147 lines
19 KiB
XML
147 lines
19 KiB
XML
<document id="0BE068055A4BA7998EE5CA58489B6007" ID-CLB-Dataset="3695" ID-DOI="10.35929/RSZ.0050" ID-GBIF-Dataset="60a46419-793d-4cae-a0bc-a086fc235849" ID-ISSN="0035-418" ID-Zenodo-Dep="5640135" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635893751209" checkinUser="felipe" docAuthor="Balkenohl, Michael" docDate="2021" docId="014C8648FFE7BB3832F9717CFC85F8DD" docLanguage="en" docName="RevSuisseZool.128.2.357-374.pdf" docOrigin="Revue suisse de Zoologie 128 (2)" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Clypeuspinus Balkenohl 2021, gen. nov." docType="treatment" docVersion="7" lastPageNumber="370" masterDocId="FD75FE30FFEABB36313E7638FFFBFFB0" masterDocTitle="Revisiting the Oriental genus Syleter Andrewes, 1941 (Coleoptera: Carabidae: Clivinini)" masterLastPageNumber="374" masterPageNumber="357" pageNumber="369" updateTime="1699467467588" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4A134E3BA2F46AD26D9E7D75D003D01C">Revisiting the Oriental genus Syleter Andrewes, 1941 (Coleoptera: Carabidae: Clivinini)</mods:title>
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<mods:namePart id="E9EC14062B42CD8299E06FE39D67CC58">Balkenohl, Michael</mods:namePart>
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<mods:affiliation id="457D50BF261FEFF95E30BCB46F49AD00">Ligusterweg 9, CH- 8906 Bonstetten, Switzerland</mods:affiliation>
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<mods:nameIdentifier id="2A0F708BEE78C5248D56A72C0C1DE181" type="email">mike.balkenohl@bluewin.ch</mods:nameIdentifier>
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<mods:title id="296F4982D7C891FEF00E55EF1B71F605">Revue suisse de Zoologie</mods:title>
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<mods:date id="E2079ADFAB9144C65B1A7930A5978A05">2021</mods:date>
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<treatment id="014C8648FFE7BB3832F9717CFC85F8DD" ID-DOI="http://doi.org/10.5281/zenodo.5646418" ID-GBIF-Taxon="189991358" ID-Zenodo-Dep="5646418" LSID="urn:lsid:plazi:treatment:014C8648FFE7BB3832F9717CFC85F8DD" httpUri="http://treatment.plazi.org/id/014C8648FFE7BB3832F9717CFC85F8DD" lastPageId="14" lastPageNumber="370" pageId="13" pageNumber="369">
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<subSubSection id="C1FF64D5FFE7BB3B32F9717CFAEEF8EC" box="[967,1301,1860,1884]" pageId="13" pageNumber="369" type="nomenclature">
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<paragraph id="895A375EFFE7BB3B32F9717CFAEEF8EC" blockId="13.[967,1301,1860,1916]" box="[967,1301,1860,1884]" pageId="13" pageNumber="369">
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<heading id="D2128032FFE7BB3B32F9717CFAEEF8EC" bold="true" box="[967,1301,1860,1884]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="369" reason="2">
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<emphasis id="BB91EB4CFFE7BB3B32F9717CFAEEF8EC" bold="true" box="[967,1301,1860,1884]" pageId="13" pageNumber="369">
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Genus
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<taxonomicName id="4EE54CDDFFE7BB3B3528717CFB51F8EC" authority="Balkenohl, 2021" authorityName="Balkenohl" authorityYear="2021" box="[1046,1194,1860,1884]" class="Insecta" family="Carabidae" genus="Clypeuspinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="369" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="BB91EB4CFFE7BB3B3528717CFB51F8EC" bold="true" box="[1046,1194,1860,1884]" italics="true" pageId="13" pageNumber="369">Clypeuspinus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A0A25637FFE7BB3B358F717CFAEEF8EC" box="[1201,1301,1860,1884]" pageId="13" pageNumber="369" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C1FF64D5FFE7BB3B3526715CFB3EF8CC" box="[1048,1221,1892,1916]" pageId="13" pageNumber="369" type="description">
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<paragraph id="895A375EFFE7BB3B3526715CFB3EF8CC" blockId="13.[967,1301,1860,1916]" box="[1048,1221,1892,1916]" pageId="13" pageNumber="369">
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<figureCitation id="11DE2BDBFFE7BB3B3526715CFBA0F8CC" box="[1048,1115,1892,1916]" captionStart="Figs 1-3" captionStartId="4.[151,192,2024,2046]" captionTargetBox="[151,1436,1374,1975]" captionTargetId="figure-659@4.[151,1436,1366,1975]" captionTargetPageId="4" captionText="Figs 1-3. Head, dorsal view. (1) Syleter paradoxus (Putzeys, 1868).(2) Parasyleter doriae (Putzeys, 1873) comb.nov.(3) Clypeuspinus validus (Andrewes, 1936) comb. nov." figureDoi="http://doi.org/10.5281/zenodo.5640137" httpUri="https://zenodo.org/record/5640137/files/figure.png" pageId="13" pageNumber="369">Figs 3</figureCitation>
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,
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<figureCitation id="11DE2BDBFFE7BB3B3559715CFB8DF8CC" box="[1127,1142,1892,1916]" captionStart="Figs 4-6" captionStartId="5.[151,192,939,961]" captionTargetBox="[158,749,264,900]" captionTargetId="figure-983@5.[151,757,257,914]" captionTargetPageId="5" captionText="Figs 4-6. Right mesotibia. (4) Syleter paradoxus (Putzeys, 1868) showing the small preapical protuberance and its seta inserted apically (arrow). (5) Parasyleter doriae (Putzeys, 1873) comb. nov. with no tubercles at all. (6) Clypeuspinus validus (Andrewes, 1936) comb. nov. showing the long protuberance and its seta inserted laterally (arrow)." figureDoi="http://doi.org/10.5281/zenodo.5640139" httpUri="https://zenodo.org/record/5640139/files/figure.png" pageId="13" pageNumber="369">6</figureCitation>
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,
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<figureCitation id="11DE2BDBFFE7BB3B35BC715CFB3EF8CC" box="[1154,1221,1892,1916]" captionStart="Figs 31-32" captionStartId="15.[151,192,2054,2076]" captionTargetBox="[244,1330,1086,2014]" captionTargetId="figure-342@15.[151,1439,1076,2041]" captionTargetPageId="15" captionText="Figs 31-32. Clypeuspinus validus (Andrewes, 1936) comb. nov., holotype. (31) Habitus. (32) Female coxostyli with laterotergite." figureDoi="http://doi.org/10.5281/zenodo.5640150" httpUri="https://zenodo.org/record/5640150/files/figure.png" pageId="13" pageNumber="369">31, 32</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C1FF64D5FFE7BB3B327E71ACFA9FF81C" box="[832,1380,1940,1964]" pageId="13" pageNumber="369" type="type_taxon">
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<paragraph id="895A375EFFE7BB3B327E71ACFA9FF81C" blockId="13.[832,1380,1940,1964]" box="[832,1380,1940,1964]" pageId="13" pageNumber="369">
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<emphasis id="BB91EB4CFFE7BB3B327E71ACFC2DF81C" bold="true" box="[832,982,1940,1964]" pageId="13" pageNumber="369">
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<typeStatus id="565E89FCFFE7BB3B327E71ACFC83F81C" box="[832,888,1940,1964]" pageId="13" pageNumber="369">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4EE54CDDFFE7BB3B32E271ACFAA4F81C" ID-CoL="4PJY3" authority="Andrewes, 1936: 202" authorityName="Andrewes" authorityPageNumber="202" authorityYear="1936" box="[988,1375,1940,1964]" class="Insecta" family="Carabidae" genus="Psilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="369" phylum="Arthropoda" rank="species" species="validus">
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<emphasis id="BB91EB4CFFE7BB3B32E271ACFB8AF81C" box="[988,1137,1940,1964]" italics="true" pageId="13" pageNumber="369">Psilus validus</emphasis>
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<bibRefCitation id="ED744AAFFFE7BB3B354671ACFAA4F81C" author="Andrewes H. E." box="[1144,1375,1940,1964]" pageId="13" pageNumber="369" pagination="201 - 209" refId="ref13467" refString="Andrewes H. E. 1936. Descriptions of some new Carabidae from Ceylon (Part III). Spolia Zeylanica 19: 201 - 209." type="journal article" year="1936">Andrewes, 1936: 202</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C1FF64D5FFE7BB38327E71FCFB3BF90A" lastPageId="14" lastPageNumber="370" pageId="13" pageNumber="369" type="diagnosis">
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<paragraph id="895A375EFFE7BB38327E71FCFF33F7AC" blockId="13.[832,1437,1988,2077]" lastBlockId="14.[151,756,1141,2076]" lastPageId="14" lastPageNumber="370" pageId="13" pageNumber="369">
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<emphasis id="BB91EB4CFFE7BB3B327E71FCFBF8F86C" bold="true" box="[832,1027,1988,2012]" pageId="13" pageNumber="369">Genus diagnosis:</emphasis>
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<emphasis id="BB91EB4CFFE7BB3B353571FCFBCDF86B" box="[1035,1078,1988,2011]" italics="true" pageId="13" pageNumber="369">Size</emphasis>
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<quantity id="4E1D9ABBFFE7BB3B350171FCFB6DF86B" box="[1087,1174,1988,2012]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="13" pageNumber="369" unit="mm" value="4.8">4.8 mm</quantity>
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.
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<emphasis id="BB91EB4CFFE7BB3B359C71FCFB27F86C" box="[1186,1244,1988,2012]" italics="true" pageId="13" pageNumber="369">Head</emphasis>
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with clypeus and frons conspicuously elongated, convex; clypeus, clypeal wing, and supraantennal plate separated by obtuse notch; clypeus straight and sloping anteriorly, clypeal wing conspicuously projecting anteriorly; mandible distinctly elongated, narrow in apical third, laterally convex, regularly convex in apical half, bent dorsally; labrum five-setose; maxillary palpomere elongated, slightly securiform; antenna with scapus and pedicellus eccentrically attached; eye relatively flat but not reduced, not projecting laterally, gena as long as eye.
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<collectingRegion id="4B21F9BCFFE4BB3831A97343FF09FA22" box="[151,242,1403,1426]" country="Myanmar" name="Chin" pageId="14" pageNumber="370">Mentum</collectingRegion>
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moderately elongated, with lateral lobe wide and convex, median tooth obtuse angled, projecting as far as lateral lobe, surface with rough longitudinal rugae.
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<emphasis id="BB91EB4CFFE4BB3831DB73E4FEB4FA43" box="[229,335,1500,1523]" italics="true" pageId="14" pageNumber="370">Pronotum</emphasis>
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with pentagonal longitudinal outline, with reflexed lateral margin complete, running from anterior angle up to basal constriction, posterior angle with moderately acute tooth projecting laterally, lateral channel distinctly widened between the two lateral setigerous punctures, median line wide, joining broadly basal constriction, proepisternum invisible in dorsal view, flange at base carinate, as wide as basal constriction.
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<emphasis id="BB91EB4CFFE4BB38301770D8FE82F948" box="[297,377,1760,1784]" italics="true" pageId="14" pageNumber="370">Elytron</emphasis>
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with interval eight carinate from humerus to apex; lateral channel of equal width from humerus to apex; third interval with three (left side) respectively four (right side) setigerous punctures.
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<emphasis id="BB91EB4CFFE4BB3831A9715AFEEDF8CA" box="[151,278,1890,1914]" italics="true" pageId="14" pageNumber="370">Hind wings</emphasis>
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fully developed.
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<emphasis id="BB91EB4CFFE4BB3830E2715BFDF4F8CA" box="[476,527,1891,1914]" italics="true" pageId="14" pageNumber="370">Legs</emphasis>
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with tarsomeres not widened (
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<specimenCount id="9FE3FCD7FFE4BB38303F71BAFE80F82A" box="[257,379,1922,1946]" pageId="14" pageNumber="370" type="female">one female</specimenCount>
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investigated); mesotibia with long protuberance preapically furnished with seta.
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<emphasis id="BB91EB4CFFE4BB383383719BFF0CF86B" italics="true" pageId="14" pageNumber="370">Male genitalia</emphasis>
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unknown.
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<emphasis id="BB91EB4CFFE4BB38304571FBFDC3F86B" box="[379,568,1987,2011]" italics="true" pageId="14" pageNumber="370">Female genitalia</emphasis>
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with coxostylus one and two nearly completely fused, without cone-like seta.
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</paragraph>
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<caption id="DD9A67D6FFE4BB3831A975DAFDE6FB82" ID-DOI="http://doi.org/10.5281/zenodo.5640146" ID-Zenodo-Dep="5640146" httpUri="https://zenodo.org/record/5640146/files/figure.png" pageId="14" pageNumber="370" startId="14.[151,192,994,1016]" targetBox="[175,1406,262,941]" targetPageId="14">
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<paragraph id="895A375EFFE4BB3831A975DAFDE6FB82" blockId="14.[151,1437,994,1074]" pageId="14" pageNumber="370">
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Figs 28-30.
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<taxonomicName id="4EE54CDDFFE4BB38302275DAFD8CFC48" authority="(Putzeys, 1873)" authorityName="Balkenohl" authorityYear="2021" baseAuthorityName="Putzeys" baseAuthorityYear="1873" box="[284,631,994,1016]" class="Insecta" family="Carabidae" genus="Parasyleter" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="370" phylum="Arthropoda" rank="species" species="doriae" status="comb. nov.">
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<emphasis id="BB91EB4CFFE4BB38302275DAFE2FFC48" box="[284,468,994,1016]" italics="true" pageId="14" pageNumber="370">Parasyleter doriae</emphasis>
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(
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<bibRefCitation id="ED744AAFFFE4BB3830DA75DAFD8BFC48" author="Putzeys J. A. A. H." box="[484,624,994,1016]" pageId="14" pageNumber="370" pagination="10 - 18" refId="ref14650" refString="Putzeys J. A. A. H. 1873. Deuxieme supplement a la revision generale des Clivinides. Annales de la Societe Entomologique de Belgique 16: 10 - 18." type="journal article" year="1873">Putzeys, 1873</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A0A25637FFE4BB3833BE75DAFD0BFC48" box="[640,752,994,1016]" pageId="14" pageNumber="370" rank="species">comb. nov.</taxonomicNameLabel>
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, specimen from Sarawak collected by G. Doria. (28) Habitus, male. (29) Male genitalia, ventral view (above) and lateral view (below) of aedeagus with parameres in between. (30) Female coxostyli and laterotergite.
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</paragraph>
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</caption>
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<paragraph id="895A375EFFE4BB38327E724EFB3BF90A" blockId="14.[832,1437,1141,1723]" pageId="14" pageNumber="370">
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<emphasis id="BB91EB4CFFE4BB38327E724EFBC7FB3E" bold="true" box="[832,1084,1142,1166]" pageId="14" pageNumber="370">Differential diagnosis:</emphasis>
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See key to the genera. However, the most similar species are
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<taxonomicName id="4EE54CDDFFE4BB3835A472AEFC05FB7F" authority="Basilewsky, 1948" authorityName="Basilewsky" authorityYear="1948" class="Insecta" family="Carabidae" genus="Basilewskyana" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="370" phylum="Arthropoda" rank="species" species="villiersi">
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<emphasis id="BB91EB4CFFE4BB3835A472AEFA67FB1E" box="[1178,1436,1174,1198]" italics="true" pageId="14" pageNumber="370">Basilewskyana villiersi</emphasis>
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<bibRefCitation id="ED744AAFFFE4BB38327E728FFC05FB7F" author="Basilewsky P." box="[832,1022,1207,1231]" pageId="14" pageNumber="370" pagination="57 - 105" refId="ref13897" refString="Basilewsky P. 1948. Carabidae africains nouveaux. I. Memoires de la Societe entomologique de Belgique 25: 57 - 105, plus 1 Tab." type="journal article" year="1948">Basilewsky, 1948</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4EE54CDDFFE4BB3835097280FADDFB7F" authority="Kult, 1959" authorityName="Kult" authorityYear="1959" box="[1079,1318,1207,1231]" class="Insecta" family="Carabidae" genus="Basilewskyana" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="370" phylum="Arthropoda" rank="species" species="brittoni">
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<emphasis id="BB91EB4CFFE4BB3835097280FB53FB7F" box="[1079,1192,1207,1231]" italics="true" pageId="14" pageNumber="370">B. brittoni</emphasis>
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<bibRefCitation id="ED744AAFFFE4BB38358F728FFADDFB7F" author="Kult K." box="[1201,1318,1207,1231]" pageId="14" pageNumber="370" pagination="193 - 225" refId="ref14268" refString="Kult K. 1959. Revision of the African Species of the old genus Clivina Latr. (Col. Carabidae). Revue de Zoologie et de Botanique Africaines 60: 193 - 225." type="journal article" year="1959">Kult, 1959</bibRefCitation>
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</taxonomicName>
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both from West Africa. These two species are completely black and much larger (7.2 and
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<quantity id="4E1D9ABBFFE4BB38355072C0FB32FAA0" box="[1134,1225,1272,1296]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="14" pageNumber="370" unit="mm" value="7.8">7.8 mm</quantity>
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). In addition, they differ mainly by a cylindrical shape, the clypeal wings slightly projecting, a deep facial furrow with distinct facial carina, without any clypeal suture, a wider labrum, the elytron with missing humeral tooth, more convex intervals, and the elytral setigerous punctures situated in the middle of the third interval. Members of the American genus
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<taxonomicName id="4EE54CDDFFE4BB38352573E5FAD1FA45" authority="Putzeys, 1846" authorityName="Putzeys" authorityYear="1846" box="[1051,1322,1501,1526]" class="Insecta" family="Carabidae" genus="Ardistomis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="370" phylum="Arthropoda" rank="genus">
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<emphasis id="BB91EB4CFFE4BB38352573E5FB75FA45" box="[1051,1166,1501,1525]" italics="true" pageId="14" pageNumber="370">Ardistomis</emphasis>
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<bibRefCitation id="ED744AAFFFE4BB3835AB73E6FAD1FA45" author="Putzeys J. A. A. H." box="[1173,1322,1501,1526]" pageId="14" pageNumber="370" pagination="521 - 663" refId="ref14539" refString="Putzeys J. A. A. H. 1846. Monographie des Clivina et genres voisins, precedee d'un tableau synoptique des genres de la tribu des Scaritides. Memoires de la Societe Royale des Sciences de Li e ge 2: 521 - 663." type="journal article" year="1846">Putzeys, 1846</bibRefCitation>
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</taxonomicName>
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also show a considerable elongation of the anterior part of the head and their mandibles are evidently elongated. But the maxillary palpomeres and the widened tarsomeres of the front and intermediate legs are very similar to
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<taxonomicName id="4EE54CDDFFE4BB3834757058FA6EF9C8" authorityName="Balkenohl" authorityYear="2021" box="[1355,1429,1632,1656]" class="Insecta" family="Carabidae" genus="Syleter" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="370" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BB91EB4CFFE4BB3834757058FA6EF9C8" box="[1355,1429,1632,1656]" italics="true" pageId="14" pageNumber="370">Syleter</emphasis>
|
||
</taxonomicName>
|
||
. However, the pronotum, elytron and the plica at the apex of the elytron is very different.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C1FF64D5FFE4BB38327E70EAFC85F8DD" pageId="14" pageNumber="370" type="etymology">
|
||
<paragraph id="895A375EFFE4BB38327E70EAFC85F8DD" blockId="14.[832,1437,1746,1901]" pageId="14" pageNumber="370">
|
||
<emphasis id="BB91EB4CFFE4BB38327E70EAFC3FF95A" bold="true" box="[832,964,1746,1770]" pageId="14" pageNumber="370">Etymology:</emphasis>
|
||
The name is a combination of two nouns, clypeus and spinus. Clypeus points to the conspicuously lobate-like projecting clypeal wings. Spinus highlights the long protuberance at the mesotibia (Latin: spinus – spur).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |