618 lines
106 KiB
XML
618 lines
106 KiB
XML
<document id="D2B7CB1E846B8143C687C42BB7550BA2" ID-DOI="10.11646/zootaxa.5383.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10350767" ID-ZooBank="7E434B40-6346-4E6D-BC55-38EBAE24BD52" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1702283478140" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2023" docId="386ABE42FFA3FFB7FF06A8172F57F878" docLanguage="en" docName="zootaxa.5383.2.5.pdf" docOrigin="Zootaxa 5383 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.2.5/52448" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athanopsis dawa Anker 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="203" masterDocId="C453C63AFFB0FFAFFF91AE4D2E20FFA5" masterDocTitle="Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)" masterLastPageNumber="215" masterPageNumber="179" pageNumber="198" updateTime="1702658426235" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="0A9181B4FCAECA7B701926677CEF3AE1">Alpheid shrimps of the genera Athanas Leach, 1814, Athanopsis Coutière, 1897 and Pseudathanas Bruce, 1983 of the coasts of the Arabian Peninsula (Malacostraca: Decapoda: Caridea)</mods:title>
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<mods:name id="62261F73B6BA11ED4D30BE7422C06866" type="personal">
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<mods:roleTerm id="D144BA56C6C0B5D82A4672AF034DB11C">Author</mods:roleTerm>
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<mods:namePart id="243C61D01AA07395CD3CFB979A15F4E2">Anker, Arthur</mods:namePart>
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<mods:affiliation id="1F4EE25D3564941D2103B651C41C98D0">Universidade Federal de Pelotas (UFPEL), Departamento de Ecologia, Zoologia e Genética, Instituto de Biologia, Campus Universitário Capão do Leão, RS, 96010 - 610, Brazil & Red Sea Research Center, King Abdullah University of Science and Technology (KAUST), Thuwal, Saudi Arabia</mods:affiliation>
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<mods:typeOfResource id="B22DD162632C08AD434D4958CC31AE03">text</mods:typeOfResource>
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<mods:title id="F50B585ED9A7C6BBDB5988B8111FE273">Zootaxa</mods:title>
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<mods:part id="140A2D51F7DA3DFFB1BB0DC9E22A9D6D">
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<mods:date id="FF51BB723AC49DE2BEEA0FFC0C60D971">2023</mods:date>
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<mods:number id="95559211BB7EC0C9C03EC332F558553B">2023-12-11</mods:number>
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<mods:number id="9CD14F706137D44481B4692ADE6E9561">5383</mods:number>
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<mods:start id="07002E22545C114B802A22649824D512">179</mods:start>
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<mods:url id="7543A6FC96CD63A5C6B0931D1BCDD313">https://www.mapress.com/zt/article/download/zootaxa.5383.2.5/52448</mods:url>
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<mods:classification id="F3D18B64502C0A8771D3293E2A4C258D">journal article</mods:classification>
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<mods:identifier id="24E5181549E65F73B0609EFB069D4824" type="DOI">10.11646/zootaxa.5383.2.5</mods:identifier>
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<mods:identifier id="822522ADCFD04D9C200BCAEE14F7941D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E2FA2166779811567732A8E001A8228B" type="Zenodo-Dep">10350767</mods:identifier>
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</mods:mods>
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<treatment id="386ABE42FFA3FFB7FF06A8172F57F878" LSID="urn:lsid:plazi:treatment:386ABE42FFA3FFB7FF06A8172F57F878" httpUri="http://treatment.plazi.org/id/386ABE42FFA3FFB7FF06A8172F57F878" lastPageId="24" lastPageNumber="203" pageId="19" pageNumber="198">
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<subSubSection id="F8D95CDFFFA3FFBCFF06A8172FEAF9D0" box="[151,458,1626,1653]" pageId="19" pageNumber="198" type="nomenclature">
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<paragraph id="B07C0F54FFA3FFBCFF06A8172FEAF9D0" blockId="19.[151,458,1626,1689]" box="[151,458,1626,1653]" pageId="19" pageNumber="198">
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<heading id="EB34B838FFA3FFBCFF06A8172FEAF9D0" bold="true" box="[151,458,1626,1653]" fontSize="11" level="1" pageId="19" pageNumber="198" reason="1">
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<emphasis id="82B7D346FFA3FFBCFF06A8172FEAF9D0" bold="true" box="[151,458,1626,1653]" pageId="19" pageNumber="198">
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<taxonomicName id="77C374D7FFA3FFBCFF06A8172F46F9D0" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[151,358,1626,1653]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="198" phylum="Arthropoda" rank="species" species="dawa" status="sp. nov.">
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<emphasis id="82B7D346FFA3FFBCFF06A8172F46F9D0" bold="true" box="[151,358,1626,1653]" italics="true" pageId="19" pageNumber="198">Athanopsis dawa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="99846E3DFFA3FFBCFEFCA8162FEAF9D0" box="[365,458,1627,1653]" pageId="19" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="F8D95CDFFFA3FFBCFF06A8322F5FF93C" box="[151,383,1663,1689]" pageId="19" pageNumber="198" type="description">
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<paragraph id="B07C0F54FFA3FFBCFF06A8322F5FF93C" blockId="19.[151,458,1626,1689]" box="[151,383,1663,1689]" pageId="19" pageNumber="198">
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(
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<figureCitation id="28F813D1FFA3FFBCFF0EA8322EDDF93C" box="[159,253,1663,1689]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="19" pageNumber="198">Figs. 10</figureCitation>
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,
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<figureCitation id="28F813D1FFA3FFBCFE98A8322F06F93C" box="[265,294,1663,1689]" captionStart="FIGURE 11" captionStartId="21.[151,250,1771,1795]" captionTargetBox="[166,1429,197,1740]" captionTargetId="figure-19@21.[151,1436,181,1747]" captionTargetPageId="21" captionText="FIGURE 11. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), right cheliped, lateral; b, same, mesial; c, same, carpus and chela, mesial; d, same, chela fingers closed, mesial; e, same, chela fingers open, mesial; f, second pereiopod, lateral; g, third pereiopod, lateral; h, same, distal portion of carpus, propodus and dactylus, mesial; i, distal portion of propodus and dactylus, lateral; j, fourth pereiopod, lateral; k, same, distal portion of carpus, propodus and dactylus, mesial; l, fifth pereiopod, lateral; m, same, distal portion of carpus, propodus and dactylus, mesial; n, same, distal portion of propodus and dactylus, lateral" figureDoi="http://doi.org/10.5281/zenodo.10350809" httpUri="https://zenodo.org/record/10350809/files/figure.png" pageId="19" pageNumber="198">11</figureCitation>
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,
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<figureCitation id="28F813D1FFA3FFBCFEA3A8322F57F93C" box="[306,375,1663,1689]" captionStart="FIGURE 12" captionStartId="22.[151,250,1733,1757]" captionTargetBox="[151,1436,181,1709]" captionTargetId="figure-34@22.[151,1436,181,1709]" captionTargetPageId="22" captionText="FIGURE 12. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424) [a, b] and species’ habitat [c]; a, shrimp in life, lateral (left side); b, same, dorsal; c, type locality, sandflat with some seagrass and coral rubble, near Dafiyat, Masirah Island, Oman. Photographs by the author [a, b] and Gustav Paulay [c]." figureDoi="http://doi.org/10.5281/zenodo.10350813" httpUri="https://zenodo.org/record/10350813/files/figure.png" pageId="19" pageNumber="198">12a, b</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="F8D95CDFFFA3FFBCFF06A88B2F40F8D4" pageId="19" pageNumber="198" type="materials_examined">
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<paragraph id="B07C0F54FFA3FFBCFF06A88B2F40F8D4" blockId="19.[151,1437,1734,2013]" pageId="19" pageNumber="198">
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<emphasis id="82B7D346FFA3FFBCFF06A88B2F5DF945" bold="true" box="[151,381,1734,1760]" pageId="19" pageNumber="198">Material examined</emphasis>
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.
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<materialsCitation id="00AB0509FFA3FFBCFE1EA88A2F7BF8E8" collectingDate="2022-07-11" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="3" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="19" pageNumber="198" specimenCode="UF 71424" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="6F78B1F6FFA3FFBCFE1EA88A2FDCF944" box="[399,508,1735,1761]" pageId="19" pageNumber="198" type="holotype">Holotype</typeStatus>
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: ovig. female (cl
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||
<quantity id="773BA2B1FFA3FFBCFD5DA88A2D09F944" box="[716,809,1735,1761]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="19" pageNumber="198" unit="mm" value="4.4">4.4 mm</quantity>
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),
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<collectionCode id="D6D29791FFA3FFBCFCD1A88A2D82F944" box="[832,930,1735,1761]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="19" pageNumber="198" type="Museum">FLMNH</collectionCode>
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||
<specimenCode id="E065A72FFFA3FFBCFC3CA88A2A04F944" box="[941,1060,1735,1761]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="19" pageNumber="198" type="Museum">UF 71424</specimenCode>
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,
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<collectingCountry id="C8D44FC4FFA3FFBCFBA2A88A2A5AF944" box="[1075,1146,1735,1761]" name="Oman" pageId="19" pageNumber="198">Oman</collectingCountry>
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||
,
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||
<location id="B51C598FFFA3FFBCFB18A88A2B16F944" LSID="urn:lsid:plazi:treatment:386ABE42FFA3FFB7FF06A8172F57F878:B51C598FFFA3FFBCFB18A88A2B16F944" box="[1161,1334,1735,1761]" country="Oman" latitude="20.587528" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" name="Masirah Island" pageId="19" pageNumber="198">Masirah Island</location>
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, channel side,
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||
<collectingMunicipality id="5018952EFFA3FFBCFF45A8A62F0BF8A0" box="[212,299,1771,1797]" pageId="19" pageNumber="198">Dafiyat</collectingMunicipality>
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||
,
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||
<geoCoordinate id="D5F76993FFA3FFBCFEA6A8A72FF5F8A0" box="[311,469,1770,1797]" degrees="20" direction="north" minutes="35" orientation="latitude" pageId="19" pageNumber="198" precision="1" seconds="15.1" value="20.587528">20°35’15.1”N</geoCoordinate>
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||
/
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||
<geoCoordinate id="D5F76993FFA3FFBCFE7CA8A72CABF8A0" box="[493,651,1770,1797]" degrees="58" direction="east" minutes="49" orientation="longitude" pageId="19" pageNumber="198" precision="1" seconds="36.9" value="58.826916">58°49’36.9”E</geoCoordinate>
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, intertidal and shallow subtidal sandflat with patches of coral rubble, rocks and seagrass, depth at low tide
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<quantity id="773BA2B1FFA3FFBCFDADA9432C90F88C" box="[572,688,1806,1833]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="19" pageNumber="198" unit="m" value="0.3" valueMax="0.5" valueMin="0.1">
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<elevation id="3BEEE867FFA3FFBCFDADA9432C90F88C" box="[572,688,1806,1833]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="19" pageNumber="198" unit="m" value="0.3" valueMax="0.5" valueMin="0.1">0.1–0.5 m</elevation>
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</quantity>
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, suction pump, in echiuran burrow, leg.
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<collectorName id="1D366A82FFA3FFBCFBE7A9422AFFF88C" box="[1142,1247,1807,1833]" pageId="19" pageNumber="198">A. Anker</collectorName>
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,
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<date id="C47D2994FFA3FFBCFB7AA9422B47F88C" box="[1259,1383,1807,1833]" pageId="19" pageNumber="198" value="2022-07-11">
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<collectingDate id="D439D07CFFA3FFBCFB7AA9422B47F88C" box="[1259,1383,1807,1833]" pageId="19" pageNumber="198" value="2022-07-11">07.11.2022</collectingDate>
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</date>
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[fcn BOMAN-13017]
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</materialsCitation>
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.
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<materialsCitation id="00AB0509FFA3FFBCFEF4A97E2F7BF8D4" collectingDate="2022-07-11" collectionCode="FLMNH" collectorName="A. Anker" country="Oman" elevation="3" latitude="20.587528" location="Masirah Island" longLatPrecision="1" longitude="58.826916" municipality="Dafiyat" pageId="19" pageNumber="198" specimenCode="UF 71423" specimenCount="1" typeStatus="paratype">
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<typeStatus id="6F78B1F6FFA3FFBCFEF4A97E2FEBF8E8" box="[357,459,1843,1869]" pageId="19" pageNumber="198" type="paratype">Paratype</typeStatus>
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: ovig. female (cl
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<quantity id="773BA2B1FFA3FFBCFD1BA97E2CC3F8E8" box="[650,739,1843,1869]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="19" pageNumber="198" unit="mm" value="4.7">4.7 mm</quantity>
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),
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<collectionCode id="D6D29791FFA3FFBCFD64A97E2D77F8E8" box="[757,855,1843,1869]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="19" pageNumber="198" type="Museum">FLMNH</collectionCode>
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||
<specimenCode id="E065A72FFFA3FFBCFCCCA97E2DF0F8E8" box="[861,976,1843,1869]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="19" pageNumber="198" type="Museum">UF 71423</specimenCode>
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, same collection data as for holotype [fcn BOMAN-13028]
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="F8D95CDFFFA3FFB7FF56A9372F5CFEB8" lastPageId="24" lastPageNumber="203" pageId="19" pageNumber="198" type="description">
|
||
<paragraph id="B07C0F54FFA3FFBBFF56A9372ABBFF70" blockId="19.[151,1437,1734,2013]" lastBlockId="20.[151,1437,151,393]" lastPageId="20" lastPageNumber="199" pageId="19" pageNumber="198">
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<emphasis id="82B7D346FFA3FFBCFF56A9372F71F831" bold="true" box="[199,337,1914,1940]" pageId="19" pageNumber="198">Description</emphasis>
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. Carapace (
|
||
<figureCitation id="28F813D1FFA3FFBCFE40A9362C68F830" box="[465,584,1915,1941]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="19" pageNumber="198">Fig.10a–d</figureCitation>
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) glabrous, somewhat domed, with conspicuous, anteriorly directed postrostral tubercle. Rostrum (
|
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<figureCitation id="28F813D1FFA3FFBCFEFDA9D22FDEF81C" box="[364,510,1951,1977]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="19" pageNumber="198">Fig. 10b, c, d</figureCitation>
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) moderately long, overreaching distal margin of first article of antennular peduncle, subtriangular, as long as broad at base, with acute tip; rostral carina distinct, continued by postrostral carina and fading just anterior to postrostral tubercle. Extra-corneal teeth (
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<figureCitation id="28F813D1FFA4FFBBFC8AAEDA2DB9FF14" box="[795,921,151,177]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="20" pageNumber="199">Fig. 10b–d</figureCitation>
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) triangular, subacute distally. Pterygostomial angle (
|
||
<figureCitation id="28F813D1FFA4FFBBFF72AEF62F7BFF70" box="[227,347,187,213]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="20" pageNumber="199">Fig. 10a, c</figureCitation>
|
||
) bluntly or subacutely produced anteriorly. Cardiac notch (
|
||
<figureCitation id="28F813D1FFA4FFBBFC63AEF62A6FFF70" box="[1010,1103,187,213]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="20" pageNumber="199">Fig. 10a</figureCitation>
|
||
) deep.
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFA4FFBBFF56AE922D19FE2C" blockId="20.[151,1437,151,393]" pageId="20" pageNumber="199">
|
||
Pleon (
|
||
<figureCitation id="28F813D1FFA4FFBBFE85AE922F50FF5C" box="[276,368,223,249]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="20" pageNumber="199">Fig. 10e</figureCitation>
|
||
) glabrous; first to fifth pleura rounded distoventrally; sixth pleonite with subtriangular articulated plate. Telson (
|
||
<figureCitation id="28F813D1FFA4FFBBFEA2AF4E2FADFEB8" box="[307,397,259,285]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="20" pageNumber="199">Fig. 10f</figureCitation>
|
||
) moderately broad, gently tapering distally, about 2.1 times as long as proximal width; dorsal surface with two pairs of small spiniform setae situated at some distance from lateral margins, approximately at 0.50–0.55 (anterior) and 0.70–0.75 (posterior) of telson length; posterior margin slightly rounded, with two pairs of spiniform setae, mesial almost three times as long as lateral.
|
||
</paragraph>
|
||
<caption id="E4BC5FDCFFA4FFBBFF06A9502B23F804" ID-DOI="http://doi.org/10.5281/zenodo.10350807" ID-Zenodo-Dep="10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="20" pageNumber="199" startId="20.[151,250,1821,1845]" targetBox="[158,1429,431,1790]" targetPageId="20" targetType="figure">
|
||
<paragraph id="B07C0F54FFA4FFBBFF06A9502B23F804" blockId="20.[151,1437,1821,1953]" pageId="20" pageNumber="199">
|
||
<emphasis id="82B7D346FFA4FFBBFF06A9502F02F890" bold="true" box="[151,290,1821,1845]" pageId="20" pageNumber="199">FIGURE 10.</emphasis>
|
||
<taxonomicName id="77C374D7FFA4FFBBFEBBA9502FFAF890" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[298,474,1821,1845]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="199" phylum="Arthropoda" rank="species" species="dawa" status="sp. nov.">
|
||
<emphasis id="82B7D346FFA4FFBBFEBBA9502FFAF890" box="[298,474,1821,1845]" italics="true" pageId="20" pageNumber="199">Athanopsis dawa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFA4FFBBFE73A9502C14F890" bold="true" box="[482,564,1821,1845]" pageId="20" pageNumber="199">
|
||
<taxonomicNameLabel id="99846E3DFFA4FFBBFE73A9502C14F890" box="[482,564,1821,1845]" pageId="20" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E4BC5FDCFFA5FFBAFF06A8A62C12F812" ID-DOI="http://doi.org/10.5281/zenodo.10350809" ID-Zenodo-Dep="10350809" httpUri="https://zenodo.org/record/10350809/files/figure.png" pageId="21" pageNumber="200" startId="21.[151,250,1771,1795]" targetBox="[166,1429,197,1740]" targetPageId="21" targetType="figure">
|
||
<paragraph id="B07C0F54FFA5FFBAFF06A8A62C12F812" blockId="21.[151,1436,1771,1975]" pageId="21" pageNumber="200">
|
||
<emphasis id="82B7D346FFA5FFBAFF06A8A62F3FF8A6" bold="true" box="[151,287,1771,1795]" pageId="21" pageNumber="200">FIGURE 11.</emphasis>
|
||
<taxonomicName id="77C374D7FFA5FFBAFEB4A8A62FF4F8A6" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[293,468,1771,1795]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="200" phylum="Arthropoda" rank="species" species="dawa" status="sp. nov.">
|
||
<emphasis id="82B7D346FFA5FFBAFEB4A8A62FF4F8A6" box="[293,468,1771,1795]" italics="true" pageId="21" pageNumber="200">Athanopsis dawa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFA5FFBAFE4BA8A62C0BF8A6" bold="true" box="[474,555,1771,1795]" pageId="21" pageNumber="200">
|
||
<taxonomicNameLabel id="99846E3DFFA5FFBAFE4BA8A62C0BF8A6" box="[474,555,1771,1795]" pageId="21" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), right cheliped, lateral; b, same, mesial; c, same, carpus and chela, mesial; d, same, chela fingers closed, mesial; e, same, chela fingers open, mesial; f, second pereiopod, lateral; g, third pereiopod, lateral; h, same, distal portion of carpus, propodus and dactylus, mesial; i, distal portion of propodus and dactylus, lateral; j, fourth pereiopod, lateral; k, same, distal portion of carpus, propodus and dactylus, mesial; l, fifth pereiopod, lateral; m, same, distal portion of carpus, propodus and dactylus, mesial; n, same, distal portion of propodus and dactylus, lateral
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E4BC5FDCFFA6FFB9FF06A8882AB1F880" ID-DOI="http://doi.org/10.5281/zenodo.10350813" ID-Zenodo-Dep="10350813" httpUri="https://zenodo.org/record/10350813/files/figure.png" pageId="22" pageNumber="201" startId="22.[151,250,1733,1757]" targetBox="[151,1436,181,1709]" targetPageId="22" targetType="figure">
|
||
<paragraph id="B07C0F54FFA6FFB9FF06A8882AB1F880" blockId="22.[151,1437,1733,1829]" pageId="22" pageNumber="201">
|
||
<emphasis id="82B7D346FFA6FFB9FF06A8882F00F978" bold="true" box="[151,288,1733,1757]" pageId="22" pageNumber="201">FIGURE 12.</emphasis>
|
||
<taxonomicName id="77C374D7FFA6FFB9FEB6A8882FF5F978" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[295,469,1733,1757]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="201" phylum="Arthropoda" rank="species" species="dawa" status="sp. nov.">
|
||
<emphasis id="82B7D346FFA6FFB9FEB6A8882FF5F978" box="[295,469,1733,1757]" italics="true" pageId="22" pageNumber="201">Athanopsis dawa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFA6FFB9FE4AA8882C0CF978" bold="true" box="[475,556,1733,1757]" pageId="22" pageNumber="201">
|
||
<taxonomicNameLabel id="99846E3DFFA6FFB9FE4AA8882C0CF978" box="[475,556,1733,1757]" pageId="22" pageNumber="201" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424) [a, b] and species’ habitat [c]; a, shrimp in life, lateral (left side); b, same, dorsal; c, type locality, sandflat with some seagrass and coral rubble, near Dafiyat, Masirah Island, Oman. Photographs by the author [a, b] and Gustav Paulay [c].
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B07C0F54FFA6FFB9FF56A9352CC2F810" blockId="22.[151,1436,1911,2010]" pageId="22" pageNumber="201">
|
||
Eyes (
|
||
<figureCitation id="28F813D1FFA6FFB9FE9FA93A2F85F834" box="[270,421,1911,1937]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="22" pageNumber="201">Fig. 10a, b, d</figureCitation>
|
||
) largely exposed dorsally and laterally, with well pigmented but slightly reduced corneas; anteromesial margin of eyestalks rounded, unarmed.
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFA6FFB8FF56A9F22A6BFF5C" blockId="22.[151,1436,1911,2010]" lastBlockId="23.[151,1437,151,2013]" lastPageId="23" lastPageNumber="202" pageId="22" pageNumber="201">
|
||
Antennule (
|
||
<figureCitation id="28F813D1FFA6FFB9FEDFA9F22FCCF87C" box="[334,492,1983,2010]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="22" pageNumber="201">Fig. 10b, c, g</figureCitation>
|
||
) stout; stylocerite with blunt tip, reaching mid-length of third article of peduncle; ventromesial carina with large, anteriorly directed, sharp tooth; second article about 0.8 times as long as wide; lateral flagellum with fused portion composed of four units, accessory ramus well developed, subdivided into at least six poorly individualised units, each carrying group of two or three aesthetascs.
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFA7FFB8FF56AF4E2C94FE2C" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="202">
|
||
Antenna (
|
||
<figureCitation id="28F813D1FFA7FFB8FEA7AF4E2F92FEB8" box="[310,434,259,285]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="23" pageNumber="202">Fig. 10b, c</figureCitation>
|
||
) with basicerite stout, its distolateral margin armed with large, sharp tooth; scaphocerite broad, short, reaching slightly beyond end of antennular peduncle; lateral margin straight; anterior margin of blade rounded, greatly exceeded by stout distolateral tooth; carpocerite very stout, slightly overreaching both antennular peduncle and scaphocerite; flagellum thickened.
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFA7FFB8FF56AFDE2C9BFDC4" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="202">
|
||
Mouthparts not dissected, typical for genus in external observation. Third maxilliped (
|
||
<figureCitation id="28F813D1FFA7FFB8FB36AFDE2B00FE08" box="[1191,1312,403,429]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="23" pageNumber="202">Fig. 10h, i</figureCitation>
|
||
) relatively stout; coxa with somewhat produced, distally acute lateral plate above mastigobranch; antepenultimate article noticeably broadened, with strongly convex dorsal margin, about 3.2 times as long as maximal width; penultimate article subrectangular, about 1.7 times as long as wide; ultimate article much slenderer and almost three times as long as penultimate article, tapering distally, with tip armed with one long and two short spiniform setae; exopod well developed, not reaching penultimate article.
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFA7FFB8FF56AC262B1DFC6C" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="202">
|
||
First pereiopods (= chelipeds) about equal in size, symmetrical in shape, carried folded ventrally when not in use (
|
||
<figureCitation id="28F813D1FFA7FFB8FF5DACC22F70FD0C" box="[204,336,655,681]" captionStart="FIGURE 11" captionStartId="21.[151,250,1771,1795]" captionTargetBox="[166,1429,197,1740]" captionTargetId="figure-19@21.[151,1436,181,1747]" captionTargetPageId="21" captionText="FIGURE 11. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), right cheliped, lateral; b, same, mesial; c, same, carpus and chela, mesial; d, same, chela fingers closed, mesial; e, same, chela fingers open, mesial; f, second pereiopod, lateral; g, third pereiopod, lateral; h, same, distal portion of carpus, propodus and dactylus, mesial; i, distal portion of propodus and dactylus, lateral; j, fourth pereiopod, lateral; k, same, distal portion of carpus, propodus and dactylus, mesial; l, fifth pereiopod, lateral; m, same, distal portion of carpus, propodus and dactylus, mesial; n, same, distal portion of propodus and dactylus, lateral" figureDoi="http://doi.org/10.5281/zenodo.10350809" httpUri="https://zenodo.org/record/10350809/files/figure.png" pageId="23" pageNumber="202">Figs. 11a–e</figureCitation>
|
||
,
|
||
<figureCitation id="28F813D1FFA7FFB8FECAACC22FA7FD0C" box="[347,391,655,681]" captionStart="FIGURE 12" captionStartId="22.[151,250,1733,1757]" captionTargetBox="[151,1436,181,1709]" captionTargetId="figure-34@22.[151,1436,181,1709]" captionTargetPageId="22" captionText="FIGURE 12. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424) [a, b] and species’ habitat [c]; a, shrimp in life, lateral (left side); b, same, dorsal; c, type locality, sandflat with some seagrass and coral rubble, near Dafiyat, Masirah Island, Oman. Photographs by the author [a, b] and Gustav Paulay [c]." figureDoi="http://doi.org/10.5281/zenodo.10350813" httpUri="https://zenodo.org/record/10350813/files/figure.png" pageId="23" pageNumber="202">12a</figureCitation>
|
||
); basis with rudimentary exopod; ischium stout, widening distally, with three spiniform setae on dorsal margin, ventral surface unarmed; merus not particularly swollen, about four times as long as wide, with deeply excavated ventral surface, slightly convex dorsal margin and somewhat concave ventrolateral margin, all surfaces smooth; carpus very short, cup-shaped, smooth; chela slightly longer than merus; palm feebly swollen, depressed ventrolaterally, about three times as long as high (wide) in mesial view of chela, smooth; fingers 0.7 times as long as palm, strongly curved laterally, noticeably deviating from axis of palm, slender, fitting tightly when closed except for small gape proximally; cutting edge of pollex with with broad hiatus proximally followed by low triangular tooth at about proximal third of pollex, this tooth being continued by straight edge to fingertip; cutting edge of dactylus with three subtriangular, blunt teeth opposed to hiatus on pollex, remaining edge smooth.
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFA7FFB8FF56AD9E2A0AF93C" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="202">
|
||
Second pereiopod (
|
||
<figureCitation id="28F813D1FFA7FFB8FE33AD9E2FDDFC48" box="[418,509,979,1005]" captionStart="FIGURE 11" captionStartId="21.[151,250,1771,1795]" captionTargetBox="[166,1429,197,1740]" captionTargetId="figure-19@21.[151,1436,181,1747]" captionTargetPageId="21" captionText="FIGURE 11. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), right cheliped, lateral; b, same, mesial; c, same, carpus and chela, mesial; d, same, chela fingers closed, mesial; e, same, chela fingers open, mesial; f, second pereiopod, lateral; g, third pereiopod, lateral; h, same, distal portion of carpus, propodus and dactylus, mesial; i, distal portion of propodus and dactylus, lateral; j, fourth pereiopod, lateral; k, same, distal portion of carpus, propodus and dactylus, mesial; l, fifth pereiopod, lateral; m, same, distal portion of carpus, propodus and dactylus, mesial; n, same, distal portion of propodus and dactylus, lateral" figureDoi="http://doi.org/10.5281/zenodo.10350809" httpUri="https://zenodo.org/record/10350809/files/figure.png" pageId="23" pageNumber="202">Fig. 11f</figureCitation>
|
||
) relatively short and rather stout; ischium about 2.6 times as long as distal width, distally widening, somewhat depressed on ventromesial surface; merus about 1.6 times as long as ischium, stout, 3.5 times as long as wide; carpus with five subarticles, proximal as long as sum of four others; chela longer than distal-most carpal subarticle, simple, with fingers equal to palm. Third to fifth pereiopods relatively stout, similar in length. Third pereiopod (
|
||
<figureCitation id="28F813D1FFA7FFB8FE40AA2E2C66FBD8" box="[465,582,1123,1149]" captionStart="FIGURE 11" captionStartId="21.[151,250,1771,1795]" captionTargetBox="[166,1429,197,1740]" captionTargetId="figure-19@21.[151,1436,181,1747]" captionTargetPageId="21" captionText="FIGURE 11. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), right cheliped, lateral; b, same, mesial; c, same, carpus and chela, mesial; d, same, chela fingers closed, mesial; e, same, chela fingers open, mesial; f, second pereiopod, lateral; g, third pereiopod, lateral; h, same, distal portion of carpus, propodus and dactylus, mesial; i, distal portion of propodus and dactylus, lateral; j, fourth pereiopod, lateral; k, same, distal portion of carpus, propodus and dactylus, mesial; l, fifth pereiopod, lateral; m, same, distal portion of carpus, propodus and dactylus, mesial; n, same, distal portion of propodus and dactylus, lateral" figureDoi="http://doi.org/10.5281/zenodo.10350809" httpUri="https://zenodo.org/record/10350809/files/figure.png" pageId="23" pageNumber="202">
|
||
Fig.
|
||
<quantity id="773BA2B1FFA7FFB8FD97AA2E2C0EFBD8" box="[518,558,1123,1149]" metricMagnitude="-2" metricUnit="kg" metricValue="1.1" pageId="23" pageNumber="202" unit="g" value="11.0">11g</quantity>
|
||
–i
|
||
</figureCitation>
|
||
) with ischium armed with two spiniform setae on ventrolateral surface and one spiniform seta on distodorsal margin; merus about 1.6 times as long as ischium, 3.5 times as long as maximal width, not particularly swollen, ventrolateral margin distally with small subacute tooth; carpus noticeably more slender than merus, about half-length of merus, with two spiniform setae on distoventral margin; propodus about 1.8 times as long as carpus, slenderer than carpus, with seven spiniform setae on ventral or ventromesial margin, and one pair of spiniform setae distally, flanking base of dactylus; dactylus simple, slender, strongly curved, about 0.4 length of propodus. Fourth pereiopod (
|
||
<figureCitation id="28F813D1FFA7FFB8FDFCAB762CC0FAF0" box="[621,736,1339,1365]" captionStart="FIGURE 11" captionStartId="21.[151,250,1771,1795]" captionTargetBox="[166,1429,197,1740]" captionTargetId="figure-19@21.[151,1436,181,1747]" captionTargetPageId="21" captionText="FIGURE 11. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), right cheliped, lateral; b, same, mesial; c, same, carpus and chela, mesial; d, same, chela fingers closed, mesial; e, same, chela fingers open, mesial; f, second pereiopod, lateral; g, third pereiopod, lateral; h, same, distal portion of carpus, propodus and dactylus, mesial; i, distal portion of propodus and dactylus, lateral; j, fourth pereiopod, lateral; k, same, distal portion of carpus, propodus and dactylus, mesial; l, fifth pereiopod, lateral; m, same, distal portion of carpus, propodus and dactylus, mesial; n, same, distal portion of propodus and dactylus, lateral" figureDoi="http://doi.org/10.5281/zenodo.10350809" httpUri="https://zenodo.org/record/10350809/files/figure.png" pageId="23" pageNumber="202">Fig. 11j, k</figureCitation>
|
||
) generally similar to third pereiopod, slenderer; ischium armed with two spiniform setae on ventrolateral surface and one spiniform seta on distodorsal margin; merus 3.3 times as long as maximal width, ventrolateral margin distally with small subacute tooth; carpus about half-length of merus, with two spiniform setae on distoventral margin; propodus with seven spiniform setae on ventral or ventromesial margin, and one pair of spiniform setae distally, flanking base of dactylus; dactylus similar to that of third pereiopod. Fifth pereiopod (
|
||
<figureCitation id="28F813D1FFA7FFB8FEC7ABA22FEBF9AC" box="[342,459,1519,1545]" captionStart="FIGURE 11" captionStartId="21.[151,250,1771,1795]" captionTargetBox="[166,1429,197,1740]" captionTargetId="figure-19@21.[151,1436,181,1747]" captionTargetPageId="21" captionText="FIGURE 11. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), right cheliped, lateral; b, same, mesial; c, same, carpus and chela, mesial; d, same, chela fingers closed, mesial; e, same, chela fingers open, mesial; f, second pereiopod, lateral; g, third pereiopod, lateral; h, same, distal portion of carpus, propodus and dactylus, mesial; i, distal portion of propodus and dactylus, lateral; j, fourth pereiopod, lateral; k, same, distal portion of carpus, propodus and dactylus, mesial; l, fifth pereiopod, lateral; m, same, distal portion of carpus, propodus and dactylus, mesial; n, same, distal portion of propodus and dactylus, lateral" figureDoi="http://doi.org/10.5281/zenodo.10350809" httpUri="https://zenodo.org/record/10350809/files/figure.png" pageId="23" pageNumber="202">Fig. 11l–n</figureCitation>
|
||
) shortest and slenderest of walking legs; ischium unarmed; merus about twice as long as ischium, about 3.2 times as long as wide, ventrolateral margin distally with small subacute tooth; carpus 0.6 length of merus, with two spiniform setae on distoventral margin; propodus with six spiniform setae on ventral or ventromesial margin, one pair of spiniform setae distally, flanking base of dactylus, and three rows of microserrulate setae on distolateral surface; dactylus similar to that of third or fourth pereiopods.
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFA7FFB8FF56A8EE2ED1F88C" blockId="23.[151,1437,151,2013]" pageId="23" pageNumber="202">
|
||
Uropod (
|
||
<figureCitation id="28F813D1FFA7FFB8FEBDA8EE2FA6F918" box="[300,390,1699,1725]" captionStart="FIGURE 10" captionStartId="20.[151,250,1821,1845]" captionTargetBox="[158,1429,431,1790]" captionTargetId="figure-138@20.[151,1436,417,1798]" captionTargetPageId="20" captionText="FIGURE 10. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), a, carapace, lateral; b, frontal region, dorsal; c, same, lateral; d, rostrum, orbital teeth, postrostral tubercle and eye, lateral; e, posterior portion of fifth pleonite and sixth pleonite, lateral; f, telson, dorsal; g, tooth of ventromesial carina of first article of antennular peduncle, lateral; h, third maxilliped, lateral; i, same, tip of ultimate article, lateral; j, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10350807" httpUri="https://zenodo.org/record/10350807/files/figure.png" pageId="23" pageNumber="202">Fig. 10j</figureCitation>
|
||
) with lateral lobe of protopod enlarged, produced distally, with larger, triangular lateral tooth and smaller, blunt mesial tooth; exopod with stout distolateral tooth and long adjacent spiniform seta, latter almost reaching level of distal margin; diaeresis straight for most part, with low lateral lobe; endopod ovoid, as long as exopod.
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFA7FFB8FF56A97E2CA9F8E8" blockId="23.[151,1437,151,2013]" box="[199,649,1843,1869]" pageId="23" pageNumber="202">Gill-exopod formula as given for genus.</paragraph>
|
||
<paragraph id="B07C0F54FFA7FFB7FF56A91B2F5CFEB8" blockId="23.[151,1437,151,2013]" lastBlockId="24.[151,1437,151,2013]" lastPageId="24" lastPageNumber="203" pageId="23" pageNumber="202">
|
||
<emphasis id="82B7D346FFA7FFB8FF56A91B2F4AF8D5" bold="true" box="[199,362,1878,1904]" pageId="23" pageNumber="202">Colour in life</emphasis>
|
||
. Background translucent whitish, with pale yellow tinge in females; carapace with bright red patch on rostro-orbital area, one narrow red band fringing each anterolateral margin, one much broader, more irregular transverse band extending from each branchial margin to cardiac region, not joining dorsally, and one very broad, white, transverse band covering almost entire area adjacent to posterior margin from dorsal mid-line to just above posterior-most portion of branchiostegial margin; pleon with four bright red, moderately narrow, transverse bands, more precisely on first, second, fourth and fifth pleonites, third pleonite with conspicuous, white, transverse band, sixth pleonite with broader, ring-like, white band; remaining appendages hyaline whitish; eggs / ovaries yolky yellow (
|
||
<figureCitation id="28F813D1FFA8FFB7FF62AF4E2F4FFEB8" box="[243,367,259,285]" captionStart="FIGURE 12" captionStartId="22.[151,250,1733,1757]" captionTargetBox="[151,1436,181,1709]" captionTargetId="figure-34@22.[151,1436,181,1709]" captionTargetPageId="22" captionText="FIGURE 12. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424) [a, b] and species’ habitat [c]; a, shrimp in life, lateral (left side); b, same, dorsal; c, type locality, sandflat with some seagrass and coral rubble, near Dafiyat, Masirah Island, Oman. Photographs by the author [a, b] and Gustav Paulay [c]." figureDoi="http://doi.org/10.5281/zenodo.10350813" httpUri="https://zenodo.org/record/10350813/files/figure.png" pageId="24" pageNumber="203">Fig. 12a, b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8D95CDFFFA8FFB7FF56AF6A2C6EFE2C" pageId="24" pageNumber="203" type="etymology">
|
||
<paragraph id="B07C0F54FFA8FFB7FF56AF6A2C6EFE2C" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="203">
|
||
<emphasis id="82B7D346FFA8FFB7FF56AF6A2F68FEE4" bold="true" box="[199,328,295,321]" pageId="24" pageNumber="203">Etymology</emphasis>
|
||
. The new species name refers to dhow (Arabic داو, latinised:
|
||
<taxonomicName id="77C374D7FFA8FFB7FC6AAF6A2A18FEE4" authorityName="Anker" authorityYear="2023" box="[1019,1080,295,321]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="dawa">
|
||
<emphasis id="82B7D346FFA8FFB7FC6AAF6A2A18FEE4" box="[1019,1080,295,321]" italics="true" pageId="24" pageNumber="203">dāwa</emphasis>
|
||
</taxonomicName>
|
||
), a generic name for traditional sailing vessels commonly seen in the western Indian Ocean from the Red Sea to the Arabian-Persian Gulf and
|
||
<collectingCountry id="C8D44FC4FFA8FFB7FF06AF222EDEFE2C" box="[151,254,367,393]" name="Tanzania" pageId="24" pageNumber="203">Zanzibar</collectingCountry>
|
||
; used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8D95CDFFFA8FFB7FF56AFDE2C4FFE08" box="[199,623,402,429]" pageId="24" pageNumber="203" type="materials_examined">
|
||
<paragraph id="B07C0F54FFA8FFB7FF56AFDE2C4FFE08" blockId="24.[151,1437,151,2013]" box="[199,623,402,429]" pageId="24" pageNumber="203">
|
||
<materialsCitation id="00AB0509FFA8FFB7FF56AFDE2C4BFE08" box="[199,619,402,429]" country="Oman" location="Masirah Island" pageId="24" pageNumber="203" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="82B7D346FFA8FFB7FF56AFDE2F40FE09" bold="true" box="[199,352,402,429]" pageId="24" pageNumber="203">
|
||
<typeStatus id="6F78B1F6FFA8FFB7FF56AFDE2F22FE08" box="[199,258,403,429]" pageId="24" pageNumber="203">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
.
|
||
<location id="B51C598FFFA8FFB7FEFFAFDE2C38FE08" LSID="urn:lsid:plazi:treatment:386ABE42FFA3FFB7FF06A8172F57F878:B51C598FFFA8FFB7FEFFAFDE2C38FE08" box="[366,536,403,429]" country="Oman" name="Masirah Island" pageId="24" pageNumber="203">Masirah Island</location>
|
||
,
|
||
<collectingCountry id="C8D44FC4FFA8FFB7FDB2AFDE2C4BFE08" box="[547,619,403,429]" name="Oman" pageId="24" pageNumber="203">Oman</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8D95CDFFFA8FFB7FF56AFFB2D9AFE74" box="[199,954,438,465]" pageId="24" pageNumber="203" type="distribution">
|
||
<paragraph id="B07C0F54FFA8FFB7FF56AFFB2D9AFE74" blockId="24.[151,1437,151,2013]" box="[199,954,438,465]" pageId="24" pageNumber="203">
|
||
<emphasis id="82B7D346FFA8FFB7FF56AFFB2F79FE75" bold="true" box="[199,345,438,464]" pageId="24" pageNumber="203">Distribution</emphasis>
|
||
. Currently known only from Masirah Island in
|
||
<collectingCountry id="C8D44FC4FFA8FFB7FCFFAFFA2D96FE74" box="[878,950,439,465]" name="Oman" pageId="24" pageNumber="203">Oman</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8D95CDFFFA8FFB7FF56AF962A6FFDBC" pageId="24" pageNumber="203" type="biology_ecology">
|
||
<paragraph id="B07C0F54FFA8FFB7FF56AF962A6FFDBC" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="203">
|
||
<emphasis id="82B7D346FFA8FFB7FF56AF962F06FE50" bold="true" box="[199,294,475,501]" pageId="24" pageNumber="203">Ecology</emphasis>
|
||
. Both
|
||
<typeStatus id="6F78B1F6FFA8FFB7FEE4AF962F85FE50" box="[373,421,475,501]" pageId="24" pageNumber="203">type</typeStatus>
|
||
specimens were extracted from presumably echiuran burrows on a shallow sandflat with patches of seagrass and coral rubble (
|
||
<figureCitation id="28F813D1FFA8FFB7FDABAFB22CB9FDBC" box="[570,665,511,537]" captionStart="FIGURE 12" captionStartId="22.[151,250,1733,1757]" captionTargetBox="[151,1436,181,1709]" captionTargetId="figure-34@22.[151,1436,181,1709]" captionTargetPageId="22" captionText="FIGURE 12. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424) [a, b] and species’ habitat [c]; a, shrimp in life, lateral (left side); b, same, dorsal; c, type locality, sandflat with some seagrass and coral rubble, near Dafiyat, Masirah Island, Oman. Photographs by the author [a, b] and Gustav Paulay [c]." figureDoi="http://doi.org/10.5281/zenodo.10350813" httpUri="https://zenodo.org/record/10350813/files/figure.png" pageId="24" pageNumber="203">Fig. 12c</figureCitation>
|
||
), at a depth less than
|
||
<quantity id="773BA2B1FFA8FFB7FC1BAFB32DEAFDBC" box="[906,970,510,537]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="203" unit="m" value="0.5">0.5 m</quantity>
|
||
at low tide.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8D95CDFFFA8FFB7FF56AC6E2F57F878" pageId="24" pageNumber="203" type="discussion">
|
||
<paragraph id="B07C0F54FFA8FFB7FF56AC6E2AFEFCB0" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="203">
|
||
<emphasis id="82B7D346FFA8FFB7FF56AC6E2F14FD98" bold="true" box="[199,308,547,573]" pageId="24" pageNumber="203">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="77C374D7FFA8FFB7FED2AC6E2C25FD98" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[323,517,547,573]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="dawa" status="sp. nov.">
|
||
<emphasis id="82B7D346FFA8FFB7FED2AC6E2C25FD98" box="[323,517,547,573]" italics="true" pageId="24" pageNumber="203">Athanopsis dawa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFA8FFB7FD9CAC6E2C49FD98" bold="true" box="[525,617,547,573]" pageId="24" pageNumber="203">
|
||
<taxonomicNameLabel id="99846E3DFFA8FFB7FD9CAC6E2C49FD98" box="[525,617,547,573]" pageId="24" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be easily distinguished from all other species of
|
||
<taxonomicName id="77C374D7FFA8FFB7FB58AC6E2B65FD98" authorityName="Coutiere" authorityYear="1897" box="[1225,1349,547,573]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFA8FFB7FB58AC6E2B65FD98" box="[1225,1349,547,573]" italics="true" pageId="24" pageNumber="203">Athanopsis</emphasis>
|
||
</taxonomicName>
|
||
by two features, both being novel for this genus. The rather conspicuous postrostral tubercle on the carapace is a unique and highly diagnostic feature of the new species, as are the almost equal and subsymmetrical chelipeds (
|
||
<figureCitation id="28F813D1FFA8FFB7FABFAC262E87FD0C" captionStart="FIGURE 11" captionStartId="21.[151,250,1771,1795]" captionTargetBox="[166,1429,197,1740]" captionTargetId="figure-19@21.[151,1436,181,1747]" captionTargetPageId="21" captionText="FIGURE 11. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424), right cheliped, lateral; b, same, mesial; c, same, carpus and chela, mesial; d, same, chela fingers closed, mesial; e, same, chela fingers open, mesial; f, second pereiopod, lateral; g, third pereiopod, lateral; h, same, distal portion of carpus, propodus and dactylus, mesial; i, distal portion of propodus and dactylus, lateral; j, fourth pereiopod, lateral; k, same, distal portion of carpus, propodus and dactylus, mesial; l, fifth pereiopod, lateral; m, same, distal portion of carpus, propodus and dactylus, mesial; n, same, distal portion of propodus and dactylus, lateral" figureDoi="http://doi.org/10.5281/zenodo.10350809" httpUri="https://zenodo.org/record/10350809/files/figure.png" pageId="24" pageNumber="203">Figs. 11a, b</figureCitation>
|
||
,
|
||
<figureCitation id="28F813D1FFA8FFB7FF25ACC22EC0FD0C" box="[180,224,655,681]" captionStart="FIGURE 12" captionStartId="22.[151,250,1733,1757]" captionTargetBox="[151,1436,181,1709]" captionTargetId="figure-34@22.[151,1436,181,1709]" captionTargetPageId="22" captionText="FIGURE 12. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424) [a, b] and species’ habitat [c]; a, shrimp in life, lateral (left side); b, same, dorsal; c, type locality, sandflat with some seagrass and coral rubble, near Dafiyat, Masirah Island, Oman. Photographs by the author [a, b] and Gustav Paulay [c]." figureDoi="http://doi.org/10.5281/zenodo.10350813" httpUri="https://zenodo.org/record/10350813/files/figure.png" pageId="24" pageNumber="203">12a</figureCitation>
|
||
). All other species of
|
||
<taxonomicName id="77C374D7FFA8FFB7FE44ACC22C71FD0C" authorityName="Coutiere" authorityYear="1897" box="[469,593,655,681]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFA8FFB7FE44ACC22C71FD0C" box="[469,593,655,681]" italics="true" pageId="24" pageNumber="203">Athanopsis</emphasis>
|
||
</taxonomicName>
|
||
lack such a tubercle and have very unequal and asymmetrical chelipeds. It must be noted that both
|
||
<typeStatus id="6F78B1F6FFA8FFB7FE33ACFE2FF2FD68" box="[418,466,691,717]" pageId="24" pageNumber="203">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="77C374D7FFA8FFB7FDE0ACFE2CECFD68" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[625,716,691,717]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="dawa" status="sp. nov.">
|
||
<emphasis id="82B7D346FFA8FFB7FDE0ACFE2CECFD68" box="[625,716,691,717]" italics="true" pageId="24" pageNumber="203">A. dawa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFA8FFB7FD42ACFE2D0BFD68" bold="true" box="[723,811,691,717]" pageId="24" pageNumber="203">
|
||
<taxonomicNameLabel id="99846E3DFFA8FFB7FD42ACFE2D0BFD68" box="[723,811,691,717]" pageId="24" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are females and that the male condition of the chelipeds is currently unknown. Nevertheless, in other species of
|
||
<taxonomicName id="77C374D7FFA8FFB7FC83AC9A2DAEFD54" authorityName="Coutiere" authorityYear="1897" box="[786,910,727,753]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFA8FFB7FC83AC9A2DAEFD54" box="[786,910,727,753]" italics="true" pageId="24" pageNumber="203">Athanopsis</emphasis>
|
||
</taxonomicName>
|
||
, the relative development of the right and left cheliped is similar between males and females (e.g.,
|
||
<bibRefCitation id="D45272A5FFA8FFB7FD75ACB62D4DFCB0" author="Anker, A." box="[740,877,763,789]" pageId="24" pageNumber="203" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
|
||
: fig. 6;
|
||
<bibRefCitation id="D45272A5FFA8FFB7FC50ACB62AA8FCB0" author="Marin, I. & Sheibani, R. & Sari, A." box="[961,1160,763,789]" pageId="24" pageNumber="203" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. & Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
|
||
Marin
|
||
<emphasis id="82B7D346FFA8FFB7FB9CACB62A65FCB0" box="[1037,1093,763,789]" italics="true" pageId="24" pageNumber="203">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
: fig. 6).
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFA8FFB7FF56AD522A52F9F4" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="203">
|
||
The lack of armature (in the form of spiniform setae) on the merus of the third and fourth pereiopods places
|
||
<taxonomicName id="77C374D7FFA8FFB7FA15AD522EF4FCF8" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="dawa" status="sp. nov.">
|
||
<emphasis id="82B7D346FFA8FFB7FA15AD522EF4FCF8" italics="true" pageId="24" pageNumber="203">A. dawa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFA8FFB7FF4BAD0E2F12FCF8" bold="true" box="[218,306,835,861]" pageId="24" pageNumber="203">
|
||
<taxonomicNameLabel id="99846E3DFFA8FFB7FF4BAD0E2F12FCF8" box="[218,306,835,861]" pageId="24" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
closest to
|
||
<taxonomicName id="77C374D7FFA8FFB7FE34AD0E2C64FCF8" authorityName="Banner & Banner" authorityYear="1981" box="[421,580,835,861]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
<emphasis id="82B7D346FFA8FFB7FE34AD0E2C64FCF8" box="[421,580,835,861]" italics="true" pageId="24" pageNumber="203">A. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="77C374D7FFA8FFB7FDC0AD0E2C9AFCF9" authorityName="Anker" authorityYear="2011" box="[593,698,835,861]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="saurus">
|
||
<emphasis id="82B7D346FFA8FFB7FDC0AD0E2C9AFCF9" box="[593,698,835,861]" italics="true" pageId="24" pageNumber="203">A. saurus</emphasis>
|
||
</taxonomicName>
|
||
(see above),
|
||
<taxonomicName id="77C374D7FFA8FFB7FCDAAD0E2A95FCF8" authority="Coutiere, 1897" authorityName="Coutiere" authorityYear="1897" box="[843,1205,835,861]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="platyrhynchus">
|
||
<emphasis id="82B7D346FFA8FFB7FCDAAD0E2A27FCF8" box="[843,1031,835,861]" italics="true" pageId="24" pageNumber="203">A. platyrhynchus</emphasis>
|
||
<bibRefCitation id="D45272A5FFA8FFB7FB9CAD0E2A95FCF8" author="Coutiere, H." box="[1037,1205,835,861]" pageId="24" pageNumber="203" pagination="301 - 303" refId="ref21484" refString="Coutiere, H. (1897) Note sur un nouveau genre d'Alpheides. Bulletin du Museum d'Histoire Naturelle, 1 (3), 301 - 303. https: // doi. org / 10.5962 / bhl. part. 19260" type="journal article" year="1897">Coutière, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="77C374D7FFA8FFB7FB2EAD0E2FB6FC24" authority="Marin, Sheibani & Sari, 2014" authorityName="Marin, Sheibani & Sari" authorityYear="2014" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="tarahomii">
|
||
<emphasis id="82B7D346FFA8FFB7FB2EAD0E2B6BFCF8" box="[1215,1355,835,861]" italics="true" pageId="24" pageNumber="203">A. tarahomii</emphasis>
|
||
<bibRefCitation id="D45272A5FFA8FFB7FAC0AD0E2FB6FC24" author="Marin, I. & Sheibani, R. & Sari, A." pageId="24" pageNumber="203" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. & Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">Marin, Sheibani & Sari, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFA8FFB7FE41AD2A2CE3FC24" authority="Anker, 2012" authorityName="Anker" authorityYear="2012" box="[464,707,871,897]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="gotoi">
|
||
<emphasis id="82B7D346FFA8FFB7FE41AD2A2C0BFC25" box="[464,555,871,897]" italics="true" pageId="24" pageNumber="203">A. gotoi</emphasis>
|
||
<bibRefCitation id="D45272A5FFA8FFB7FDA3AD2A2CE3FC24" author="Anker, A." box="[562,707,871,897]" pageId="24" pageNumber="203" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker, 2012</bibRefCitation>
|
||
</taxonomicName>
|
||
. However, all of them present several significant differences, in addition to lacking the post-rostral tubercle on the carapace and having very unequal and asymmetrical chelipeds.
|
||
<taxonomicName id="77C374D7FFA8FFB7FF06ADE22F75FC6C" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[151,341,943,969]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="dawa" status="sp. nov.">
|
||
<emphasis id="82B7D346FFA8FFB7FF06ADE22F75FC6C" box="[151,341,943,969]" italics="true" pageId="24" pageNumber="203">Athanopsis dawa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFA8FFB7FEC8ADE22F90FC6C" bold="true" box="[345,432,943,969]" pageId="24" pageNumber="203">
|
||
<taxonomicNameLabel id="99846E3DFFA8FFB7FEC8ADE22F90FC6C" box="[345,432,943,969]" pageId="24" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be separated from
|
||
<taxonomicName id="77C374D7FFA8FFB7FD37ADE22D64FC6C" authorityName="Banner & Banner" authorityYear="1981" box="[678,836,943,969]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
<emphasis id="82B7D346FFA8FFB7FD37ADE22D64FC6C" box="[678,836,943,969]" italics="true" pageId="24" pageNumber="203">A. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
by the relatively longer second article of the antennular peduncle, which is as long as wide (
|
||
<emphasis id="82B7D346FFA8FFB7FDD1AD9E2C7EFC49" box="[576,606,979,1004]" italics="true" pageId="24" pageNumber="203">vs.</emphasis>
|
||
much wider than long in
|
||
<taxonomicName id="77C374D7FFA8FFB7FC01AD9E2A13FC48" authorityName="Banner & Banner" authorityYear="1981" box="[912,1075,979,1005]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
<emphasis id="82B7D346FFA8FFB7FC01AD9E2A13FC48" box="[912,1075,979,1005]" italics="true" pageId="24" pageNumber="203">A. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="77C374D7FFA8FFB7FB1FAD9E2ADBFC49" authorityName="Anker" authorityYear="2011" box="[1166,1275,979,1005]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="saurus">
|
||
<emphasis id="82B7D346FFA8FFB7FB1FAD9E2ADBFC49" box="[1166,1275,979,1005]" italics="true" pageId="24" pageNumber="203">A. saurus</emphasis>
|
||
</taxonomicName>
|
||
by the lateral margins of the rostrum shallowly concave throughout their length (
|
||
<emphasis id="82B7D346FFA8FFB7FC01ADBA2D87FBB5" box="[912,935,1015,1040]" italics="true" pageId="24" pageNumber="203">vs</emphasis>
|
||
. distinctly convex proximally in
|
||
<taxonomicName id="77C374D7FFA8FFB7FAB1ADBA2BABFBB5" authorityName="Anker" authorityYear="2011" box="[1312,1419,1015,1041]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="saurus">
|
||
<emphasis id="82B7D346FFA8FFB7FAB1ADBA2BABFBB5" box="[1312,1419,1015,1041]" italics="true" pageId="24" pageNumber="203">A. saurus</emphasis>
|
||
</taxonomicName>
|
||
); from both
|
||
<taxonomicName id="77C374D7FFA8FFB7FE80AA562F93FB90" authorityName="Banner & Banner" authorityYear="1981" box="[273,435,1051,1077]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
<emphasis id="82B7D346FFA8FFB7FE80AA562F93FB90" box="[273,435,1051,1077]" italics="true" pageId="24" pageNumber="203">A. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFA8FFB7FE7DAA562C78FB91" authorityName="Anker" authorityYear="2011" box="[492,600,1051,1077]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="saurus">
|
||
<emphasis id="82B7D346FFA8FFB7FE7DAA562C78FB91" box="[492,600,1051,1077]" italics="true" pageId="24" pageNumber="203">A. saurus</emphasis>
|
||
</taxonomicName>
|
||
by the absence of dense setal cover on parts of the major chela (
|
||
<emphasis id="82B7D346FFA8FFB7FAD2AA562B7AFB91" box="[1347,1370,1051,1076]" italics="true" pageId="24" pageNumber="203">vs</emphasis>
|
||
. with fields of setae on the fingers and distal portion of the palm in
|
||
<taxonomicName id="77C374D7FFA8FFB7FCD4AA722DC5FBFC" authorityName="Banner & Banner" authorityYear="1981" box="[837,997,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
<emphasis id="82B7D346FFA8FFB7FCD4AA722DC5FBFC" box="[837,997,1087,1113]" italics="true" pageId="24" pageNumber="203">A. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFA8FFB7FB8BAA722AA4FBFD" authorityName="Anker" authorityYear="2011" box="[1050,1156,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="saurus">
|
||
<emphasis id="82B7D346FFA8FFB7FB8BAA722AA4FBFD" box="[1050,1156,1087,1113]" italics="true" pageId="24" pageNumber="203">A. saurus</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="77C374D7FFA8FFB7FB49AA722BB5FBFC" authorityName="Coutiere" authorityYear="1897" box="[1240,1429,1087,1113]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="platyrhynchus">
|
||
<emphasis id="82B7D346FFA8FFB7FB49AA722BB5FBFC" box="[1240,1429,1087,1113]" italics="true" pageId="24" pageNumber="203">A. platyrhynchus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="77C374D7FFA8FFB7FF06AA2E2F06FBD8" authorityName="Marin, Sheibani & Sari" authorityYear="2014" box="[151,294,1123,1149]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="tarahomii">
|
||
<emphasis id="82B7D346FFA8FFB7FF06AA2E2F06FBD8" box="[151,294,1123,1149]" italics="true" pageId="24" pageNumber="203">A. tarahomii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFA8FFB7FECEAA2E2F99FBD9" authorityName="Anker" authorityYear="2012" box="[351,441,1123,1149]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="gotoi">
|
||
<emphasis id="82B7D346FFA8FFB7FECEAA2E2F99FBD9" box="[351,441,1123,1149]" italics="true" pageId="24" pageNumber="203">A. gotoi</emphasis>
|
||
</taxonomicName>
|
||
by the much longer stylocerite, by far exceeding the distal margin of the second article of the antennular peduncle (
|
||
<emphasis id="82B7D346FFA8FFB7FE4AAACA2FD2FB05" box="[475,498,1159,1184]" italics="true" pageId="24" pageNumber="203">vs</emphasis>
|
||
. reaching only to the mid-length of the second article in
|
||
<taxonomicName id="77C374D7FFA8FFB7FB18AACA2B69FB04" authorityName="Coutiere" authorityYear="1897" box="[1161,1353,1159,1185]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="platyrhynchus">
|
||
<emphasis id="82B7D346FFA8FFB7FB18AACA2B69FB04" box="[1161,1353,1159,1185]" italics="true" pageId="24" pageNumber="203">A. platyrhynchus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFA8FFB7FA15AACA2F25FB60" authorityName="Marin, Sheibani & Sari" authorityYear="2014" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="tarahomii">
|
||
<emphasis id="82B7D346FFA8FFB7FA15AACA2F25FB60" italics="true" pageId="24" pageNumber="203">A. tarahomii</emphasis>
|
||
</taxonomicName>
|
||
, or falling short of this article’s distal margin in
|
||
<taxonomicName id="77C374D7FFA8FFB7FCB9AAE62DA2FB61" authorityName="Anker" authorityYear="2012" box="[808,898,1195,1221]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="gotoi">
|
||
<emphasis id="82B7D346FFA8FFB7FCB9AAE62DA2FB61" box="[808,898,1195,1221]" italics="true" pageId="24" pageNumber="203">A. gotoi</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="77C374D7FFA8FFB7FC46AAE62A45FB60" authorityName="Marin, Sheibani & Sari" authorityYear="2014" box="[983,1125,1195,1221]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="tarahomii">
|
||
<emphasis id="82B7D346FFA8FFB7FC46AAE62A45FB60" box="[983,1125,1195,1221]" italics="true" pageId="24" pageNumber="203">A. tarahomii</emphasis>
|
||
</taxonomicName>
|
||
by the noticeably slenderer third and fourth pereiopods (which are very stout in
|
||
<taxonomicName id="77C374D7FFA8FFB7FD4DAA822D49FB4C" authorityName="Marin, Sheibani & Sari" authorityYear="2014" box="[732,873,1231,1257]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="tarahomii">
|
||
<emphasis id="82B7D346FFA8FFB7FD4DAA822D49FB4C" box="[732,873,1231,1257]" italics="true" pageId="24" pageNumber="203">A. tarahomii</emphasis>
|
||
</taxonomicName>
|
||
), the third maxilliped with a proportionally longer and more rectangular-shaped penultimate article (
|
||
<emphasis id="82B7D346FFA8FFB7FD2FAABE2CF5FAA9" box="[702,725,1267,1292]" italics="true" pageId="24" pageNumber="203">vs</emphasis>
|
||
. with a very short, almost rounded, dorsally convex penultimate article in
|
||
<taxonomicName id="77C374D7FFA8FFB7FE92AB5A2FB1FA94" authorityName="Marin, Sheibani & Sari" authorityYear="2014" box="[259,401,1303,1329]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="tarahomii">
|
||
<emphasis id="82B7D346FFA8FFB7FE92AB5A2FB1FA94" box="[259,401,1303,1329]" italics="true" pageId="24" pageNumber="203">A. tarahomii</emphasis>
|
||
</taxonomicName>
|
||
), and the rostrum pointing forwards (
|
||
<emphasis id="82B7D346FFA8FFB7FCAFAB5A2D75FA95" box="[830,853,1303,1328]" italics="true" pageId="24" pageNumber="203">vs</emphasis>
|
||
. descending in
|
||
<taxonomicName id="77C374D7FFA8FFB7FB99AB5A2AB6FA94" authorityName="Marin, Sheibani & Sari" authorityYear="2014" box="[1032,1174,1303,1329]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="tarahomii">
|
||
<emphasis id="82B7D346FFA8FFB7FB99AB5A2AB6FA94" box="[1032,1174,1303,1329]" italics="true" pageId="24" pageNumber="203">A. tarahomii</emphasis>
|
||
</taxonomicName>
|
||
); and from
|
||
<taxonomicName id="77C374D7FFA8FFB7FA8EAB5A2B59FA95" authorityName="Anker" authorityYear="2012" box="[1311,1401,1303,1329]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="gotoi">
|
||
<emphasis id="82B7D346FFA8FFB7FA8EAB5A2B59FA95" box="[1311,1401,1303,1329]" italics="true" pageId="24" pageNumber="203">A. gotoi</emphasis>
|
||
</taxonomicName>
|
||
by the dorsally partly exposed eyes (which are dorsally concealed by the orbital hoods in
|
||
<taxonomicName id="77C374D7FFA8FFB7FBF3AB762A9CFAF1" authorityName="Anker" authorityYear="2012" box="[1122,1212,1339,1365]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="gotoi">
|
||
<emphasis id="82B7D346FFA8FFB7FBF3AB762A9CFAF1" box="[1122,1212,1339,1365]" italics="true" pageId="24" pageNumber="203">A. gotoi</emphasis>
|
||
</taxonomicName>
|
||
) and the absence of tubercles on the ventral surface of the chelae (which are present in the major chela of
|
||
<taxonomicName id="77C374D7FFA8FFB7FBCCAB122A97FADD" authorityName="Anker" authorityYear="2012" box="[1117,1207,1375,1401]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="gotoi">
|
||
<emphasis id="82B7D346FFA8FFB7FBCCAB122A97FADD" box="[1117,1207,1375,1401]" italics="true" pageId="24" pageNumber="203">A. gotoi</emphasis>
|
||
</taxonomicName>
|
||
) (cf.
|
||
<bibRefCitation id="D45272A5FFA8FFB7FB64AB122BB7FADC" author="Coutiere, H." box="[1269,1431,1375,1401]" pageId="24" pageNumber="203" pagination="1 - 6" refId="ref21527" refString="Coutiere, H. (1899) Les " Alpheidae ", morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie, Serie 8, Vol. 9. Masson, Paris, 559 pp., pls. 1 - 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="book chapter" year="1899">Coutière 1899</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D45272A5FFA8FFB7FF06ABCE2F83FA38" author="Banner, D. M. & Banner, A. H." box="[151,419,1411,1437]" pageId="24" pageNumber="203" pagination="1 - 99" refId="ref20938" refString="Banner, D. M. & Banner, A. H. (1981) Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner & Banner 1981</bibRefCitation>
|
||
; Anker 2011, 2012;
|
||
<bibRefCitation id="D45272A5FFA8FFB7FD1CABCE2D79FA38" author="Marin, I. & Sheibani, R. & Sari, A." box="[653,857,1411,1437]" pageId="24" pageNumber="203" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. & Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
|
||
Marin
|
||
<emphasis id="82B7D346FFA8FFB7FD4BABCE2D2DFA38" box="[730,781,1411,1437]" italics="true" pageId="24" pageNumber="203">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
). The colour patterns are known for
|
||
<taxonomicName id="77C374D7FFA8FFB7FB6AABCE2BBDFA38" authorityName="Banner & Banner" authorityYear="1981" box="[1275,1437,1411,1437]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
<emphasis id="82B7D346FFA8FFB7FB6AABCE2BBDFA38" box="[1275,1437,1411,1437]" italics="true" pageId="24" pageNumber="203">A. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="D45272A5FFA8FFB7FF0EABEA2F09FA64" author="Anker, A." box="[159,297,1447,1473]" pageId="24" pageNumber="203" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
|
||
; see also
|
||
<figureCitation id="28F813D1FFA8FFB7FE03ABEA2FF4FA64" box="[402,468,1447,1473]" captionStart="FIGURE 8" captionStartId="17.[151,250,1909,1933]" captionTargetBox="[151,1435,181,1885]" captionTargetId="figure-19@17.[151,1435,181,1885]" captionTargetPageId="17" captionText="FIGURE 8. Athanopsis brevirostris Banner & Banner, 1981, ovigerous female (cl 5.0 mm) from Thuwal, Saudi Arabia (FLMNH UF 71444) [a, b] and its host [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, burrowing echiuran, probably Ochetostoma sp. or Listriolobus sp. (Thalassematidae), photographed shortly after death. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350797" httpUri="https://zenodo.org/record/10350797/files/figure.png" pageId="24" pageNumber="203">Fig. 8</figureCitation>
|
||
),
|
||
<taxonomicName id="77C374D7FFA8FFB7FE7AABEA2C75FA65" authorityName="Anker" authorityYear="2011" box="[491,597,1447,1473]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="saurus">
|
||
<emphasis id="82B7D346FFA8FFB7FE7AABEA2C75FA65" box="[491,597,1447,1473]" italics="true" pageId="24" pageNumber="203">A. saurus</emphasis>
|
||
</taxonomicName>
|
||
(Anker 2011, 2012; see also
|
||
<figureCitation id="28F813D1FFA8FFB7FC0AABEA2DFEFA64" box="[923,990,1447,1473]" captionStart="FIGURE 9" captionStartId="18.[151,250,1878,1902]" captionTargetBox="[151,1436,151,1860]" captionTargetId="figure-34@18.[151,1436,151,1860]" captionTargetPageId="18" captionText="FIGURE 9. Athanopsis saurus Anker, 2011, male (cl 4.3 mm) from Pouébo, New Caledonia (MNHN-IU-2018-974) [a–e] and its host [f]; a, shrimp in life, lateral (left side); b, same, dorsal; c, d, e, shrimp on various areas of the host (see f); f, burrowing echiuran, probably Ochetostoma sp. (Thalassematidae), with commensal bivalves attached to the trunk (two of them also visible in c). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350801" httpUri="https://zenodo.org/record/10350801/files/figure.png" pageId="24" pageNumber="203">Fig. 9</figureCitation>
|
||
),
|
||
<taxonomicName id="77C374D7FFA8FFB7FC62ABEA2AA0FA64" authorityName="Marin, Sheibani & Sari" authorityYear="2014" box="[1011,1152,1447,1473]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="tarahomii">
|
||
<emphasis id="82B7D346FFA8FFB7FC62ABEA2AA0FA64" box="[1011,1152,1447,1473]" italics="true" pageId="24" pageNumber="203">A. tarahomii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="28F813D1FFA8FFB7FB01ABEA2AC3FA64" box="[1168,1251,1447,1473]" captionStart="FIGURE 14" captionStartId="27.[151,250,1909,1933]" captionTargetBox="[170,1417,181,1884]" captionTargetId="figure-19@27.[170,1417,181,1884]" captionTargetPageId="27" captionText="FIGURE 14. Athanopsis tarahomii Marin, Sheibani & Sari, 2014, male (cl 4.6 mm) [a, b] from Thuwal, Saudi Arabia (FLMNH UF 71425); ovigerous female (cl 4.7 mm) from the same locality (same burrow) (FLMNH UF 71425) [c, d]; a, shrimp in life (male), lateral (left side); b, same, dorsal; c, shrimp in life (female), lateral (right side); d, same, dorsal. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350819" httpUri="https://zenodo.org/record/10350819/files/figure.png" pageId="24" pageNumber="203">Fig. 14</figureCitation>
|
||
; not
|
||
<bibRefCitation id="D45272A5FFA8FFB7FA88ABEA2EF3FA40" author="Marin, I. & Sheibani, R. & Sari, A." pageId="24" pageNumber="203" pagination="398 - 410" refId="ref22937" refString="Marin, I., Sheibani, R. & Sari, A. (2014) A re-description of rare alpheid shrimp Amphibetaeus jousseaumei (Coutiere, 1896) and description of a new species of the genus Athanopsis Coutiere, 1897 (Crustacea: Decapoda: Alpheidae) from Iranian coast of the Persian Gulf. Zootaxa, 3846 (3), 398 - 410. https: // doi. org / 10.11646 / zootaxa. 3846.3.4" type="journal article" year="2014">
|
||
Marin
|
||
<emphasis id="82B7D346FFA8FFB7FAF5ABEA2BB5FA64" box="[1380,1429,1447,1473]" italics="true" pageId="24" pageNumber="203">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
: fig. 7, see below), and
|
||
<taxonomicName id="77C374D7FFA8FFB7FE49AB862CEEFA40" authority="(Anker 2012)" baseAuthorityName="Anker" baseAuthorityYear="2012" box="[472,718,1483,1509]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="gotoi">
|
||
<emphasis id="82B7D346FFA8FFB7FE49AB862C11FA41" box="[472,561,1483,1509]" italics="true" pageId="24" pageNumber="203">A. gotoi</emphasis>
|
||
(
|
||
<bibRefCitation id="D45272A5FFA8FFB7FDAFAB862CE6FA40" author="Anker, A." box="[574,710,1483,1509]" pageId="24" pageNumber="203" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and are different from that of
|
||
<taxonomicName id="77C374D7FFA8FFB7FB8AAB862A56FA40" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1051,1142,1483,1509]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="dawa" status="sp. nov.">
|
||
<emphasis id="82B7D346FFA8FFB7FB8AAB862A56FA40" box="[1051,1142,1483,1509]" italics="true" pageId="24" pageNumber="203">A. dawa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFA8FFB7FBEDAB862AF5FA40" bold="true" box="[1148,1237,1483,1509]" pageId="24" pageNumber="203">
|
||
<taxonomicNameLabel id="99846E3DFFA8FFB7FBEDAB862AF5FA40" box="[1148,1237,1483,1509]" pageId="24" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="28F813D1FFA8FFB7FB72AB862B7EFA40" box="[1251,1374,1483,1509]" captionStart="FIGURE 12" captionStartId="22.[151,250,1733,1757]" captionTargetBox="[151,1436,181,1709]" captionTargetId="figure-34@22.[151,1436,181,1709]" captionTargetPageId="22" captionText="FIGURE 12. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424) [a, b] and species’ habitat [c]; a, shrimp in life, lateral (left side); b, same, dorsal; c, type locality, sandflat with some seagrass and coral rubble, near Dafiyat, Masirah Island, Oman. Photographs by the author [a, b] and Gustav Paulay [c]." figureDoi="http://doi.org/10.5281/zenodo.10350813" httpUri="https://zenodo.org/record/10350813/files/figure.png" pageId="24" pageNumber="203">Fig. 12a, b</figureCitation>
|
||
). The colour pattern of
|
||
<taxonomicName id="77C374D7FFA8FFB7FECAABA22C39F9AC" authorityName="Coutiere" authorityYear="1897" box="[347,537,1519,1545]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="platyrhynchus">
|
||
<emphasis id="82B7D346FFA8FFB7FECAABA22C39F9AC" box="[347,537,1519,1545]" italics="true" pageId="24" pageNumber="203">A. platyrhynchus</emphasis>
|
||
</taxonomicName>
|
||
remains known only from
|
||
<bibRefCitation id="D45272A5FFA8FFB7FCDFABA22A2DF9AC" author="Coutiere, H." box="[846,1037,1519,1545]" pageId="24" pageNumber="203" pagination="301 - 303" refId="ref21484" refString="Coutiere, H. (1897) Note sur un nouveau genre d'Alpheides. Bulletin du Museum d'Histoire Naturelle, 1 (3), 301 - 303. https: // doi. org / 10.5962 / bhl. part. 19260" type="journal article" year="1897">Coutière’s (1897</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="D45272A5FFA8FFB7FB88ABA22A72F9AC" author="Coutiere, H." box="[1049,1106,1519,1545]" pageId="24" pageNumber="203" pagination="1 - 6" refId="ref21527" refString="Coutiere, H. (1899) Les " Alpheidae ", morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie, Serie 8, Vol. 9. Masson, Paris, 559 pp., pls. 1 - 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="book chapter" year="1899">1899</bibRefCitation>
|
||
) brief description (see
|
||
<bibRefCitation id="D45272A5FFA8FFB7FAC7ABA22EEFF988" author="Anker, A." pageId="24" pageNumber="203" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
|
||
for translation). Generally, the taxonomic identity of
|
||
<taxonomicName id="77C374D7FFA8FFB7FCA3A85E2DD0F988" authorityName="Coutiere" authorityYear="1897" box="[818,1008,1555,1581]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="platyrhynchus">
|
||
<emphasis id="82B7D346FFA8FFB7FCA3A85E2DD0F988" box="[818,1008,1555,1581]" italics="true" pageId="24" pageNumber="203">A. platyrhynchus</emphasis>
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="6F78B1F6FFA8FFB7FBBBA85E2A7AF988" box="[1066,1114,1555,1581]" pageId="24" pageNumber="203">type</typeStatus>
|
||
species of the genus known with certainty only from
|
||
<collectingCountry id="C8D44FC4FFA8FFB7FE3EA87A2C2FF9F4" box="[431,527,1591,1617]" name="Djibouti" pageId="24" pageNumber="203">Djibouti</collectingCountry>
|
||
, remains problematic, as pointed out by
|
||
<bibRefCitation id="D45272A5FFA8FFB7FC42A87A2A4EF9F4" author="Anker, A." box="[979,1134,1591,1617]" pageId="24" pageNumber="203" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker (2012)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B07C0F54FFA8FFB7FF56A8162F57F878" blockId="24.[151,1437,151,2013]" pageId="24" pageNumber="203">
|
||
The remaining species of
|
||
<taxonomicName id="77C374D7FFA8FFB7FE7AA8162C47F9D0" authorityName="Coutiere" authorityYear="1897" box="[491,615,1627,1653]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="82B7D346FFA8FFB7FE7AA8162C47F9D0" box="[491,615,1627,1653]" italics="true" pageId="24" pageNumber="203">Athanopsis</emphasis>
|
||
</taxonomicName>
|
||
, i.e.,
|
||
<taxonomicName id="77C374D7FFA8FFB7FD36A8162D91F9D0" authority="Miya, 1980" authorityName="Miya" authorityYear="1980" box="[679,945,1627,1653]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="dentipes">
|
||
<emphasis id="82B7D346FFA8FFB7FD36A8162D04F9D0" box="[679,804,1627,1653]" italics="true" pageId="24" pageNumber="203">A. dentipes</emphasis>
|
||
<bibRefCitation id="D45272A5FFA8FFB7FCBDA8162D91F9D0" author="Miya, Y." box="[812,945,1627,1653]" pageId="24" pageNumber="203" pagination="117 - 131" refId="ref23018" refString="Miya, Y. (1980) Two new records of the genera, Athanopsis and Prionalpheus, from Japan, with description of a new species (Crustacea, Decapoda Alpheidae). Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 5, 117 - 131." type="journal article" year="1980">Miya, 1980</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="77C374D7FFA8FFB7FC2DA8162B6CF9D0" authority="Banner & Banner 1982" authorityName="Banner & Banner" authorityYear="1982" box="[956,1356,1627,1653]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="82B7D346FFA8FFB7FC2DA8162A1EF9D0" box="[956,1086,1627,1653]" italics="true" pageId="24" pageNumber="203">A. australis</emphasis>
|
||
<bibRefCitation id="D45272A5FFA8FFB7FBD7A8162B6CF9D0" author="Banner, D. M. & Banner, A. H." box="[1094,1356,1627,1653]" pageId="24" pageNumber="203" pagination="1 - 357" refId="ref20980" refString="Banner, D. M. & Banner, A. H. (1982) The alpheid shrimp of Australia, Part III: The remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner & Banner 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFA8FFB7FA15A8162FC2F93C" authority="Berggren, 1991" authorityName="Berggren" authorityYear="1991" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="rubricinctuta">
|
||
<emphasis id="82B7D346FFA8FFB7FA15A8162F0BF93C" italics="true" pageId="24" pageNumber="203">A. rubricinctuta</emphasis>
|
||
<bibRefCitation id="D45272A5FFA8FFB7FEA2A8322FC2F93C" author="Berggren, M." box="[307,482,1663,1689]" pageId="24" pageNumber="203" pagination="166 - 178" refId="ref21169" refString="Berggren, M. (1991) Athanopsis rubricinctuta, new species (Decapoda: Natantia: Alpheidae), a shrimp associated with an echiuroid at Inhaca Island, Mocambique. Journal of Crustacean Biology, 11, 166 - 178. https: // doi. org / 10.2307 / 1548555" type="journal article" year="1991">Berggren, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
(see below), are characterised by the presence of spiniform setae on the meri of the third, fourth and fifth pereiopods and present several other morphological differences with the new species (cf.
|
||
<bibRefCitation id="D45272A5FFA8FFB7FF06A88A2F35F944" author="Miya, Y." box="[151,277,1735,1761]" pageId="24" pageNumber="203" pagination="117 - 131" refId="ref23018" refString="Miya, Y. (1980) Two new records of the genera, Athanopsis and Prionalpheus, from Japan, with description of a new species (Crustacea, Decapoda Alpheidae). Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 5, 117 - 131." type="journal article" year="1980">Miya 1980</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D45272A5FFA8FFB7FEB0A88A2C09F944" author="Banner, D. M. & Banner, A. H." box="[289,553,1735,1761]" pageId="24" pageNumber="203" pagination="1 - 357" refId="ref20980" refString="Banner, D. M. & Banner, A. H. (1982) The alpheid shrimp of Australia, Part III: The remaining alpheids, principally the genus Alpheus, and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner & Banner 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D45272A5FFA8FFB7FDA4A88A2CFFF944" author="Berggren, M." box="[565,735,1735,1761]" pageId="24" pageNumber="203" pagination="166 - 178" refId="ref21169" refString="Berggren, M. (1991) Athanopsis rubricinctuta, new species (Decapoda: Natantia: Alpheidae), a shrimp associated with an echiuroid at Inhaca Island, Mocambique. Journal of Crustacean Biology, 11, 166 - 178. https: // doi. org / 10.2307 / 1548555" type="journal article" year="1991">Berggren 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D45272A5FFA8FFB7FD7BA88A2DD1F944" author="Anker, A. & Ahyong, S. T." box="[746,1009,1735,1761]" pageId="24" pageNumber="203" pagination="685 - 697" refId="ref19649" refString="Anker, A. & Ahyong, S. T. (2007) A rediagnosis of Athanopsis australis Banner & Banner, 1982, a rare alpheid shrimp from southern Australia, with a phylogeny of Athanopsis Coutiere, 1897 and remarks on antitropical distributions in the Alpheidae (Decapoda, Caridea). Crustaceana, 80, 685 - 697. https: // doi. org / 10.1163 / 156854007781360612" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
). In addition, the colour patterns of
|
||
<taxonomicName id="77C374D7FFA8FFB7FA15A88A2ED4F8A0" authorityName="Miya" authorityYear="1980" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="dentipes">
|
||
<emphasis id="82B7D346FFA8FFB7FA15A88A2ED4F8A0" italics="true" pageId="24" pageNumber="203">A. dentipes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="77C374D7FFA8FFB7FEBBA8A62F8AF8A0" authorityName="Banner & Banner" authorityYear="1982" box="[298,426,1771,1797]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="82B7D346FFA8FFB7FEBBA8A62F8AF8A0" box="[298,426,1771,1797]" italics="true" pageId="24" pageNumber="203">A. australis</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="D45272A5FFA8FFB7FE4FA8A62C7AF8A0" author="Miya, Y." box="[478,602,1771,1797]" pageId="24" pageNumber="203" pagination="117 - 131" refId="ref23018" refString="Miya, Y. (1980) Two new records of the genera, Athanopsis and Prionalpheus, from Japan, with description of a new species (Crustacea, Decapoda Alpheidae). Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 5, 117 - 131." type="journal article" year="1980">Miya 1980</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D45272A5FFA8FFB7FDF5A8A62CCAF8A0" author="Anker, A." box="[612,746,1771,1797]" pageId="24" pageNumber="203" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
|
||
) are very different from that of
|
||
<taxonomicName id="77C374D7FFA8FFB7FBDAA8A62A86F8A0" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[1099,1190,1771,1797]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="dawa" status="sp. nov.">
|
||
<emphasis id="82B7D346FFA8FFB7FBDAA8A62A86F8A0" box="[1099,1190,1771,1797]" italics="true" pageId="24" pageNumber="203">A. dawa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFA8FFB7FB3DA8A62B25F8A0" bold="true" box="[1196,1285,1771,1797]" pageId="24" pageNumber="203">
|
||
<taxonomicNameLabel id="99846E3DFFA8FFB7FB3DA8A62B25F8A0" box="[1196,1285,1771,1797]" pageId="24" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="28F813D1FFA8FFB7FA85A8A62BAFF8A0" box="[1300,1423,1771,1797]" captionStart="FIGURE 12" captionStartId="22.[151,250,1733,1757]" captionTargetBox="[151,1436,181,1709]" captionTargetId="figure-34@22.[151,1436,181,1709]" captionTargetPageId="22" captionText="FIGURE 12. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424) [a, b] and species’ habitat [c]; a, shrimp in life, lateral (left side); b, same, dorsal; c, type locality, sandflat with some seagrass and coral rubble, near Dafiyat, Masirah Island, Oman. Photographs by the author [a, b] and Gustav Paulay [c]." figureDoi="http://doi.org/10.5281/zenodo.10350813" httpUri="https://zenodo.org/record/10350813/files/figure.png" pageId="24" pageNumber="203">Fig. 12a, b</figureCitation>
|
||
). On the other hand, the colour patterns of
|
||
<taxonomicName id="77C374D7FFA8FFB7FDF0A9422C9CF88C" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[609,700,1807,1833]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="dawa" status="sp. nov.">
|
||
<emphasis id="82B7D346FFA8FFB7FDF0A9422C9CF88C" box="[609,700,1807,1833]" italics="true" pageId="24" pageNumber="203">A. dawa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFA8FFB7FD52A9422D3CF88C" bold="true" box="[707,796,1807,1833]" pageId="24" pageNumber="203">
|
||
<taxonomicNameLabel id="99846E3DFFA8FFB7FD52A9422D3CF88C" box="[707,796,1807,1833]" pageId="24" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="28F813D1FFA8FFB7FCBAA9422D83F88C" box="[811,931,1807,1833]" captionStart="FIGURE 12" captionStartId="22.[151,250,1733,1757]" captionTargetBox="[151,1436,181,1709]" captionTargetId="figure-34@22.[151,1436,181,1709]" captionTargetPageId="22" captionText="FIGURE 12. Athanopsis dawa sp. nov., holotype, female (cl 4.4 mm) from Masirah Island, Oman (FLMNH UF 71424) [a, b] and species’ habitat [c]; a, shrimp in life, lateral (left side); b, same, dorsal; c, type locality, sandflat with some seagrass and coral rubble, near Dafiyat, Masirah Island, Oman. Photographs by the author [a, b] and Gustav Paulay [c]." figureDoi="http://doi.org/10.5281/zenodo.10350813" httpUri="https://zenodo.org/record/10350813/files/figure.png" pageId="24" pageNumber="203">Fig. 12a, b</figureCitation>
|
||
) and
|
||
<taxonomicName id="77C374D7FFA8FFB7FC70A9422AB4F88C" authorityName="Berggren" authorityYear="1991" box="[993,1172,1807,1833]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="rubricinctuta">
|
||
<emphasis id="82B7D346FFA8FFB7FC70A9422AB4F88C" box="[993,1172,1807,1833]" italics="true" pageId="24" pageNumber="203">A. rubricinctuta</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="28F813D1FFA8FFB7FB33A9422ACEF88C" box="[1186,1262,1807,1833]" captionStart="FIGURE 13" captionStartId="26.[151,250,1900,1924]" captionTargetBox="[151,1436,181,1876]" captionTargetId="figure-34@26.[151,1436,181,1876]" captionTargetPageId="26" captionText="FIGURE 13. Athanopsis rubricinctuta Berggren, 1991, male (cl 5.4 mm) from Masirah Island, Oman (FLMNH UF 71437) [a, b]; male (cl 5.8 mm) from the same locality (FLMNH UF 71443) [c]; a, shrimp in life, lateral (right side); b, same, dorsal; c, shrimp in life (different individual), lateral (left side). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.10350817" httpUri="https://zenodo.org/record/10350817/files/figure.png" pageId="24" pageNumber="203">Fig 13</figureCitation>
|
||
; see also
|
||
<bibRefCitation id="D45272A5FFA8FFB7FAC7A9422EF3F8E8" author="Anker, A." pageId="24" pageNumber="203" pagination="48 - 61" refId="ref19400" refString="Anker, A. (2012) Notes on the Indo-West Pacific shrimp genus Athanopsis Coutiere, 1897 (Crustacea, Decapoda, Alpheidae), with the description of a new species associated with echiurans (Annelida, Thalassematidae). Zootaxa, 3307 (1), 48 - 61. https: // doi. org / 10.11646 / zootaxa. 3307.1.3" type="journal article" year="2012">Anker 2012</bibRefCitation>
|
||
: fig. 7F, G) are very similar, to the extent that the two species may be easily confused in the field. However, at a closer inspection,
|
||
<taxonomicName id="77C374D7FFA8FFB7FEE5A91A2FEFF8D4" authority="Anker, 2023" authorityName="Anker" authorityYear="2023" box="[372,463,1879,1905]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="dawa" status="sp. nov.">
|
||
<emphasis id="82B7D346FFA8FFB7FEE5A91A2FEFF8D4" box="[372,463,1879,1905]" italics="true" pageId="24" pageNumber="203">A. dawa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82B7D346FFA8FFB7FE44A91A2C0EF8D4" bold="true" box="[469,558,1879,1905]" pageId="24" pageNumber="203">
|
||
<taxonomicNameLabel id="99846E3DFFA8FFB7FE44A91A2C0EF8D4" box="[469,558,1879,1905]" pageId="24" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="77C374D7FFA8FFB7FCCAA91A2A2DF8D4" authorityName="Berggren" authorityYear="1991" box="[859,1037,1879,1905]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="rubricinctuta">
|
||
<emphasis id="82B7D346FFA8FFB7FCCAA91A2A2DF8D4" box="[859,1037,1879,1905]" italics="true" pageId="24" pageNumber="203">A. rubricinctuta</emphasis>
|
||
</taxonomicName>
|
||
by the white transverse band on the carapace descending from the dorsal area almost to the branchiostegial margin (
|
||
<emphasis id="82B7D346FFA8FFB7FB88A9362A10F831" box="[1049,1072,1915,1940]" italics="true" pageId="24" pageNumber="203">vs</emphasis>
|
||
. being limited to the dorsal area in
|
||
<taxonomicName id="77C374D7FFA8FFB7FF25A9D22F47F81C" authorityName="Berggren" authorityYear="1991" box="[180,359,1951,1977]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="rubricinctuta">
|
||
<emphasis id="82B7D346FFA8FFB7FF25A9D22F47F81C" box="[180,359,1951,1977]" italics="true" pageId="24" pageNumber="203">A. rubricinctuta</emphasis>
|
||
</taxonomicName>
|
||
) and the presence of an additional white transverse band on the third pleonite (which is missing in
|
||
<taxonomicName id="77C374D7FFA8FFB7FF25A98E2F47F878" authorityName="Berggren" authorityYear="1991" box="[180,359,1987,2013]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="24" pageNumber="203" phylum="Arthropoda" rank="species" species="rubricinctuta">
|
||
<emphasis id="82B7D346FFA8FFB7FF25A98E2F47F878" box="[180,359,1987,2013]" italics="true" pageId="24" pageNumber="203">A. rubricinctuta</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |