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<document id="521541017D5900DBE6E2C62E098C2B2C" ID-DOI="10.11646/zootaxa.5383.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10350767" ID-ZooBank="7E434B40-6346-4E6D-BC55-38EBAE24BD52" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702283478140" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2023" docId="386ABE42FFB3FFA7FF06AFB32CE0FC9C" docLanguage="en" docName="zootaxa.5383.2.5.pdf" docOrigin="Zootaxa 5383 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.2.5/52448" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athanas claereboudti Anker 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="186" masterDocId="C453C63AFFB0FFAFFF91AE4D2E20FFA5" masterDocTitle="Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)" masterLastPageNumber="215" masterPageNumber="179" pageNumber="182" updateTime="1702284319693" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="4A271EECF3F878D726D97311A96F24BE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D7EBCC8E06155A1DB733DD6446B7C423">
<mods:title id="EDF3D748305FDCA1D2FE710652E2A3C6">Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)</mods:title>
</mods:titleInfo>
<mods:name id="80111984B180468AAC2E5A6916BE8083" type="personal">
<mods:role id="67CFBBC5858219BF798FFA74DAAC767A">
<mods:roleTerm id="7B61EAF4EEAE8F66A8F1FDF1A7F4F200">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8483895F23B44AF20809ECBAB51CFDEA">Anker, Arthur</mods:namePart>
<mods:affiliation id="40FB8747311A07A3B6EBA0052962C5A3">Universidade Federal de Pelotas (UFPEL), Departamento de Ecologia, Zoologia e Genética, Instituto de Biologia, Campus Universitário Capão do Leão, RS, 96010 - 610, Brazil &amp; Red Sea Research Center, King Abdullah University of Science and Technology (KAUST), Thuwal, Saudi Arabia</mods:affiliation>
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<mods:title id="29DF287BDC8DC2A35DD694E44E829C7C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="58EB583DE0DAEAE63C24F2F4D1E47EAE">
<mods:date id="920BACEC11773FB4E5E617EB9B014CDF">2023</mods:date>
<mods:detail id="C54093089415900BA80FD145D0B24B22" type="pubDate">
<mods:number id="AB04AF7E65088864F2FB8B812D19D5DD">2023-12-11</mods:number>
</mods:detail>
<mods:detail id="BCF3DFB3ABC90C80532AE69AF668ABE4" type="volume">
<mods:number id="6ACA2FBB169534F65D7946790BE2E5EB">5383</mods:number>
</mods:detail>
<mods:detail id="9C2D333D6FE3D8F4F47FCAA963F33384" type="issue">
<mods:number id="6BDF74E476C4463CACC7939A527A032A">2</mods:number>
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<mods:start id="3A7099BF56DF390FDFC7517E0A4AAD79">179</mods:start>
<mods:end id="B8D912679D588F062787B1E205BAED99">215</mods:end>
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<mods:url id="C68FE8F9D5566ECA984806563426E474">https://www.mapress.com/zt/article/download/zootaxa.5383.2.5/52448</mods:url>
</mods:location>
<mods:classification id="EECE7E24390C63EE38FCFF4C59F01B00">journal article</mods:classification>
<mods:identifier id="5CFEE3EE4C714857C99E6CE5071B3EB3" type="DOI">10.11646/zootaxa.5383.2.5</mods:identifier>
<mods:identifier id="2D2EE3884AA87D906B4408865FD0EF7C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C4435A0F8BB04B9DFA458B76D2D16353" type="Zenodo-Dep">10350767</mods:identifier>
<mods:identifier id="03259C48DFE0572176B12001513924C2" type="ZooBank">7E434B40-6346-4E6D-BC55-38EBAE24BD52</mods:identifier>
</mods:mods>
<treatment id="386ABE42FFB3FFA7FF06AFB32CE0FC9C" ID-DOI="http://doi.org/10.5281/zenodo.10350881" ID-Zenodo-Dep="10350881" LSID="urn:lsid:plazi:treatment:386ABE42FFB3FFA7FF06AFB32CE0FC9C" httpUri="http://treatment.plazi.org/id/386ABE42FFB3FFA7FF06AFB32CE0FC9C" lastPageId="8" lastPageNumber="186" pageId="3" pageNumber="182">
<subSubSection id="F8D95CDFFFB3FFACFF06AFB32FDEFDBC" box="[151,510,510,537]" pageId="3" pageNumber="182" type="nomenclature">
<paragraph id="B07C0F54FFB3FFACFF06AFB32FDEFDBC" blockId="3.[151,510,510,573]" box="[151,510,510,537]" pageId="3" pageNumber="182">
<heading id="EB34B838FFB3FFACFF06AFB32FDEFDBC" bold="true" box="[151,510,510,537]" fontSize="11" level="1" pageId="3" pageNumber="182" reason="1">
<emphasis id="82B7D346FFB3FFACFF06AFB32FDEFDBC" bold="true" box="[151,510,510,537]" pageId="3" pageNumber="182">
<taxonomicName id="77C374D7FFB3FFACFF06AFB32FBAFDBC" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[151,410,510,537]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
<emphasis id="82B7D346FFB3FFACFF06AFB32FBAFDBC" bold="true" box="[151,410,510,537]" italics="true" pageId="3" pageNumber="182">Athanas claereboudti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="99846E3DFFB3FFACFE30AFB22FDEFDBC" box="[417,510,511,537]" pageId="3" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB3FFACFF06AC6E2F32FD98" box="[151,274,547,573]" pageId="3" pageNumber="182" type="description">
<paragraph id="B07C0F54FFB3FFACFF06AC6E2F32FD98" blockId="3.[151,510,510,573]" box="[151,274,547,573]" pageId="3" pageNumber="182">
(
<figureCitation id="28F813D1FFB3FFACFF0EAC6E2F29FD98" box="[159,265,547,573]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="4.[151,250,1726,1750]" captionStartId-1="5.[151,250,1789,1813]" captionStartId-2="6.[151,250,1883,1907]" captionTargetBox-0="[158,1429,195,1694]" captionTargetBox-1="[159,1428,189,1758]" captionTargetBox-2="[151,1435,181,1859]" captionTargetId-0="figure-34@4.[151,1436,181,1702]" captionTargetId-1="figure-19@5.[151,1436,181,1766]" captionTargetId-2="figure-34@6.[151,1435,181,1859]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." captionText-1="FIGURE 2. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930) [a–d]; paratype, male (cl 4.9 mm) from Masirah Island, Oman (FLMNH UF 71414) [e–h]; a, left (minor) cheliped, mesial; b, same, lateral; c, same, chela with fingers open, mesial; d, right (major) cheliped, coxa and basis, lateral; e, right (major) cheliped (drawn in situ), mesial; f, same, lateral; g, distal portion of merus, carpus and chela, lateral; h, same, chela fingers, lateral." captionText-2="FIGURE 3. Athanas claereboudti sp. nov., holotype, male (cl 5.0 mm) from Masirah Island, Oman (FLMNH UF 71417); a, shrimp in life, lateral (left side); b, same, lateral (right side); c, same, dorsal. Photographs by the author." figureDoi-0="http://doi.org/10.5281/zenodo.10350769" figureDoi-1="http://doi.org/10.5281/zenodo.10350775" figureDoi-2="http://doi.org/10.5281/zenodo.10350777" httpUri-0="https://zenodo.org/record/10350769/files/figure.png" httpUri-1="https://zenodo.org/record/10350775/files/figure.png" httpUri-2="https://zenodo.org/record/10350777/files/figure.png" pageId="3" pageNumber="182">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB3FFACFF06AC292B73FC97" pageId="3" pageNumber="182" type="materials_examined">
<paragraph id="B07C0F54FFB3FFACFF06AC292B73FC97" blockId="3.[151,1437,612,2043]" pageId="3" pageNumber="182">
<emphasis id="82B7D346FFB3FFACFF06AC292F58FDDB" bold="true" box="[151,376,612,638]" pageId="3" pageNumber="182">Material examined</emphasis>
.
<materialsCitation id="00AB0509FFB3FFACFE14AC292C39FD63" collectingDate="2022-08-11" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.462223" location="Masirah Island" longLatPrecision="1" longitude="58.77772" pageId="3" pageNumber="182" specimenCode="UF 71417" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="6F78B1F6FFB3FFACFE14AC292FD2FDDB" box="[389,498,612,638]" pageId="3" pageNumber="182" type="holotype">Holotype</typeStatus>
:
<specimenCount id="A6C5C4DDFFB3FFACFE6AAC292C65FDDB" box="[507,581,612,638]" count="1" pageId="3" pageNumber="182" type="male">1 male</specimenCount>
(cl 5.0 mm),
<collectionCode id="D6D29791FFB3FFACFD4BAC282D1CFDDA" box="[730,828,613,639]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="3" pageNumber="182" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB3FFACFCD3AC282D95FDDB" box="[834,949,612,639]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="3" pageNumber="182" type="Museum">UF 71417</specimenCode>
,
<collectingCountry id="C8D44FC4FFB3FFACFC2EAC292A26FDDB" box="[959,1030,612,638]" name="Oman" pageId="3" pageNumber="182">Oman</collectingCountry>
,
<location id="B51C598FFFB3FFACFB81AC292A99FDDB" LSID="urn:lsid:plazi:treatment:386ABE42FFB3FFA7FF06AFB32CE0FC9C:B51C598FFFB3FFACFB81AC292A99FDDB" box="[1040,1209,612,638]" country="Oman" latitude="20.462223" longLatPrecision="1" longitude="58.77772" name="Masirah Island" pageId="3" pageNumber="182">Masirah Island</location>
, west coast (channel side),
<geoCoordinate id="D5F76993FFB3FFACFF4EACC52F5EFD07" box="[223,382,648,674]" degrees="20" direction="north" minutes="27" orientation="latitude" pageId="3" pageNumber="182" precision="1" seconds="44.0" value="20.462223">20°27’44.0”N</geoCoordinate>
/
<geoCoordinate id="D5F76993FFB3FFACFE0AACC52C18FD06" box="[411,568,648,675]" degrees="58" direction="east" minutes="46" orientation="longitude" pageId="3" pageNumber="182" precision="1" seconds="39.8" value="58.77772">58°46’39.8”E</geoCoordinate>
, muddy sandflat with seagrass, depth
<quantity id="773BA2B1FFB3FFACFC66ACC52A1DFD06" box="[1015,1085,648,675]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="182" unit="m" value="0.5">
<elevation id="3BEEE867FFB3FFACFC66ACC52A1DFD06" box="[1015,1085,648,675]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="182" unit="m" value="0.5">0.5 m</elevation>
</quantity>
, suction pump, leg.
<collectorName id="1D366A82FFB3FFACFABDACC52BB8FD07" box="[1324,1432,648,674]" pageId="3" pageNumber="182">A. Anker</collectorName>
,
<date id="C47D2994FFB3FFACFF06ACE12F34FD63" box="[151,276,684,710]" pageId="3" pageNumber="182" value="2022-08-11">
<collectingDate id="D439D07CFFB3FFACFF06ACE12F34FD63" box="[151,276,684,710]" pageId="3" pageNumber="182" value="2022-08-11">08.11.2022</collectingDate>
</date>
[fcn BOMAN-13043]
</materialsCitation>
.
<materialsCitation id="00AB0509FFB3FFACFDB6ACE02D10FD4F" collectingDate="2022-08-11" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.462223" location="Masirah Island" longLatPrecision="1" longitude="58.77772" pageId="3" pageNumber="182" specimenCode="UF 71414" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="6F78B1F6FFB3FFACFDB6ACE02CB8FD62" box="[551,664,685,711]" pageId="3" pageNumber="182" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="A6C5C4DDFFB3FFACFD34ACE12CD3FD63" box="[677,755,684,710]" count="1" pageId="3" pageNumber="182" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFB3FFACFCB4ACE12DA0FD62" box="[805,896,684,711]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="3" pageNumber="182" unit="mm" value="4.9">4.9 mm</quantity>
, missing minor cheliped),
<collectionCode id="D6D29791FFB3FFACFB27ACE02B38FD62" box="[1206,1304,685,711]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="3" pageNumber="182" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB3FFACFAB0ACE02BB8FD63" box="[1313,1432,684,711]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="3" pageNumber="182" type="Museum">UF 71414</specimenCode>
, same collection data as for holotype [fcn BOMAN-13042]
</materialsCitation>
;
<materialsCitation id="00AB0509FFB3FFACFCADAC9D2B6FFC97" collectingDate="2022-04-02" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="23.509027" location="Bandar Khayran" longLatPrecision="1" longitude="58.73153" pageId="3" pageNumber="182" specimenCode="UF 63930" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A6C5C4DDFFB3FFACFCADAC9D2DA8FD4F" box="[828,904,720,746]" count="1" pageId="3" pageNumber="182" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFB3FFACFC27AC9D2A2FFD4E" box="[950,1039,720,747]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="182" unit="mm" value="4.1">4.1 mm</quantity>
, missing major cheliped),
<collectionCode id="D6D29791FFB3FFACFAABAC9C2BBCFD4E" box="[1338,1436,721,747]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="3" pageNumber="182" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFB3FFACFF06ACB82F2AFCAB" box="[151,266,756,783]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="3" pageNumber="182" type="Museum">UF 63930</specimenCode>
,
<collectingCountry id="C8D44FC4FFB3FFACFE84ACB92F7CFCAB" box="[277,348,756,782]" name="Oman" pageId="3" pageNumber="182">Oman</collectingCountry>
, east of Muscat,
<location id="B51C598FFFB3FFACFD87ACB92CF0FCAB" LSID="urn:lsid:plazi:treatment:386ABE42FFB3FFA7FF06AFB32CE0FC9C:B51C598FFFB3FFACFD87ACB92CF0FCAB" box="[534,720,756,782]" country="Oman" latitude="23.509027" longLatPrecision="1" longitude="58.73153" name="Bandar Khayran" pageId="3" pageNumber="182">Bandar Khayran</location>
,
<geoCoordinate id="D5F76993FFB3FFACFD4AACB92D58FCAA" box="[731,888,756,783]" degrees="23" direction="north" minutes="30" orientation="latitude" pageId="3" pageNumber="182" precision="1" seconds="32.5" value="23.509027">23°30’32.5”N</geoCoordinate>
/
<geoCoordinate id="D5F76993FFB3FFACFC1CACB92A0BFCAA" box="[909,1067,756,783]" degrees="58" direction="east" minutes="43" orientation="longitude" pageId="3" pageNumber="182" precision="1" seconds="53.5" value="58.73153">58°43’53.5”E</geoCoordinate>
, mangrove and adjacent mudflat, at night, depth at low tide
<quantity id="773BA2B1FFB3FFACFE2DAD552C25FC96" box="[444,517,792,819]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="3" pageNumber="182" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">
<elevation id="3BEEE867FFB3FFACFE2DAD552C25FC96" box="[444,517,792,819]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="3" pageNumber="182" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 m</elevation>
</quantity>
, suction pump, in burrow, leg.
<collectorName id="1D366A82FFB3FFACFCCEAD552DE8FC97" box="[863,968,792,818]" pageId="3" pageNumber="182">A. Anker</collectorName>
,
<date id="C47D2994FFB3FFACFC42AD552A71FC97" box="[979,1105,792,818]" pageId="3" pageNumber="182" value="2022-04-02">
<collectingDate id="D439D07CFFB3FFACFC42AD552A71FC97" box="[979,1105,792,818]" pageId="3" pageNumber="182" value="2022-04-02">04.02.2022</collectingDate>
</date>
[fcn BOMAN-11363]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB3FFA8FF56AD712CD5F988" lastPageId="7" lastPageNumber="186" pageId="3" pageNumber="182" type="description">
<paragraph id="B07C0F54FFB3FFACFF56AD712B2CFC67" blockId="3.[151,1437,612,2043]" pageId="3" pageNumber="182">
<emphasis id="82B7D346FFB3FFACFF56AD712F71FCF3" bold="true" box="[199,337,828,854]" pageId="3" pageNumber="182">Description</emphasis>
. Carapace (
<figureCitation id="28F813D1FFB3FFACFE71AD712C6FFCF3" box="[480,591,828,854]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="3" pageNumber="182">Fig. 1a–c</figureCitation>
) covered with short erect setae, especially dorsally, without post-rostral tubercle. Rostrum (
<figureCitation id="28F813D1FFB3FFACFEE2AD2D2FC3FCDF" box="[371,483,864,890]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="3" pageNumber="182">Fig. 1b, c</figureCitation>
) short, reaching to mid-length of first article of antennular peduncle, subtriangular, with subacute tip; rostral carina absent. Extra-corneal teeth (
<figureCitation id="28F813D1FFB3FFACFCABADC92D85FC3B" box="[826,933,900,926]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="3" pageNumber="182">Fig. 1b, c</figureCitation>
) reduced to small bumps; infra-corneal teeth absent. Pterygostomial angle (
<figureCitation id="28F813D1FFB3FFACFE78ADE52C18FC67" box="[489,568,936,962]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="3" pageNumber="182">Fig. 1c</figureCitation>
) rounded, not produced anteriorly. Cardiac notch (
<figureCitation id="28F813D1FFB3FFACFBE0ADE52A9FFC67" box="[1137,1215,936,962]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="3" pageNumber="182">Fig. 1a</figureCitation>
) deep.
</paragraph>
<paragraph id="B07C0F54FFB3FFACFF56AD812D4DFBD3" blockId="3.[151,1437,612,2043]" pageId="3" pageNumber="182">
Pleon (
<figureCitation id="28F813D1FFB3FFACFE82AD812F41FC43" box="[275,353,972,998]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="3" pageNumber="182">Fig. 1d</figureCitation>
) mostly glabrous; first to fourth pleura rounded distoventrally; fifth pleonite angular distoventrally; sixth pleonite with subtriangular articulated plate. Telson (
<figureCitation id="28F813D1FFB3FFACFC8FADBD2D4BFBAF" box="[798,875,1008,1034]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="3" pageNumber="182">Fig. 1e</figureCitation>
) strongly tapering distally, about 1.8 times as long as proximal width; dorsal surface with two pairs of stout spiniform setae situated at some distance from lateral margins, approximately at 0.35 (anterior) and 0.65 (posterior) of telson length; posterior margin slightly concave, with two pairs of spiniform setae, mesial twice as long as lateral.
</paragraph>
<paragraph id="B07C0F54FFB3FFACFF56AACC2CC2FB1B" blockId="3.[151,1437,612,2043]" pageId="3" pageNumber="182">
Eyes (
<figureCitation id="28F813D1FFB3FFACFE80AACD2FA5FB3F" box="[273,389,1152,1178]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="3" pageNumber="182">Fig. 1b, c</figureCitation>
) partly exposed dorsally and laterally, with well pigmented but slightly reduced corneas; anteromesial margin of eyestalks rounded, unarmed.
</paragraph>
<paragraph id="B07C0F54FFB3FFACFF56AA852B79FAEB" blockId="3.[151,1437,612,2043]" pageId="3" pageNumber="182">
Antennule (
<figureCitation id="28F813D1FFB3FFACFEDCAA852FF8FB47" box="[333,472,1224,1251]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="3" pageNumber="182">Fig. 1b, c, f</figureCitation>
) moderately stout; stylocerite with acute tip, reaching or slightly overreaching mid-length of second article of peduncle; ventromesial carina with large, anteriorly directed, blunt tooth; second article about 1.3 times as long as wide; lateral flagellum with fused portion composed of five units, accessory ramus well developed, subdivided into four poorly individualised units, each carrying group of three or four aesthetascs.
</paragraph>
<paragraph id="B07C0F54FFB3FFACFF56AB152DF9FA7B" blockId="3.[151,1437,612,2043]" pageId="3" pageNumber="182">
Antenna (
<figureCitation id="28F813D1FFB3FFACFEA7AB152FE2FAD7" box="[310,450,1368,1395]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="3" pageNumber="182">Fig. 1b, c, g</figureCitation>
) with basicerite very stout, its distolateral margin armed with blunt tooth; scaphocerite broad, reaching to end of antennular peduncle; lateral margin straight to faintly convex; blade broadly convex anteriorly, by far overreaching small distolateral tooth; carpocerite very stout, significantly overreaching both antennular peduncle and scaphocerite; flagellum conspicuously thickened.
</paragraph>
<paragraph id="B07C0F54FFB3FFACFF56ABA52D81F937" blockId="3.[151,1437,612,2043]" pageId="3" pageNumber="182">
Mouthparts not dissected, typical for genus in external observation. Third maxilliped (
<figureCitation id="28F813D1FFB3FFACFB30ABA52AD4F9A7" box="[1185,1268,1512,1538]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="3" pageNumber="182">Fig. 1h</figureCitation>
) slender; coxa with somewhat produced, distally subacute lateral plate above mastigobranch; antepenultimate article somewhat curved, tapering distally, about six times as long as maximal width; penultimate article slender, about five times as long as wide; ultimate article slender, twice as long as penultimate article, tapering only apically, with unarmed tip; exopod well developed, about same length as antepenultimate article.
</paragraph>
<paragraph id="B07C0F54FFB3FFA8FF56A8D12ED8FD98" blockId="3.[151,1437,612,2043]" lastBlockId="7.[151,1437,151,2013]" lastPageId="7" lastPageNumber="186" pageId="3" pageNumber="182">
First pereiopods (= chelipeds) about 20% unequal in size, asymmetrical in shape, carried folded ventrally when not in use (
<figureCitation id="28F813D1FFB3FFACFF74A88D2F11F97F" box="[229,305,1728,1754]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1813]" captionTargetBox="[159,1428,189,1758]" captionTargetId="figure-19@5.[151,1436,181,1766]" captionTargetPageId="5" captionText="FIGURE 2. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930) [a–d]; paratype, male (cl 4.9 mm) from Masirah Island, Oman (FLMNH UF 71414) [e–h]; a, left (minor) cheliped, mesial; b, same, lateral; c, same, chela with fingers open, mesial; d, right (major) cheliped, coxa and basis, lateral; e, right (major) cheliped (drawn in situ), mesial; f, same, lateral; g, distal portion of merus, carpus and chela, lateral; h, same, chela fingers, lateral." figureDoi="http://doi.org/10.5281/zenodo.10350775" httpUri="https://zenodo.org/record/10350775/files/figure.png" pageId="3" pageNumber="182">Figs. 2</figureCitation>
,
<figureCitation id="28F813D1FFB3FFACFEADA88D2F6BF97F" box="[316,331,1728,1754]" captionStart="FIGURE 3" captionStartId="6.[151,250,1883,1907]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-34@6.[151,1435,181,1859]" captionTargetPageId="6" captionText="FIGURE 3. Athanas claereboudti sp. nov., holotype, male (cl 5.0 mm) from Masirah Island, Oman (FLMNH UF 71417); a, shrimp in life, lateral (left side); b, same, lateral (right side); c, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350777" httpUri="https://zenodo.org/record/10350777/files/figure.png" pageId="3" pageNumber="182">3</figureCitation>
). Minor cheliped (
<figureCitation id="28F813D1FFB3FFACFD80A88D2C59F97F" box="[529,633,1728,1754]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1813]" captionTargetBox="[159,1428,189,1758]" captionTargetId="figure-19@5.[151,1436,181,1766]" captionTargetPageId="5" captionText="FIGURE 2. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930) [a–d]; paratype, male (cl 4.9 mm) from Masirah Island, Oman (FLMNH UF 71414) [e–h]; a, left (minor) cheliped, mesial; b, same, lateral; c, same, chela with fingers open, mesial; d, right (major) cheliped, coxa and basis, lateral; e, right (major) cheliped (drawn in situ), mesial; f, same, lateral; g, distal portion of merus, carpus and chela, lateral; h, same, chela fingers, lateral." figureDoi="http://doi.org/10.5281/zenodo.10350775" httpUri="https://zenodo.org/record/10350775/files/figure.png" pageId="3" pageNumber="182">Fig. 2a–d</figureCitation>
) robust; basis with rudimentary exopod; ischium stout, somewhat flattened ventrally, widening distally, with one spiniform seta on ventromesial margin and row of four unevenly spaced spiniform setae on dorsal margin; merus greatly swollen, with strongly convex dorsal and ventral margins, about 2.5 times as long as greatest width, excavated ventrally, dorsal margin with small tubercles in distal half, distodorsal surface with conspicuously protruding, slightly curved, blunt process, ventrolateral margin furnished with small teeth, distal-most largest; carpus pear-shaped, narrow proximally, widening distally, ventromesial margin with small lobe, rest of surface and margins smooth; chela slightly longer than merus; palm swollen, depressed ventrolaterally, about twice as long as high (wide) in mesial view of chela, ventral margin with row of small tubercles; fingers about 0.8 times as long as palm, strongly curved, noticeably deviating from axis of palm, slender, with unarmed, blade-like cutting edges. Major cheliped (
<figureCitation id="28F813D1FFB7FFA8FE94AEDA2F51FF14" box="[261,369,151,177]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1813]" captionTargetBox="[159,1428,189,1758]" captionTargetId="figure-19@5.[151,1436,181,1766]" captionTargetPageId="5" captionText="FIGURE 2. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930) [a–d]; paratype, male (cl 4.9 mm) from Masirah Island, Oman (FLMNH UF 71414) [e–h]; a, left (minor) cheliped, mesial; b, same, lateral; c, same, chela with fingers open, mesial; d, right (major) cheliped, coxa and basis, lateral; e, right (major) cheliped (drawn in situ), mesial; f, same, lateral; g, distal portion of merus, carpus and chela, lateral; h, same, chela fingers, lateral." figureDoi="http://doi.org/10.5281/zenodo.10350775" httpUri="https://zenodo.org/record/10350775/files/figure.png" pageId="7" pageNumber="186">Fig. 2e–h</figureCitation>
) robust; basis with rudimentary exopod; ischium stout, somewhat flattened ventrally, widening distally, with row of seven equidistant spiniform setae on dorsal margin, ventrolateral margin rugose, ending distally in subacute tooth; merus less swollen than in minor cheliped, with strongly convex dorsal margin and almost straight ventral margin, about 2.8 times as long as greatest width, excavated ventrally, dorsal margin smooth, distodorsal surface with conspicuously protruding, slightly curved, blunt process similar to that of minor cheliped, ventrolateral margin evenly serrated, i.e., with small rounded teeth; carpus cup-shaped, very short, widening and with blunt lobes distally, smooth; chela distinctly longer than merus; palm swollen, depressed ventrolaterally, about 2.2 times as long as high (wide) in mesial view of chela, ventral margin with row of small tubercles; fingers about half-length of palm, somewhat curved, noticeably deviating from axis of palm, stout, gaping distally; pollex with four stout teeth on cutting edge, with large gap between two distal-most teeth; dactylus with strongly bent tip, cutting edge with two subtriangular teeth, one smaller proximal tooth and one much larger tooth at about 0.4 dactylar length, distal half of dactylus unarmed, concave.
</paragraph>
<caption id="E4BC5FDCFFB4FFABFF06A8F32B29F8C3" ID-DOI="http://doi.org/10.5281/zenodo.10350769" ID-Zenodo-Dep="10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="4" pageNumber="183" startId="4.[151,250,1726,1750]" targetBox="[158,1429,195,1694]" targetPageId="4" targetType="figure">
<paragraph id="B07C0F54FFB4FFABFF06A8F32B29F8C3" blockId="4.[151,1437,1726,1894]" pageId="4" pageNumber="183">
<emphasis id="82B7D346FFB4FFABFF06A8F32F34F973" bold="true" box="[151,276,1726,1750]" pageId="4" pageNumber="183">FIGURE 1.</emphasis>
<taxonomicName id="77C374D7FFB4FFABFE8AA8F32FD3F973" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[283,499,1726,1750]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="183" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
<emphasis id="82B7D346FFB4FFABFE8AA8F32FD3F973" box="[283,499,1726,1750]" italics="true" pageId="4" pageNumber="183">Athanas claereboudti</emphasis>
</taxonomicName>
<emphasis id="82B7D346FFB4FFABFE6BA8F32C6CF973" bold="true" box="[506,588,1726,1750]" pageId="4" pageNumber="183">
<taxonomicNameLabel id="99846E3DFFB4FFABFE6BA8F32C6CF973" box="[506,588,1726,1750]" pageId="4" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted).
</paragraph>
</caption>
<caption id="E4BC5FDCFFB5FFAAFF06A8B02B49F827" ID-DOI="http://doi.org/10.5281/zenodo.10350775" ID-Zenodo-Dep="10350775" httpUri="https://zenodo.org/record/10350775/files/figure.png" pageId="5" pageNumber="184" startId="5.[151,250,1789,1813]" targetBox="[159,1428,189,1758]" targetPageId="5" targetType="figure">
<paragraph id="B07C0F54FFB5FFAAFF06A8B02B49F827" blockId="5.[151,1436,1789,1922]" pageId="5" pageNumber="184">
<emphasis id="82B7D346FFB5FFAAFF06A8B02F35F8B3" bold="true" box="[151,277,1789,1814]" pageId="5" pageNumber="184">FIGURE 2.</emphasis>
<taxonomicName id="77C374D7FFB5FFAAFE8FA8B32FD9F8B3" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[286,505,1790,1814]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="184" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
<emphasis id="82B7D346FFB5FFAAFE8FA8B32FD9F8B3" box="[286,505,1790,1814]" italics="true" pageId="5" pageNumber="184">Athanas claereboudti</emphasis>
</taxonomicName>
<emphasis id="82B7D346FFB5FFAAFD93A8B32C75F8B3" bold="true" box="[514,597,1790,1814]" pageId="5" pageNumber="184">
<taxonomicNameLabel id="99846E3DFFB5FFAAFD93A8B32C75F8B3" box="[514,597,1790,1814]" pageId="5" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930) [a–d]; paratype, male (cl 4.9 mm) from Masirah Island, Oman (FLMNH UF 71414) [e–h]; a, left (minor) cheliped, mesial; b, same, lateral; c, same, chela with fingers open, mesial; d, right (major) cheliped, coxa and basis, lateral; e, right (major) cheliped (drawn
<emphasis id="82B7D346FFB5FFAAFF74A9272F04F824" box="[229,292,1898,1921]" italics="true" pageId="5" pageNumber="184">in situ</emphasis>
), mesial; f, same, lateral; g, distal portion of merus, carpus and chela, lateral; h, same, chela fingers, lateral.
</paragraph>
</caption>
<caption id="E4BC5FDCFFB6FFA9FF06A9162A92F832" ID-DOI="http://doi.org/10.5281/zenodo.10350777" ID-Zenodo-Dep="10350777" httpUri="https://zenodo.org/record/10350777/files/figure.png" pageId="6" pageNumber="185" startId="6.[151,250,1883,1907]" targetBox="[151,1435,181,1859]" targetPageId="6" targetType="figure">
<paragraph id="B07C0F54FFB6FFA9FF06A9162A92F832" blockId="6.[151,1436,1883,1943]" pageId="6" pageNumber="185">
<emphasis id="82B7D346FFB6FFA9FF06A9162F34F8D6" bold="true" box="[151,276,1883,1907]" pageId="6" pageNumber="185">FIGURE 3.</emphasis>
<taxonomicName id="77C374D7FFB6FFA9FE8DA9162FD6F8D6" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[284,502,1883,1907]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="185" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
<emphasis id="82B7D346FFB6FFA9FE8DA9162FD6F8D6" box="[284,502,1883,1907]" italics="true" pageId="6" pageNumber="185">Athanas claereboudti</emphasis>
</taxonomicName>
<emphasis id="82B7D346FFB6FFA9FE6CA9162C6FF8D6" bold="true" box="[509,591,1883,1907]" pageId="6" pageNumber="185">
<taxonomicNameLabel id="99846E3DFFB6FFA9FE6CA9162C6FF8D6" box="[509,591,1883,1907]" pageId="6" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 5.0 mm) from Masirah Island, Oman (FLMNH UF 71417); a, shrimp in life, lateral (left side); b, same, lateral (right side); c, same, dorsal. Photographs by the author.
</paragraph>
</caption>
<paragraph id="B07C0F54FFB7FFA8FF56AC0A2F3AFBD8" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="186">
Second pereiopod (
<figureCitation id="28F813D1FFB7FFA8FE37AC0A2FD2FDC4" box="[422,498,583,609]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="7" pageNumber="186">Fig. 1i</figureCitation>
) slender; ischium at most four times as long as maximal width; merus about 1.3 times as long as ischium; carpus with five subarticles, proximal slightly longer than sum of four others; chela longer than distal-most carpal subarticle, simple, with fingers much longer than palm. Third to fifth pereiopods relatively slender, similar in length. Third pereiopod (
<figureCitation id="28F813D1FFB7FFA8FD14ACFE2CEFFD68" box="[645,719,691,717]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="7" pageNumber="186">Fig. 1j</figureCitation>
) with ischium armed with two spiniform setae on ventrolateral surface; merus about 1.8 times as long as ischium, 5.2 times as long as maximal width, feebly convex dorsally; carpus noticeably more slender than merus, about 0.6 length of merus, with one stiff seta on distoventral margin; propodus about 1.2 times as long as carpus, with two widely spaced, stiff setae on ventral margin and one pair of spiniform setae distally, flanking base of dactylus; dactylus simple, slender, strongly curved in distal half, almost sickle-shaped, about 0.6 length of propodus. Fourth pereiopod (
<figureCitation id="28F813D1FFB7FFA8FCC2AD2A2D82FC24" box="[851,930,871,897]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="7" pageNumber="186">Fig. 1k</figureCitation>
) generally similar to third pereiopod; ischium armed with two spiniform setae on ventrolateral surface; merus noticeably slenderer than in third pereiopod, 5.8 times as long as maximal width; carpus about 0.6 times as long as merus; propodus with one stiff seta on ventral margin and one pair of spiniform setae distally, flanking base of dactylus; dactylus similar to that of third pereiopod. Fifth pereiopod (
<figureCitation id="28F813D1FFB7FFA8FECDADBA2F8AFBB4" box="[348,426,1015,1041]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="7" pageNumber="186">Fig. 1l</figureCitation>
) slenderest of walking legs; ischium with one spiniform seta on ventrolateral surface; merus about twice as long as ischium; carpus almost 0.7 length of merus; propodus distinctly longer than merus, with six rows of microserrulate setae on distolateral surface; dactylus slightly less curved than in third or fourth pereiopods.
</paragraph>
<paragraph id="B07C0F54FFB7FFA8FF56AACA2F09FB4C" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="186">
Uropod (
<figureCitation id="28F813D1FFB7FFA8FEBFAACA2FA0FB04" box="[302,384,1159,1185]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="7" pageNumber="186">Fig. 1n</figureCitation>
) with lateral lobe of protopod subacute distally; exopod with small, blunt, distolateral tooth and small adjacent spiniform seta; diaeresis straight for most part; endopod ovoid, slightly shorter and narrower than exopod.
</paragraph>
<paragraph id="B07C0F54FFB7FFA8FF56AABE2F3AFAF0" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="186">Gill-exopod formula as given for genus; basis of third maxilliped with rudimentary exopod; mastigobranchs present on coxae of third maxilliped and first to third pereiopods; setobranchs present on coxae of first to fourth pereiopods.</paragraph>
<paragraph id="B07C0F54FFB7FFA8FF56AB132CD5F988" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="186">
<emphasis id="82B7D346FFB7FFA8FF56AB132F4AFADD" bold="true" box="[199,362,1374,1400]" pageId="7" pageNumber="186">Colour in life</emphasis>
. Background hyaline whitish with slight yellowish tinge; carapace and pleon largely covered by irregular blotches of red chromatophores; mid-dorsal line of carapace, pleon and entire telson without red chromatophores, instead with patches of buff white chromatophores; carapace flanks and ventral surface of anterior pleonites mostly colourless; antennular and antennal peduncles patchily covered with red chromatophores, flagella whitish; chelipeds hyaline white with occasional red blotch on merus; remaining pereiopods and pleopods whitish; uropods with blotches of red chromatophores (
<figureCitation id="28F813D1FFB7FFA8FD33A85E2CC7F988" box="[674,743,1555,1581]" captionStart="FIGURE 3" captionStartId="6.[151,250,1883,1907]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-34@6.[151,1435,181,1859]" captionTargetPageId="6" captionText="FIGURE 3. Athanas claereboudti sp. nov., holotype, male (cl 5.0 mm) from Masirah Island, Oman (FLMNH UF 71417); a, shrimp in life, lateral (left side); b, same, lateral (right side); c, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350777" httpUri="https://zenodo.org/record/10350777/files/figure.png" pageId="7" pageNumber="186">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB7FFA8FF56A87A2AA2F9D0" pageId="7" pageNumber="186" type="etymology">
<paragraph id="B07C0F54FFB7FFA8FF56A87A2AA2F9D0" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="186">
<emphasis id="82B7D346FFB7FFA8FF56A87A2F68F9F4" bold="true" box="[199,328,1591,1617]" pageId="7" pageNumber="186">Etymology</emphasis>
. The species is named after Dr. Michel Claereboudt (formerly at Sultan Qaboos University,
<collectingCountry id="C8D44FC4FFB7FFA8FAD2A87A2BB8F9F4" box="[1347,1432,1591,1617]" name="Oman" pageId="7" pageNumber="186">Muscat</collectingCountry>
,
<collectingCountry id="C8D44FC4FFB7FFA8FF06A8162EFDF9D0" box="[151,221,1627,1653]" name="Oman" pageId="7" pageNumber="186">Oman</collectingCountry>
) for his invaluable help in the organisation of the
<collectingCountry id="C8D44FC4FFB7FFA8FC9CA8162D71F9D0" box="[781,849,1627,1653]" name="Oman" pageId="7" pageNumber="186">Oman</collectingCountry>
BioBlitz expedition series.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB7FFA8FF56A8322C4FF93C" box="[199,623,1662,1689]" pageId="7" pageNumber="186" type="materials_examined">
<paragraph id="B07C0F54FFB7FFA8FF56A8322C4FF93C" blockId="7.[151,1437,151,2013]" box="[199,623,1662,1689]" pageId="7" pageNumber="186">
<materialsCitation id="00AB0509FFB7FFA8FF56A8322C4BF93C" box="[199,619,1662,1689]" country="Oman" location="Masirah Island" pageId="7" pageNumber="186" specimenCount="1" typeStatus="holotype">
<emphasis id="82B7D346FFB7FFA8FF56A8322F40F93D" bold="true" box="[199,352,1662,1689]" pageId="7" pageNumber="186">
<typeStatus id="6F78B1F6FFB7FFA8FF56A8322F22F93C" box="[199,258,1663,1689]" pageId="7" pageNumber="186">Type</typeStatus>
locality
</emphasis>
.
<location id="B51C598FFFB7FFA8FEFFA8322C38F93C" LSID="urn:lsid:plazi:treatment:386ABE42FFB3FFA7FF06AFB32CE0FC9C:B51C598FFFB7FFA8FEFFA8322C38F93C" box="[366,536,1663,1689]" country="Oman" name="Masirah Island" pageId="7" pageNumber="186">Masirah Island</location>
,
<collectingCountry id="C8D44FC4FFB7FFA8FDB2A8322C4BF93C" box="[547,619,1663,1689]" name="Oman" pageId="7" pageNumber="186">Oman</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB7FFA8FF56A8EF2ED1F944" pageId="7" pageNumber="186" type="distribution">
<paragraph id="B07C0F54FFB7FFA8FF56A8EF2ED1F944" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="186">
<emphasis id="82B7D346FFB7FFA8FF56A8EF2F79F919" bold="true" box="[199,345,1698,1724]" pageId="7" pageNumber="186">Distribution</emphasis>
. Currently known only from two localities in
<collectingCountry id="C8D44FC4FFB7FFA8FCF1A8EE2D89F918" box="[864,937,1699,1725]" name="Oman" pageId="7" pageNumber="186">Oman</collectingCountry>
: Masirah Island and Bandar Khayran east of
<collectingCountry id="C8D44FC4FFB7FFA8FF06A88A2ECDF944" box="[151,237,1735,1761]" name="Oman" pageId="7" pageNumber="186">Muscat</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB7FFA8FF56A8A62BBBF8D4" pageId="7" pageNumber="186" type="biology_ecology">
<paragraph id="B07C0F54FFB7FFA8FF56A8A62BBBF8D4" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="186">
<emphasis id="82B7D346FFB7FFA8FF56A8A62F06F8A0" bold="true" box="[199,294,1771,1797]" pageId="7" pageNumber="186">Ecology</emphasis>
. All
<specimenCount id="A6C5C4DDFFB7FFA8FECEA8A62C34F8A0" box="[351,532,1771,1797]" count="3" pageId="7" pageNumber="186" type="generic">three specimens</specimenCount>
were collected from burrows of unknown hosts on very shallow (less than
<quantity id="773BA2B1FFB7FFA8FAF6A8A62BB4F8A0" box="[1383,1428,1771,1797]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="186" unit="m" value="1.0">1 m</quantity>
) subtidal mudflats or muddy sandflats, close to mangroves (Bandar Khayran) or seagrass beds (Masirah). The larger syntopic infauna, which is still being sorted and studied, includes large burrowing snapping shrimps (
<taxonomicName id="77C374D7FFB7FFA8FA97A97E2BB8F8E8" box="[1286,1432,1843,1869]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="186" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="82B7D346FFB7FFA8FA97A97E2B40F8E8" box="[1286,1376,1843,1869]" italics="true" pageId="7" pageNumber="186">Alpheus</emphasis>
spp.
</taxonomicName>
, including
<taxonomicName id="77C374D7FFB7FFA8FE99A91A2C5BF8D4" authority="Coutiere, 1897" authorityName="Coutiere" authorityYear="1897" box="[264,635,1879,1905]" class="Malacostraca" family="Alpheidae" genus="Alpheus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="186" phylum="Arthropoda" rank="species" species="djeddensis">
<emphasis id="82B7D346FFB7FFA8FE99A91A2F00F8D4" box="[264,288,1879,1905]" italics="true" pageId="7" pageNumber="186">A.</emphasis>
aff.
<emphasis id="82B7D346FFB7FFA8FEC5A91A2FEAF8D4" box="[340,458,1879,1905]" italics="true" pageId="7" pageNumber="186">djeddensis</emphasis>
<bibRefCitation id="D45272A5FFB7FFA8FE40A91A2C5BF8D4" author="Coutiere, H." box="[465,635,1879,1905]" pageId="7" pageNumber="186" pagination="301 - 303" refId="ref21484" refString="Coutiere, H. (1897) Note sur un nouveau genre d'Alpheides. Bulletin du Museum d'Histoire Naturelle, 1 (3), 301 - 303. https: // doi. org / 10.5962 / bhl. part. 19260" type="journal article" year="1897">Coutière, 1897</bibRefCitation>
</taxonomicName>
), callianassid ghost-shrimps, upogebiid mud-shrimps and stomatopods.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFB7FFA7FF56A9362CE0FC9C" lastPageId="8" lastPageNumber="187" pageId="7" pageNumber="186" type="discussion">
<paragraph id="B07C0F54FFB7FFA7FF56A9362CE0FC9C" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1437,151,827]" lastPageId="8" lastPageNumber="187" pageId="7" pageNumber="186">
<emphasis id="82B7D346FFB7FFA8FF56A9362F14F830" bold="true" box="[199,308,1915,1941]" pageId="7" pageNumber="186">Remarks</emphasis>
.
<taxonomicName id="77C374D7FFB7FFA8FED3A9362C12F830" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[322,562,1915,1941]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="186" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
<emphasis id="82B7D346FFB7FFA8FED3A9362C12F830" box="[322,562,1915,1941]" italics="true" pageId="7" pageNumber="186">Athanas claereboudti</emphasis>
</taxonomicName>
<emphasis id="82B7D346FFB7FFA8FDA8A9362CB3F830" bold="true" box="[569,659,1915,1941]" pageId="7" pageNumber="186">
<taxonomicNameLabel id="99846E3DFFB7FFA8FDA8A9362CB3F830" box="[569,659,1915,1941]" pageId="7" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
possesses a unique and conspicuous character not found in any other member of the genus
<taxonomicName id="77C374D7FFB7FFA8FE05A9D22FD0F81C" authorityName="Leach" authorityYear="1814" box="[404,496,1951,1977]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB7FFA8FE05A9D22FD0F81C" box="[404,496,1951,1977]" italics="true" pageId="7" pageNumber="186">Athanas</emphasis>
</taxonomicName>
. The merus of both the major and the minor cheliped of
<taxonomicName id="77C374D7FFB7FFA8FB18A9D22B17F81C" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1161,1335,1951,1977]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="186" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
<emphasis id="82B7D346FFB7FFA8FB18A9D22B17F81C" box="[1161,1335,1951,1977]" italics="true" pageId="7" pageNumber="186">A. claereboudti</emphasis>
</taxonomicName>
<emphasis id="82B7D346FFB7FFA8FAD1A9D22BBCF81C" bold="true" box="[1344,1436,1951,1977]" pageId="7" pageNumber="186">
<taxonomicNameLabel id="99846E3DFFB7FFA8FAD1A9D22BBCF81C" box="[1344,1436,1951,1977]" pageId="7" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
presents a very prominent, blunt, somewhat curved process in its distodorsal part (
<figureCitation id="28F813D1FFB7FFA8FBC2A98E2B22F878" box="[1107,1282,1987,2013]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1813]" captionTargetBox="[159,1428,189,1758]" captionTargetId="figure-19@5.[151,1436,181,1766]" captionTargetPageId="5" captionText="FIGURE 2. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930) [a–d]; paratype, male (cl 4.9 mm) from Masirah Island, Oman (FLMNH UF 71414) [e–h]; a, left (minor) cheliped, mesial; b, same, lateral; c, same, chela with fingers open, mesial; d, right (major) cheliped, coxa and basis, lateral; e, right (major) cheliped (drawn in situ), mesial; f, same, lateral; g, distal portion of merus, carpus and chela, lateral; h, same, chela fingers, lateral." figureDoi="http://doi.org/10.5281/zenodo.10350775" httpUri="https://zenodo.org/record/10350775/files/figure.png" pageId="7" pageNumber="186">Fig. 2a, b, e, g</figureCitation>
). The overall combination of morphological characters of the new species, especially those of the frontal region and chelipeds, suggests that it may be most closely related to
<taxonomicName id="77C374D7FFB8FFA7FD37AEF62AB5FF70" authority="Anker, Naderloo &amp; Marin, 2010" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" box="[678,1173,187,213]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFB8FFA7FD37AEF62D01FF71" box="[678,801,187,213]" italics="true" pageId="8" pageNumber="187">A. iranicus</emphasis>
<bibRefCitation id="D45272A5FFB8FFA7FCB6AEF62AB5FF70" author="Anker, A. &amp; Naderloo, R. &amp; Marin, I." box="[807,1173,187,213]" pageId="8" pageNumber="187" pagination="53 - 60" refId="ref20396" refString="Anker, A., Naderloo, R. &amp; Marin, I. (2010) On a new species of the shrimp genus Athanas Leach, 1814 (Crustacea, Decapoda, Alpheidae) from Iran. Zootaxa, 2372 (1), 53 - 60. https: // doi. org / 10.11646 / zootaxa. 2372.1.6" type="journal article" year="2010">Anker, Naderloo &amp; Marin, 2010</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="77C374D7FFB8FFA7FB30AEF62BB8FF70" authority="Anker, 2011" authorityName="Anker" authorityYear="2011" box="[1185,1432,187,213]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="82B7D346FFB8FFA7FB30AEF62B24FF70" box="[1185,1284,187,213]" italics="true" pageId="8" pageNumber="187">A. daviei</emphasis>
Anker, 2011
</taxonomicName>
, and
<taxonomicName id="77C374D7FFB8FFA7FF5BAE922C44FF5C" authority="Duris &amp; Anker 2014" authorityName="Duris &amp; Anker" authorityYear="2014" box="[202,612,223,251]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="manticolus">
<emphasis id="82B7D346FFB8FFA7FF5BAE922F47FF5C" box="[202,359,223,249]" italics="true" pageId="8" pageNumber="187">A. manticolus</emphasis>
<bibRefCitation id="D45272A5FFB8FFA7FEE3AEAD2C44FF5C" author="Duris, Z. &amp; Anker, A." box="[370,612,223,251]" pageId="8" pageNumber="187" pagination="550 - 558" refId="ref22106" refString="Duris, Z. &amp; Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš &amp; Anker 2014</bibRefCitation>
</taxonomicName>
, all of them also being infaunal, symbiotic species (Anker
<emphasis id="82B7D346FFB8FFA7FA86AE922B6BFF5C" box="[1303,1355,223,249]" italics="true" pageId="8" pageNumber="187">et al</emphasis>
. 2010;
<bibRefCitation id="D45272A5FFB8FFA7FF06AF4E2F0EFEB8" author="Anker, A." box="[151,302,259,285]" pageId="8" pageNumber="187" pagination="1 - 22" refId="ref19338" refString="Anker, A. (2011 b) Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia. Zootaxa, 2734 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2734.1.1" type="journal article" year="2011">Anker 2011b</bibRefCitation>
;
<bibRefCitation id="D45272A5FFB8FFA7FEAAAF492C0BFEB8" author="Duris, Z. &amp; Anker, A." box="[315,555,259,287]" pageId="8" pageNumber="187" pagination="550 - 558" refId="ref22106" refString="Duris, Z. &amp; Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš &amp; Anker 2014</bibRefCitation>
). However,
<taxonomicName id="77C374D7FFB8FFA7FD24AF4E2D43FEB8" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[693,867,259,285]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
<emphasis id="82B7D346FFB8FFA7FD24AF4E2D43FEB8" box="[693,867,259,285]" italics="true" pageId="8" pageNumber="187">A. claereboudti</emphasis>
</taxonomicName>
<emphasis id="82B7D346FFB8FFA7FCFAAF4E2DE6FEB8" bold="true" box="[875,966,259,285]" pageId="8" pageNumber="187">
<taxonomicNameLabel id="99846E3DFFB8FFA7FCFAAF4E2DE6FEB8" box="[875,966,259,285]" pageId="8" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be easily separated from each of the three aforementioned species by at least three morphological characters, in addition to the bulging distodorsal process on the cheliped meri. For instance,
<taxonomicName id="77C374D7FFB8FFA7FDECAF062D09FEC0" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[637,809,331,357]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
<emphasis id="82B7D346FFB8FFA7FDECAF062D09FEC0" box="[637,809,331,357]" italics="true" pageId="8" pageNumber="187">A. claereboudti</emphasis>
</taxonomicName>
<emphasis id="82B7D346FFB8FFA7FCA1AF062DA9FEC0" bold="true" box="[816,905,331,357]" pageId="8" pageNumber="187">
<taxonomicNameLabel id="99846E3DFFB8FFA7FCA1AF062DA9FEC0" box="[816,905,331,357]" pageId="8" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="77C374D7FFB8FFA7FB8CAF062AB8FEC1" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" box="[1053,1176,331,357]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFB8FFA7FB8CAF062AB8FEC1" box="[1053,1176,331,357]" italics="true" pageId="8" pageNumber="187">A. iranicus</emphasis>
</taxonomicName>
by the absence of setal brushes on the carpus and chela (which are characteristic of
<taxonomicName id="77C374D7FFB8FFA7FCA5AF222D8EFE2D" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" box="[820,942,367,393]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFB8FFA7FCA5AF222D8EFE2D" box="[820,942,367,393]" italics="true" pageId="8" pageNumber="187">A. iranicus</emphasis>
</taxonomicName>
, see also below); the posterior margin of the telson slightly concave (
<emphasis id="82B7D346FFB8FFA7FE36AFDE2F9EFE09" box="[423,446,403,428]" italics="true" pageId="8" pageNumber="187">vs</emphasis>
. broadly rounded in
<taxonomicName id="77C374D7FFB8FFA7FD39AFDE2D03FE09" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" box="[680,803,403,429]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFB8FFA7FD39AFDE2D03FE09" box="[680,803,403,429]" italics="true" pageId="8" pageNumber="187">A. iranicus</emphasis>
</taxonomicName>
); and the antennal scaphocerite reaching or very slightly overreaching the end of the antennular peduncle (
<emphasis id="82B7D346FFB8FFA7FD53AFFA2CC1FE75" box="[706,737,439,464]" italics="true" pageId="8" pageNumber="187">vs.</emphasis>
reaching far beyond it in
<taxonomicName id="77C374D7FFB8FFA7FB90AFFA2A5CFE75" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" box="[1025,1148,439,465]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFB8FFA7FB90AFFA2A5CFE75" box="[1025,1148,439,465]" italics="true" pageId="8" pageNumber="187">A. iranicus</emphasis>
</taxonomicName>
) (cf.
<figureCitation id="28F813D1FFB8FFA7FB28AFFA2B11FE74" box="[1209,1329,439,465]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1813]" captionTargetBox="[159,1428,189,1758]" captionTargetId="figure-19@5.[151,1436,181,1766]" captionTargetPageId="5" captionText="FIGURE 2. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930) [a–d]; paratype, male (cl 4.9 mm) from Masirah Island, Oman (FLMNH UF 71414) [e–h]; a, left (minor) cheliped, mesial; b, same, lateral; c, same, chela with fingers open, mesial; d, right (major) cheliped, coxa and basis, lateral; e, right (major) cheliped (drawn in situ), mesial; f, same, lateral; g, distal portion of merus, carpus and chela, lateral; h, same, chela fingers, lateral." figureDoi="http://doi.org/10.5281/zenodo.10350775" httpUri="https://zenodo.org/record/10350775/files/figure.png" pageId="8" pageNumber="187">Figs. 2e–h</figureCitation>
; Anker
<emphasis id="82B7D346FFB8FFA7FA19AFFA2E8DFE50" italics="true" pageId="8" pageNumber="187">et al</emphasis>
. 2010: figs. 2B, E, 3A, B, E). Similarly, the new species differs from
<taxonomicName id="77C374D7FFB8FFA7FC26AF962A3AFE50" authorityName="Anker" authorityYear="2011" box="[951,1050,475,501]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="82B7D346FFB8FFA7FC26AF962A3AFE50" box="[951,1050,475,501]" italics="true" pageId="8" pageNumber="187">A. daviei</emphasis>
</taxonomicName>
by the noticeably shorter rostrum; the longer stylocerite; the much more swollen cheliped meri; and the slenderer second pereiopod (cf.
<figureCitation id="28F813D1FFB8FFA7FA8FAFB22B4BFDBC" box="[1310,1387,511,537]" captionStart="FIGURE 1" captionStartId="4.[151,250,1726,1750]" captionTargetBox="[158,1429,195,1694]" captionTargetId="figure-34@4.[151,1436,181,1702]" captionTargetPageId="4" captionText="FIGURE 1. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, posterior portion of pleon and telson, lateral; e, telson, dorsal; f, tooth of ventromesial carina of first article of antennular peduncle, lateral; g, antennal scaphocerite, dorsal; h, third maxilliped, lateral; i, second pereiopod, lateral; j, third pereiopod, lateral; k, fourth pereiopod, lateral; l, fifth pereiopod, lateral; m, appendix masculina and appendix interna of second pleopod, anterior (lateral); n, uropod, dorsal (marginal setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.10350769" httpUri="https://zenodo.org/record/10350769/files/figure.png" pageId="8" pageNumber="187">Figs. 1</figureCitation>
,
<figureCitation id="28F813D1FFB8FFA7FAEAAFB22E88FD98" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1813]" captionTargetBox="[159,1428,189,1758]" captionTargetId="figure-19@5.[151,1436,181,1766]" captionTargetPageId="5" captionText="FIGURE 2. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930) [a–d]; paratype, male (cl 4.9 mm) from Masirah Island, Oman (FLMNH UF 71414) [e–h]; a, left (minor) cheliped, mesial; b, same, lateral; c, same, chela with fingers open, mesial; d, right (major) cheliped, coxa and basis, lateral; e, right (major) cheliped (drawn in situ), mesial; f, same, lateral; g, distal portion of merus, carpus and chela, lateral; h, same, chela fingers, lateral." figureDoi="http://doi.org/10.5281/zenodo.10350775" httpUri="https://zenodo.org/record/10350775/files/figure.png" pageId="8" pageNumber="187">2a, b</figureCitation>
; Anker 2011: figs. 3A, 4A, B, D–F). Finally,
<taxonomicName id="77C374D7FFB8FFA7FD37AC6E2D72FD98" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[678,850,547,573]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
<emphasis id="82B7D346FFB8FFA7FD37AC6E2D72FD98" box="[678,850,547,573]" italics="true" pageId="8" pageNumber="187">A. claereboudti</emphasis>
</taxonomicName>
<emphasis id="82B7D346FFB8FFA7FCC8AC6E2D93FD98" bold="true" box="[857,947,547,573]" pageId="8" pageNumber="187">
<taxonomicNameLabel id="99846E3DFFB8FFA7FCC8AC6E2D93FD98" box="[857,947,547,573]" pageId="8" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be easily separated from
<taxonomicName id="77C374D7FFB8FFA7FA93AC6E2BBCFD98" authorityName="Duris &amp; Anker" authorityYear="2014" box="[1282,1436,547,573]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="manticolus">
<emphasis id="82B7D346FFB8FFA7FA93AC6E2BBCFD98" box="[1282,1436,547,573]" italics="true" pageId="8" pageNumber="187">A. manticolus</emphasis>
</taxonomicName>
by the dorsally non-dentate rostrum; the absence of postrostral tubercle; and the more unequal and asymmetrical chelipeds, each with a much more swollen merus (cf.
<figureCitation id="28F813D1FFB8FFA7FD68AC262D68FD20" box="[761,840,619,645]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1813]" captionTargetBox="[159,1428,189,1758]" captionTargetId="figure-19@5.[151,1436,181,1766]" captionTargetPageId="5" captionText="FIGURE 2. Athanas claereboudti sp. nov., paratype, male (cl 4.1 mm) from Bandar Khayran, Oman (FLMNH UF 63930) [a–d]; paratype, male (cl 4.9 mm) from Masirah Island, Oman (FLMNH UF 71414) [e–h]; a, left (minor) cheliped, mesial; b, same, lateral; c, same, chela with fingers open, mesial; d, right (major) cheliped, coxa and basis, lateral; e, right (major) cheliped (drawn in situ), mesial; f, same, lateral; g, distal portion of merus, carpus and chela, lateral; h, same, chela fingers, lateral." figureDoi="http://doi.org/10.5281/zenodo.10350775" httpUri="https://zenodo.org/record/10350775/files/figure.png" pageId="8" pageNumber="187">Figs. 2</figureCitation>
,
<figureCitation id="28F813D1FFB8FFA7FCC5AC262D44FD20" box="[852,868,619,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1883,1907]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-34@6.[151,1435,181,1859]" captionTargetPageId="6" captionText="FIGURE 3. Athanas claereboudti sp. nov., holotype, male (cl 5.0 mm) from Masirah Island, Oman (FLMNH UF 71417); a, shrimp in life, lateral (left side); b, same, lateral (right side); c, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350777" httpUri="https://zenodo.org/record/10350777/files/figure.png" pageId="8" pageNumber="187">3</figureCitation>
;
<bibRefCitation id="D45272A5FFB8FFA7FCE3AC212A7EFD20" author="Duris, Z. &amp; Anker, A." box="[882,1118,619,647]" pageId="8" pageNumber="187" pagination="550 - 558" refId="ref22106" refString="Duris, Z. &amp; Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš &amp; Anker 2014</bibRefCitation>
: figs. 2A, C, 4). In addition,
<taxonomicName id="77C374D7FFB8FFA7FF06ACC22F63FD0C" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[151,323,655,681]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
<emphasis id="82B7D346FFB8FFA7FF06ACC22F63FD0C" box="[151,323,655,681]" italics="true" pageId="8" pageNumber="187">A. claereboudti</emphasis>
</taxonomicName>
<emphasis id="82B7D346FFB8FFA7FEDBACC22F83FD0C" bold="true" box="[330,419,655,681]" pageId="8" pageNumber="187">
<taxonomicNameLabel id="99846E3DFFB8FFA7FEDBACC22F83FD0C" box="[330,419,655,681]" pageId="8" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="77C374D7FFB8FFA7FD42ACC22D6EFD0D" authorityName="Anker, Naderloo &amp; Marin" authorityYear="2010" box="[723,846,655,681]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="82B7D346FFB8FFA7FD42ACC22D6EFD0D" box="[723,846,655,681]" italics="true" pageId="8" pageNumber="187">A. iranicus</emphasis>
</taxonomicName>
,
<taxonomicName id="77C374D7FFB8FFA7FCCDACC22D9FFD0C" authorityName="Anker" authorityYear="2011" box="[860,959,655,681]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="82B7D346FFB8FFA7FCCDACC22D9FFD0C" box="[860,959,655,681]" italics="true" pageId="8" pageNumber="187">A. daviei</emphasis>
</taxonomicName>
and
<taxonomicName id="77C374D7FFB8FFA7FC64ACC22AAFFD0C" authorityName="Duris &amp; Anker" authorityYear="2014" box="[1013,1167,655,681]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="manticolus">
<emphasis id="82B7D346FFB8FFA7FC64ACC22AAFFD0C" box="[1013,1167,655,681]" italics="true" pageId="8" pageNumber="187">A. manticolus</emphasis>
</taxonomicName>
by its diagnostic colour pattern (cf.
<figureCitation id="28F813D1FFB8FFA7FE89ACFE2F44FD68" box="[280,356,691,717]" captionStart="FIGURE 3" captionStartId="6.[151,250,1883,1907]" captionTargetBox="[151,1435,181,1859]" captionTargetId="figure-34@6.[151,1435,181,1859]" captionTargetPageId="6" captionText="FIGURE 3. Athanas claereboudti sp. nov., holotype, male (cl 5.0 mm) from Masirah Island, Oman (FLMNH UF 71417); a, shrimp in life, lateral (left side); b, same, lateral (right side); c, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350777" httpUri="https://zenodo.org/record/10350777/files/figure.png" pageId="8" pageNumber="187">Figs. 3</figureCitation>
,
<figureCitation id="28F813D1FFB8FFA7FEE0ACFF2FA1FD68" box="[369,385,690,717]" captionStart="FIGURE 5" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,453,1677]" captionTargetId="figure-138@11.[151,1436,453,1677]" captionTargetPageId="11" captionText="FIGURE 5. Athanas iranicus Anker, Naderloo &amp; Marin, 2010, ovigerous female (cl 3.6 mm) from Bandar Khayran, Oman (FLMNH UF 68858); a, shrimp in life, right antennal flagellum broken, dorsal; b, same, lateral (left side). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350783" httpUri="https://zenodo.org/record/10350783/files/figure.png" pageId="8" pageNumber="187">5</figureCitation>
, Anker 2011: fig. 8E, F;
<bibRefCitation id="D45272A5FFB8FFA7FD02ACF92D5AFD68" author="Duris, Z. &amp; Anker, A." box="[659,890,691,719]" pageId="8" pageNumber="187" pagination="550 - 558" refId="ref22106" refString="Duris, Z. &amp; Anker, A. (2014) Athanas manticolus sp. nov., a new stomatopod-associated alpheid shrimp from Vietnam (Crustacea, Decapoda). Zootaxa, 3784 (5), 550 - 558. https: // doi. org / 10.11646 / zootaxa. 3784.5.4" type="journal article" year="2014">Ďuriš &amp; Anker 2014</bibRefCitation>
: fig. 6A, B). All other known species of
<taxonomicName id="77C374D7FFB8FFA7FAA8ACFE2BB5FD68" authorityName="Leach" authorityYear="1814" box="[1337,1429,691,717]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="genus">
<emphasis id="82B7D346FFB8FFA7FAA8ACFE2BB5FD68" box="[1337,1429,691,717]" italics="true" pageId="8" pageNumber="187">Athanas</emphasis>
</taxonomicName>
, including all infaunal species, present more significant morphological differences with
<taxonomicName id="77C374D7FFB8FFA7FBF2AC9A2B2DFD54" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1123,1293,727,753]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="187" phylum="Arthropoda" rank="species" species="claereboudti" status="sp. nov.">
<emphasis id="82B7D346FFB8FFA7FBF2AC9A2B2DFD54" box="[1123,1293,727,753]" italics="true" pageId="8" pageNumber="187">A. claereboudti</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="99846E3DFFB8FFA7FA85AC9A2B4DFD54" box="[1300,1389,727,753]" pageId="8" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and (where known) also differ from the new species by their colour patterns (e.g.,
<bibRefCitation id="D45272A5FFB8FFA7FC6BACB62AB8FCB0" author="Hayashi, K. I." box="[1018,1176,763,789]" pageId="8" pageNumber="187" pagination="395 - 403" refId="ref22261" refString="Hayashi, K. I. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of mantis shrimp. Crustaceana, 75, 395 - 403. https: // doi. org / 10.1163 / 156854002760095462" type="journal article" year="2002">Hayashi 2002</bibRefCitation>
;
<bibRefCitation id="D45272A5FFB8FFA7FB30ACB62BB8FCB0" author="Anker, A. &amp; Komai, T." box="[1185,1432,763,789]" pageId="8" pageNumber="187" pagination="45 - 54" refId="ref20339" refString="Anker, A. &amp; Komai, T. (2010) Description of a new species of Athanas Leach, 1814 (Crustacea, Decapoda, Alpheidae) from Madagascar and Japan. Zootaxa, 2680 (1), 45 - 54. https: // doi. org / 10.11646 / zootaxa. 2680.1.4" type="journal article" year="2010">Anker &amp; Komai 2010</bibRefCitation>
;
<bibRefCitation id="D45272A5FFB8FFA7FF06AD522F0CFC9C" author="Anker, A." box="[151,300,799,825]" pageId="8" pageNumber="187" pagination="1 - 22" refId="ref19338" refString="Anker, A. (2011 b) Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia. Zootaxa, 2734 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2734.1.1" type="journal article" year="2011">Anker 2011b</bibRefCitation>
;
<bibRefCitation id="D45272A5FFB8FFA7FEA9AD522F9EFC9C" author="Marin, I." box="[312,446,799,825]" pageId="8" pageNumber="187" pagination="50 - 62" refId="ref22879" refString="Marin, I. (2017) Athanas alpheusophilus sp. nov. (Decapoda: Alpheidae) - a new Alpheus - associated shrimp from the Russian coast of the Sea of Japan. Zootaxa, 4324 (1), 50 - 62. https: // doi. org / 10.11646 / zootaxa. 4324.1.3" type="journal article" year="2017">Marin 2017</bibRefCitation>
;
<bibRefCitation id="D45272A5FFB8FFA7FE58AD522C95FC9C" author="Anker, A. &amp; Duris, Z." box="[457,693,799,827]" pageId="8" pageNumber="187" pagination="288 - 300" refId="ref20021" refString="Anker, A. &amp; Duris, Z. (2022) Athanas philippei n. sp., a new alpheid shrimp associated with stomatopods in New Caledonia (Decapoda: Caridea). Zootaxa, 5094 (2), 288 - 300. https: // doi. org / 10.11646 / zootaxa. 5094.2.4" type="journal article" year="2022">Anker &amp; Ďuriš 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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