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<document id="4B9EF02606AA781C4B1266FBCF4861BF" ID-DOI="10.5852/ejt.2019.526" ID-GBIF-Dataset="2da65c0a-9c48-43f5-ada3-e088de28dec1" ID-ISSN="2118-9773" ID-Zenodo-Dep="3152217" ID-ZooBank="3FFC527D-E9CA-4D9F-BF59-076FA60171AC" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1558535825802" checkinUser="plazi" docAuthor="Nurinsiyah, Ayu Savitri, Neiber, Marco T. &amp; Hausdorf, Bernhard" docDate="2019" docId="03B1DB4C8B144E224C34FBAE3853FDFE" docLanguage="en" docName="ej-526_nurinsiyah_neiber_hausdorf.pdf.imf" docOrigin="European Journal of Taxonomy 526" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria leucochila Nurinsiyah &amp; Neiber &amp; Hausdorf 2019, comb. nov." docType="treatment" docVersion="16" lastPageNumber="60" masterDocId="FF88A3348B2D4E194E35FFFC3C0DFFDB" masterDocTitle="Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java" masterLastPageNumber="73" masterPageNumber="1" pageNumber="58" updateTime="1716321918122" updateUser="carolina">
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<mods:title id="27922D9F44582557F5F8262F4D71818A">Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java</mods:title>
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<mods:name id="7FCEB81580210116B82F70DD702A4EFE" type="personal">
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<mods:namePart id="CE0DBF25E0EACA0BC933A0810218C96D">Nurinsiyah, Ayu Savitri</mods:namePart>
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<mods:affiliation id="4DD94B29A31E1AD8B02252790942EB1C">Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia.</mods:affiliation>
<mods:nameIdentifier id="EAC97F31A8268252BCD83D84AFC34DF7" type="email">ayu_nurinsiyah@yahoo.com &amp; ayus002@lipi.go.id</mods:nameIdentifier>
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<mods:namePart id="FCBC264565D2A196B680E874BB960134">Neiber, Marco T.</mods:namePart>
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<mods:affiliation id="5B7BE342EB8F5E6E3F954DE0694984F0">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier id="BEFEE3BCDB0AD0384E0F72EF17B9A96D" type="email">mneiber@hotmail.de</mods:nameIdentifier>
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<mods:namePart id="B9B8C5243488879DF03DA0FF244AA697">Hausdorf, Bernhard</mods:namePart>
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<mods:affiliation id="457E8EE159AE648C9C2FDE72E84CEE59">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.</mods:affiliation>
<mods:nameIdentifier id="A6A2E8EF7A4711570E2FAFE607A43DAA" type="email">hausdorf@zoologie.uni-hamburg.de</mods:nameIdentifier>
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<mods:title id="D51C08DBA36C394C5A37819F098EAE4A">European Journal of Taxonomy</mods:title>
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<mods:date id="1E6A63606627DA89364E09035B7978EE">2019</mods:date>
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<mods:number id="B3D1A8C53600FE4D5ED5A933A1BC1AC8">2019-05-21</mods:number>
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<mods:number id="3859E0D4A004A4051655293F22121CBF">526</mods:number>
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<mods:start id="6723D2AF3D309BFAC2F80DA9B86F9E84">1</mods:start>
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<treatment id="03B1DB4C8B144E224C34FBAE3853FDFE" ID-GBIF-Taxon="157239591" LSID="urn:lsid:plazi:treatment:03B1DB4C8B144E224C34FBAE3853FDFE" httpUri="http://treatment.plazi.org/id/03B1DB4C8B144E224C34FBAE3853FDFE" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58">
<subSubSection id="C30239D18B144E204C34FBAE383EFBB6" box="[513,1075,1106,1134]" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="8BA76A5A8B144E204C34FBAE383EFBB6" blockId="57.[513,1075,1106,1168]" box="[513,1075,1106,1134]" pageId="57" pageNumber="58">
<heading id="D0EFDD368B144E204C34FBAE383EFBB6" box="[513,1075,1106,1134]" centered="true" fontSize="11" level="2" pageId="57" pageNumber="58" reason="2">
<taxonomicName id="4C1811D98B144E204C34FBAE3FA7FBB5" ID-CoL="7RTHL" authority="(Gude, 1905)" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" baseAuthorityName="Gude" baseAuthorityYear="1905" box="[513,938,1106,1134]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="leucochila" status="comb. nov.">
<emphasis id="B96CB6488B144E204C34FBAE3F0BFBB6" bold="true" box="[513,774,1106,1133]" italics="true" pageId="57" pageNumber="58">Landouria leucochila</emphasis>
(
<bibRefCitation id="EF8917AB8B144E204D23FBAF3FAFFBB5" author="Gude" box="[790,930,1107,1134]" firstAuthor="Gude" pageId="57" pageNumber="58" pagination="11 - 14" refId="ref33098" refString="Gude G. K. 1905. Description of nine new species of helicoid land shells. Journal of Malacology 12: 11 - 14, pls 3 - 4." type="journal article" year="1905">Gude, 1905</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B144E204D84FBAF383EFBB6" box="[945,1075,1107,1133]" pageId="57" pageNumber="58" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B144E204C9CFB8A3F87FB4B" box="[681,906,1142,1168]" pageId="57" pageNumber="58" type="description">
<paragraph id="8BA76A5A8B144E204C9CFB8A3F87FB4B" blockId="57.[513,1075,1106,1168]" box="[681,906,1142,1168]" pageId="57" pageNumber="58">
<figureCitation id="132376DF8B144E204C9CFB8A3F09FB4B" box="[681,772,1142,1168]" captionStart="Figs 81–84" captionStartId="57.[189,241,1859,1885]" captionTargetBox="[243,1343,265,2019]" captionTargetId="figure@58.[243,1343,265,2019]" captionTargetPageId="58" captionText="Figs 81–84 (opposite page). Shells of species of Landouria from Java. 81. L. leucochila (Gude, 1905), Java (holotype, NHM 1922.8.29.87). 82. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 83. L. ciliocincta (Möllendorff, 1897), East Java, Zuider Mountains (lectotype, SMF 9176a). 84. L. ciliocincta, East Java, Malang, Kondangmerak, protected forest (MZB 19211). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152300/files/figure.png" pageId="57" pageNumber="58">Fig. 81;</figureCitation>
<tableCitation id="C69A5FE18B144E204D3EFB8A3F87FB4B" box="[779,906,1142,1168]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="57" pageNumber="58" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B144E204E88FB443880FAC8" pageId="57" pageNumber="58" type="reference_group">
<paragraph id="8BA76A5A8B144E204E88FB443FB8FB09" blockId="57.[189,949,1207,1234]" box="[189,949,1207,1234]" pageId="57" pageNumber="58">
<treatmentCitationGroup id="AB084D748B144E204E88FB443FB8FB09" box="[189,949,1207,1234]" pageId="57" pageNumber="58">
<taxonomicName id="4C1811D98B144E204E88FB443E81FB09" ID-CoL="7SRFX" authority="Gude, 1905: 14" authorityName="Gude" authorityPageNumber="14" authorityYear="1905" box="[189,652,1207,1234]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="leucochila">
<emphasis id="B96CB6488B144E204E88FB443DC0FB09" box="[189,461,1208,1234]" italics="true" pageId="57" pageNumber="58">Plectotropis leucochila</emphasis>
<treatmentCitation id="0AB94C4B8B144E204FE1FB4B3E81FB09" author="Gude G. K." box="[468,652,1207,1234]" page="14" pageId="57" pageNumber="58" year="1905">
<bibRefCitation id="EF8917AB8B144E204FE1FB4B3E81FB09" author="Gude G. K." box="[468,652,1207,1234]" firstAuthor="Gude" pageId="57" pageNumber="58" pagination="11 - 14" refId="ref33098" refString="Gude G. K. 1905. Description of nine new species of helicoid land shells. Journal of Malacology 12: 11 - 14, pls 3 - 4." type="journal article" year="1905">Gude, 1905: 14</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, pl. 3, figs 1a–c (“
<collectingRegion id="49DCA4B88B144E204D55FB443F91FB09" box="[864,924,1208,1234]" country="Indonesia" name="Jawa Timur" pageId="57" pageNumber="58">Java</collectingRegion>
”).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA76A5A8B144E204E88FB053880FAC8" blockId="57.[189,1165,1273,1300]" box="[189,1165,1273,1300]" pageId="57" pageNumber="58">
<treatmentCitationGroup id="AB084D748B144E204E88FB053880FAC8" box="[189,1165,1273,1300]" pageId="57" pageNumber="58">
<taxonomicName id="4C1811D98B144E204E88FB053F3AFAC8" ID-CoL="3S6S5" authority="- van Benthem Jutting 1950: 461" authorityName="- van Benthem Jutting" authorityPageNumber="461" authorityYear="1950" box="[189,823,1273,1300]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="rotatoria">
<emphasis id="B96CB6488B144E204E88FB053DA4FAC8" box="[189,425,1273,1299]" italics="true" pageId="57" pageNumber="58">Landouria rotatoria</emphasis>
–
<treatmentCitation id="0AB94C4B8B144E204FF3FB063F3AFAC8" author="van Benthem Jutting W. S. S." box="[454,823,1273,1300]" page="461" pageId="57" pageNumber="58" year="1950">
<bibRefCitation id="EF8917AB8B144E204FF3FB063F3AFAC8" author="van Benthem Jutting W. S. S." box="[454,823,1273,1300]" firstAuthor="van Benthem Jutting" pageId="57" pageNumber="58" pagination="381 - 505" refId="ref35442" refString="van Benthem Jutting W. S. S. 1950. Systematic studies on the non-marine Mollusca of the Indo-Australian archipelago. II. Critical revision of the Javanese pulmonate land-shells of the families Helicarionidae, Pleurodontidae, Fruticicolidae and Streptaxidae. Treubia 20: 381 - 505." type="journal article" year="1950">van Benthem Jutting 1950: 461</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(in part). (not
<bibRefCitation id="EF8917AB8B144E204DD6FB053888FAC8" author="Pfeiffer" box="[995,1157,1273,1299]" firstAuthor="Pfeiffer" pageId="57" pageNumber="58" refId="ref34385" refString="Pfeiffer L. 1842. Symbolae ad Historiam Heliceorum. Sectio alterna. T. Fischer, Cassel [Kassel, Germany]. https: // doi. org / 10.5962 / bhl. title. 11903" type="book" year="1842">Pfeiffer, 1842</bibRefCitation>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B144E204E88FAC63FBEFA7D" pageId="57" pageNumber="58" type="diagnosis">
<paragraph id="8BA76A5A8B144E204E88FAC63D3BFA8F" blockId="57.[189,310,1338,1364]" box="[189,310,1338,1364]" pageId="57" pageNumber="58">
<heading id="D0EFDD368B144E204E88FAC63D3BFA8F" bold="true" box="[189,310,1338,1364]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="B96CB6488B144E204E88FAC63D3BFA8F" bold="true" box="[189,310,1338,1364]" pageId="57" pageNumber="58">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B144E204E88FA953FBEFA7D" blockId="57.[189,1398,1385,1447]" pageId="57" pageNumber="58">
<taxonomicName id="4C1811D98B144E204E88FA953DB0FA58" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" baseAuthorityName="Gude" baseAuthorityYear="1905" box="[189,445,1385,1411]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="leucochila">
<emphasis id="B96CB6488B144E204E88FA953DB0FA58" box="[189,445,1385,1411]" italics="true" pageId="57" pageNumber="58">Landouria leucochila</emphasis>
</taxonomicName>
is characterized by a large, depressed conical, yellowish shell with an angular body whorl, a wide umbilicus and scales all over the teleoconch.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B144E204E88FA313F16F9E2" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="8BA76A5A8B144E204E88FA313DA7FA3C" blockId="57.[189,426,1485,1511]" box="[189,426,1485,1511]" pageId="57" pageNumber="58">
<heading id="D0EFDD368B144E204E88FA313DA7FA3C" bold="true" box="[189,426,1485,1511]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="B96CB6488B144E204E88FA313DA7FA3C" bold="true" box="[189,426,1485,1511]" pageId="57" pageNumber="58">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B144E204EDFFA003D51F9CD" blockId="57.[189,795,1532,1594]" box="[234,348,1532,1558]" pageId="57" pageNumber="58">
<typeStatus id="54A3D4F88B144E204EDFFA003D51F9CD" box="[234,348,1532,1558]" pageId="57" pageNumber="58" type="holotype">
<emphasis id="B96CB6488B144E204EDFFA003D51F9CD" bold="true" box="[234,348,1532,1558]" pageId="57" pageNumber="58">Holotype</emphasis>
</typeStatus>
</paragraph>
<paragraph id="8BA76A5A8B144E204E88F9E33F16F9E2" blockId="57.[189,795,1532,1594]" box="[189,795,1567,1594]" pageId="57" pageNumber="58">
<materialsCitation id="3B7060078B144E204E88F9E33F16F9E2" ID-GBIF-Occurrence="2243137506" box="[189,795,1567,1594]" collectionCode="NHM" country="Indonesia" location="Java" pageId="57" pageNumber="58" specimenCode="NHM 1922.8.29.87" specimenCount="1" stateProvince="Java" typeStatus="holotype">
<collectingCountry id="F30F2ACA8B144E204E88F9E33D56F9E2" box="[189,347,1567,1593]" name="Indonesia" pageId="57" pageNumber="58">INDONESIA</collectingCountry>
•
<typeStatus id="54A3D4F88B144E204F41F9E33DD3F9E2" box="[372,478,1567,1593]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
;
<collectingRegion id="49DCA4B88B144E204FDFF9DC3E2FF9E1" box="[490,546,1568,1594]" country="Indonesia" name="Jawa Timur" pageId="57" pageNumber="58">Java</collectingRegion>
;
<specimenCode id="DBBEC2218B144E204C1BF9DC3F1AF9E2" box="[558,791,1567,1594]" pageId="57" pageNumber="58">NHM 1922.8.29.87</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B144E224E88F99D3DB6FE76" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58" type="description">
<paragraph id="8BA76A5A8B144E204E88F99D3D43F9A0" blockId="57.[189,334,1633,1659]" box="[189,334,1633,1659]" pageId="57" pageNumber="58">
<heading id="D0EFDD368B144E204E88F99D3D43F9A0" bold="true" box="[189,334,1633,1659]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="B96CB6488B144E204E88F99D3D43F9A0" bold="true" box="[189,334,1633,1659]" pageId="57" pageNumber="58">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B144E224E88F9733ED0FEB7" blockId="57.[189,1399,1679,1776]" lastBlockId="59.[189,1399,267,364]" lastPageId="59" lastPageNumber="60" pageId="57" pageNumber="58">
SHELL (
<figureCitation id="132376DF8B144E204F20F9733D62F972" box="[277,367,1679,1705]" captionStart="Figs 81–84" captionStartId="57.[189,241,1859,1885]" captionTargetBox="[243,1343,265,2019]" captionTargetId="figure@58.[243,1343,265,2019]" captionTargetPageId="58" captionText="Figs 81–84 (opposite page). Shells of species of Landouria from Java. 81. L. leucochila (Gude, 1905), Java (holotype, NHM 1922.8.29.87). 82. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 83. L. ciliocincta (Möllendorff, 1897), East Java, Zuider Mountains (lectotype, SMF 9176a). 84. L. ciliocincta, East Java, Malang, Kondangmerak, protected forest (MZB 19211). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152300/files/figure.png" pageId="57" pageNumber="58">Fig. 81</figureCitation>
;
<tableCitation id="C69A5FE18B144E204F4BF9733E09F972" box="[382,516,1679,1705]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="57" pageNumber="58" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 3–4</tableCitation>
). Depressed conical, with 5.25 hardly convex whorls; protoconch almost smooth; teleoconch with irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes all over teleoconch; tuberculate around umbilicus; yellowish; body whorl angular; aperture oval; upper insertion of peristome descending; peristome expanded, reflexed and distinctly thickened; umbilicus eccentric, wide, comprising 30% of shell diameter, hardly obscured by columellar edge.
</paragraph>
<caption id="DF673AD28B144E204E88F8BF3D89F831" httpUri="https://zenodo.org/record/3152300/files/figure.png" pageId="57" pageNumber="58" targetBox="[243,1343,265,2019]" targetPageId="58">
<paragraph id="8BA76A5A8B144E204E88F8BF3D89F831" blockId="57.[189,1399,1859,2027]" pageId="57" pageNumber="58">
<emphasis id="B96CB6488B144E204E88F8BF3D4FF886" bold="true" box="[189,322,1859,1885]" pageId="57" pageNumber="58">Figs 81–84</emphasis>
(opposite page). Shells of species of
<taxonomicName id="4C1811D98B144E204CC9F8BF3F75F886" authorityName="Godwin-Austen" authorityYear="1918" box="[764,888,1859,1885]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="58" phylum="Mollusca" rank="genus">
<emphasis id="B96CB6488B144E204CC9F8BF3F75F886" box="[764,888,1859,1885]" italics="true" pageId="57" pageNumber="58">Landouria</emphasis>
</taxonomicName>
from
<collectingRegion id="49DCA4B88B144E204DF4F8B83FF4F885" box="[961,1017,1860,1886]" country="Indonesia" name="Jawa Timur" pageId="57" pageNumber="58">Java</collectingRegion>
.
<emphasis id="B96CB6488B144E204A31F8BF382CF886" bold="true" box="[1028,1057,1859,1885]" pageId="57" pageNumber="58">81</emphasis>
.
<taxonomicName id="4C1811D98B144E204A04F8B8397FF885" authority="(Gude, 1905)" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" baseAuthorityName="Gude" baseAuthorityYear="1905" box="[1073,1394,1859,1886]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="leucochila">
<emphasis id="B96CB6488B144E204A04F8B838C7F886" box="[1073,1226,1859,1885]" italics="true" pageId="57" pageNumber="58">L. leucochila</emphasis>
(
<bibRefCitation id="EF8917AB8B144E204AEFF8BF3964F885" author="Gude" box="[1242,1385,1859,1886]" firstAuthor="Gude" pageId="57" pageNumber="58" pagination="11 - 14" refId="ref33098" refString="Gude G. K. 1905. Description of nine new species of helicoid land shells. Journal of Malacology 12: 11 - 14, pls 3 - 4." type="journal article" year="1905">Gude, 1905</bibRefCitation>
)
</taxonomicName>
,
<collectingRegion id="49DCA4B88B144E204E88F89B3CFCF85A" box="[189,241,1895,1921]" country="Indonesia" name="Jawa Timur" pageId="57" pageNumber="58">Java</collectingRegion>
(
<typeStatus id="54A3D4F88B144E204F32F89A3D7FF85B" box="[263,370,1894,1920]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
, NHM 1922.8.29.87).
<emphasis id="B96CB6488B144E204CA4F89A3EA3F85B" bold="true" box="[657,686,1894,1920]" pageId="57" pageNumber="58">82</emphasis>
.
<taxonomicName id="4C1811D98B144E204CF1F89B3F47F85A" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[708,842,1895,1921]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="zonifera" status="sp. nov.">
<emphasis id="B96CB6488B144E204CF1F89B3F47F85A" box="[708,842,1895,1921]" italics="true" pageId="57" pageNumber="58">L. zonifera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B144E204D6DF89B3FB1F85A" box="[856,956,1895,1921]" pageId="57" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B144E204DFBF89A385AF85B" box="[974,1111,1894,1920]" country="Indonesia" name="Yogyakarta" pageId="57" pageNumber="58">Yogyakarta</collectingRegion>
, Kulonprogo, Sibolong (
<typeStatus id="54A3D4F88B144E204EF0F8753D3DF878" box="[197,304,1929,1955]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
, MZB 17727).
<emphasis id="B96CB6488B144E204FD7F8753DF2F878" bold="true" box="[482,511,1929,1955]" pageId="57" pageNumber="58">83</emphasis>
.
<taxonomicName id="4C1811D98B144E204C38F8763F96F87F" authority="(Mollendorff, 1897)" authorityName="Mollendorff" authorityYear="1897" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[525,923,1929,1956]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="ciliocincta">
<emphasis id="B96CB6488B144E204C38F8763EAAF87F" box="[525,679,1930,1956]" italics="true" pageId="57" pageNumber="58">L. ciliocincta</emphasis>
(
<bibRefCitation id="EF8917AB8B144E204C80F8753F9FF87F" author="Mollendorff" box="[693,914,1929,1956]" firstAuthor="Mollendorff" pageId="57" pageNumber="58" pagination="89 - 97" refId="ref33803" refString="Mollendorff O. F. von. 1897. Neue Landschnecken von Java. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 29: 57 - 72, 89 - 97." type="journal article" year="1897">Möllendorff, 1897</bibRefCitation>
)
</taxonomicName>
,
<collectingRegion id="49DCA4B88B144E204D93F876381BF87F" box="[934,1046,1930,1956]" country="Indonesia" name="Jawa Timur" pageId="57" pageNumber="58">East Java</collectingRegion>
, Zuider Mountains (
<typeStatus id="54A3D4F88B144E204B37F875397FF878" box="[1282,1394,1929,1955]" pageId="57" pageNumber="58" type="lectotype">lectotype</typeStatus>
, SMF 9176a).
<emphasis id="B96CB6488B144E204F52F8513D89F81C" bold="true" box="[359,388,1965,1991]" pageId="57" pageNumber="58">84</emphasis>
.
<taxonomicName id="4C1811D98B144E204FA2F8523E3BF81C" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[407,566,1965,1991]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="ciliocincta">
<emphasis id="B96CB6488B144E204FA2F8523E3BF81C" box="[407,566,1965,1991]" italics="true" pageId="57" pageNumber="58">L. ciliocincta</emphasis>
</taxonomicName>
,
<collectingRegion id="49DCA4B88B144E204C7CF8513EB3F81C" box="[585,702,1965,1991]" country="Indonesia" name="Jawa Timur" pageId="57" pageNumber="58">East Java</collectingRegion>
, Malang, Kondangmerak, protected forest (MZB 19211). Scale bar:
<quantity id="4CE0C7BF8B144E204F0CF82C3D8DF831" box="[313,384,2000,2027]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="57" pageNumber="58" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
</caption>
<paragraph id="8BA76A5A8B164E224E88FE6F3DB6FE76" blockId="59.[189,443,403,430]" box="[189,443,403,430]" pageId="59" pageNumber="60">GENITALIA. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30239D18B164E224E88FE203853FDFE" box="[189,1118,476,549]" pageId="59" pageNumber="60" type="distribution">
<paragraph id="8BA76A5A8B164E224E88FE203D5BFE2D" blockId="59.[189,342,476,502]" box="[189,342,476,502]" pageId="59" pageNumber="60">
<heading id="D0EFDD368B164E224E88FE203D5BFE2D" bold="true" box="[189,342,476,502]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
<emphasis id="B96CB6488B164E224E88FE203D5BFE2D" bold="true" box="[189,342,476,502]" pageId="59" pageNumber="60">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B164E224E88FDF73853FDFE" blockId="59.[189,1118,523,549]" box="[189,1118,523,549]" pageId="59" pageNumber="60">
The type locality and the distribution of
<taxonomicName id="4C1811D98B164E224CA3FDF03F22FDFE" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" baseAuthorityName="Gude" baseAuthorityYear="1905" box="[662,815,523,549]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="60" phylum="Mollusca" rank="species" species="leucochila">
<emphasis id="B96CB6488B164E224CA3FDF03F22FDFE" box="[662,815,523,549]" italics="true" pageId="59" pageNumber="60">L. leucochila</emphasis>
</taxonomicName>
in
<collectingRegion id="49DCA4B88B164E224D60FDF73F84FDFE" box="[853,905,523,549]" country="Indonesia" name="Jawa Timur" pageId="59" pageNumber="60">Java</collectingRegion>
remain unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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