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<mods:title id="B7FE8A95A45720A8958D34687D03F528">Faunal composition, diversity, and distribution of ants (Hymenoptera: Formicidae) of Dhofar Governorate, Oman, with updated list of the Omani species and remarks on zoogeography</mods:title>
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<mods:namePart id="CEACAFD45C7D075A24423C376F525977">Sharaf, Mostafa R.</mods:namePart>
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<emphasis id="AEBC8146FFB0FF98FE75000E12A8FBEA" bold="true" box="[485,756,1055,1081]" italics="true" pageId="32" pageNumber="33">Lepisiota spinisquama</emphasis>
(
<bibRefCitation id="F85920A5FFB0FF98FC94000E141AFBEA" author="Kuznetsov-Ugamsky N. N." box="[772,1094,1055,1081]" pageId="32" pageNumber="33" pagination="477 - 492" refId="ref26957" refString="Kuznetsov-Ugamsky N. N. 1929. Die Gattung Acantholepis in Turkestan. Zoologischer Anzeiger 82: 477 - 492." type="journal article" year="1929">Kuznetsov-Ugamsky, 1929</bibRefCitation>
)
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<figureCitation id="04F341D1FFB0FF98FD600053131FFB8F" box="[752,835,1090,1116]" captionStart="Fig" captionStartId="34.[189,232,1930,1956]" captionTargetBox="[527,1060,265,1859]" captionTargetId="figure-15@34.[527,1060,265,1893]" captionTargetPageId="34" captionText="Fig. 21. Lepisiota spinisquama (Kuznetsov-Ugamsky, 1929), worker (CASENT0922842, AntWeb.org (Michele Esposito)). A. Body in profile. B. Head in full-face view. C. Distribution map." figureDoi="http://doi.org/10.5281/zenodo.7094371" httpUri="https://zenodo.org/record/7094371/files/figure.png" pageId="32" pageNumber="33">Fig. 21</figureCitation>
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<paragraph id="9C775D54FFB0FF98FF2D0092113AFB12" blockId="32.[189,1399,1155,1217]" pageId="32" pageNumber="33">
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<treatmentCitation id="1D697B45FFB0FF98FF2D009214CAFB4D" author="Kuznetsov-Ugamsky N. N." box="[189,1174,1155,1182]" page="483" pageId="32" pageNumber="33" year="1929">
<taxonomicName id="5BC826D7FFB0FF98FF2D009214CAFB4D" authority="Kuznetsov-Ugamsky, 1929: 483" authorityName="Kuznetsov-Ugamsky" authorityPageNumber="483" authorityYear="1929" box="[189,1174,1155,1182]" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="frauenfeldi" subSpecies="spinisquama">
<emphasis id="AEBC8146FFB0FF98FF2D009211B7FB4E" box="[189,491,1155,1181]" italics="true" pageId="32" pageNumber="33">Acantholepis frauenfeldi</emphasis>
subsp.
<emphasis id="AEBC8146FFB0FF98FDCC009512ACFB4E" box="[604,752,1156,1181]" italics="true" pageId="32" pageNumber="33">spinisquama</emphasis>
<bibRefCitation id="F85920A5FFB0FF98FC93009214CAFB4D" author="Kuznetsov-Ugamsky N. N." box="[771,1174,1155,1182]" pageId="32" pageNumber="33" pagination="477 - 492" refId="ref26957" refString="Kuznetsov-Ugamsky N. N. 1929. Die Gattung Acantholepis in Turkestan. Zoologischer Anzeiger 82: 477 - 492." type="journal article" year="1929">Kuznetsov-Ugamsky, 1929: 483</bibRefCitation>
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(w)
<collectingCountry id="E4DF1DC4FFB0FF98FB760092152FFB4E" box="[1254,1395,1155,1181]" name="Kazakhstan" pageId="32" pageNumber="33">Kazakhstan</collectingCountry>
. Palearctic.
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</paragraph>
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<emphasis id="AEBC8146FFB0FF98FF2D00F9116AFAD1" bold="true" box="[189,310,1256,1282]" pageId="32" pageNumber="33">Diagnosis</emphasis>
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<paragraph id="9C775D54FFB0FF98FF2D0106117EFAA4" blockId="32.[189,1399,1303,1400]" pageId="32" pageNumber="33">Body dark brown; propodeal spines shorter and slightly curved, in profile appearing at level of the petiolar spines; body slightly shining; petiolar height lower in profile; appressed pubescence abundant on body.</paragraph>
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<emphasis id="AEBC8146FFB0FF98FF2D018F11F6FA6B" bold="true" box="[189,426,1438,1464]" pageId="32" pageNumber="33">Material examined</emphasis>
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<collectingRegion id="5E0C93B6FFB0FF98FEA101DC11D6FA34" box="[305,394,1485,1511]" country="Oman" name="Zufar" pageId="32" pageNumber="33">Dhofar</collectingRegion>
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• 1 w;
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;
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,
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<collectingDate id="F832827CFFB0FF98FB9E01DC14F1FA34" box="[1038,1197,1485,1512]" pageId="32" pageNumber="33" value="2017-11-16">16 Nov. 2017</collectingDate>
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leg.;
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3 w;
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;
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,
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; alt.
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leg.;
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<materialsCitation id="2CA05709FFB0FF98FEB902051434F9FD" ID-GBIF-Occurrence="3909999342" box="[297,1128,1555,1582]" collectingDate="2017-11-19" collectionCode="KSMA" collectorName="M. R. Sharaf" country="Oman" elevation="34" latitude="16.707" location="Dhalkout" longLatPrecision="76" longitude="53.251" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Zufar">
1 w; same collection data as for preceding; CASENT0922822;
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.
</paragraph>
</subSubSection>
<subSubSection id="D4D20EDFFFB0FF98FF2D02441488F94E" pageId="32" pageNumber="33" type="biology_ecology">
<paragraph id="9C775D54FFB0FF98FF2D02441261F9BC" blockId="32.[189,573,1621,1647]" box="[189,573,1621,1647]" pageId="32" pageNumber="33">
<heading id="C73FEA38FFB0FF98FF2D02441261F9BC" bold="true" box="[189,573,1621,1647]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="AEBC8146FFB0FF98FF2D02441261F9BC" bold="true" box="[189,573,1621,1647]" pageId="32" pageNumber="33">Ecological and biological notes</emphasis>
</heading>
</paragraph>
<paragraph id="9C775D54FFB0FF98FF2D02921488F94E" blockId="32.[189,1236,1667,1694]" box="[189,1236,1667,1694]" pageId="32" pageNumber="33">This species was found foraging on small shrubs and was collected using a beating sheet.</paragraph>
</subSubSection>
<subSubSection id="D4D20EDFFFB0FF98FF2D02D4128EF8A4" pageId="32" pageNumber="33" type="distribution">
<paragraph id="9C775D54FFB0FF98FF2D02D411C3F90C" blockId="32.[189,415,1733,1759]" box="[189,415,1733,1759]" pageId="32" pageNumber="33">
<heading id="C73FEA38FFB0FF98FF2D02D411C3F90C" bold="true" box="[189,415,1733,1759]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="AEBC8146FFB0FF98FF2D02D411C3F90C" bold="true" box="[189,415,1733,1759]" pageId="32" pageNumber="33">Geographic range</emphasis>
</heading>
</paragraph>
<paragraph id="9C775D54FFB0FF98FF2D02E2128EF8A4" blockId="32.[189,1399,1779,1911]" pageId="32" pageNumber="33">
This species was originally described from
<collectingCountry id="E4DF1DC4FFB0FF98FD6102E21326F8DE" box="[753,890,1779,1805]" name="Kazakhstan" pageId="32" pageNumber="33">Kazakhstan</collectingCountry>
and recorded subsequently from
<collectingCountry id="E4DF1DC4FFB0FF98FABF02E5152BF8DD" box="[1327,1399,1780,1806]" name="Oman" pageId="32" pageNumber="33">Oman</collectingCountry>
(
<bibRefCitation id="F85920A5FFB0FF98FF550306127DF8E2" author="Collingwood C. A. &amp; Agosti D." box="[197,545,1815,1841]" pageId="32" pageNumber="33" pagination="300 - 385" refId="ref24426" refString="Collingwood C. A. &amp; Agosti D. 1996. Formicidae (Insecta: Hymenoptera) of Saudi Arabia (part 2). Fauna of Saudi Arabia 15: 300 - 385." type="journal article" year="1996">Collingwood &amp; Agosti 1996</bibRefCitation>
) and the
<collectingCountry id="E4DF1DC4FFB0FF98FD09030612AFF8E2" box="[665,755,1815,1841]" name="Yemen" pageId="32" pageNumber="33">Socotra</collectingCountry>
Archipelago (
<bibRefCitation id="F85920A5FFB0FF98FC30030614D8F8E2" author="Collingwood C. A." box="[928,1156,1814,1841]" pageId="32" pageNumber="33" pagination="230 - 302" refId="ref24400" refString="Collingwood C. A. 1985. Hymenoptera: Fam. Formicidae of Saudi Arabia. Fauna of Saudi Arabia 7: 230 - 302." type="journal article" year="1985">Collingwood 1985</bibRefCitation>
;
<bibRefCitation id="F85920A5FFB0FF98FB04030610A7F887" author="Collingwood C. A. &amp; Pohl H. &amp; Guesten R. &amp; Wranik W. &amp; van Harten A." pageId="32" pageNumber="33" pagination="473 - 495" refId="ref24544" refString="Collingwood C. A., Pohl H., Guesten R., Wranik W. &amp; van Harten A. 2004. The ants (Insecta: Hymenoptera: Formicidae) of the Socotra Archipelago. Fauna of Arabia 20: 473 - 495." type="journal article" year="2004">
Collingwood
<emphasis id="AEBC8146FFB0FF98FAA80306152BF8E2" box="[1336,1399,1815,1841]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2004
</bibRefCitation>
;
<bibRefCitation id="F85920A5FFB0FF98FE98032B11B1F887" author="Sharaf M. R. &amp; Fisher B. L. &amp; Collingwood C. A. &amp; Aldawood S. A." box="[264,493,1850,1876]" pageId="32" pageNumber="33" pagination="317 - 378" refId="ref29256" refString="Sharaf M. R., Fisher B. L., Collingwood C. A. &amp; Aldawood S. A. 2017 c. Ant fauna (Hymenoptera: Formicidae) of the Socotra Archipelago (Yemen): zoogeography, distribution and description of a new species. Journal of Natural History 51 (5 - 6): 317 - 378. https: // doi. org / 10.1080 / 00222933.2016.1271157" type="journal article" year="2017">
Sharaf
<emphasis id="AEBC8146FFB0FF98FECD032A11C4F887" box="[349,408,1850,1876]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2017c
</bibRefCitation>
).
<bibRefCitation id="F85920A5FFB0FF98FD91032B1296F887" author="Rad S. P. &amp; Taylor B. &amp; Torabi R. &amp; Aram E. &amp; Abolfathi G. &amp; Afshari R. &amp; Borjali F. &amp; Ghatei M. &amp; Hediary H. &amp; Jazini F. &amp; Kiah V. H. &amp; Mahmoudi F. &amp; Safariyan F. &amp; Seiri M." box="[513,714,1850,1876]" pageId="32" pageNumber="33" pagination="145 - 149" refId="ref27941" refString="Rad S. P., Taylor B., Torabi R., Aram E., Abolfathi G., Afshari R., Borjali F., Ghatei M., Hediary H., Jazini F., Kiah V. H., Mahmoudi F., Safariyan F. &amp; Seiri M. 2018. Further records of ants (Hymenoptera: Formicidae) from Iran. Zoology in the Middle East 64: 145 - 149. https: // doi. org / 10.1080 / 09397140.2018.1442301" type="journal article" year="2018">
Rad
<emphasis id="AEBC8146FFB0FF98FDA9032A1228F887" box="[569,628,1850,1876]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
(2018)
</bibRefCitation>
recorded it from pitfall traps in
<collectingCountry id="E4DF1DC4FFB0FF98FBD8032B1427F887" box="[1096,1147,1850,1876]" name="Iran" pageId="32" pageNumber="33">Iran</collectingCountry>
.
<bibRefCitation id="F85920A5FFB0FF98FB17032B152BF886" author="Collingwood C. A." box="[1159,1399,1850,1877]" pageId="32" pageNumber="33" pagination="230 - 302" refId="ref24400" refString="Collingwood C. A. 1985. Hymenoptera: Fam. Formicidae of Saudi Arabia. Fauna of Saudi Arabia 7: 230 - 302." type="journal article" year="1985">Collingwood (1985)</bibRefCitation>
previously reported this species from Dhofar.
</paragraph>
</subSubSection>
</treatment>
</document>