box="[1012,1066,1801,1827]" italics="true" pageId="8" pageNumber="187">et al.</emphasis>
(2001)
</bibRefCitation>
; some taxonomic remarks on the Omani material tentatively identified herein as
<taxonomicName id="77C374D7FFB8FFA7FD6AA9632D9FF8E2" authorityName="Anker et al. 2001:" authorityYear="1049" box="[763,959,1837,1864]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFB8FFA7FD6AA9632D33F8ED" box="[763,787,1838,1864]" italics="true" pageId="8" pageNumber="187">A.</emphasis>
cf.
<emphasis id="82B7D346FFB8FFA7FCAEA9602D9FF8E2" box="[831,959,1837,1863]" italics="true" pageId="8" pageNumber="187">dentirostris</emphasis>
</taxonomicName>
are provided below, including description and illustration of the shrimps conspicuous colour pattern (
<figureCitation id="28F813D1FFB8FFA7FCA3A91C2D58F8CE" box="[818,888,1873,1899]" captionStart="FIGURE 4" captionStartId="9.[151,250,1533,1557]" captionTargetBox="[151,1435,181,1510]" captionTargetId="figure-19@9.[151,1435,181,1510]" captionTargetPageId="9" captionText="FIGURE 4. Athanas cf. dentirostris Anker, Jeng &amp; Chan, 2001, male (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71432); a, shrimp in life, lateral (right side); b, same, lateral (left side); c, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350781" httpUri="https://zenodo.org/record/10350781/files/figure.png" pageId="8" pageNumber="187">Fig. 4</figureCitation>
).
</paragraph>
<caption id="E4BC5FDCFFB9FFA6FF06ABB02B3FF99F" ID-DOI="http://doi.org/10.5281/zenodo.10350781" ID-Zenodo-Dep="10350781" httpUri="https://zenodo.org/record/10350781/files/figure.png" pageId="9" pageNumber="188" startId="9.[151,250,1533,1557]" targetBox="[151,1435,181,1510]" targetPageId="9" targetType="figure">
<paragraph id="B07C0F54FFB9FFA6FF06ABB02B3FF99F" blockId="9.[151,1436,1533,1594]" pageId="9" pageNumber="188">
<emphasis id="82B7D346FFB9FFA6FF06ABB02F34F9B0" bold="true" box="[151,276,1533,1557]" pageId="9" pageNumber="188">FIGURE 4.</emphasis>
<taxonomicName id="77C374D7FFB9FFA6FE8DABB32D09F9B3" authority="Anker, Jeng &amp; Chan, 2001" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[284,809,1534,1558]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="188" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFB9FFA6FE8DABB32F4FF9B3" box="[284,367,1534,1558]" italics="true" pageId="9" pageNumber="188">Athanas</emphasis>
cf.
<emphasis id="82B7D346FFB9FFA6FE09ABB32C2CF9B3" box="[408,524,1534,1558]" italics="true" pageId="9" pageNumber="188">dentirostris</emphasis>
<bibRefCitation id="D45272A5FFB9FFA6FD82ABB32D09F9B3" author="Anker, A. &amp; Jeng, M. S. &amp; Chan, T. Y." box="[531,809,1534,1558]" pageId="9" pageNumber="188" pagination="1049 - 1061" refId="ref20272" refString="Anker, A., Jeng, M. S. &amp; Chan, T. Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea), associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049 - 1061. https: // doi. org / 10.1163 / 20021975 - 99990197" type="journal article" year="2001">Anker, Jeng &amp; Chan, 2001</bibRefCitation>
</taxonomicName>
, male (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71432); a, shrimp in life, lateral (right side); b, same, lateral (left side); c, same, dorsal. Photographs by the author.
</paragraph>
</caption>
<paragraph id="B07C0F54FFB9FFA6FF56A8C12E95F84F" blockId="9.[151,1437,1676,2026]" pageId="9" pageNumber="188">
<emphasis id="82B7D346FFB9FFA6FF56A8C12F45F903" bold="true" box="[199,357,1676,1702]" pageId="9" pageNumber="188">Colour in life</emphasis>
. Background translucent whitish; carapace and pleon irregularly speckled with numerous, large, dark red dots, each composed of several red chromatophores; some white chromatophores interspersed among red ones, forming large star-shaped patches; dorsal surface of carapace and pleon with larger transverse bands formed by white chromatophores, most conspicuous on third and fourth pleonites; telson with proximal half translucent with red dots and distal half conspicuously white; antennular and antennal peduncles mostly covered with red dots, flagella with scarce red dots proximally, colourless distally; third maxillipeds and second pereiopods whitish with red dots; chelipeds hyaline white with patches of red dots, denser on merus and chelae, carpus additionally with some white patches, fingers ivory white; remaining pereiopods mostly colourless; uropods with endopod and proximal and central portion of exopod translucent with red dots, distal portion with conspicuous white patch (
<figureCitation id="28F813D1FFB9FFA6FAFEA9E12E88F84F" captionStart="FIGURE 4" captionStartId="9.[151,250,1533,1557]" captionTargetBox="[151,1435,181,1510]" captionTargetId="figure-19@9.[151,1435,181,1510]" captionTargetPageId="9" captionText="FIGURE 4. Athanas cf. dentirostris Anker, Jeng &amp; Chan, 2001, male (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71432); a, shrimp in life, lateral (right side); b, same, lateral (left side); c, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350781" httpUri="https://zenodo.org/record/10350781/files/figure.png" pageId="9" pageNumber="188">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBAFFA5FF56AEDB2C4EFF70" pageId="10" pageNumber="189" type="distribution">
<paragraph id="B07C0F54FFBAFFA5FF56AEDB2C4EFF70" blockId="10.[151,1437,150,1005]" pageId="10" pageNumber="189">
<emphasis id="82B7D346FFBAFFA5FF56AEDB2F79FF15" bold="true" box="[199,345,150,176]" pageId="10" pageNumber="189">Distribution</emphasis>
.
<collectingCountry id="C8D44FC4FFBAFFA5FEF6AEDA2FEAFF14" box="[359,458,151,177]" name="Vietnam" pageId="10" pageNumber="189">Vietnam</collectingCountry>
: Cua Luc (
<typeStatus id="6F78B1F6FFBAFFA5FDD2AEDA2C55FF14" box="[579,629,151,177]" pageId="10" pageNumber="189">type</typeStatus>
locality of
<taxonomicName id="77C374D7FFBAFFA5FD65AEDA2DB4FF14" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[756,916,151,177]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FD65AEDA2DB4FF14" box="[756,916,151,177]" italics="true" pageId="10" pageNumber="189">A. dentirostris</emphasis>
</taxonomicName>
) (
<bibRefCitation id="D45272A5FFBAFFA5FC3AAEDA2A55FF14" author="Anker, A. &amp; Jeng, M. S. &amp; Chan, T. Y." box="[939,1141,151,177]" pageId="10" pageNumber="189" pagination="1049 - 1061" refId="ref20272" refString="Anker, A., Jeng, M. S. &amp; Chan, T. Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea), associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049 - 1061. https: // doi. org / 10.1163 / 20021975 - 99990197" type="journal article" year="2001">
Anker
<emphasis id="82B7D346FFBAFFA5FC6BAEDA2A12FF14" box="[1018,1074,151,177]" italics="true" pageId="10" pageNumber="189">et al.</emphasis>
2001
</bibRefCitation>
); possibly
<collectingCountry id="C8D44FC4FFBAFFA5FB7CAEDA2B16FF14" box="[1261,1334,151,177]" name="Oman" pageId="10" pageNumber="189">Oman</collectingCountry>
: Masirah Island (
<taxonomicName id="77C374D7FFBAFFA5FF7DAEF62F8FFF70" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[236,431,187,213]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FF7DAEF62F24FF70" box="[236,260,187,213]" italics="true" pageId="10" pageNumber="189">A.</emphasis>
cf.
<emphasis id="82B7D346FFBAFFA5FEBEAEF62F8FFF70" box="[303,431,187,213]" italics="true" pageId="10" pageNumber="189">dentirostris</emphasis>
</taxonomicName>
) (present study).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBAFFA5FF56AE922B43FE08" pageId="10" pageNumber="189" type="biology_ecology">
<paragraph id="B07C0F54FFBAFFA5FF56AE922B43FE08" blockId="10.[151,1437,150,1005]" pageId="10" pageNumber="189">
<emphasis id="82B7D346FFBAFFA5FF56AE922F06FF5C" bold="true" box="[199,294,223,249]" pageId="10" pageNumber="189">Ecology</emphasis>
. All specimens of
<taxonomicName id="77C374D7FFBAFFA5FE66AE922C9AFF5C" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[503,698,223,249]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FE66AE922C30FF5C" box="[503,528,223,249]" italics="true" pageId="10" pageNumber="189">A.</emphasis>
cf.
<emphasis id="82B7D346FFBAFFA5FDABAE922C9AFF5C" box="[570,698,223,249]" italics="true" pageId="10" pageNumber="189">dentirostris</emphasis>
</taxonomicName>
were collected from burrows in the sandy-muddy intertidal or the adjacent shallow subtidal (less than
<quantity id="773BA2B1FFBAFFA5FDBDAF4E2C7BFEB8" box="[556,603,259,285]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="189" unit="m" value="1.0">1 m</quantity>
), with large patches of seagrass. The syntopic infauna, which is still being studied, included large snapping shrimps (
<taxonomicName id="77C374D7FFBAFFA5FDE2AF6A2D23FEE4" box="[627,771,295,321]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="82B7D346FFBAFFA5FDE2AF6A2CEDFEE4" box="[627,717,295,321]" italics="true" pageId="10" pageNumber="189">Alpheus</emphasis>
spp.
</taxonomicName>
) and upogebiid mud-shrimps. At least
<specimenCount id="A6C5C4DDFFBAFFA5FB29AF6A2B7FFEE4" box="[1208,1375,295,321]" count="2" pageId="10" pageNumber="189" type="generic">two specimens</specimenCount>
of
<taxonomicName id="77C374D7FFBAFFA5FA15AF6A2F1AFEC0" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FA15AF6A2BBDFEE4" box="[1412,1437,295,321]" italics="true" pageId="10" pageNumber="189">A.</emphasis>
cf.
<emphasis id="82B7D346FFBAFFA5FF2BAF062F1AFEC0" box="[186,314,331,357]" italics="true" pageId="10" pageNumber="189">dentirostris</emphasis>
</taxonomicName>
were collected from burrows together with the snapping shrimp
<taxonomicName id="77C374D7FFBAFFA5FB96AF062BB8FEC0" authority="De Haan, 1849" authorityName="De Haan" authorityYear="1849" box="[1031,1432,331,357]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="lobidens">
<emphasis id="82B7D346FFBAFFA5FB96AF062A41FEC0" box="[1031,1121,331,357]" italics="true" pageId="10" pageNumber="189">Alpheus</emphasis>
cf.
<emphasis id="82B7D346FFBAFFA5FB1BAF062AC9FEC0" box="[1162,1257,331,357]" italics="true" pageId="10" pageNumber="189">lobidens</emphasis>
<bibRefCitation id="D45272A5FFBAFFA5FB7FAF062BB8FEC0" author="De Haan, W." box="[1262,1432,331,357]" pageId="10" pageNumber="189" pagination="1 - 55" refId="ref21779" refString="De Haan, W. (1849) [1833 - 1850]. Crustacea. In: von Siebold, P. F. (Ed.), Fauna Japonica sive descriptio animalium, quae in itinere per Japoniam, jussu et auspicilis superiorum, qui summum in India Batava Imperium tenent, suspecto, annis 1823 - 1830 collegit, notis, observationibus et adumbrationibus illustravit. Apud Auctorem, Lugduni Batavorum, i - xxxi + ix - xvi + 1 - 243, pls. A - J + L - Q + 1 - 55." type="book chapter" year="1849">De Haan, 1849</bibRefCitation>
</taxonomicName>
, in the muddier part of the intertidal. The
<typeStatus id="6F78B1F6FFBAFFA5FDF1AF222CE2FE2C" box="[608,706,367,393]" pageId="10" pageNumber="189" type="holotype">holotype</typeStatus>
of
<taxonomicName id="77C374D7FFBAFFA5FD76AF222DA6FE2C" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[743,902,367,393]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FD76AF222DA6FE2C" box="[743,902,367,393]" italics="true" pageId="10" pageNumber="189">A. dentirostris</emphasis>
</taxonomicName>
was collected, according to the specimen label, together with
<taxonomicName id="77C374D7FFBAFFA5FEA2AFDE2C50FE08" baseAuthorityName="Sakai &amp; Türkay" baseAuthorityYear="1995" box="[307,624,403,429]" class="Malacostraca" family="Upogebiidae" genus="Austinogebia" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="takaoensis">
<emphasis id="82B7D346FFBAFFA5FEA2AFDE2FE7FE08" box="[307,455,403,429]" italics="true" pageId="10" pageNumber="189">Austinogebia</emphasis>
aff.
<emphasis id="82B7D346FFBAFFA5FE6BAFDE2C50FE08" box="[506,624,403,429]" italics="true" pageId="10" pageNumber="189">takaoensis</emphasis>
</taxonomicName>
(Sakai &amp; Türkay, 1995) “in fine mud and grit” (Anker
<emphasis id="82B7D346FFBAFFA5FB4AAFDE2B33FE08" box="[1243,1299,403,429]" italics="true" pageId="10" pageNumber="189">et al.</emphasis>
2021).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFBAFFA5FF56AFFA2CC3FC48" pageId="10" pageNumber="189" type="discussion">
<paragraph id="B07C0F54FFBAFFA5FF56AFFA2CC3FC48" blockId="10.[151,1437,150,1005]" pageId="10" pageNumber="189">
<emphasis id="82B7D346FFBAFFA5FF56AFFA2F14FE74" bold="true" box="[199,308,439,465]" pageId="10" pageNumber="189">Remarks</emphasis>
.
<taxonomicName id="77C374D7FFBAFFA5FED1AFFA2C02FE74" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[320,546,439,465]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FED1AFFA2C02FE74" box="[320,546,439,465]" italics="true" pageId="10" pageNumber="189">Athanas dentirostris</emphasis>
</taxonomicName>
is presently known with certainty only from the
<typeStatus id="6F78B1F6FFBAFFA5FBA7AFFA2AB8FE74" box="[1078,1176,439,465]" pageId="10" pageNumber="189" type="holotype">holotype</typeStatus>
from northern
<collectingCountry id="C8D44FC4FFBAFFA5FAACAFFA2BBCFE74" box="[1341,1436,439,465]" name="Vietnam" pageId="10" pageNumber="189">Vietnam</collectingCountry>
(
<bibRefCitation id="D45272A5FFBAFFA5FF0EAF962F53FE50" author="Anker, A. &amp; Jeng, M. S. &amp; Chan, T. Y." box="[159,371,475,501]" pageId="10" pageNumber="189" pagination="1049 - 1061" refId="ref20272" refString="Anker, A., Jeng, M. S. &amp; Chan, T. Y. (2001) Two unusual species of Alpheidae (Decapoda: Caridea), associated with upogebiid mudshrimps in the mudflats of Taiwan and Vietnam. Journal of Crustacean Biology, 21, 1049 - 1061. https: // doi. org / 10.1163 / 20021975 - 99990197" type="journal article" year="2001">
Anker
<emphasis id="82B7D346FFBAFFA5FF60AF962F0DFE50" box="[241,301,475,501]" italics="true" pageId="10" pageNumber="189">et al.</emphasis>
2001
</bibRefCitation>
). The abundant Omani material may well be conspecific with
<taxonomicName id="77C374D7FFBAFFA5FBDEAF962AD2FE50" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[1103,1266,475,501]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FBDEAF962AD2FE50" box="[1103,1266,475,501]" italics="true" pageId="10" pageNumber="189">A. dentirostris</emphasis>
</taxonomicName>
or represent a closely related species. The dorsal teeth of the rostrum appear to be more pronounced in the Omani material of
<taxonomicName id="77C374D7FFBAFFA5FF06AC6E2F41FD98" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[151,353,547,573]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FF06AC6E2E90FD98" box="[151,176,547,573]" italics="true" pageId="10" pageNumber="189">A.</emphasis>
cf.
<emphasis id="82B7D346FFBAFFA5FF70AC6E2F41FD98" box="[225,353,547,573]" italics="true" pageId="10" pageNumber="189">dentirostris</emphasis>
</taxonomicName>
compared to those of the
<typeStatus id="6F78B1F6FFBAFFA5FD09AC6E2CDAFD98" box="[664,762,547,573]" pageId="10" pageNumber="189" type="holotype">holotype</typeStatus>
of
<taxonomicName id="77C374D7FFBAFFA5FCB7AC6E2DE9FD98" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[806,969,547,573]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FCB7AC6E2DE9FD98" box="[806,969,547,573]" italics="true" pageId="10" pageNumber="189">A. dentirostris</emphasis>
</taxonomicName>
. In fact, in some Omani specimens, the distal-most teeth are very conspicuous and more closely positioned to each other, compared to the smaller and more widely spaced teeth of the
<typeStatus id="6F78B1F6FFBAFFA5FD95AC262C4AFD20" box="[516,618,619,645]" pageId="10" pageNumber="189" type="holotype">holotype</typeStatus>
. Furthermore, the armature of the finger cutting edges of the major chela of the Omani male specimens appears to be stronger than in the Vietnamese male
<typeStatus id="6F78B1F6FFBAFFA5FBD5ACC22A89FD0C" box="[1092,1193,655,681]" pageId="10" pageNumber="189" type="holotype">holotype</typeStatus>
, with one some teeth also present on the dactylus. The cutting edge of the major chela dactylus was illustrated as unarmed in Anker
<emphasis id="82B7D346FFBAFFA5FA19ACFE2E94FD54" italics="true" pageId="10" pageNumber="189">et al.</emphasis>
(2021: fig. 3c); however, it must be noted that the cutting edges and adjacent areas of both fingers are largely concealed by a dense setal cover. In all other important details, the Omani specimens seem to correspond well to the
<typeStatus id="6F78B1F6FFBAFFA5FF06AD522ED9FC9C" box="[151,249,799,825]" pageId="10" pageNumber="189" type="holotype">holotype</typeStatus>
of
<taxonomicName id="77C374D7FFBAFFA5FE8CAD522F9CFC9C" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[285,444,799,825]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FE8CAD522F9CFC9C" box="[285,444,799,825]" italics="true" pageId="10" pageNumber="189">A. dentirostris</emphasis>
</taxonomicName>
. Since no intraspecific variation can be presently assessed for the Vietnamese population of
<taxonomicName id="77C374D7FFBAFFA5FF26AD0E2F77FCF8" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[183,343,835,861]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FF26AD0E2F77FCF8" box="[183,343,835,861]" italics="true" pageId="10" pageNumber="189">A. dentirostris</emphasis>
</taxonomicName>
and considering the significantly smaller size of the
<typeStatus id="6F78B1F6FFBAFFA5FC21AD0E2A32FCF8" box="[944,1042,835,861]" pageId="10" pageNumber="189" type="holotype">holotype</typeStatus>
male (cl
<quantity id="773BA2B1FFBAFFA5FBEFAD0E2AF6FCF8" box="[1150,1238,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="189" unit="mm" value="3.8">3.8 mm</quantity>
) compared to the largest males from Masirah Island (cl 5.0
<quantity id="773BA2B1FFBAFFA5FDEFAD2B2CF9FC24" box="[638,729,870,897]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="10" pageNumber="189" unit="mm" value="5.2">5.2 mm</quantity>
), which may explain differences in the cheliped armature, the author prefers to report the Omani material as
<taxonomicName id="77C374D7FFBAFFA5FD30ADC62D45FC00" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[673,869,907,933]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FD30ADC62C9AFC00" box="[673,698,907,933]" italics="true" pageId="10" pageNumber="189">A.</emphasis>
cf.
<emphasis id="82B7D346FFBAFFA5FD74ADC62D45FC00" box="[741,869,907,933]" italics="true" pageId="10" pageNumber="189">dentirostris</emphasis>
</taxonomicName>
. The collection of more material of
<taxonomicName id="77C374D7FFBAFFA5FB6CADC62BBCFC00" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[1277,1436,907,933]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FB6CADC62BBCFC00" box="[1277,1436,907,933]" italics="true" pageId="10" pageNumber="189">A. dentirostris</emphasis>
</taxonomicName>
in
<collectingCountry id="C8D44FC4FFBAFFA5FF22ADE22F32FC6C" box="[179,274,943,969]" name="Vietnam" pageId="10" pageNumber="189">Vietnam</collectingCountry>
and across South-East Asia is needed to solve the identity of
<taxonomicName id="77C374D7FFBAFFA5FC50ADE22AA4FC6C" authorityName="Anker, Jeng &amp; Chan" authorityYear="2001" box="[961,1156,943,969]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="189" phylum="Arthropoda" rank="species" species="dentirostris">
<emphasis id="82B7D346FFBAFFA5FC50ADE22DFAFC6C" box="[961,986,943,969]" italics="true" pageId="10" pageNumber="189">A.</emphasis>
cf.
<emphasis id="82B7D346FFBAFFA5FB95ADE22AA4FC6C" box="[1028,1156,943,969]" italics="true" pageId="10" pageNumber="189">dentirostris</emphasis>
</taxonomicName>
, which may also involve comparisons of DNA sequences and colour patterns.
</paragraph>
</subSubSection>
</treatment>
</document>