treatments-xml/data/38/6A/BE/386ABE42FFA9FFB6FF06AEDA2D41FAD1.xml
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<mods:title id="70924F5036AFE04F655A034863AA937A">Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)</mods:title>
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<mods:namePart id="D300A9F3C2A4088CA8D9A9E42441338F">Anker, Arthur</mods:namePart>
<mods:affiliation id="02959B0BD37561A4AC906F5F418A9963">Universidade Federal de Pelotas (UFPEL), Departamento de Ecologia, Zoologia e Genética, Instituto de Biologia, Campus Universitário Capão do Leão, RS, 96010 - 610, Brazil &amp; Red Sea Research Center, King Abdullah University of Science and Technology (KAUST), Thuwal, Saudi Arabia</mods:affiliation>
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<taxonomicName id="77C374D7FFA9FFB6FF06AEDA2CABFF14" ID-CoL="85MDG" authority="Berggren, 1991" authorityName="Berggren" authorityYear="1991" box="[151,651,151,178]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="204" phylum="Arthropoda" rank="species" species="rubricinctuta">
<emphasis id="82B7D346FFA9FFB6FF06AEDA2CABFF14" bold="true" box="[151,651,151,178]" pageId="25" pageNumber="204">
<emphasis id="82B7D346FFA9FFB6FF06AEDA2FE6FF17" bold="true" box="[151,454,151,178]" italics="true" pageId="25" pageNumber="204">Athanopsis rubricinctuta</emphasis>
<bibRefCitation id="D45272A5FFA9FFB6FE5CAEDA2CABFF14" author="Berggren, M." box="[461,651,151,177]" pageId="25" pageNumber="204" pagination="166 - 178" refId="ref21169" refString="Berggren, M. (1991) Athanopsis rubricinctuta, new species (Decapoda: Natantia: Alpheidae), a shrimp associated with an echiuroid at Inhaca Island, Mocambique. Journal of Crustacean Biology, 11, 166 - 178. https: // doi. org / 10.2307 / 1548555" type="journal article" year="1991">Berggren, 1991</bibRefCitation>
</emphasis>
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<paragraph id="B07C0F54FFA9FFB6FF06AEF62ED9FF70" blockId="25.[151,651,151,213]" box="[151,249,187,213]" pageId="25" pageNumber="204">
(
<figureCitation id="28F813D1FFA9FFB6FF0EAEF62ED0FF70" box="[159,240,187,213]" captionStart="FIGURE 13" captionStartId="26.[151,250,1900,1924]" captionTargetBox="[151,1436,181,1876]" captionTargetId="figure-34@26.[151,1436,181,1876]" captionTargetPageId="26" captionText="FIGURE 13. Athanopsis rubricinctuta Berggren, 1991, male (cl 5.4 mm) from Masirah Island, Oman (FLMNH UF 71437) [a, b]; male (cl 5.8 mm) from the same locality (FLMNH UF 71443) [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, shrimp in life (different individual), lateral (left side). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350817" httpUri="https://zenodo.org/record/10350817/files/figure.png" pageId="25" pageNumber="204">Fig. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFA9FFB6FF06AF4D2CF0FEB2" box="[151,720,255,280]" pageId="25" pageNumber="204" type="reference_group">
<paragraph id="B07C0F54FFA9FFB6FF06AF4D2CF0FEB2" blockId="25.[151,720,255,280]" box="[151,720,255,280]" pageId="25" pageNumber="204">
<treatmentCitationGroup id="90D3287AFFA9FFB6FF06AF4D2CF0FEB2" box="[151,720,255,280]" pageId="25" pageNumber="204">
<treatmentCitation id="31622945FFA9FFB6FF06AF4D2C46FEB2" author="Berggren, M." box="[151,614,255,280]" page="166" pageId="25" pageNumber="204" year="1991">
<taxonomicName id="77C374D7FFA9FFB6FF06AF4D2C46FEB2" ID-CoL="85MDG" authority="Berggren 1991: 166" authorityName="Berggren" authorityPageNumber="166" authorityYear="1991" box="[151,614,255,280]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="204" phylum="Arthropoda" rank="species" species="rubricinctuta">
<emphasis id="82B7D346FFA9FFB6FF06AF4D2FB4FEBD" box="[151,404,256,280]" italics="true" pageId="25" pageNumber="204">Athanopsis rubricinctuta</emphasis>
<bibRefCitation id="D45272A5FFA9FFB6FE0BAF4D2C46FEB2" author="Berggren, M." box="[410,614,255,280]" pageId="25" pageNumber="204" pagination="166 - 178" refId="ref21169" refString="Berggren, M. (1991) Athanopsis rubricinctuta, new species (Decapoda: Natantia: Alpheidae), a shrimp associated with an echiuroid at Inhaca Island, Mocambique. Journal of Crustacean Biology, 11, 166 - 178. https: // doi. org / 10.2307 / 1548555" type="journal article" year="1991">Berggren 1991: 166</bibRefCitation>
</taxonomicName>
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, figs. 16.
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</paragraph>
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<emphasis id="82B7D346FFA9FFB6FF06AF082F5EFEFA" bold="true" box="[151,382,325,351]" pageId="25" pageNumber="204">Material examined</emphasis>
.
<materialsCitation id="00AB0509FFA9FFB6FE03AF0B2DE8FE69" collectingDate="2022-07-11" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="25" pageNumber="204" specimenCode="UF 71437" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFA9FFB6FE03AF0B2FC2FEC5" box="[402,482,326,352]" count="1" pageId="25" pageNumber="204" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFA9FFB6FD89AF0B2C57FEC5" box="[536,631,326,353]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="25" pageNumber="204" unit="mm" value="5.5">5.5 mm</quantity>
),
<collectionCode id="D6D29791FFA9FFB6FD1EAF0B2CD1FEC5" box="[655,753,326,352]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="25" pageNumber="204" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFA9FFB6FD6FAF0B2D57FEC5" box="[766,887,326,352]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="25" pageNumber="204" type="Museum">UF 71437</specimenCode>
,
<collectingCountry id="C8D44FC4FFA9FFB6FC16AF0B2DEEFEC5" box="[903,974,326,352]" name="Oman" pageId="25" pageNumber="204">Oman</collectingCountry>
,
<location id="B51C598FFFA9FFB6FC4FAF0B2AADFEC5" LSID="urn:lsid:plazi:treatment:386ABE42FFA9FFB6FF06AEDA2D41FAD1:B51C598FFFA9FFB6FC4FAF0B2AADFEC5" box="[990,1165,326,352]" country="Oman" latitude="20.587528" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" name="Masirah Island" pageId="25" pageNumber="204">Masirah Island</location>
, channel side,
<collectingMunicipality id="5018952EFFA9FFB6FAD0AF0B2BB8FEC5" box="[1345,1432,326,352]" pageId="25" pageNumber="204">Dafiyat</collectingMunicipality>
,
<geoCoordinate id="D5F76993FFA9FFB6FF06AF272F16FE20" box="[151,310,362,389]" degrees="20" direction="north" minutes="35" orientation="latitude" pageId="25" pageNumber="204" precision="1" seconds="15.1" value="20.587528">20°3515.1”N</geoCoordinate>
/
<geoCoordinate id="D5F76993FFA9FFB6FEC5AF272FD1FE20" box="[340,497,362,389]" degrees="58" direction="east" minutes="49" orientation="longitude" pageId="25" pageNumber="204" precision="1" seconds="36.9" value="58.826916">58°4936.9”E</geoCoordinate>
, intertidal and shallow subtidal sandflat with patches of coral rubble, rocks and seagrass, depth at low tide less than
<quantity id="773BA2B1FFA9FFB6FDD8AFC32CAFFE0D" box="[585,655,398,425]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="204" unit="m" value="0.5">
<elevation id="3BEEE867FFA9FFB6FDD8AFC32CAFFE0D" box="[585,655,398,425]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="204" unit="m" value="0.5">0.5 m</elevation>
</quantity>
, suction pump, in echiuran burrow, leg.
<collectorName id="1D366A82FFA9FFB6FBFDAFC32AF8FE0D" box="[1132,1240,398,424]" pageId="25" pageNumber="204">A. Anker</collectorName>
,
<date id="C47D2994FFA9FFB6FB76AFC32B44FE0D" box="[1255,1380,398,424]" pageId="25" pageNumber="204" value="2022-07-11">
<collectingDate id="D439D07CFFA9FFB6FB76AFC32B44FE0D" box="[1255,1380,398,424]" pageId="25" pageNumber="204" value="2022-07-11">07.11.2022</collectingDate>
</date>
[fcn BOMAN-13029, specimen infested with a pair of hemiarthrine isopods]
</materialsCitation>
;
<materialsCitation id="00AB0509FFA9FFB6FC42AFFF2DB3FE54" collectingDate="2022-07-11" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="25" pageNumber="204" specimenCode="UF 71437" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFA9FFB6FC42AFFF2A3FFE69" box="[979,1055,434,460]" count="1" pageId="25" pageNumber="204" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFA9FFB6FBDDAFFF2A87FE69" box="[1100,1191,434,461]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="25" pageNumber="204" unit="mm" value="5.4">5.4 mm</quantity>
),
<collectionCode id="D6D29791FFA9FFB6FB2BAFFF2B3CFE69" box="[1210,1308,434,460]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="25" pageNumber="204" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFA9FFB6FAB5AFFF2BB8FE69" box="[1316,1432,434,460]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="25" pageNumber="204" type="Museum">UF 71437</specimenCode>
, same collection data as for previous specimen [fcn BOMAN-13015]
</materialsCitation>
;
<materialsCitation id="00AB0509FFA9FFB6FC0CAF9B2C33FDF9" collectingDate="2022-11-19" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="5" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="25" pageNumber="204" specimenCode="UF 71443, UF 71440" specimenCount="1" specimenCount-male="1">
<specimenCount id="A6C5C4DDFFA9FFB6FC0CAF9B2DC8FE55" box="[925,1000,470,496]" count="1" pageId="25" pageNumber="204" type="male">1 male</specimenCount>
(cl
<quantity id="773BA2B1FFA9FFB6FB83AF9B2A4BFE55" box="[1042,1131,470,497]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="25" pageNumber="204" unit="mm" value="5.8">5.8 mm</quantity>
),
<collectionCode id="D6D29791FFA9FFB6FBECAF9B2AFFFE55" box="[1149,1247,470,496]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="25" pageNumber="204" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFA9FFB6FB74AF9B2B78FE55" box="[1253,1368,470,496]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="25" pageNumber="204" type="Museum">UF 71443</specimenCode>
, same collection data as for previous specimen [fcn BOMAN-13016]; 1 ovig. female (cl 5.0 mm),
<collectionCode id="D6D29791FFA9FFB6FB29AFB72B3AFDB1" box="[1208,1306,506,532]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="25" pageNumber="204" type="Museum">FLMNH</collectionCode>
<specimenCode id="E065A72FFFA9FFB6FAB2AFB72BB8FDB1" box="[1315,1432,506,532]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="25" pageNumber="204" type="Museum">UF 71440</specimenCode>
, same collection locality as for previous specimens, in echiuran burrow with numerous
<taxonomicName id="77C374D7FFA9FFB6FBF8AC522AD3FD9D" box="[1129,1267,542,568]" class="Gastropoda" family="Rissoinidae" genus="Rissoina" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="204" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="82B7D346FFA9FFB6FBF8AC522AE9FD9D" box="[1129,1225,543,568]" italics="true" pageId="25" pageNumber="204">Rissoina</emphasis>
sp.
</taxonomicName>
, leg.
<collectorName id="1D366A82FFA9FFB6FABEAC532BB8FD9D" box="[1327,1432,542,568]" pageId="25" pageNumber="204">A. Anker</collectorName>
,
<date id="C47D2994FFA9FFB6FF06AC0F2F34FDF9" box="[151,276,578,604]" pageId="25" pageNumber="204" value="2022-11-19">
<collectingDate id="D439D07CFFA9FFB6FF06AC0F2F34FDF9" box="[151,276,578,604]" pageId="25" pageNumber="204" value="2022-11-19">19.11.2022</collectingDate>
</date>
[fcn BOMAN-14802]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFA9FFB6FF56AC282A31FCD9" pageId="25" pageNumber="204" type="description">
<paragraph id="B07C0F54FFA9FFB6FF56AC282C3AFD01" blockId="25.[151,1437,325,1396]" pageId="25" pageNumber="204">
<emphasis id="82B7D346FFA9FFB6FF56AC282F71FDDA" bold="true" box="[199,337,613,639]" pageId="25" pageNumber="204">Description</emphasis>
. See
<bibRefCitation id="D45272A5FFA9FFB6FE00AC2B2C6BFD25" author="Berggren, M." box="[401,587,614,640]" pageId="25" pageNumber="204" pagination="166 - 178" refId="ref21169" refString="Berggren, M. (1991) Athanopsis rubricinctuta, new species (Decapoda: Natantia: Alpheidae), a shrimp associated with an echiuroid at Inhaca Island, Mocambique. Journal of Crustacean Biology, 11, 166 - 178. https: // doi. org / 10.2307 / 1548555" type="journal article" year="1991">Berggren (1991)</bibRefCitation>
for original description and illustrations; colour photographs of the Omani specimens are provided in
<figureCitation id="28F813D1FFA9FFB6FE52ACC72C36FD01" box="[451,534,650,676]" captionStart="FIGURE 13" captionStartId="26.[151,250,1900,1924]" captionTargetBox="[151,1436,181,1876]" captionTargetId="figure-34@26.[151,1436,181,1876]" captionTargetPageId="26" captionText="FIGURE 13. Athanopsis rubricinctuta Berggren, 1991, male (cl 5.4 mm) from Masirah Island, Oman (FLMNH UF 71437) [a, b]; male (cl 5.8 mm) from the same locality (FLMNH UF 71443) [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, shrimp in life (different individual), lateral (left side). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350817" httpUri="https://zenodo.org/record/10350817/files/figure.png" pageId="25" pageNumber="204">Fig. 13</figureCitation>
.
</paragraph>
<paragraph id="B07C0F54FFA9FFB6FF56ACE02A31FCD9" blockId="25.[151,1437,325,1396]" pageId="25" pageNumber="204">
<emphasis id="82B7D346FFA9FFB6FF56ACE02F4DFD62" bold="true" box="[199,365,685,711]" pageId="25" pageNumber="204">Colour in life</emphasis>
. Background translucent whitish; carapace with bright red patch on rostro-orbital area, one narrow red band fringing each anterolateral margin, one much broader, more irregular transverse band extending from each branchial margin to cardiac region, however, not joining dorsally, and white transverse band on dorsal surface, near posterior margin; pleon with four bright red, moderately narrow, transverse bands, more precisely on first, second, fourth and fifth pleonites, sixth pleonite with white dorsal patch; remaining appendages hyaline whitish (
<figureCitation id="28F813D1FFA9FFB6FF68AD2F2F6BFCD9" box="[249,331,866,892]" captionStart="FIGURE 13" captionStartId="26.[151,250,1900,1924]" captionTargetBox="[151,1436,181,1876]" captionTargetId="figure-34@26.[151,1436,181,1876]" captionTargetPageId="26" captionText="FIGURE 13. Athanopsis rubricinctuta Berggren, 1991, male (cl 5.4 mm) from Masirah Island, Oman (FLMNH UF 71437) [a, b]; male (cl 5.8 mm) from the same locality (FLMNH UF 71443) [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, shrimp in life (different individual), lateral (left side). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350817" httpUri="https://zenodo.org/record/10350817/files/figure.png" pageId="25" pageNumber="204">Fig. 13</figureCitation>
); see also
<bibRefCitation id="D45272A5FFA9FFB6FE51AD2F2C55FCD9" author="Berggren, M." box="[448,629,866,892]" pageId="25" pageNumber="204" pagination="166 - 178" refId="ref21169" refString="Berggren, M. (1991) Athanopsis rubricinctuta, new species (Decapoda: Natantia: Alpheidae), a shrimp associated with an echiuroid at Inhaca Island, Mocambique. Journal of Crustacean Biology, 11, 166 - 178. https: // doi. org / 10.2307 / 1548555" type="journal article" year="1991">Berggren (1991</bibRefCitation>
: fig. 6) and
<bibRefCitation id="D45272A5FFA9FFB6FD69AD2F2DABFCD9" author="Anker, A." box="[760,907,866,892]" pageId="25" pageNumber="204" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker (2012</bibRefCitation>
: fig. 7F, G).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFA9FFB6FF56ADCB2C8DFC05" box="[199,685,901,928]" pageId="25" pageNumber="204" type="materials_examined">
<paragraph id="B07C0F54FFA9FFB6FF56ADCB2C8DFC05" blockId="25.[151,1437,325,1396]" box="[199,685,901,928]" pageId="25" pageNumber="204">
<materialsCitation id="00AB0509FFA9FFB6FF56ADCB2C8AFC05" box="[199,682,901,928]" country="Mozambique" location="Inhaca Island" pageId="25" pageNumber="204" specimenCount="1" typeStatus="holotype">
<emphasis id="82B7D346FFA9FFB6FF56ADCB2F40FC3A" bold="true" box="[199,352,901,928]" pageId="25" pageNumber="204">
<typeStatus id="6F78B1F6FFA9FFB6FF56ADCB2F22FC05" box="[199,258,902,928]" pageId="25" pageNumber="204">Type</typeStatus>
locality
</emphasis>
.
<location id="B51C598FFFA9FFB6FEFFADCB2C26FC05" LSID="urn:lsid:plazi:treatment:386ABE42FFA9FFB6FF06AEDA2D41FAD1:B51C598FFFA9FFB6FEFFADCB2C26FC05" box="[366,518,902,928]" country="Mozambique" name="Inhaca Island" pageId="25" pageNumber="204">Inhaca Island</location>
,
<collectingCountry id="C8D44FC4FFA9FFB6FD83ADCB2C8AFC05" box="[530,682,902,928]" name="Mozambique" pageId="25" pageNumber="204">Mozambique</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFA9FFB6FF56ADE42FB1FC4D" pageId="25" pageNumber="204" type="distribution">
<paragraph id="B07C0F54FFA9FFB6FF56ADE42FB1FC4D" blockId="25.[151,1437,325,1396]" pageId="25" pageNumber="204">
<emphasis id="82B7D346FFA9FFB6FF56ADE42F79FC66" bold="true" box="[199,345,937,963]" pageId="25" pageNumber="204">Distribution</emphasis>
. Currently known only from the
<typeStatus id="6F78B1F6FFA9FFB6FD53ADE72CD2FC61" box="[706,754,938,964]" pageId="25" pageNumber="204">type</typeStatus>
locality in
<collectingCountry id="C8D44FC4FFA9FFB6FCFDADE72A20FC61" box="[876,1024,938,964]" name="Mozambique" pageId="25" pageNumber="204">Mozambique</collectingCountry>
(
<bibRefCitation id="D45272A5FFA9FFB6FB9CADE72A94FC61" author="Berggren, M." box="[1037,1204,938,964]" pageId="25" pageNumber="204" pagination="166 - 178" refId="ref21169" refString="Berggren, M. (1991) Athanopsis rubricinctuta, new species (Decapoda: Natantia: Alpheidae), a shrimp associated with an echiuroid at Inhaca Island, Mocambique. Journal of Crustacean Biology, 11, 166 - 178. https: // doi. org / 10.2307 / 1548555" type="journal article" year="1991">Berggren 1991</bibRefCitation>
) and Masirah Island,
<collectingCountry id="C8D44FC4FFA9FFB6FF06AD832EFBFC4D" box="[151,219,974,1000]" name="Oman" pageId="25" pageNumber="204">Oman</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFA9FFB6FF56ADBF2AB1FBF1" pageId="25" pageNumber="204" type="biology_ecology">
<paragraph id="B07C0F54FFA9FFB6FF56ADBF2AB1FBF1" blockId="25.[151,1437,325,1396]" pageId="25" pageNumber="204">
<emphasis id="82B7D346FFA9FFB6FF56ADBF2F06FBA9" bold="true" box="[199,294,1010,1036]" pageId="25" pageNumber="204">Ecology</emphasis>
. All specimens were extracted from echiuran (probably
<taxonomicName id="77C374D7FFA9FFB6FC1CADBF2A66FBA9" box="[909,1094,1010,1036]" class="Polychaeta" family="Thalassematidae" genus="Ochetostoma" kingdom="Animalia" order="Echiuroidea" pageId="25" pageNumber="204" phylum="Annelida" rank="species" species="undetermined">
<emphasis id="82B7D346FFA9FFB6FC1CADBF2A01FBA9" box="[909,1057,1010,1036]" italics="true" pageId="25" pageNumber="204">Ochetostoma</emphasis>
sp.
</taxonomicName>
) burrows on a shallow sandflat with patches of seagrass and rubble, at a depth of
<quantity id="773BA2B1FFA9FFB6FD5CAA5B2D2EFB95" box="[717,782,1046,1073]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="204" unit="m" value="0.5">0.5 m</quantity>
or less at low tide. The
<typeStatus id="6F78B1F6FFA9FFB6FBB1AA5B2A70FB95" box="[1056,1104,1046,1072]" pageId="25" pageNumber="204">type</typeStatus>
specimens were found in the intertidal, in association with “echiuroids of the family
<taxonomicName id="77C374D7FFA9FFB6FC93AA772D9FFBF1" authorityName="Forbes &amp; Goodsir" authorityYear="1841" box="[770,959,1082,1108]" class="Polychaeta" family="Thalassematidae" kingdom="Animalia" order="Echiuroidea" pageId="25" pageNumber="204" phylum="Annelida" rank="family">Thalassematidae</taxonomicName>
” (
<bibRefCitation id="D45272A5FFA9FFB6FC49AA772AA4FBF1" author="Berggren, M." box="[984,1156,1082,1108]" pageId="25" pageNumber="204" pagination="166 - 178" refId="ref21169" refString="Berggren, M. (1991) Athanopsis rubricinctuta, new species (Decapoda: Natantia: Alpheidae), a shrimp associated with an echiuroid at Inhaca Island, Mocambique. Journal of Crustacean Biology, 11, 166 - 178. https: // doi. org / 10.2307 / 1548555" type="journal article" year="1991">Berggren 1991</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F8D95CDFFFA9FFB6FF56AA132D41FAD1" pageId="25" pageNumber="204" type="discussion">
<paragraph id="B07C0F54FFA9FFB6FF56AA132D41FAD1" blockId="25.[151,1437,325,1396]" pageId="25" pageNumber="204">
<emphasis id="82B7D346FFA9FFB6FF56AA132F14FBDD" bold="true" box="[199,308,1118,1144]" pageId="25" pageNumber="204">Remarks</emphasis>
.
<taxonomicName id="77C374D7FFA9FFB6FEDBAA132C42FBDD" authorityName="Berggren" authorityYear="1991" box="[330,610,1118,1144]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="204" phylum="Arthropoda" rank="species" species="rubricinctua">
<emphasis id="82B7D346FFA9FFB6FEDBAA132C42FBDD" box="[330,610,1118,1144]" italics="true" pageId="25" pageNumber="204">Athanopsis rubricinctua</emphasis>
</taxonomicName>
was hitherto known only from the type series from Inhaca Island,
<collectingCountry id="C8D44FC4FFA9FFB6FF06AACF2F0FFB39" box="[151,303,1154,1180]" name="Mozambique" pageId="25" pageNumber="204">Mozambique</collectingCountry>
. The finding of several specimens of
<taxonomicName id="77C374D7FFA9FFB6FD4CAACF2DB1FB39" authorityName="Berggren" authorityYear="1991" box="[733,913,1154,1180]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="204" phylum="Arthropoda" rank="species" species="rubricinctuta">
<emphasis id="82B7D346FFA9FFB6FD4CAACF2DB1FB39" box="[733,913,1154,1180]" italics="true" pageId="25" pageNumber="204">A. rubricinctuta</emphasis>
</taxonomicName>
in
<collectingCountry id="C8D44FC4FFA9FFB6FC29AACF2DDCFB39" box="[952,1020,1154,1180]" name="Oman" pageId="25" pageNumber="204">Oman</collectingCountry>
considerably extends the previously known range of the species in the western Indian Ocean. The Omani material agrees very well with
<bibRefCitation id="D45272A5FFA9FFB6FAB3AAEB2EC2FB41" author="Berggren, M." pageId="25" pageNumber="204" pagination="166 - 178" refId="ref21169" refString="Berggren, M. (1991) Athanopsis rubricinctuta, new species (Decapoda: Natantia: Alpheidae), a shrimp associated with an echiuroid at Inhaca Island, Mocambique. Journal of Crustacean Biology, 11, 166 - 178. https: // doi. org / 10.2307 / 1548555" type="journal article" year="1991">Berggrens (1991)</bibRefCitation>
type material, including the dorsally partly exposed eyes due to the reduction of orbital teeth, with only minor variation observed, for instance, in the armature of the merus of the third, fourth and fifth pereiopods (with one or two spiniform setae in the Omani specimens
<emphasis id="82B7D346FFA9FFB6FD4DAB5E2CD3FA89" box="[732,755,1299,1324]" italics="true" pageId="25" pageNumber="204">vs</emphasis>
. with one spiniform seta in the
<typeStatus id="6F78B1F6FFA9FFB6FBC4AB5F2A9CFA89" box="[1109,1212,1298,1324]" pageId="25" pageNumber="204" type="holotype">holotype</typeStatus>
). The colour pattern of the Omani specimens (
<figureCitation id="28F813D1FFA9FFB6FE2FAB7B2C2FFAF5" box="[446,527,1334,1360]" captionStart="FIGURE 11" captionStartId="21.[151,250,1771,1795]" captionTargetBox="[166,1429,197,1740]" captionTargetId="figure-19@21.[151,1436,181,1747]" captionTargetPageId="21" captionText="FIGURE 11. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), right cheliped, lateral; b, same, mesial; c, same, carpus and chela, mesial; d, same, chela fingers closed, mesial; e, same, chela fingers open, mesial; f, second pereiopod, lateral; g, third pereiopod, lateral; h, same, distal portion of carpus, propodus and dactylus, mesial; i, distal portion of propodus and dactylus, lateral; j, fourth pereiopod, lateral; k, same, distal portion of carpus, propodus and dactylus, mesial; l, fifth pereiopod, lateral; m, same, distal portion of carpus, propodus and dactylus, mesial; n, same, distal portion of propodus and dactylus, lateral" figureDoi="http://doi.org/10.5281/zenodo.10350809" httpUri="https://zenodo.org/record/10350809/files/figure.png" pageId="25" pageNumber="204">Fig. 11</figureCitation>
) is very similar to that of the Mozambiquan specimens (
<bibRefCitation id="D45272A5FFA9FFB6FB0DAB7B2B6BFAF5" author="Berggren, M." box="[1180,1355,1334,1360]" pageId="25" pageNumber="204" pagination="166 - 178" refId="ref21169" refString="Berggren, M. (1991) Athanopsis rubricinctuta, new species (Decapoda: Natantia: Alpheidae), a shrimp associated with an echiuroid at Inhaca Island, Mocambique. Journal of Crustacean Biology, 11, 166 - 178. https: // doi. org / 10.2307 / 1548555" type="journal article" year="1991">Berggren 1991</bibRefCitation>
: fig. 6;
<bibRefCitation id="D45272A5FFA9FFB6FF06AB172F00FAD1" author="Anker, A." box="[151,288,1370,1396]" pageId="25" pageNumber="204" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
: fig. 7F, G) and is highly diagnostic for the species.
</paragraph>
</subSubSection>
</treatment>
</document>