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<document id="5CBA7FE49C389F0FB90EBD8D03AF6C20" ID-DOI="10.5852/ejt.2019.526" ID-GBIF-Dataset="2da65c0a-9c48-43f5-ada3-e088de28dec1" ID-ISSN="2118-9773" ID-Zenodo-Dep="3152217" ID-ZooBank="3FFC527D-E9CA-4D9F-BF59-076FA60171AC" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1558535825802" checkinUser="plazi" docAuthor="Nurinsiyah, Ayu Savitri, Neiber, Marco T. &amp; Hausdorf, Bernhard" docDate="2019" docId="03B1DB4C8B374E054C48FB0C3D94FBA4" docLanguage="en" docName="ej-526_nurinsiyah_neiber_hausdorf.pdf.imf" docOrigin="European Journal of Taxonomy 526" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria naggsi Nurinsiyah &amp; Neiber &amp; Hausdorf 2019, sp. nov." docType="treatment" docUuid="4EF9E468-DDEF-460D-939C-7D9C86254CF4" docUuidSource="ZooBank" docVersion="14" lastPageNumber="29" masterDocId="FF88A3348B2D4E194E35FFFC3C0DFFDB" masterDocTitle="Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java" masterLastPageNumber="73" masterPageNumber="1" pageNumber="27" updateTime="1716316918745" updateUser="carolina">
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<mods:title id="47CC3901E3552D6D8DB93C2643F22753">Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java</mods:title>
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<mods:name id="C725415D19873215FF1F8A83CA03D8B8" type="personal">
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<mods:namePart id="B1D6ECEEDD4ECDE1A9AEBCB271B4C7BA">Nurinsiyah, Ayu Savitri</mods:namePart>
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<mods:affiliation id="4DB82346ADF812ECF4D5EF9FB111D3B5">Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia.</mods:affiliation>
<mods:nameIdentifier id="4688B6948A734E47B70DD7EDBE2EAA5D" type="email">ayu_nurinsiyah@yahoo.com &amp; ayus002@lipi.go.id</mods:nameIdentifier>
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<mods:namePart id="868B73E60E7F8895D89A5BFE33DE5C18">Neiber, Marco T.</mods:namePart>
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<mods:affiliation id="92831762B3C7F7B148C2C5A205D2C429">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier id="FB9E6D2DF4F996C4B9A3D6894C1B0DCF" type="email">mneiber@hotmail.de</mods:nameIdentifier>
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<mods:namePart id="A55BA774184CF8BE0C3D0035A0924C77">Hausdorf, Bernhard</mods:namePart>
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<mods:affiliation id="3E233DDCA21996D9694BFFD3E44B90A0">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.</mods:affiliation>
<mods:nameIdentifier id="040CE08BC7E7F6F93DC1D7A383B159FF" type="email">hausdorf@zoologie.uni-hamburg.de</mods:nameIdentifier>
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<mods:title id="B42ED0270DFFC6AD7F7EA8B8A59DABE8">European Journal of Taxonomy</mods:title>
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<mods:date id="9C0444BA41BA4A18ABA5819ECA384751">2019</mods:date>
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<mods:number id="C1AA1EF1E92B08826FC650DC9DF09E88">2019-05-21</mods:number>
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<mods:number id="ACB3A5078A5F613D082462F6E90AEE0C">526</mods:number>
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<treatment id="03B1DB4C8B374E054C48FB0C3D94FBA4" ID-GBIF-Taxon="157239582" LSID="urn:lsid:zoobank.org:act:4EF9E468-DDEF-460D-939C-7D9C86254CF4" httpUri="http://treatment.plazi.org/id/03B1DB4C8B374E054C48FB0C3D94FBA4" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
<subSubSection id="C30239D18B374E034C48FB0C38CAFAF4" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BA76A5A8B374E034C48FB0C3FBAFAD0" blockId="26.[365,1223,1264,1362]" box="[637,951,1264,1291]" pageId="26" pageNumber="27">
<heading id="D0EFDD368B374E034C48FB0C3FBAFAD0" box="[637,951,1264,1291]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="27" reason="2">
<taxonomicName id="4C1811D98B374E034C48FB0C3F58FAD0" ID-CoL="7RTHS" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[637,853,1264,1291]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="naggsi" status="sp. nov.">
<emphasis id="B96CB6488B374E034C48FB0C3F58FAD0" bold="true" box="[637,853,1264,1291]" italics="true" pageId="26" pageNumber="27">Landouria naggsi</emphasis>
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<taxonomicNameLabel id="A25F0B338B374E034D69FB0D3FBAFAD0" box="[860,951,1265,1291]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B374E034CA4FACB3FAEFA8A" box="[657,931,1335,1362]" pageId="26" pageNumber="27" type="description">
<paragraph id="8BA76A5A8B374E034CA4FACB3FAEFA8A" blockId="26.[365,1223,1264,1362]" box="[657,931,1335,1362]" pageId="26" pageNumber="27">
Figs
<figureCitation id="132376DF8B374E034CFEFACB3EE6FA89" box="[715,747,1335,1362]" captionStart="Figs 25–28" captionStartId="28.[189,241,1858,1884]" captionTargetBox="[221,1366,265,2019]" captionTargetId="figure@27.[221,1366,265,2019]" captionTargetPageId="27" captionText="Figs 25–28 (opposite page). Shells of species of Landouria from Java. 25. L. naggsi sp. nov., West Java, Cibitung, near Cikaso Waterfall (holotype, MZB 20424). 26. L. epiplatia (Möllendorff, 1897), West Java, Jampang (lectotype of Plectotropis epiplatia, SMF 8909). 27. L. epiplatia, West Java, Jampang (lectotype of Plectotropis trichotrochium Möllendorff, 1897, SMF 8918). 28. L. epiplatia?, West Java, Bogor, Gunung Gede (ZMH 79946). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152245/files/figure.png" pageId="26" pageNumber="27">25</figureCitation>
,
<figureCitation id="132376DF8B374E034CC2FACB3F15FA89" box="[759,792,1335,1362]" captionStart="Figs 52–55" captionStartId="33.[189,241,1717,1743]" captionTargetBox="[189,1398,265,1993]" captionTargetId="figure@34.[189,1398,265,1993]" captionTargetPageId="34" captionText="Figs 52–55 (opposite page). Distribution of species of Landouria in Java (10′ grid). 52. L. winteriana (Pfeiffer, 1842), L. naggsi sp. nov., L. nusakambangensis sp. nov., L. petrukensis sp. nov., L. tholiformis sp. nov., L. tonywhitteni sp. nov., L. smimensis (Mousson, 1848). 53. L. conoidea (Leschke, 1914), L. abdidalem sp. nov., L. sewuensis sp. nov., L. sukoliloensis sp. nov., L. pacitanensis sp. nov., L. intumescens (Martens, 1867), L. madurensis sp. nov. 54. L. rotatoria (Pfeiffer, 1842), L. nodifera sp. nov., L. parahyangensis sp. nov., L. schepmani (Möllendorff, 1897), L. zonifera sp. nov., L. ciliocincta (Möllendorff, 1897), L. pakidulan sp. nov. 55. L. epiplatia (Möllendorff, 1897), L. monticola van Benthem Jutting, 1950, L. davini Dharma, 2015, L. menorehensis sp. nov., L. dharmai sp. nov., L. moussoniana (Martens, 1867)." httpUri="https://zenodo.org/record/3152265/files/figure.png" pageId="26" pageNumber="27">52</figureCitation>
;
<tableCitation id="C69A5FE18B374E034D16FACB3FAEFA8A" box="[803,931,1335,1361]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="26" pageNumber="27" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B374E034E88FA7C3DC2FA37" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8BA76A5A8B374E034E88FA7C3D3BFA41" blockId="26.[189,310,1408,1434]" box="[189,310,1408,1434]" pageId="26" pageNumber="27">
<heading id="D0EFDD368B374E034E88FA7C3D3BFA41" bold="true" box="[189,310,1408,1434]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B96CB6488B374E034E88FA7C3D3BFA41" bold="true" box="[189,310,1408,1434]" pageId="26" pageNumber="27">Diagnosis</emphasis>
</heading>
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<paragraph id="8BA76A5A8B374E034E88FA533DC2FA37" blockId="26.[189,1398,1455,1516]" pageId="26" pageNumber="27">
<taxonomicName id="4C1811D98B374E034E88FA533D82FA12" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,399,1455,1481]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="naggsi" status="sp. nov.">
<emphasis id="B96CB6488B374E034E88FA533D82FA12" box="[189,399,1455,1481]" italics="true" pageId="26" pageNumber="27">Landouria naggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B374E034FADFA533DF9FA12" box="[408,500,1455,1481]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by an angular, tightly coiled, high conical shell with scales all over the teleoconch.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B374E034E88F9E73E14F95D" pageId="26" pageNumber="27" type="etymology">
<paragraph id="8BA76A5A8B374E034E88F9E73D49F9EE" blockId="26.[189,324,1563,1589]" box="[189,324,1563,1589]" pageId="26" pageNumber="27">
<heading id="D0EFDD368B374E034E88F9E73D49F9EE" bold="true" box="[189,324,1563,1589]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B96CB6488B374E034E88F9E73D49F9EE" bold="true" box="[189,324,1563,1589]" pageId="26" pageNumber="27">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B374E034E88F9B53E14F95D" blockId="26.[189,1399,1609,1670]" pageId="26" pageNumber="27">
<taxonomicName id="4C1811D98B374E034E88F9B53D83F9B8" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,398,1609,1635]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="naggsi" status="sp. nov.">
<emphasis id="B96CB6488B374E034E88F9B53D83F9B8" box="[189,398,1609,1635]" italics="true" pageId="26" pageNumber="27">Landouria naggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B374E034FA3F9B63DFDF9BF" box="[406,496,1610,1636]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
is named in honour of the British malacologist Fred Naggs, who brought the authors of this study together.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B374E054E88F9523D10FE92" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8BA76A5A8B374E034E88F9523DA7F913" blockId="26.[189,426,1710,1736]" box="[189,426,1710,1736]" pageId="26" pageNumber="27">
<heading id="D0EFDD368B374E034E88F9523DA7F913" bold="true" box="[189,426,1710,1736]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B96CB6488B374E034E88F9523DA7F913" bold="true" box="[189,426,1710,1736]" pageId="26" pageNumber="27">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B374E034EDFF9213D51F92C" blockId="26.[189,1399,1757,1889]" box="[234,348,1757,1783]" pageId="26" pageNumber="27">
<typeStatus id="54A3D4F88B374E034EDFF9213D51F92C" box="[234,348,1757,1783]" pageId="26" pageNumber="27" type="holotype">
<emphasis id="B96CB6488B374E034EDFF9213D51F92C" bold="true" box="[234,348,1757,1783]" pageId="26" pageNumber="27">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BA76A5A8B374E034E88F8FC3D58F8BB" blockId="26.[189,1399,1757,1889]" pageId="26" pageNumber="27">
<materialsCitation id="3B7060078B374E034E88F8FC3D58F8BB" ID-GBIF-Occurrence="2243137342" collectedFrom="teak agroforestry" collectingDate="2015-07-05" collectionCode="MZB" collectorName="A. S. Nurinsiyah &amp; T. Setiadi" country="Indonesia" elevation="47" latitude="-7.3613887" location="near Cikaso Waterfall" longLatPrecision="21" longitude="106.619446" municipality="Cibitung" pageId="26" pageNumber="27" specimenCode="20424" specimenCount="1" stateProvince="West Java" typeStatus="holotype">
<collectingCountry id="F30F2ACA8B374E034E88F8FC3D56F8C1" box="[189,347,1792,1818]" name="Indonesia" pageId="26" pageNumber="27">INDONESIA</collectingCountry>
•
<typeStatus id="54A3D4F88B374E034F4EF8FC3DE8F8C1" box="[379,485,1792,1818]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
;
<collectingRegion id="49DCA4B88B374E034FC1F8FC3E62F8C1" box="[500,623,1792,1818]" country="Indonesia" name="Jawa Barat" pageId="26" pageNumber="27">West Java</collectingRegion>
,
<collectingMunicipality id="6BC3F0208B374E034C4AF8FC3EE4F8C1" box="[639,745,1792,1818]" pageId="26" pageNumber="27">Cibitung</collectingMunicipality>
,
<location id="8EC73C818B374E034CC2F8FC380DF8C1" LSID="urn:lsid:plazi:treatment:03B1DB4C8B374E054C48FB0C3D94FBA4:8EC73C818B374E034CC2F8FC380DF8C1" box="[759,1024,1792,1818]" country="Indonesia" latitude="-7.3613887" longLatPrecision="21" longitude="106.619446" municipality="Cibitung" name="near Cikaso Waterfall" pageId="26" pageNumber="27" stateProvince="West Java">near Cikaso Waterfall</location>
,
<collectedFrom id="380429A78B374E034A3AF8FC38D0F8C1" box="[1039,1245,1792,1818]" pageId="26" pageNumber="27">teak agroforestry</collectedFrom>
;
<geoCoordinate id="EE2C0C9D8B374E034AD9F902397CF8C1" box="[1260,1393,1790,1818]" degrees="7" direction="south" minutes="21" orientation="latitude" pageId="26" pageNumber="27" precision="15" seconds="41" value="-7.3613887">7°21′41″ S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B374E034E88F8DD3D52F8E6" box="[189,351,1825,1853]" degrees="106" direction="east" minutes="37" orientation="longitude" pageId="26" pageNumber="27" precision="15" seconds="10" value="106.619446">106°37′10″ E</geoCoordinate>
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<elevation id="00358D698B374E034F59F8DF3DE5F8E6" box="[364,488,1827,1853]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="26" pageNumber="27" unit="m" value="47.0">
<quantity id="4CE0C7BF8B374E034F59F8DF3DA5F8E6" box="[364,424,1827,1853]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="26" pageNumber="27" unit="m" value="47.0">47 m</quantity>
a.s.l.
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;
<collectingDate id="EFE2B5728B374E034FC3F8DF3E71F8E5" box="[502,636,1827,1854]" pageId="26" pageNumber="27" value="2015-07-05">
<date id="FFA64C9A8B374E034FC3F8DF3E71F8E5" box="[502,636,1827,1854]" pageId="26" pageNumber="27" value="2015-07-05">5 Jul. 2015</date>
</collectingDate>
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<collectorName id="26ED0F8C8B374E034CB2F8DF3F48F8E6" box="[647,837,1827,1853]" pageId="26" pageNumber="27">A.S. Nurinsiyah</collectorName>
and
<collectorName id="26ED0F8C8B374E034D4AF8DF3FE2F8E6" box="[895,1007,1827,1853]" pageId="26" pageNumber="27">T. Setiadi</collectorName>
leg.; D =
<quantity id="4CE0C7BF8B374E034A5DF8DF38DEF8E6" box="[1128,1235,1827,1853]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="26" pageNumber="27" unit="mm" value="11.8">11.8 mm</quantity>
, H =
<quantity id="4CE0C7BF8B374E034B21F8DF397FF8E6" box="[1300,1394,1827,1853]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="26" pageNumber="27" unit="mm" value="9.8">9.8 mm</quantity>
;
<collectionCode id="ED09F29F8B374E034E88F8BB3CF0F8BA" box="[189,253,1863,1889]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="26" pageNumber="27" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B374E034F31F8BA3D5CF8BB" box="[260,337,1862,1888]" pageId="26" pageNumber="27">20424</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA76A5A8B374E034EDFF874397BF832" blockId="26.[189,1399,1928,2025]" pageId="26" pageNumber="27">
<materialsCitation id="3B7060078B374E034EDFF8743F32F833" ID-GBIF-Occurrence="2243137487" collectedFrom="riparian (high shrubs)" collectionCode="MZB" country="Indonesia" elevation="61" latitude="-7.361111" location="near Cikaso Waterfall" longLatPrecision="21" longitude="106.61806" municipality="Cibitung" pageId="26" pageNumber="27" specimenCode="20426" specimenCount="1" stateProvince="West Java" typeStatus="paratype">
<typeStatus id="54A3D4F88B374E034EDFF8743D6AF879" box="[234,359,1928,1954]" pageId="26" pageNumber="27" type="paratype">
<emphasis id="B96CB6488B374E034EDFF8743D6AF879" bold="true" box="[234,359,1928,1954]" pageId="26" pageNumber="27">Paratypes</emphasis>
</typeStatus>
<collectingCountry id="F30F2ACA8B374E034E88F8573D56F81E" box="[189,347,1963,1989]" name="Indonesia" pageId="26" pageNumber="27">INDONESIA</collectingCountry>
–
<collectingRegion id="49DCA4B88B374E034FBEF8573E18F81E" box="[395,533,1963,1989]" country="Indonesia" name="Jawa Barat" pageId="26" pageNumber="27">
<emphasis id="B96CB6488B374E034FBEF8573E18F81E" bold="true" box="[395,533,1963,1989]" pageId="26" pageNumber="27">West Java</emphasis>
</collectingRegion>
•
<specimenCount id="9D1EA1D38B374E034C74F8573E92F81E" box="[577,671,1963,1989]" count="1" pageId="26" pageNumber="27" type="generic">1 spec.</specimenCount>
;
<collectingMunicipality id="6BC3F0208B374E034C80F8573F12F81E" box="[693,799,1963,1989]" pageId="26" pageNumber="27">Cibitung</collectingMunicipality>
,
<location id="8EC73C818B374E034D01F8573844F81E" LSID="urn:lsid:plazi:treatment:03B1DB4C8B374E054C48FB0C3D94FBA4:8EC73C818B374E034D01F8573844F81E" box="[820,1097,1963,1989]" country="Indonesia" latitude="-7.361111" longLatPrecision="21" longitude="106.61806" municipality="Cibitung" name="near Cikaso Waterfall" pageId="26" pageNumber="27" stateProvince="West Java">near Cikaso Waterfall</location>
,
<collectedFrom id="380429A78B374E034A68F857397FF81D" box="[1117,1394,1963,1990]" pageId="26" pageNumber="27">riparian (high shrubs)</collectedFrom>
;
<geoCoordinate id="EE2C0C9D8B374E034E88F8303D49F833" box="[189,324,1996,2024]" degrees="7" direction="south" minutes="21" orientation="latitude" pageId="26" pageNumber="27" precision="15" seconds="40" value="-7.361111">7°21′40″ S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B374E034F63F8303DF3F832" box="[342,510,1996,2025]" degrees="106" direction="east" minutes="37" orientation="longitude" pageId="26" pageNumber="27" precision="15" seconds="05" value="106.61806">106°37′05″ E</geoCoordinate>
;
<elevation id="00358D698B374E034C25F8323E9AF833" box="[528,663,1998,2025]" metricMagnitude="1" metricUnit="m" metricValue="6.1" pageId="26" pageNumber="27" unit="m" value="61.0">
<quantity id="4CE0C7BF8B374E034C25F8323E5CF832" box="[528,593,1998,2025]" metricMagnitude="1" metricUnit="m" metricValue="6.1" pageId="26" pageNumber="27" unit="m" value="61.0">61 m</quantity>
a.s.l.
</elevation>
;
<collectionCode id="ED09F29F8B374E034C9FF8333EE7F832" box="[682,746,1999,2025]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="26" pageNumber="27" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B374E034CC3F8323F32F833" box="[758,831,1998,2024]" pageId="26" pageNumber="27">20426</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B7060078B374E054D51F8323D18FEFE" ID-GBIF-Occurrence="2243137455" collectionCode="ZMH" country="Indonesia" elevation="61" lastPageId="28" lastPageNumber="29" latitude="-7.361111" location="Cikaso Waterfall" longLatPrecision="21" longitude="106.61806" municipality="Cibitung" pageId="26" pageNumber="27" specimenCode="148184" specimenCount="1" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="9D1EA1D38B374E034D51F8323FB2F832" box="[868,959,1998,2025]" count="1" pageId="26" pageNumber="27" type="generic">1 spec.</specimenCount>
; same data as for preceding;
<collectionCode id="ED09F29F8B374E034B00F833397BF832" box="[1333,1398,1999,2025]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="26" pageNumber="27" type="Museum">ZMH</collectionCode>
<paragraph id="8BA76A5A8B314E054E88FEF73D10FE92" blockId="28.[189,1398,267,329]" pageId="28" pageNumber="29">
<specimenCode id="DBBEC2218B314E054E88FEF73D18FEFE" box="[189,277,267,293]" pageId="28" pageNumber="29">148184</specimenCode>
•
<materialsCitation id="3B7060078B314E054F07FEF73F63FEFD" ID-GBIF-Occurrence="2243137557" box="[306,878,267,294]" collectingDate="2015-07-05" collectionCode="MZB" collectorName="A. S. Nurinsiyah &amp; T. Setiadi" country="Indonesia" elevation="47" latitude="-7.3613887" location="Cikaso Waterfall" longLatPrecision="21" longitude="106.619446" municipality="Cibitung" pageId="28" pageNumber="29" specimenCode="20425" specimenCount="3" stateProvince="Jawa Barat">
<specimenCount id="9D1EA1D38B314E054F07FEF73D84FEFD" box="[306,393,267,294]" count="3" pageId="28" pageNumber="29" type="generic">3 spec.</specimenCount>
; same data as for holotype;
<collectionCode id="ED09F29F8B314E054CE9FEF03F11FEFD" box="[732,796,268,294]" collectionName="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="28" pageNumber="29" type="Museum">MZB</collectionCode>
<specimenCode id="DBBEC2218B314E054D10FEF73F63FEFD" box="[805,878,267,294]" pageId="28" pageNumber="29">20425</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B7060078B314E054DBEFEF73D14FE92" ID-GBIF-Occurrence="2243137321" collectingDate="2015-07-05" collectionCode="ZMH" collectorName="A. S. Nurinsiyah &amp; T. Setiadi" country="Indonesia" elevation="47" latitude="-7.3613887" location="Cikaso Waterfall" longLatPrecision="21" longitude="106.619446" municipality="Cibitung" pageId="28" pageNumber="29" specimenCode="ZMH 148183" specimenCount="2" stateProvince="Jawa Barat">
<specimenCount id="9D1EA1D38B314E054DBEFEF73FEFFEFD" box="[907,994,267,294]" count="2" pageId="28" pageNumber="29" type="generic">2 spec.</specimenCount>
; same data as for holotype;
<specimenCode id="DBBEC2218B314E054B00FEF03D14FE92" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="28" pageNumber="29" type="Museum">ZMH 148183</specimenCode>
</materialsCitation>
.
</paragraph>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B314E054E88FE8B3DB6FD4D" pageId="28" pageNumber="29" type="description">
<paragraph id="8BA76A5A8B314E054E88FE8B3D43FE4A" blockId="28.[189,334,375,401]" box="[189,334,375,401]" pageId="28" pageNumber="29">
<heading id="D0EFDD368B314E054E88FE8B3D43FE4A" bold="true" box="[189,334,375,401]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B96CB6488B314E054E88FE8B3D43FE4A" bold="true" box="[189,334,375,401]" pageId="28" pageNumber="29">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B314E054E88FE5A38FDFD96" blockId="28.[189,1399,422,589]" pageId="28" pageNumber="29">
SHELL (Fig. 25;
<tableCitation id="C69A5FE18B314E054F4CFE5A3DF3FE1B" box="[377,510,422,448]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="28" pageNumber="29" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
). Conical, with 6.25–7 slightly convex whorls; protoconch almost smooth; teleoconch with irregular wrinkles; incised spiral lines not discernible; scales all over teleoconch; tuberculate around umbilicus; brownish-corneous; body whorl angular; aperture oval; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus hardly eccentric, wide, comprising 28–33% of shell diameter, hardly obscured by columellar edge.
</paragraph>
<paragraph id="8BA76A5A8B314E054E88FD803DB6FD4D" blockId="28.[189,443,636,662]" box="[189,443,636,662]" pageId="28" pageNumber="29">GENITALIA. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30239D18B314E054E88FD393EE7FC38" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8BA76A5A8B314E054E88FD393D22FD04" blockId="28.[189,303,709,735]" box="[189,303,709,735]" pageId="28" pageNumber="29">
<heading id="D0EFDD368B314E054E88FD393D22FD04" bold="true" box="[189,303,709,735]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B96CB6488B314E054E88FD393D22FD04" bold="true" box="[189,303,709,735]" pageId="28" pageNumber="29">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B314E054E88FD0F397AFCAC" blockId="28.[189,1399,755,888]" pageId="28" pageNumber="29">
<taxonomicName id="4C1811D98B314E054E88FD0F3D82FCD6" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,399,755,781]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="naggsi" status="sp. nov.">
<emphasis id="B96CB6488B314E054E88FD0F3D82FCD6" box="[189,399,755,781]" italics="true" pageId="28" pageNumber="29">Landouria naggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B314E054FADFD083DFEFCD5" box="[408,499,756,782]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="4C1811D98B314E054C4CFD083F5EFCD6" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[633,851,755,781]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="parahyangensis" status="sp. nov.">
<emphasis id="B96CB6488B314E054C4CFD083F5EFCD6" box="[633,851,755,781]" italics="true" pageId="28" pageNumber="29">L. parahyangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B314E054D6EFD083FBBFCD5" box="[859,950,756,782]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
(Fig. 48), from which it differs in the smaller (D 11.6–14.0 mm vs
<quantity id="4CE0C7BF8B314E054C20FCEA3ECDFCEA" box="[533,704,790,817]" metricMagnitude="-2" metricUnit="m" metricValue="1.5799999999999998" metricValueMax="1.69" metricValueMin="1.47" pageId="28" pageNumber="29" unit="mm" value="15.799999999999999" valueMax="16.9" valueMin="14.7">14.7–16.9 mm</quantity>
in
<taxonomicName id="4C1811D98B314E054CD0FCEB3FB2FCEA" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[741,959,791,817]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="parahyangensis" status="sp. nov.">
<emphasis id="B96CB6488B314E054CD0FCEB3FB2FCEA" box="[741,959,791,817]" italics="true" pageId="28" pageNumber="29">L. parahyangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B314E054DF3FCEB3829FCEA" box="[966,1060,791,817]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
), higher conical (D/H: 1.21– 1.37 vs
<quantity id="4CE0C7BF8B314E054F29FCC63DBCFC8F" box="[284,433,826,852]" metricMagnitude="-2" metricUnit="m" metricValue="4.6101" metricValueMax="4.9276" metricValueMin="4.2926" pageId="28" pageNumber="29" unit="in" value="1.815" valueMax="1.94" valueMin="1.69">1.69–1.94 in</quantity>
<taxonomicName id="4C1811D98B314E054F8FFCC73E98FC8F" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[442,661,826,852]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="parahyangensis" status="sp. nov.">
<emphasis id="B96CB6488B314E054F8FFCC73E98FC8F" box="[442,661,826,852]" italics="true" pageId="28" pageNumber="29">L. parahyangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B314E054CA8FCC63EF0FC8F" box="[669,765,826,852]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
), less sharply keeled shell with a narrower umbilicus (U/D 0.28–0.33 vs
<quantity id="4CE0C7BF8B314E054FAEFCA13E20FCAC" box="[411,557,861,888]" metricMagnitude="-2" metricUnit="m" metricValue="1.0668" metricValueMax="1.143" metricValueMin="0.9906" pageId="28" pageNumber="29" unit="in" value="0.42" valueMax="0.45" valueMin="0.39">0.39–0.45 in</quantity>
<taxonomicName id="4C1811D98B314E054C06FCA23F06FCAC" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[563,779,861,887]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="parahyangensis" status="sp. nov.">
<emphasis id="B96CB6488B314E054C06FCA23F06FCAC" box="[563,779,861,887]" italics="true" pageId="28" pageNumber="29">L. parahyangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B314E054D24FCA23F66FCA3" box="[785,875,862,888]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
) and in having scales all over the teleoconch.
</paragraph>
<paragraph id="8BA76A5A8B314E054E88FC5A3EE7FC38" blockId="28.[189,1398,934,996]" pageId="28" pageNumber="29">
With regard to the conical shell form
<taxonomicName id="4C1811D98B314E054C44FC5B3ED3FC1B" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[625,734,935,960]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="naggsi" status="sp. nov.">
<emphasis id="B96CB6488B314E054C44FC5B3ED3FC1B" box="[625,734,935,960]" italics="true" pageId="28" pageNumber="29">L. naggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B314E054CD0FC5B3F32FC1A" box="[741,831,935,961]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="4C1811D98B314E054DF7FC5B391DFC1B" authority="(Leschke, 1914)" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" baseAuthorityName="Leschke" baseAuthorityYear="1914" box="[962,1296,934,960]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="B96CB6488B314E054DF7FC5B3841FC1B" box="[962,1100,934,960]" italics="true" pageId="28" pageNumber="29">L. conoidea</emphasis>
(
<bibRefCitation id="EF8917AB8B314E054A6EFC5A3905FC1B" author="Leschke" box="[1115,1288,934,960]" firstAuthor="Leschke" pageId="28" pageNumber="29" pagination="205 - 284" refId="ref33668" refString="Leschke M. 1914. Zur Molluskenfauna von Java und Celebes. Mitteilungen aus dem naturhistorischen Museum in Hamburg 31: 205 - 284, 1 pl." type="journal article" year="1914">Leschke, 1914</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="132376DF8B314E054B15FC5A3964FC1B" box="[1312,1385,934,960]" captionStart="Figs 2–5" captionStartId="11.[189,241,1860,1886]" captionTargetBox="[231,1357,265,1834]" captionTargetId="figure@11.[231,1357,265,1834]" captionTargetPageId="11" captionText="Figs 2–5. Shells of species of Landouria from Java. 2. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype of Helix winteriana, ZMA 376193a). 3. L. winteriana, West Java, Cibodas (syntype of Plectotropis kraepelini Leschke, 1914, ZMH 98416). 4. L. conoidea (Leschke, 1914), West Java, Bogor (syntype, ZMH 98144). 5. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak (neotype of Helix rotatoria, MZB 19205). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152223/files/figure.png" pageId="28" pageNumber="29">Fig. 4</figureCitation>
). For differences, see remarks under that species.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B314E054E88FBEE3D94FBA4" pageId="28" pageNumber="29" type="distribution">
<paragraph id="8BA76A5A8B314E054E88FBEE3D5BFBF7" blockId="28.[189,342,1042,1068]" box="[189,342,1042,1068]" pageId="28" pageNumber="29">
<heading id="D0EFDD368B314E054E88FBEE3D5BFBF7" bold="true" box="[189,342,1042,1068]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B96CB6488B314E054E88FBEE3D5BFBF7" bold="true" box="[189,342,1042,1068]" pageId="28" pageNumber="29">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B314E054E88FBBD3D94FBA4" blockId="28.[189,1398,1089,1151]" pageId="28" pageNumber="29">
<taxonomicName id="4C1811D98B314E054E88FBBD3D9EFB80" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[189,403,1089,1115]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="naggsi" status="sp. nov.">
<emphasis id="B96CB6488B314E054E88FBBD3D9EFB80" box="[189,403,1089,1115]" italics="true" pageId="28" pageNumber="29">Landouria naggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B314E054F95FBBD3DF2FB80" box="[416,511,1089,1115]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
is known only from the surroundings of Cikaso Waterfall in Sukabumi Regency (Fig. 52).
</paragraph>
</subSubSection>
</treatment>
</document>
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