152 lines
22 KiB
XML
152 lines
22 KiB
XML
<document id="64096EF906BDD75B74F0A86A88733D7B" ID-CLB-Dataset="3695" ID-DOI="10.35929/RSZ.0050" ID-GBIF-Dataset="60a46419-793d-4cae-a0bc-a086fc235849" ID-ISSN="0035-418" ID-Zenodo-Dep="5640135" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635893751209" checkinUser="felipe" docAuthor="Balkenohl, Michael" docDate="2021" docId="014C8648FFE7BB3B30147132FBB1F94B" docLanguage="en" docName="RevSuisseZool.128.2.357-374.pdf" docOrigin="Revue suisse de Zoologie 128 (2)" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Parasyleter Balkenohl 2021, gen. nov." docType="treatment" docVersion="7" lastPageNumber="369" masterDocId="FD75FE30FFEABB36313E7638FFFBFFB0" masterDocTitle="Revisiting the Oriental genus Syleter Andrewes, 1941 (Coleoptera: Carabidae: Clivinini)" masterLastPageNumber="374" masterPageNumber="357" pageNumber="369" updateTime="1699467467588" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="5F15CBE50E366699417B831DB7B06F9A">
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<mods:title id="DAFED0B2761B85F866693D479D72F5F1">Revisiting the Oriental genus Syleter Andrewes, 1941 (Coleoptera: Carabidae: Clivinini)</mods:title>
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<mods:name id="2A239A0EB1D58E75F8E577CF7B1175CC" type="personal">
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<mods:namePart id="339189AA57983900508CDC8BEB235F47">Balkenohl, Michael</mods:namePart>
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<mods:affiliation id="CD78E34B53DD2BD0E2F5F3832E049CC6">Ligusterweg 9, CH- 8906 Bonstetten, Switzerland</mods:affiliation>
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<mods:nameIdentifier id="6CFE4F5796A203DFB6146B635300C22D" type="email">mike.balkenohl@bluewin.ch</mods:nameIdentifier>
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<mods:typeOfResource id="66F5B18178F4237F6FC928194CF26331">text</mods:typeOfResource>
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<mods:title id="E6EECFB9129474DEB1F72682564289A2">Revue suisse de Zoologie</mods:title>
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<mods:part id="878B37184B1876D24B416E9980BB5479">
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<mods:date id="08EB839E88BF1DFC1D128609CE4F86EC">2021</mods:date>
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<mods:detail id="CE52681448AC9F6B3528EE8E4CC0849A" type="volume">
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<mods:number id="594B39FFA1A256B31374421554B165E0">128</mods:number>
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<mods:start id="14A6203CA5FE8C837530DDF508C46AB2">357</mods:start>
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<mods:end id="F49C3011861C48FF324D6910F3E55B4F">374</mods:end>
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<mods:classification id="08747875C97E6B12A0F799814269B19D">journal article</mods:classification>
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<mods:identifier id="4897AB5BDA9BC3A5ED898E34C1513919" type="DOI">10.35929/RSZ.0050</mods:identifier>
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<mods:identifier id="025930711340C46DCF8775972237C426" type="ISSN">0035-418</mods:identifier>
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<treatment id="014C8648FFE7BB3B30147132FBB1F94B" ID-DOI="http://doi.org/10.5281/zenodo.5640152" ID-GBIF-Taxon="189991357" ID-Zenodo-Dep="5640152" LSID="urn:lsid:plazi:treatment:014C8648FFE7BB3B30147132FBB1F94B" httpUri="http://treatment.plazi.org/id/014C8648FFE7BB3B30147132FBB1F94B" lastPageNumber="369" pageId="13" pageNumber="369">
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<subSubSection id="C1FF64D5FFE7BB3B30147132FD9AF892" box="[298,609,1802,1826]" pageId="13" pageNumber="369" type="nomenclature">
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<paragraph id="895A375EFFE7BB3B30147132FD9AF892" blockId="13.[298,609,1802,1857]" box="[298,609,1802,1826]" pageId="13" pageNumber="369">
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<heading id="D2128032FFE7BB3B30147132FD9AF892" bold="true" box="[298,609,1802,1826]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="369" reason="2">
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<emphasis id="BB91EB4CFFE7BB3B30147132FD9AF892" bold="true" box="[298,609,1802,1826]" pageId="13" pageNumber="369">
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Genus
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<taxonomicName id="4EE54CDDFFE7BB3B30447132FE0DF892" authority="Balkenohl, 2021" authorityName="Balkenohl" authorityYear="2021" box="[378,502,1802,1826]" class="Insecta" family="Carabidae" genus="Parasyleter" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="369" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="BB91EB4CFFE7BB3B30447132FE0DF892" bold="true" box="[378,502,1802,1826]" italics="true" pageId="13" pageNumber="369">Parasyleter</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A0A25637FFE7BB3B30C37132FD9AF892" box="[509,609,1802,1826]" pageId="13" pageNumber="369" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C1FF64D5FFE7BB3B30657111FDCBF8F1" box="[347,560,1833,1857]" pageId="13" pageNumber="369" type="description">
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<paragraph id="895A375EFFE7BB3B30657111FDCBF8F1" blockId="13.[298,609,1802,1857]" box="[347,560,1833,1857]" pageId="13" pageNumber="369">
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<figureCitation id="11DE2BDBFFE7BB3B30657111FE64F8F1" box="[347,415,1833,1857]" captionStart="Figs 1-3" captionStartId="4.[151,192,2024,2046]" captionTargetBox="[151,1436,1374,1975]" captionTargetId="figure-659@4.[151,1436,1366,1975]" captionTargetPageId="4" captionText="Figs 1-3. Head, dorsal view. (1) Syleter paradoxus (Putzeys, 1868).(2) Parasyleter doriae (Putzeys, 1873) comb.nov.(3) Clypeuspinus validus (Andrewes, 1936) comb. nov." figureDoi="http://doi.org/10.5281/zenodo.5640137" httpUri="https://zenodo.org/record/5640137/files/figure.png" pageId="13" pageNumber="369">Figs 2</figureCitation>
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,
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<figureCitation id="11DE2BDBFFE7BB3B30957111FE41F8F1" box="[427,442,1833,1857]" captionStart="Figs 4-6" captionStartId="5.[151,192,939,961]" captionTargetBox="[158,749,264,900]" captionTargetId="figure-983@5.[151,757,257,914]" captionTargetPageId="5" captionText="Figs 4-6. Right mesotibia. (4) Syleter paradoxus (Putzeys, 1868) showing the small preapical protuberance and its seta inserted apically (arrow). (5) Parasyleter doriae (Putzeys, 1873) comb. nov. with no tubercles at all. (6) Clypeuspinus validus (Andrewes, 1936) comb. nov. showing the long protuberance and its seta inserted laterally (arrow)." figureDoi="http://doi.org/10.5281/zenodo.5640139" httpUri="https://zenodo.org/record/5640139/files/figure.png" pageId="13" pageNumber="369">5</figureCitation>
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,
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<figureCitation id="11DE2BDBFFE7BB3B30FB7111FDCBF8F1" box="[453,560,1833,1857]" captionStart="Figs 28-30" captionStartId="14.[151,192,994,1016]" captionTargetBox="[175,1406,262,941]" captionTargetId="figure-485@14.[151,1439,241,968]" captionTargetPageId="14" captionText="Figs 28-30. Parasyleter doriae (Putzeys, 1873) comb. nov., specimen from Sarawak collected by G. Doria. (28) Habitus, male. (29) Male genitalia, ventral view (above) and lateral view (below) of aedeagus with parameres in between. (30) Female coxostyli and laterotergite." figureDoi="http://doi.org/10.5281/zenodo.5640146" httpUri="https://zenodo.org/record/5640146/files/figure.png" pageId="13" pageNumber="369">28, 29, 30</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C1FF64D5FFE7BB3B31A97160FD44F8C0" box="[151,703,1880,1905]" pageId="13" pageNumber="369" type="type_taxon">
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<paragraph id="895A375EFFE7BB3B31A97160FD44F8C0" blockId="13.[151,703,1880,1905]" box="[151,703,1880,1905]" pageId="13" pageNumber="369">
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<emphasis id="BB91EB4CFFE7BB3B31A97160FED6F8C0" bold="true" box="[151,301,1880,1904]" pageId="13" pageNumber="369">
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<typeStatus id="565E89FCFFE7BB3B31A97160FF34F8C0" box="[151,207,1880,1904]" pageId="13" pageNumber="369">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4EE54CDDFFE7BB3B300A7160FD47F8C0" ID-CoL="8M5J7" authority="Putzeys, 1873: 14" authorityName="Putzeys" authorityPageNumber="14" authorityYear="1873" box="[308,700,1880,1905]" class="Insecta" family="Carabidae" genus="Dyschirius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="369" phylum="Arthropoda" rank="species" species="doriae">
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<emphasis id="BB91EB4CFFE7BB3B300A7160FE0FF8C0" box="[308,500,1880,1904]" italics="true" pageId="13" pageNumber="369">Dyschirius doriae</emphasis>
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<bibRefCitation id="ED744AAFFFE7BB3B30C47161FD47F8C0" author="Putzeys J. A. A. H." box="[506,700,1880,1905]" pageId="13" pageNumber="369" pagination="10 - 18" refId="ref14650" refString="Putzeys J. A. A. H. 1873. Deuxieme supplement a la revision generale des Clivinides. Annales de la Societe Entomologique de Belgique 16: 10 - 18." type="journal article" year="1873">Putzeys, 1873: 14</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C1FF64D5FFE7BB3B31A971BFFAD5FA09" pageId="13" pageNumber="369" type="diagnosis">
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<paragraph id="895A375EFFE7BB3B31A971BFFB28FA39" blockId="13.[151,756,1926,2077]" lastBlockId="13.[832,1437,257,1417]" pageId="13" pageNumber="369">
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<emphasis id="BB91EB4CFFE7BB3B31A971BFFEA4F82F" bold="true" box="[151,351,1927,1951]" pageId="13" pageNumber="369">Genus diagnosis:</emphasis>
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<emphasis id="BB91EB4CFFE7BB3B305571BFFE6DF82E" box="[363,406,1927,1950]" italics="true" pageId="13" pageNumber="369">Size</emphasis>
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<quantity id="4E1D9ABBFFE7BB3B309C71BEFDB1F82E" box="[418,586,1926,1950]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="13" pageNumber="369" unit="mm" value="2.05" valueMax="2.3" valueMin="1.8">1.8 to 2.3 mm</quantity>
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.
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<emphasis id="BB91EB4CFFE7BB3B336771BEFD68F82E" box="[601,659,1926,1950]" italics="true" pageId="13" pageNumber="369">Head</emphasis>
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(
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<figureCitation id="11DE2BDBFFE7BB3B339871BEFD17F82E" box="[678,748,1926,1950]" captionStart="Figs 1-3" captionStartId="4.[151,192,2024,2046]" captionTargetBox="[151,1436,1374,1975]" captionTargetId="figure-659@4.[151,1436,1366,1975]" captionTargetPageId="4" captionText="Figs 1-3. Head, dorsal view. (1) Syleter paradoxus (Putzeys, 1868).(2) Parasyleter doriae (Putzeys, 1873) comb.nov.(3) Clypeuspinus validus (Andrewes, 1936) comb. nov." figureDoi="http://doi.org/10.5281/zenodo.5640137" httpUri="https://zenodo.org/record/5640137/files/figure.png" pageId="13" pageNumber="369">Fig. 2</figureCitation>
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) with clypeus and frons distinctly elongated and flattened anteriorly; clypeus and clypeal wing fused, delimitation between clypeal wing and supraantennal plate observable as indistinct obtuse notch in two of the species; clypeus straight or slightly convex anteriorly, as far as or slightly more projecting than clypeal wing; mandible elongated, narrow in apical third, laterally nearly straight or convex, hooked at apical tip; labrum three- or five-setose; maxillary palpomere distinctly narrowed and slender towards apex, nearly straight, two apical segments securiform; antenna with scapus and pedicellus eccentrically attached; eye without or with distinct reduction, gena small or extended posteriorly.
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<collectingRegion id="4B21F9BCFFE7BB3B32F9741EFBD9FD8D" box="[967,1058,550,573]" country="Myanmar" name="Chin" pageId="13" pageNumber="369">Mentum</collectingRegion>
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not elongated, with lateral lobe convex, median tooth as wide as lateral lobe, acute, distinctly more projecting than lateral lobe, surface with slight transverse rugae.
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<emphasis id="BB91EB4CFFE7BB3B359274BFFAECFD2E" box="[1196,1303,647,670]" italics="true" pageId="13" pageNumber="369">Pronotum</emphasis>
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square or indistinctly trapezoid, with reflexed lateral margin incomplete, running from anterior angle up to posterior setigerous puncture, posterior angle rounded off without tooth, lateral channel of equal width, median line narrow, finely or just not joining basal constriction; proepisternum visible basally in dorsal view; flange at base convex, as narrow as or up to two times as wide as basal constriction.
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<emphasis id="BB91EB4CFFE7BB3B354D75B2FB38FC12" box="[1139,1219,906,930]" italics="true" pageId="13" pageNumber="369">Elytron</emphasis>
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with interval eight convex, lateral channel distinctly flared apically, with interval eight distinctly emarginated apically and row of umbilical setigerous punctures distinctly following the emargination, interval eight with distinct emarginated carina at apex; third interval without setigerous punctures.
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<emphasis id="BB91EB4CFFE7BB3B32887275FBCAFBD5" box="[950,1073,1101,1125]" italics="true" pageId="13" pageNumber="369">Hind wings</emphasis>
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fully developed or reduced.
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<emphasis id="BB91EB4CFFE7BB3B34547276FA67FBD5" box="[1386,1436,1102,1125]" italics="true" pageId="13" pageNumber="369">Legs</emphasis>
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with front and intermediate tarsomeres slightly widened, more distinct in males; mesotibia with four setae laterally, preapical seta either without any protuberance or arising from minute tubercle (
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<figureCitation id="11DE2BDBFFE7BB3B359B72F6FB13FB56" box="[1189,1256,1230,1254]" captionStart="Figs 4-6" captionStartId="5.[151,192,939,961]" captionTargetBox="[158,749,264,900]" captionTargetId="figure-983@5.[151,757,257,914]" captionTargetPageId="5" captionText="Figs 4-6. Right mesotibia. (4) Syleter paradoxus (Putzeys, 1868) showing the small preapical protuberance and its seta inserted apically (arrow). (5) Parasyleter doriae (Putzeys, 1873) comb. nov. with no tubercles at all. (6) Clypeuspinus validus (Andrewes, 1936) comb. nov. showing the long protuberance and its seta inserted laterally (arrow)." figureDoi="http://doi.org/10.5281/zenodo.5640139" httpUri="https://zenodo.org/record/5640139/files/figure.png" pageId="13" pageNumber="369">Fig. 5</figureCitation>
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).
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<emphasis id="BB91EB4CFFE7BB3B35C072F6FA67FB56" box="[1278,1436,1230,1254]" italics="true" pageId="13" pageNumber="369">Male genitalia</emphasis>
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(
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<figureCitation id="11DE2BDBFFE7BB3B327672D7FC60FAB7" box="[840,923,1263,1287]" captionStart="Figs 28-30" captionStartId="14.[151,192,994,1016]" captionTargetBox="[175,1406,262,941]" captionTargetId="figure-485@14.[151,1439,241,968]" captionTargetPageId="14" captionText="Figs 28-30. Parasyleter doriae (Putzeys, 1873) comb. nov., specimen from Sarawak collected by G. Doria. (28) Habitus, male. (29) Male genitalia, ventral view (above) and lateral view (below) of aedeagus with parameres in between. (30) Female coxostyli and laterotergite." figureDoi="http://doi.org/10.5281/zenodo.5640146" httpUri="https://zenodo.org/record/5640146/files/figure.png" pageId="13" pageNumber="369">Fig. 29</figureCitation>
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) with aedeagus distinctly curved, apical part elongated, apex more or less spatulate or securiform.
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<emphasis id="BB91EB4CFFE7BB3B327E7308FC0CFAF8" box="[832,1015,1328,1352]" italics="true" pageId="13" pageNumber="369">Female genitalia</emphasis>
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(
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<figureCitation id="11DE2BDBFFE7BB3B35387308FBADFAF8" box="[1030,1110,1328,1352]" captionStart="Figs 28-30" captionStartId="14.[151,192,994,1016]" captionTargetBox="[175,1406,262,941]" captionTargetId="figure-485@14.[151,1439,241,968]" captionTargetPageId="14" captionText="Figs 28-30. Parasyleter doriae (Putzeys, 1873) comb. nov., specimen from Sarawak collected by G. Doria. (28) Habitus, male. (29) Male genitalia, ventral view (above) and lateral view (below) of aedeagus with parameres in between. (30) Female coxostyli and laterotergite." figureDoi="http://doi.org/10.5281/zenodo.5640146" httpUri="https://zenodo.org/record/5640146/files/figure.png" pageId="13" pageNumber="369">Fig. 30</figureCitation>
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) with coxostylus one and two completely fused, with evident cone-like seta in apical third; laterotergite with isolated setae.
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</paragraph>
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<paragraph id="895A375EFFE7BB3B327E7399FAD5FA09" blockId="13.[832,1326,1440,1465]" box="[832,1326,1440,1465]" pageId="13" pageNumber="369">
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<emphasis id="BB91EB4CFFE7BB3B327E7399FBC7FA09" bold="true" box="[832,1084,1441,1465]" pageId="13" pageNumber="369">Differential diagnosis:</emphasis>
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See key to the genera.
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</paragraph>
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</subSubSection>
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<subSubSection id="C1FF64D5FFE7BB3B327E73E8FA84F9B8" pageId="13" pageNumber="369" type="etymology">
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<paragraph id="895A375EFFE7BB3B327E73E8FA84F9B8" blockId="13.[832,1436,1488,1545]" pageId="13" pageNumber="369">
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<emphasis id="BB91EB4CFFE7BB3B327E73E8FC3FFA58" bold="true" box="[832,964,1488,1512]" pageId="13" pageNumber="369">Etymology:</emphasis>
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The name is a combination of the Greek adjective para- (besides) and the original genus name.
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</paragraph>
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</subSubSection>
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<subSubSection id="C1FF64D5FFE7BB3B327E7018FBB1F94B" pageId="13" pageNumber="369" type="synonymic_list">
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<paragraph id="895A375EFFE7BB3B327E7018FBB1F94B" blockId="13.[832,1437,1568,1787]" pageId="13" pageNumber="369">
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<emphasis id="BB91EB4CFFE7BB3B327E7018FC67F988" bold="true" box="[832,924,1568,1592]" pageId="13" pageNumber="369">Species:</emphasis>
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Three described species belong into the genus:
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<taxonomicName id="4EE54CDDFFE7BB3B327E7078FB5BF9E8" authority="Putzeys, 1873" authorityName="Putzeys" authorityYear="1873" box="[832,1184,1600,1625]" class="Insecta" family="Carabidae" genus="Dyschirius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="369" phylum="Arthropoda" rank="species" species="doriae">
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<emphasis id="BB91EB4CFFE7BB3B327E7078FBFAF9E8" box="[832,1025,1600,1624]" italics="true" pageId="13" pageNumber="369">Dyschirius doriae</emphasis>
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<bibRefCitation id="ED744AAFFFE7BB3B35377079FB5BF9E8" author="Putzeys J. A. A. H." box="[1033,1184,1600,1625]" pageId="13" pageNumber="369" pagination="10 - 18" refId="ref14650" refString="Putzeys J. A. A. H. 1873. Deuxieme supplement a la revision generale des Clivinides. Annales de la Societe Entomologique de Belgique 16: 10 - 18." type="journal article" year="1873">Putzeys, 1873</bibRefCitation>
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</taxonomicName>
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known from
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<collectingCountry id="F1F277CEFFE7BB3B340B7078FA66F9E8" box="[1333,1437,1600,1624]" name="Indonesia" pageId="13" pageNumber="369">Indonesia</collectingCountry>
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(
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<figureCitation id="11DE2BDBFFE7BB3B32797059FC6FF9C9" box="[839,916,1633,1657]" captionStart="Figs 1-3" captionStartId="4.[151,192,2024,2046]" captionTargetBox="[151,1436,1374,1975]" captionTargetId="figure-659@4.[151,1436,1366,1975]" captionTargetPageId="4" captionText="Figs 1-3. Head, dorsal view. (1) Syleter paradoxus (Putzeys, 1868).(2) Parasyleter doriae (Putzeys, 1873) comb.nov.(3) Clypeuspinus validus (Andrewes, 1936) comb. nov." figureDoi="http://doi.org/10.5281/zenodo.5640137" httpUri="https://zenodo.org/record/5640137/files/figure.png" pageId="13" pageNumber="369">Figs 2</figureCitation>
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,
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<figureCitation id="11DE2BDBFFE7BB3B32987059FC4EF9C9" box="[934,949,1633,1657]" captionStart="Figs 4-6" captionStartId="5.[151,192,939,961]" captionTargetBox="[158,749,264,900]" captionTargetId="figure-983@5.[151,757,257,914]" captionTargetPageId="5" captionText="Figs 4-6. Right mesotibia. (4) Syleter paradoxus (Putzeys, 1868) showing the small preapical protuberance and its seta inserted apically (arrow). (5) Parasyleter doriae (Putzeys, 1873) comb. nov. with no tubercles at all. (6) Clypeuspinus validus (Andrewes, 1936) comb. nov. showing the long protuberance and its seta inserted laterally (arrow)." figureDoi="http://doi.org/10.5281/zenodo.5640139" httpUri="https://zenodo.org/record/5640139/files/figure.png" pageId="13" pageNumber="369">5</figureCitation>
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,
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<figureCitation id="11DE2BDBFFE7BB3B32F67059FBBEF9C9" box="[968,1093,1633,1657]" captionStart="Figs 28-30" captionStartId="14.[151,192,994,1016]" captionTargetBox="[175,1406,262,941]" captionTargetId="figure-485@14.[151,1439,241,968]" captionTargetPageId="14" captionText="Figs 28-30. Parasyleter doriae (Putzeys, 1873) comb. nov., specimen from Sarawak collected by G. Doria. (28) Habitus, male. (29) Male genitalia, ventral view (above) and lateral view (below) of aedeagus with parameres in between. (30) Female coxostyli and laterotergite." figureDoi="http://doi.org/10.5281/zenodo.5640146" httpUri="https://zenodo.org/record/5640146/files/figure.png" pageId="13" pageNumber="369">28, 29, 30</figureCitation>
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),
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<taxonomicName id="4EE54CDDFFE7BB3B35617059FC8DF929" authority="Andrewes, 1927" authorityName="Andrewes" authorityYear="1927" class="Insecta" family="Carabidae" genus="Psilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="369" phylum="Arthropoda" rank="species" species="malayicus">
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<emphasis id="BB91EB4CFFE7BB3B35617059FAE7F9C9" box="[1119,1308,1633,1657]" italics="true" pageId="13" pageNumber="369">Psilus malayicus</emphasis>
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<bibRefCitation id="ED744AAFFFE7BB3B34167059FC8DF929" author="Andrewes H. E." pageId="13" pageNumber="369" pagination="263 - 272" refId="ref13362" refString="Andrewes H. E. 1927. Papers on Oriental Carabidae. - XX. The Annals and Magazine of Natural History, including Zoology, Botany, and Geology, Ninth Series 20: 263 - 272." type="book chapter" year="1927">Andrewes, 1927</bibRefCitation>
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</taxonomicName>
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distributed in
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<collectingCountry id="F1F277CEFFE7BB3B351970B9FB76F929" box="[1063,1165,1665,1689]" name="Malaysia" pageId="13" pageNumber="369">Malaysia</collectingCountry>
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, and
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<taxonomicName id="4EE54CDDFFE7BB3B35ED70B9FC03F90A" authority="Andrewes, 1923" authorityName="Andrewes" authorityYear="1923" class="Insecta" family="Carabidae" genus="Psilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="369" phylum="Arthropoda" rank="species" species="porphyreus">
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<emphasis id="BB91EB4CFFE7BB3B35ED70B9FA67F929" box="[1235,1436,1665,1689]" italics="true" pageId="13" pageNumber="369">Psilus porphyreus</emphasis>
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<bibRefCitation id="ED744AAFFFE7BB3B327E709AFC03F90A" author="Andrewes H. E." box="[832,1016,1698,1722]" pageId="13" pageNumber="369" pagination="223 - 251" refId="ref13311" refString="Andrewes H. E. 1923. Descriptions of some new Carabidae from Ceylon. Spolia Zeylanica 12: 223 - 251." type="journal article" year="1923">Andrewes, 1923</bibRefCitation>
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</taxonomicName>
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known from
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<collectingCountry id="F1F277CEFFE7BB3B359E709AFAEEF90A" box="[1184,1301,1698,1722]" name="Sri Lanka" pageId="13" pageNumber="369">Sri Lanka</collectingCountry>
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. The genus includes also species not described yet and is currently in revision by the author.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |