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<mods:title>An integrative taxonomic revision and redefinition of Gephyromantis (Laurentomantis) malagasius based on archival DNA analysis reveals four new mantellid frog species from Madagascar</mods:title>
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<mods:namePart>Vences, Miguel</mods:namePart>
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<mods:namePart>Koehler, Joern</mods:namePart>
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<mods:namePart>Crottini, Angelica</mods:namePart>
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<mods:affiliation>CIBIO, Centro de Investigacao em Biodiversidade e Recursos Geneticos, InBIO Laboratorio Associado, Campus de Vairao, Universidade do Porto, 4485 - 661 Vairao, Portugal &amp; Departamento de Biologia, Faculdade de Ciencias, Universidade do Porto, 4099 - 002 Porto, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairao, 4485 - 661 Vairao, Portugal</mods:affiliation>
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<mods:namePart>Hofreiter, Michael</mods:namePart>
<mods:affiliation>Institut fuer Biochemie und Biologie, Universitaet Potsdam, Karl-Liebknecht-Str. 24 - 25, 14476 Potsdam, Germany</mods:affiliation>
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<mods:namePart>Hutter, Carl R.</mods:namePart>
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<mods:namePart>du Preez, Louis</mods:namePart>
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<mods:affiliation>Unit for Environmental Sciences and Development ,, North-West University, Potchefstroom campus, Private Bag X 6001, Potchefstroom 2520, South Africa &amp; South African Institute for Aquatic Biodiversity, Private Bag 1015, Makhanda, 6140, South Africa</mods:affiliation>
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<mods:namePart>Preick, Michaela</mods:namePart>
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<mods:namePart>Rakotoarison, Andolalao</mods:namePart>
<mods:affiliation>Mention Zoologie et Biodiversite Animale, Universite d'Antananarivo, BP 906, 101 Antananarivo, Madagascar</mods:affiliation>
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<mods:namePart>Rancilhac, Lois</mods:namePart>
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<mods:namePart>Raselimanana, Achille P.</mods:namePart>
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<mods:namePart>Rosa, Goncalo M.</mods:namePart>
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<mods:affiliation>Institute of Zoology, Zoological Society of London, London NW 1 4 RY, UK &amp; Centre for Ecology, Evolution and Environmental Changes (cE 3 c), Faculdade de Ciencias, Universidade de Lisboa, 1749 - 016 Lisboa, Portugal</mods:affiliation>
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<mods:namePart>Scherz, Mark D.</mods:namePart>
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<mods:namePart>Glaw, Frank</mods:namePart>
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<mods:affiliation>Zoologische Staatssammlung Muenchen (ZSM-SNSB), Muenchhausenstr. 21, 81247 Muenchen, Germany</mods:affiliation>
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<taxonomicName LSID="A7AA4313-5887-5A51-906A-280F924D96A4" authority="(Boettger, 1880)" baseAuthorityName="Boettger" baseAuthorityYear="1880" class="Amphibia" family="Mantellidae" genus="Gephyromantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gephyromantis horridus" order="Anura" pageId="0" pageNumber="271" phylum="Chordata" rank="species" species="horridus">Gephyromantis horridus (Boettger, 1880)</taxonomicName>
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<taxonomicName authorityName="Boettger" authorityYear="1880" class="Amphibia" family="Mantellidae" genus="Hemimantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemimantis horrida" order="Anura" pageId="0" pageNumber="271" phylum="Chordata" rank="species" species="horrida">Hemimantis horrida</taxonomicName>
Boettger, 1880: 282
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<paragraph pageId="0" pageNumber="271">Note.</paragraph>
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This species was described based on a juvenile specimen from Nosy Be, and a second specimen (an adult female) from the type locality has previously been reported by 
<bibRefCitation author="Glaw, F" journalOrPublisher="Spixiana" pageId="0" pageNumber="271" pagination="121 - 127" refId="B24" refString="Glaw, F, Vences, M, 2011. Description of a new frog species of Gephyromantis (subgenus Laurentomantis) with tibial glands from Madagascar (Amphibia, Mantellidae). Spixiana 34: 121 - 127" title="Description of a new frog species of Gephyromantis (subgenus Laurentomantis) with tibial glands from Madagascar (Amphibia, Mantellidae)." volume="34" year="2011">Glaw and Vences (2011)</bibRefCitation>
. Pictures of both specimens are shown in Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Preserved specimens of Gephyromantis horridus from the type locality Nosy Be. A, B Holotype SMF 7177 (juvenile) in dorsal and ventral views. C, D Specimen NMW 3643 (female) in dorsal and ventral views." figureDoi="10.3897/vz.72.e78830.figure4" httpUri="https://binary.pensoft.net/fig/688674" pageId="0" pageNumber="271">4</figureCitation>
; measurements are given in 
<bibRefCitation author="Vences, M" journalOrPublisher="Contributions to Zoology" pageId="0" pageNumber="271" pagination="191 - 212" refId="B58" refString="Vences, M, Glaw, F, Andreone, F, Jesu, R, Schimmenti, G, 2002. Systematic revision of the enigmatic Malagasy broad-headed frogs (Laurentomantis Dubois, 1980), and their phylogenetic position within the endemic mantellid radiation of Madagascar. Contributions to Zoology 70: 191 - 212" title="Systematic revision of the enigmatic Malagasy broad-headed frogs (Laurentomantis Dubois, 1980), and their phylogenetic position within the endemic mantellid radiation of Madagascar." volume="70" year="2002">Vences et al. (2002)</bibRefCitation>
and 
<bibRefCitation author="Glaw, F" journalOrPublisher="Spixiana" pageId="0" pageNumber="271" pagination="121 - 127" refId="B24" refString="Glaw, F, Vences, M, 2011. Description of a new frog species of Gephyromantis (subgenus Laurentomantis) with tibial glands from Madagascar (Amphibia, Mantellidae). Spixiana 34: 121 - 127" title="Description of a new frog species of Gephyromantis (subgenus Laurentomantis) with tibial glands from Madagascar (Amphibia, Mantellidae)." volume="34" year="2011">Glaw and Vences (2011)</bibRefCitation>
, respectively. The Tsaratanana population included in this species by 
<bibRefCitation author="Vences, M" journalOrPublisher="Contributions to Zoology" pageId="0" pageNumber="271" pagination="191 - 212" refId="B58" refString="Vences, M, Glaw, F, Andreone, F, Jesu, R, Schimmenti, G, 2002. Systematic revision of the enigmatic Malagasy broad-headed frogs (Laurentomantis Dubois, 1980), and their phylogenetic position within the endemic mantellid radiation of Madagascar. Contributions to Zoology 70: 191 - 212" title="Systematic revision of the enigmatic Malagasy broad-headed frogs (Laurentomantis Dubois, 1980), and their phylogenetic position within the endemic mantellid radiation of Madagascar." volume="70" year="2002">Vences et al. (2002)</bibRefCitation>
is assigned to 
<taxonomicName genus="G." lsidName="G. ranjomavo" pageId="0" pageNumber="271" rank="species" species="ranjomavo">
<emphasis italics="true" pageId="0" pageNumber="271">G. ranjomavo</emphasis>
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based on genetic data, and the only locality beside Nosy Be currently accepted for 
<taxonomicName genus="G." lsidName="G. horridus" pageId="0" pageNumber="271" rank="species" species="horridus">
<emphasis italics="true" pageId="0" pageNumber="271">G. horridus</emphasis>
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is therefore Montagne 
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. Measurements of the first male known from the Montagne 
<normalizedToken originalValue="d’Ambre">d'Ambre</normalizedToken>
population are given in Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric measurements (all in mm) of voucher specimens of Gephyromantis belonging to the subgenus Gephyromantis Laurentomantis newly examined for this study and identified on the basis of DNA sequence data. See Vences et al. (2002) and Glaw and Vences (2011) for measurements of additional individuals. For abbreviations of measurements, see Materials and Methods. HT, holotype." httpUri="http://table.plazi.org/id/F9637F48C90A59E74CFCAD26567B92BE" pageId="0" pageNumber="271" tableUuid="F9637F48C90A59E74CFCAD26567B92BE">1</tableCitation>
and photos in life of various individuals from this locality are included in Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Specimens assigned to Gephyromantis horridus from Montagne d'Ambre National Park in life. A, B Adult female photographed in March 1994 (ZFMK 57433) in dorsolateral and ventral views. C - E Adult male (ZSM 126 / 2018) in dorsolateral, dorsal and ventral views. F, G Adult female discovered under rotten wood in May 2014 (not collected) in dorsolateral and dorsal views. Note the absence of tibial glands in all specimens. Not to scale." figureDoi="10.3897/vz.72.e78830.figure5" httpUri="https://binary.pensoft.net/fig/688675" pageId="0" pageNumber="271">5</figureCitation>
. The male is characterized by very prominent femoral glands of the same color as the surrounding skin of the ventral thigh, composed of approximately six large gland granules as recognizable in external view (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Specimens assigned to Gephyromantis horridus from Montagne d'Ambre National Park in life. A, B Adult female photographed in March 1994 (ZFMK 57433) in dorsolateral and ventral views. C - E Adult male (ZSM 126 / 2018) in dorsolateral, dorsal and ventral views. F, G Adult female discovered under rotten wood in May 2014 (not collected) in dorsolateral and dorsal views. Note the absence of tibial glands in all specimens. Not to scale." figureDoi="10.3897/vz.72.e78830.figure5" httpUri="https://binary.pensoft.net/fig/688675" pageId="0" pageNumber="271">5</figureCitation>
). The male has no tibial gland. According to the measurements reported in 
<bibRefCitation author="Vences, M" journalOrPublisher="Contributions to Zoology" pageId="0" pageNumber="271" pagination="191 - 212" refId="B58" refString="Vences, M, Glaw, F, Andreone, F, Jesu, R, Schimmenti, G, 2002. Systematic revision of the enigmatic Malagasy broad-headed frogs (Laurentomantis Dubois, 1980), and their phylogenetic position within the endemic mantellid radiation of Madagascar. Contributions to Zoology 70: 191 - 212" title="Systematic revision of the enigmatic Malagasy broad-headed frogs (Laurentomantis Dubois, 1980), and their phylogenetic position within the endemic mantellid radiation of Madagascar." volume="70" year="2002">Vences et al. (2002)</bibRefCitation>
, 
<bibRefCitation author="Glaw, F" journalOrPublisher="Spixiana" pageId="0" pageNumber="271" pagination="121 - 127" refId="B24" refString="Glaw, F, Vences, M, 2011. Description of a new frog species of Gephyromantis (subgenus Laurentomantis) with tibial glands from Madagascar (Amphibia, Mantellidae). Spixiana 34: 121 - 127" title="Description of a new frog species of Gephyromantis (subgenus Laurentomantis) with tibial glands from Madagascar (Amphibia, Mantellidae)." volume="34" year="2011">Glaw and Vences (2011)</bibRefCitation>
and this study, SVL is 33.0-35.4 mm in two females and 33.5 in a male from Montagne 
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, and 33.7 mm in a female from Nosy Be. The advertisement call of this species was previously unknown, and is described in the following for the Montagne 
<normalizedToken originalValue="d’Ambre">d'Ambre</normalizedToken>
population.
</paragraph>
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<emphasis bold="true" pageId="0" pageNumber="271">Table 1.</emphasis>
Morphometric measurements (all in mm) of voucher specimens of 
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belonging to the subgenus 
<taxonomicName class="Amphibia" family="Mantellidae" genus="Gephyromantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gephyromantis (Laurentomantis)" order="Anura" pageId="0" pageNumber="271" phylum="Chordata" rank="subGenus" subGenus="Laurentomantis">Gephyromantis Laurentomantis</taxonomicName>
newly examined for this study and identified on the basis of DNA sequence data. See 
<bibRefCitation author="Vences, M" journalOrPublisher="Contributions to Zoology" pageId="0" pageNumber="271" pagination="191 - 212" refId="B58" refString="Vences, M, Glaw, F, Andreone, F, Jesu, R, Schimmenti, G, 2002. Systematic revision of the enigmatic Malagasy broad-headed frogs (Laurentomantis Dubois, 1980), and their phylogenetic position within the endemic mantellid radiation of Madagascar. Contributions to Zoology 70: 191 - 212" title="Systematic revision of the enigmatic Malagasy broad-headed frogs (Laurentomantis Dubois, 1980), and their phylogenetic position within the endemic mantellid radiation of Madagascar." volume="70" year="2002">Vences et al. (2002)</bibRefCitation>
and 
<bibRefCitation author="Glaw, F" journalOrPublisher="Spixiana" pageId="0" pageNumber="271" pagination="121 - 127" refId="B24" refString="Glaw, F, Vences, M, 2011. Description of a new frog species of Gephyromantis (subgenus Laurentomantis) with tibial glands from Madagascar (Amphibia, Mantellidae). Spixiana 34: 121 - 127" title="Description of a new frog species of Gephyromantis (subgenus Laurentomantis) with tibial glands from Madagascar (Amphibia, Mantellidae)." volume="34" year="2011">Glaw and Vences (2011)</bibRefCitation>
for measurements of additional individuals. For abbreviations of measurements, see Materials and Methods. HT, holotype.
</paragraph>
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<th colspan="1" pageId="0" pageNumber="271" rowspan="1">HL</th>
<th colspan="1" pageId="0" pageNumber="271" rowspan="1">ED</th>
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<th colspan="1" pageId="0" pageNumber="271" rowspan="1">NND</th>
<th colspan="1" pageId="0" pageNumber="271" rowspan="1">FORL</th>
<th colspan="1" pageId="0" pageNumber="271" rowspan="1">HAL</th>
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<th colspan="1" pageId="0" pageNumber="271" rowspan="1">FOL</th>
<th colspan="1" pageId="0" pageNumber="271" rowspan="1">TIBL</th>
<th colspan="1" pageId="0" pageNumber="271" rowspan="1">FGL</th>
<th colspan="1" pageId="0" pageNumber="271" rowspan="1">FGW</th>
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<taxonomicName genus="G." lsidName="G. horridus" pageId="0" pageNumber="271" rank="species" species="horridus">
<emphasis italics="true" pageId="0" pageNumber="271">G. horridus</emphasis>
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<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 126/2018</td>
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<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
Montagne 
<normalizedToken originalValue="d‘Ambre">d'Ambre</normalizedToken>
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">33.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">13.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">13.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">4.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">24.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">11.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">53.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">23.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">15.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">15.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">7.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">4.8</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. ranjomavo" pageId="0" pageNumber="271" rank="species" species="ranjomavo">
<emphasis italics="true" pageId="0" pageNumber="271">G. ranjomavo</emphasis>
</taxonomicName>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 60/2016</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">MSZC 0163</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Ampotsidy</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">26.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">10.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">10.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">4.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">19.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">41.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">18.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">11.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">12.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.6</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. malagasius" pageId="0" pageNumber="271" rank="species" species="malagasius">
<emphasis italics="true" pageId="0" pageNumber="271">G. malagasius</emphasis>
</taxonomicName>
(= 
<taxonomicName baseAuthorityName="Angel" baseAuthorityYear="1935" class="Amphibia" family="Mantellidae" genus="Gephyromantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gephyromantis ventrimaculatus" order="Anura" pageId="0" pageNumber="271" phylum="Chordata" rank="species" species="ventrimaculatus">
<emphasis italics="true" pageId="0" pageNumber="271">Gephyromantis ventrimaculatus</emphasis>
</taxonomicName>
)
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 2464/2007</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZCMV 5497</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Manombo</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">27.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">10.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">11.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">4.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">19.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">44.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">18.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">12.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">14.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">5.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.4</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 2463/2007</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZCMV 5486</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">F</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Manombo</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">29.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">11.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">13.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">4.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">24.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">11.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">51.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">22.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">14.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">16.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">NA</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">NA</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 537/2006</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZCMV 3362</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Ranomafana</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">23.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">10.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">19.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">40.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">17.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">11.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">12.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">5.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.0</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. matsilo" pageId="0" pageNumber="271" rank="species" species="matsilo">
<emphasis italics="true" pageId="0" pageNumber="271">G. matsilo</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="271">sp. nov.</emphasis>
(lineage A)
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 711/2009</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZCMV 7234</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">HT</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Ambatoroma, Befanjana</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">21.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">17.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">37.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">16.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">10.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">12.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">5.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.0</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 712/2009</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZCMV 7300</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">PT</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">F</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Babitanety, Befanjana</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">24.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">10.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">4.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">18.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">42.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">17.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">11.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">13.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">NA</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">NA</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZMA 20247</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZCMV 90</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">PT</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Ambohitsara</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">21.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">18.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">39.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">17.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">11.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">11.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">5.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.3</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. fiharimpe" pageId="0" pageNumber="271" rank="species" species="fiharimpe">
<emphasis italics="true" pageId="0" pageNumber="271">G. fiharimpe</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="271">sp. nov.</emphasis>
(lineage B)
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 164/2016</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">FGZC 5181</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">HT</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Mandraka</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">23.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">17.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">41.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">18.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">12.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">13.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.6</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZMA 19421</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">FGMV 2002.415</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">PT</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">F?</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Ranomafana</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">25.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">10.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">18.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">43.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">19.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">12.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">13.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">NA</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">NA</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. oelkrugi" pageId="0" pageNumber="271" rank="species" species="oelkrugi">
<emphasis italics="true" pageId="0" pageNumber="271">G. oelkrugi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="271">sp. nov.</emphasis>
(lineage C)
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 271/2016</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">FGZC 5300</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">PT</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">F?</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Masoala</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">21.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">14.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">7.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">32.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">14.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">10.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">NA</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">NA</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 315/2010</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">NA</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">PT</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Ambodivoangy</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">21.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">16.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">17.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">38.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">16.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">11.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">12.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.6</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 314/2010</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">FGZC 4220</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">HT</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Ambodivoangy</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">21.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">15.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">6.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">33.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">14.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">11.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.0</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. portonae" pageId="0" pageNumber="271" rank="species" species="portonae">
<emphasis italics="true" pageId="0" pageNumber="271">G. portonae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="271">sp. nov.</emphasis>
(lineage D)
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="271" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 115/2021</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ACZCV 0032</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">HT</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Betampona</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">22.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">10.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">17.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">36.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">16.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">10.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">11.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">5.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.5</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 118/2021</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ACZCV 0223</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">PT</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">F</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Betampona</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">26.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">10.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">4.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">19.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">45.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">20.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">12.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">13.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">NA</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">NA</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 116/2021</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ACZCV 0030</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">PT</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">F</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Betampona</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">25.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">10.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">18.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">40.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">17.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">11.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">12.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">NA</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">NA</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ZSM 117/2021</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">ACZCV 0025</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">PT</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">M</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Betampona</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">22.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">8.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">3.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.5</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1.9</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.4</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">19.3</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">9.0</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">41.2</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">17.7</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">12.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">12.6</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">5.1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">2.4</td>
</tr>
</table>
</paragraph>
<caption ID-Table-UUID="667E7BF5450183C822A2EB6CF2A49BD0" httpUri="http://table.plazi.org/id/667E7BF5450183C822A2EB6CF2A49BD0" pageId="0" pageNumber="271" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="271">
<emphasis bold="true" pageId="0" pageNumber="271">Table 2.</emphasis>
Comparative numerical parameters of analyzed advertisement calls of frogs in the subgenus 
<taxonomicName class="Amphibia" family="Mantellidae" genus="Gephyromantis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gephyromantis (Laurentomantis)" order="Anura" pageId="0" pageNumber="271" phylum="Chordata" rank="subGenus" subGenus="Laurentomantis">Gephyromantis Laurentomantis</taxonomicName>
. Values are provided as range followed by mean 
<normalizedToken originalValue="±">+/-</normalizedToken>
standard deviation in parentheses.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="271">
<table pageId="0" pageNumber="271">
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="271">species</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="271">locality</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="271">call duration [ms</emphasis>
]
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="271">note duration [ms</emphasis>
]
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="271">notes per call</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="271">note repetition rate within call [notes/second</emphasis>
]
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="271">pulses per note</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="271">dominant frequency [Hz</emphasis>
]
</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. horridus" pageId="0" pageNumber="271" rank="species" species="horridus">
<emphasis italics="true" pageId="0" pageNumber="271">G. horridus</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
Montagne 
<normalizedToken originalValue="d’Ambre">d'Ambre</normalizedToken>
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
543-618 (585.1 
<normalizedToken originalValue="±">+/-</normalizedToken>
27.4)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
6-12 (9.4 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.7)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
17-19 (18.1 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.8)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
29.4-31.0 (29.9 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.7)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
2-4 (3.0 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.5)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
1172-1369 (1264 
<normalizedToken originalValue="±">+/-</normalizedToken>
71)
</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. ranjomavo" pageId="0" pageNumber="271" rank="species" species="ranjomavo">
<emphasis italics="true" pageId="0" pageNumber="271">G. ranjomavo</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Manarikoba Forest</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
1780-2526 (2237.9 
<normalizedToken originalValue="±">+/-</normalizedToken>
255.6)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
10-19 (13.6 
<normalizedToken originalValue="±">+/-</normalizedToken>
3.1)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
23-33 (29.0 
<normalizedToken originalValue="±">+/-</normalizedToken>
3.2)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
10.8-14.2 (12.7 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
2348-3204 (2909 
<normalizedToken originalValue="±">+/-</normalizedToken>
355)
</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. striatus" pageId="0" pageNumber="271" rank="species" species="striatus">
<emphasis italics="true" pageId="0" pageNumber="271">G. striatus</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Marojejy</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
439-1360 (848.5 
<normalizedToken originalValue="±">+/-</normalizedToken>
319.1)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
3-7 (5.1 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.8)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
17-45 (29.8 
<normalizedToken originalValue="±">+/-</normalizedToken>
8.4)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
25.6-33.5 (31.1 
<normalizedToken originalValue="±">+/-</normalizedToken>
3.2)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
3768-4153 (3983 
<normalizedToken originalValue="±">+/-</normalizedToken>
161)
</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. marokoroko" pageId="0" pageNumber="271" rank="species" species="marokoroko">
<emphasis italics="true" pageId="0" pageNumber="271">G. marokoroko</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Vohidrazana</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
557-1203 (968.7 
<normalizedToken originalValue="±">+/-</normalizedToken>
357.7)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
10-26 (17.4 
<normalizedToken originalValue="±">+/-</normalizedToken>
5.0)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
12-28 (21.0 
<normalizedToken originalValue="±">+/-</normalizedToken>
8.2)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
18.2-19.6 (18.8 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.6)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
2-5 (3.3 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
2411-2476 (2444 
<normalizedToken originalValue="±">+/-</normalizedToken>
28)
</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. malagasius" pageId="0" pageNumber="271" rank="species" species="malagasius">
<emphasis italics="true" pageId="0" pageNumber="271">G. malagasius</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Vohiparara/ Ranomafana</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
360-465 (428.4 
<normalizedToken originalValue="±">+/-</normalizedToken>
37.2)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
8-17 (12.6 
<normalizedToken originalValue="±">+/-</normalizedToken>
2.9)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
8-10 (9.6 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.8)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
19.8-21.1 (20.5 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.5)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
2-10 (6.2 
<normalizedToken originalValue="±">+/-</normalizedToken>
2.1)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
2606-3516 (3127 
<normalizedToken originalValue="±">+/-</normalizedToken>
254)
</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. malagasius" pageId="0" pageNumber="271" rank="species" species="malagasius">
<emphasis italics="true" pageId="0" pageNumber="271">G. malagasius</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Manombo</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
348-446 (386.3 
<normalizedToken originalValue="±">+/-</normalizedToken>
34.5)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
9-16 (12.2 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.9)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
16-20 (17.5 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.6)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
42.7-45.5 (44.7 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
2-9 (5.5 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.9)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
2916-3192 (3041 
<normalizedToken originalValue="±">+/-</normalizedToken>
107)
</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. matsilo" pageId="0" pageNumber="271" rank="species" species="matsilo">
<emphasis italics="true" pageId="0" pageNumber="271">G. matsilo</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="271">sp. nov.</emphasis>
(lineage A)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Betampona</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
355-660 (552.3 
<normalizedToken originalValue="±">+/-</normalizedToken>
88.0)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
13-21 (16.2 
<normalizedToken originalValue="±">+/-</normalizedToken>
2.5)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
8-15 (12.6 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.9)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
20.8-21.6 (21.2 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.3)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
6-10 (6.9 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.1)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
3090-3434 (3200 
<normalizedToken originalValue="±">+/-</normalizedToken>
116)
</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. fiharimpe" pageId="0" pageNumber="271" rank="species" species="fiharimpe">
<emphasis italics="true" pageId="0" pageNumber="271">G. fiharimpe</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="271">sp. nov.</emphasis>
(lineage B)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Ankeniheny</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
609-935 (798.8 
<normalizedToken originalValue="±">+/-</normalizedToken>
110.8)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
7-9 (7.8 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.8)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
11-19 (15.5 
<normalizedToken originalValue="±">+/-</normalizedToken>
2.4)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
14.9-25.6 (18.3 
<normalizedToken originalValue="±">+/-</normalizedToken>
3.9)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
3638-3723 (3676 
<normalizedToken originalValue="±">+/-</normalizedToken>
23)
</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. fiharimpe" pageId="0" pageNumber="271" rank="species" species="fiharimpe">
<emphasis italics="true" pageId="0" pageNumber="271">G. fiharimpe</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="271">sp. nov.</emphasis>
(lineage B)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Andasibe</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
840-1503 (1179.3 
<normalizedToken originalValue="±">+/-</normalizedToken>
232.7)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
5-12 (7.6 
<normalizedToken originalValue="±">+/-</normalizedToken>
2.2)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
26-43 (36.9 
<normalizedToken originalValue="±">+/-</normalizedToken>
7.3)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
18.2-37.4 (25.5 
<normalizedToken originalValue="±">+/-</normalizedToken>
6.5)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
3649-4078 (3796 
<normalizedToken originalValue="±">+/-</normalizedToken>
157)
</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. fiharimpe" pageId="0" pageNumber="271" rank="species" species="fiharimpe">
<emphasis italics="true" pageId="0" pageNumber="271">G. fiharimpe</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="271">sp. nov.</emphasis>
(lineage B)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Vohidrazana</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
950-1764 (1405.6 
<normalizedToken originalValue="±">+/-</normalizedToken>
290.4)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
2-7 (4.3 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.7)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
26-52 (40.7 
<normalizedToken originalValue="±">+/-</normalizedToken>
11.5)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
18.8-54.1 (32.1 
<normalizedToken originalValue="±">+/-</normalizedToken>
13.1)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
3726-3913 (3781 
<normalizedToken originalValue="±">+/-</normalizedToken>
68)
</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. fiharimpe" pageId="0" pageNumber="271" rank="species" species="fiharimpe">
<emphasis italics="true" pageId="0" pageNumber="271">G. fiharimpe</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="271">sp. nov.</emphasis>
(lineage B)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Ambatolahy/Ranomafana</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
916-1162 (1073.9 
<normalizedToken originalValue="±">+/-</normalizedToken>
86.0)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
4-7 (6.2 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.9)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
27-34 (30.6 
<normalizedToken originalValue="±">+/-</normalizedToken>
2.3)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
25.6-28.0 (26.8 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.0)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
3305-3736 (3523 
<normalizedToken originalValue="±">+/-</normalizedToken>
145)
</td>
</tr>
<tr pageId="0" pageNumber="271">
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
<taxonomicName genus="G." lsidName="G. oelkrugi" pageId="0" pageNumber="271" rank="species" species="oelkrugi">
<emphasis italics="true" pageId="0" pageNumber="271">G. oelkrugi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="271">sp. nov.</emphasis>
(lineage C)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">Ambodivoangy</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
279-504 (350.9 
<normalizedToken originalValue="±">+/-</normalizedToken>
88.4)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
3-10 (6.8 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.3)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
13-25 (16.9 
<normalizedToken originalValue="±">+/-</normalizedToken>
4.4)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
43.5-54.1 (46.5 
<normalizedToken originalValue="±">+/-</normalizedToken>
4.0)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
1-4 (2.9 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.7)
</td>
<td colspan="1" pageId="0" pageNumber="271" rowspan="1">
3266-3882 (3539 
<normalizedToken originalValue="±">+/-</normalizedToken>
190)
</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="271" type="call">
<paragraph pageId="0" pageNumber="271">Call.</paragraph>
<paragraph pageId="0" pageNumber="271">
The advertisement call of specimen ZSM 126/2018 was recorded at 20:20-21:00 on 11 December 2017 on a muddy bank beside a very slow segment of a stream on the west slope of Montagne 
<normalizedToken originalValue="d’Ambre">d'Ambre</normalizedToken>
(
<geoCoordinate degrees="12.5915" direction="south" orientation="latitude" precision="5" value="-12.5915">12.5915°S</geoCoordinate>
, 
<geoCoordinate degrees="049.1372" direction="east" orientation="longitude" precision="5" value="49.1372">049.1372°E</geoCoordinate>
, 939 m a.s.l.; air temperature not available). It consists of a series of short, pulsed notes (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Audiospectrogram and corresponding oscillogram of one advertisement call of Gephyromantis horridus (ZSM 126 / 2018) recorded on 11 December 2017 in Montagne d'Ambre National Park." figureDoi="10.3897/vz.72.e78830.figure6" httpUri="https://binary.pensoft.net/fig/688676" pageId="0" pageNumber="271">6</figureCitation>
). They were extremely difficult to localize in the field due to their low amplitude. There is amplidude modulation within each call, with call energy being greatest at approximately 70% of the 
<normalizedToken originalValue="call’s">call's</normalizedToken>
duration, with initial notes being the least energetic ones. Within calls, notes are repeated at very constant intervals. Each note contains few clearly countable pulses repeated at an approximate rate of 500 pulses/second. Within regular call series, calls are emitted in rapid succession. Numerical parameters of 9 analyzed calls are as follows (range followed by mean 
<normalizedToken originalValue="±">+/-</normalizedToken>
standard deviation in parentheses): call duration 543-618 ms (585.1 
<normalizedToken originalValue="±">+/-</normalizedToken>
27.4 ms); note duration 6-12 ms (9.4 
<normalizedToken originalValue="±">+/-</normalizedToken>
1.7 ms); number of notes per call 17-19 (18.1 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.8); note repetition rate within calls 29.4-31.0 notes/second (29.9 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.7 notes/second); pulses per note 2-4 (3.0 
<normalizedToken originalValue="±">+/-</normalizedToken>
0.5); dominant frequency 1172-1369 Hz (1264 
<normalizedToken originalValue="±">+/-</normalizedToken>
71 Hz); prevalent bandwidth 900-3500 Hz. Calls were emitted in regular call series at a rate of approximately 41 calls/minute.
</paragraph>
</subSubSection>
</treatment>
</document>