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<document id="B3A4B62D99AC6932D185AEB577AFB8E2" ID-CLB-Dataset="298186" ID-DOI="10.3897/zookeys.1204.114932" ID-GBIF-Dataset="33ca18a1-99c6-4b56-9a8e-92fe6f6f9209" ID-ZooBank="F5134CC6-A83C-48B6-A1EA-235E6858D5CB" ID-publisher-id="114932" URI-arpha="EAF89A3A-8354-506D-AF9B-9E1C53605541" URI-zoobank="http://zoobank.org/F5134CC6-A83C-48B6-A1EA-235E6858D5CB" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1717680124246" checkinUser="pensoft" docAuthor="Nascimento, Elynton Alves &amp; Bocakova, Milada" docDate="2024" docId="C6E98B65EB055292A9814D46CFC92555" docLanguage="en" docName="ZooKeys 1204: 241-259" docOrigin="ZooKeys 1204" docSource="https://zookeys.pensoft.net/article/114932/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Gorhamium unidentatum Nascimento &amp; Bocakova 2024, sp. nov." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="259" masterDocId="EAF89A3A8354506DAF9B9E1C53605541" masterDocTitle="Phylogenetic analysis reveals a new net-winged beetle genus of Eurrhacini (Coleoptera, Lycidae) from the Pacific slopes of Central America and Ecuador" masterLastPageNumber="259" masterPageNumber="241" pageNumber="241" updateTime="1717709099581" updateUser="ExternalLinkService">
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<mods:title id="F8462ED3FEC41DF412277C21F131E6AF">Phylogenetic analysis reveals a new net-winged beetle genus of Eurrhacini (Coleoptera, Lycidae) from the Pacific slopes of Central America and Ecuador</mods:title>
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<mods:namePart id="2995F985E215B33FC1994357576F857D">Nascimento, Elynton Alves</mods:namePart>
<mods:nameIdentifier id="421C54CFCF7E5E2B8D88637C46F720BF" type="ORCID">0000-0002-9071-2823</mods:nameIdentifier>
<mods:affiliation id="F89F9ED445AB33CA8C8C5388E2CB83C9">Departamento de Engenharia Ambiental, Universidade Estadual do Centro-Oeste, Rua Professora Maria Roza Zanon de Almeida, s / n, Engenheiro Gutierrez, Irati PR, CEP 84505 - 677, Brazil</mods:affiliation>
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<mods:namePart id="C42BC8CB50D940CCD15E815FEDA66922">Bocakova, Milada</mods:namePart>
<mods:nameIdentifier id="E79696C0B793A9AF115A8F0E6DD0686B" type="ORCID">0000-0002-2507-0887</mods:nameIdentifier>
<mods:affiliation id="95D3B907BFD1D67962294BF9AA02F694">Department of Biology, Faculty of Education, Palacky University, Purkrabska 2, CZ- 77140 Olomouc, Czech Republic</mods:affiliation>
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<mods:typeOfResource id="FD0CEA33437610DC814A5C87EF1E913A">text</mods:typeOfResource>
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<mods:date id="99C9FDCCACDCDDA8022D45C4B3E922C5">2024</mods:date>
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<mods:number id="EC625469FEC01DC07672D88F73C23D76">2024-06-06</mods:number>
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<treatment id="C6E98B65EB055292A9814D46CFC92555" ID-DOI="http://doi.org/10.5281/zenodo.11505171" ID-GBIF-Taxon="232157812" ID-Zenodo-Dep="11505171" ID-arpha="C6E98B65-EB05-5292-A981-4D46CFC92555" ID-zoobank="https://zoobank.org/8CEF51B9-4C36-4654-BEB4-B3BFA4F63DC3" LSID="urn:lsid:plazi:treatment:C6E98B65EB055292A9814D46CFC92555" LSID-ZBK="urn:lsid:zoobank.org:act:8CEF51B9-4C36-4654-BEB4-B3BFA4F63DC3" httpUri="http://treatment.plazi.org/id/C6E98B65EB055292A9814D46CFC92555">
<subSubSection id="DACA6B5934B5A1F7AF6C3A35D0321310" type="nomenclature">
<paragraph id="C84EF25B2A04984EF54B14F16D1B013C">
<taxonomicName id="3AB1756E7762E2A38571949B20555F04" authority="Nascimento &amp; Bocakova, 2024" authorityName="Nascimento &amp; Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="unidentatum" status="sp. nov.">
<emphasis id="2F66D3709ED689B616EFD6BF72B3386E" italics="true">Gorhamium unidentatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ED2BE49C885774FE21CE5C2F8CCE4723" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="30B989BA65C5B74D54CA58B20CABB7B6" type="description">
<paragraph id="4716E1D608638477884FD2F33F0A5130">
<figureCitation id="2BF18289527BC2625C0912813067641D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal view A, B Gorhamium bidentatum sp. nov. A male B female C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure2" httpUri="https://binary.pensoft.net/fig/1063776">Figs 2 C</figureCitation>
,
<figureCitation id="A84EED845B42427D6894507C739EF61E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pronotum A Gorhamium bidentatum sp. nov., male B Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure3" httpUri="https://binary.pensoft.net/fig/1063777">3 B</figureCitation>
,
<figureCitation id="D41F9DB00C355974ABEDB6C6C13EDDBF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A head ventrally B, C antenna dorsally A, B Gorhamium bidentatum sp. nov., male C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure4" httpUri="https://binary.pensoft.net/fig/1063778">4 C</figureCitation>
,
<figureCitation id="F97A5C59AEFC8EE041D71C46F91E0EFD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Terminal abdominal segments A D Gorhamium bidentatum sp. nov. E H Gorhamium unidentatum sp. nov., male G Cladocalon chiriquense (Gorham, 1884) A, B, E, G male terminalia (sternum IX and tergum IX X), A ventral view; B, E, G dorsal view C Male sternum and tergum VIII, ventral view D female terminal sternum, ventral view. F, Male sternum VIII, ventral view H Male tergum VIII, dorsal view." figureDoi="10.3897/zookeys.1204.114932.figure5" httpUri="https://binary.pensoft.net/fig/1063779">5 E H</figureCitation>
,
<figureCitation id="2C52F5A914D1B60CAD8B8B7F48BFE1C5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C E Gorhamium bidentatum sp. nov. F H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 pointed anchor-shaped base of phallus, a 2 flat anchor-shaped base of phallus, b phallic ventral opening, c dorsal dent, d 1 arcuate ventrobasal parameral process, d 2 flattened ventrobasal parameral process, d 3 a ring-like ventral bridge (ventrobasal processes medially fused), e internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 F, G</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1CBG" type="type material">
<paragraph id="3F3717C379D551A94ACE6655AEBE3D89">
<heading id="24CD34E1CA1A9875F5F9F7E518CD67E4" reason="title">Type material.</heading>
</paragraph>
<paragraph id="7CD4DFA5A58936203D458F1CF6D31A54">
<materialsCitation id="B2134B7DCE5ABA38CE5C633C2A58A820" ID-GBIF-Occurrence="4873767306" collectingDate="2010-12-05" collectingDateMax="2010-12-06" collectingDateMin="2010-12-05" collectionCode="UPOL" collectorName="Bolm" country="Ecuador" county="Otonga" elevation="1500" latitude="-0.41666666" location="San Francisco de las Pampas" longLatPrecision="1308" longitude="-79.0" municipality="Otonga res" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="3AC80DC2B8CCF31A44DCFC7209FDC10D" bold="true" italics="true">
<typeStatus id="965FC858F76CF69A829CE67D598DD3B0" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="56FA2EBC413EB0EB09D25DB3E6FE709D" count="1" type="male">male</specimenCount>
, “
<collectingCountry id="2AB8B3AC0A3C4CFFE933FED1FAEABD89" name="Ecuador">Ecuador</collectingCountry>
,
<locationDeviation id="D842F8B01E231C7023C9143E9D20DB97" location="Quito">
<quantity id="3BD78060BDFFDABF85C45CD56933F1BD" metricMagnitude="4" metricUnit="m" metricValue="5.0" unit="km" value="50.0">50 km</quantity>
SW Quito
</locationDeviation>
,
<location id="2A3743C01AF15E0104B065CCA7B9F0EB" LSID="urn:lsid:plazi:treatment:C6E98B65EB055292A9814D46CFC92555:2A3743C01AF15E0104B065CCA7B9F0EB" country="Ecuador" county="Otonga" latitude="-0.41666666" longLatPrecision="1308" longitude="-79.0" municipality="Otonga res" name="San Francisco de las Pampas">San Francisco de las Pampas</location>
,
<collectingMunicipality id="0531C39BF03EA3653A23EE3ACD0D26CE">Otonga res</collectingMunicipality>
.,
<quantity id="A1CC01BDCB9563C43E9572970D3B1FC1" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">
<elevation id="5D5F2F59B56EC0CB59FFA2DEEA7DF791" metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIDBG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-79.000000,-0.416667]}">
<geoCoordinate id="DACE8DC080B6A0DD4A5CE69A38C73572" degrees="0" direction="south" minutes="25" orientation="latitude" precision="925" value="-0.41666666">0 ° 25 ' S</geoCoordinate>
,
<geoCoordinate id="44B1F541AEC485F8DF737A1337653FEC" degrees="79" direction="west" minutes="00" orientation="longitude" precision="925" value="-79.0">79 ° 00 ' W</geoCoordinate>
</named-content>
,
<date id="9C13CD128E8316C5105E58E3E0B8F659" value="2010-12-05" valueMax="2010-12-06" valueMin="2010-12-05">
<collectingDate id="3DD84CE59B270366DF28F9749A8E1DD8" value="2010-12-05" valueMax="2010-12-06" valueMin="2010-12-05">56. Dec 2010</collectingDate>
</date>
,
<collectorName id="275F0F745447C710873FF5DFE77BA678">Bolm</collectorName>
lgt. “ (
<named-content id="E9542D025BD4278B9B4B9F39ED2D0E1E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/university-palacky-olomouc" xlink_title="Palacky University Olomouc">
<collectionCode id="65AAEFD64D5C9686CFC50FE5085FD073" country="Czech Republic" httpUri="http://grbio.org/cool/zyr7-qg0p" name="University Palacky Olomouc" type="">UPOL</collectionCode>
</named-content>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESDBG" type="diagnosis">
<paragraph id="D0AD02788B10F6F09544602CFC4C961C">
<heading id="7860D7F6550B5F9C730AD29BF631687F" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="A05280F284FB339056073B2540E0E3F5">
Pronotum black. Elytra bicolor orange-black with suture, longitudinal median oval spot, and triangular apical spot black. Phallus ball-shaped apically, ventromedial opening widest in basal third. Apex of each paramere fitted with a sharp laterally projected tooth, internal sac with a series of diminutive teeth (e, Fig.
<figureCitation id="109ACEE166B92D6A2BC203B6A3F89CBF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C E Gorhamium bidentatum sp. nov. F H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 pointed anchor-shaped base of phallus, a 2 flat anchor-shaped base of phallus, b phallic ventral opening, c dorsal dent, d 1 arcuate ventrobasal parameral process, d 2 flattened ventrobasal parameral process, d 3 a ring-like ventral bridge (ventrobasal processes medially fused), e internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3DBG" type="description">
<paragraph id="58928340AE9D2DF9BC19961F5A2F6B2F">
<heading id="E6134EDBB371C009581F73B741EE67B2" reason="title">Description.</heading>
</paragraph>
<paragraph id="9066DBC0253D61E22DE99E4231CFDA78">
Body length:
<quantity id="5B49143AB16BBFEFFF77D88CA66AF743" metricMagnitude="-3" metricUnit="m" metricValue="6.4" unit="mm" value="6.4">6.4 mm</quantity>
, width across the humeri:
<quantity id="92656686F7E04C03ECDC0780B1D5A009" metricMagnitude="-3" metricUnit="m" metricValue="1.2" unit="mm" value="1.2">1.2 mm</quantity>
. Body black, only elytral sidebars orange (with whole suture, longitudinal median oval spot and triangular spot in apical quarter black, remaining sidebars orange (Fig.
<figureCitation id="0F130E5905A90870611B0FB87BCD46F5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal view A, B Gorhamium bidentatum sp. nov. A male B female C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure2" httpUri="https://binary.pensoft.net/fig/1063776">2 C</figureCitation>
). Head mostly hidden by pronotum in dorsal view. Elytra slender, 4.5 × longer than humeral width (Fig.
<figureCitation id="7126472E98A78F02DAA346DD44565328" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal view A, B Gorhamium bidentatum sp. nov. A male B female C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure2" httpUri="https://binary.pensoft.net/fig/1063776">2 C</figureCitation>
); primary costae 2 and 4 and basal 1 / 5 of primary costa 3 elevated; reticulate cells oval, strongly irregular, secondary costae 3 and 4 diminishing apically.
</paragraph>
<paragraph id="62F5F9EB5FB9979B87476887D0C8516C">
<emphasis id="0EDCE62168DE3D399732F08714094A73" bold="true">Male.</emphasis>
Eyes medium-sized, interocular distance 1.3 × longer than eye diameter (Fig.
<figureCitation id="E5322454F2186F817BEAE3E1A36A1BAD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pronotum A Gorhamium bidentatum sp. nov., male B Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure3" httpUri="https://binary.pensoft.net/fig/1063777">3 B</figureCitation>
). Antennae with antennomere 3 (= A 3) 1.3 × shorter than A 4, A 4 A 10 subequal in length; antennal branches flattened, considerably lengthening medially, A 3 lamella 1.2 × longer than antennomere A 3 length, A 4 lamella 1.35 × longer than A 4 length, A 5 lamella 1.5 × longer than A 5 length (Fig.
<figureCitation id="6C7A3F2FA14479288C4F7A4C6057E485" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A head ventrally B, C antenna dorsally A, B Gorhamium bidentatum sp. nov., male C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure4" httpUri="https://binary.pensoft.net/fig/1063778">4 C</figureCitation>
). Abdominal sternum VIII widely emarginated in distal third (Fig.
<figureCitation id="6298879208E943350B98019658B8F098" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Terminal abdominal segments A D Gorhamium bidentatum sp. nov. E H Gorhamium unidentatum sp. nov., male G Cladocalon chiriquense (Gorham, 1884) A, B, E, G male terminalia (sternum IX and tergum IX X), A ventral view; B, E, G dorsal view C Male sternum and tergum VIII, ventral view D female terminal sternum, ventral view. F, Male sternum VIII, ventral view H Male tergum VIII, dorsal view." figureDoi="10.3897/zookeys.1204.114932.figure5" httpUri="https://binary.pensoft.net/fig/1063779">5 F</figureCitation>
), its proximal margin almost straight. Tergum X elongate, 1.7 × longer than sternum IX on the sides (Fig.
<figureCitation id="28320DD5621B6B409D89D62B5B6BF94C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Terminal abdominal segments A D Gorhamium bidentatum sp. nov. E H Gorhamium unidentatum sp. nov., male G Cladocalon chiriquense (Gorham, 1884) A, B, E, G male terminalia (sternum IX and tergum IX X), A ventral view; B, E, G dorsal view C Male sternum and tergum VIII, ventral view D female terminal sternum, ventral view. F, Male sternum VIII, ventral view H Male tergum VIII, dorsal view." figureDoi="10.3897/zookeys.1204.114932.figure5" httpUri="https://binary.pensoft.net/fig/1063779">5 E</figureCitation>
). Phallus bent ventrally in distal 1 / 3, constricted subapically, apex ball-shaped; ventromedial opening widest in basal quarter (b, Fig.
<figureCitation id="223ECB7EE4676E176E6C2912EB3FEA5C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C E Gorhamium bidentatum sp. nov. F H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 pointed anchor-shaped base of phallus, a 2 flat anchor-shaped base of phallus, b phallic ventral opening, c dorsal dent, d 1 arcuate ventrobasal parameral process, d 2 flattened ventrobasal parameral process, d 3 a ring-like ventral bridge (ventrobasal processes medially fused), e internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 G</figureCitation>
); dorsal hook shifted in distal quarter (c, Fig.
<figureCitation id="B430E165D56964537616BB024174B939" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C E Gorhamium bidentatum sp. nov. F H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 pointed anchor-shaped base of phallus, a 2 flat anchor-shaped base of phallus, b phallic ventral opening, c dorsal dent, d 1 arcuate ventrobasal parameral process, d 2 flattened ventrobasal parameral process, d 3 a ring-like ventral bridge (ventrobasal processes medially fused), e internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 G</figureCitation>
). Parameres as long as 2 / 3 of phallus, with a single, laterally projected, apical tooth; ventrobasal parameral protrusions slender, joined medially in a ring-like bridge (d
<subScript id="D176895C500074E3243D6A706A2419BE">3</subScript>
, Fig.
<figureCitation id="D5785D6740F0E16F4330E2B769C1064B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C E Gorhamium bidentatum sp. nov. F H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 pointed anchor-shaped base of phallus, a 2 flat anchor-shaped base of phallus, b phallic ventral opening, c dorsal dent, d 1 arcuate ventrobasal parameral process, d 2 flattened ventrobasal parameral process, d 3 a ring-like ventral bridge (ventrobasal processes medially fused), e internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 G</figureCitation>
).
</paragraph>
<paragraph id="AA8D8FCE5C179351CB8B4E3BE2A15994">
<emphasis id="646413948DFED35243180F403D4FC4BF" bold="true">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQFBG" type="etymology">
<paragraph id="805DDDD1DDF453D3BE552B0DAB9AEC55">
<heading id="6AA47C68BD29012BB2666A8C8A34F357" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="CD7585B123072F81D86CFEC8BCD712B4">The specific name refers to the single sharp tooth at the apex of each paramere.</paragraph>
</subSubSection>
<subSubSection id="SECID0EVFBG" type="distribution">
<paragraph id="4C04EFFDD022226F40DB02D3EA638488">
<heading id="1F62EDDA3A8B8F6984E67F6861249E0A" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="22196464E829925FCED351347C7E8F29">
<collectingCountry id="150612BE873F9FC75ADCDA0FEE761AA1" name="Ecuador">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>