treatments-xml/data/5F/32/E1/5F32E155187AE727B82545CC3A6F69C2.xml

<document ID-DOI="http://dx.doi.org/10.3897/zse.94.26859" ID-GBIF-Dataset="30d1d279-f033-4868-b0c9-925a406be2c5" ID-GBIF-Taxon="182236654" ID-Pensoft-Pub="1860-0743-2-369" ID-Pensoft-UUID="83D45F0665FE5C729C47210ACD710944" ID-Zenodo-Dep="1344764" ID-ZooBank="C354D7DF198F4D4FA2A8F763FD03018D" ModsDocID="1860-0743-94-2-369" checkinTime="1621423395062" checkinUser="pensoft" docAuthor="Song, Zhi-Shun, Malenovsky, Igor, Chen, Jian-Qin, Deckert, Juergen &amp; Liang, Ai-Ping" docDate="2018" docId="5F32E155187AE727B82545CC3A6F69C2" docLanguage="en" docName="ZoosystEvol 94(2): 369-391" docOrigin="Zoosystematics and Evolution 94 (2)" docPubDate="2018-08-06" docSource="http://dx.doi.org/10.3897/zse.94.26859" docTitle="Orthopagus exoletus , comb. n. 2018, stat. rev." docType="treatment" docVersion="3" id="83D45F0665FE5C729C47210ACD710944" lastPageId="4" lastPageNumber="373" masterDocId="83D45F0665FE5C729C47210ACD710944" masterDocTitle="Taxonomic review of the planthopper genus Orthopagus (Hemiptera, Fulgoromorpha, Dictyopharidae), with descriptions of two new species" masterLastPageNumber="391" masterPageNumber="369" pageId="2" pageNumber="371" updateTime="1632016297512" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic review of the planthopper genus Orthopagus (Hemiptera, Fulgoromorpha, Dictyopharidae), with descriptions of two new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Song, Zhi-Shun</mods:namePart>
<mods:affiliation>Jiangsu Key Laboratory of Biofunctional Molecules, School of Life Sciences, Chemistry &amp; Chemical Engineering, Ji &amp; angsu Second Normal University, Nanjing 210013, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Malenovsky, Igor</mods:namePart>
<mods:affiliation>Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar</mods:affiliation>
<mods:nameIdentifier type="email">imalenovsky@mzm.cz</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Jian-Qin</mods:namePart>
<mods:affiliation>Jiangsu Key Laboratory of Biofunctional Molecules, School of Life Sciences, Chemistry &amp; Chemical Engineering, Ji</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deckert, Juergen</mods:namePart>
<mods:affiliation>Department of Botany and Zoology, Faculty of Science, Masaryk University, Kotlarska 2, 611 37 Brno, Czech Republic</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Ai-Ping</mods:namePart>
<mods:affiliation>Museum fuer Naturkunde, Leibniz Institute for Research on Evolution and Biodiversity Science, Invalidenstrasse 43, Berlin 10115, Germany &amp; Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="email">liangap@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zoosystematics and Evolution</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-08-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>94</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:extent unit="page">
<mods:start>369</mods:start>
<mods:end>391</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zse.94.26859</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.94.26859</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-2-369</mods:identifier>
<mods:identifier type="ZooBank">C354D7DF198F4D4FA2A8F763FD03018D</mods:identifier>
<mods:identifier type="Pensoft-UUID">83D45F0665FE5C729C47210ACD710944</mods:identifier>
<mods:identifier type="Zenodo-Dep">1344764</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182236654" LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2" httpUri="http://treatment.plazi.org/id/5F32E155187AE727B82545CC3A6F69C2" lastPageId="4" lastPageNumber="373" pageId="2" pageNumber="371">
<subSubSection pageId="2" pageNumber="371" type="nomenclature">
<paragraph pageId="2" pageNumber="371">
<taxonomicName authority="(Melichar, 1903), comb. n." authorityName=", comb. n." authorityYear="2018" baseAuthorityName="Melichar" baseAuthorityYear="1903" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus exoletus" order="Hemiptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="exoletus" status="stat. rev.">Orthopagus exoletus (Melichar, 1903), comb. n.</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="371">stat. rev.</taxonomicNameLabel>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Orthopagus species. (A, B) O. bartletti sp. n., male, holotype, Karnataka, India; (C, D) O. exoletus (Melichar), male and female, Sri Lanka; (E, F) O. hainanensis sp. n., male and female, paratypes, Hainan, China; (G, H) O. lunulifer Uhler, male and female, China; (I, J) O. philippinus Melichar, male and female, Philippines; (K, L) O. splendens (Germar), male and female, China." figureDoi="10.3897/zse.94.26859.figure2" httpUri="https://binary.pensoft.net/fig/221398" pageId="2" pageNumber="371">Figs 2C-D</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Forewing of Orthopagus species. (A) O. bartletti sp. n.; (B) O. exoletus (Melichar); (C) O. hainanensis sp. n.; (D) O. lunulifer Uhler; (E) O. philippinus Melichar; (F) O. splendens (Germar)." figureDoi="10.3897/zse.94.26859.figure3" httpUri="https://binary.pensoft.net/fig/221399" pageId="2" pageNumber="371">, 3B</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="2" pageNumber="371">, 5A-I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="372" type="reference_group">
<paragraph pageId="3" pageNumber="372">
<taxonomicName authority="Melichar, 1903" authorityName="Song &amp; Malenovský &amp; Chen &amp; Deckert &amp; Liang" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Udugama exoleta" order="Hemiptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="exoleta" status="stat. rev.">
<pageBreakToken pageId="3" pageNumber="372" start="start">Udugama</pageBreakToken>
exoleta
</taxonomicName>
Melichar, 1903: 28, Pl. I, figs 7, 7a-b. Syntypes: 5 females, Moruwale, Sri Lanka (not examined). Synonymized under 
<taxonomicName authority="Melichar, 1903" authorityName="Song &amp; Malenovský &amp; Chen &amp; Deckert &amp; Liang" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Udugama splendens" order="Hemiptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="splendens" status="stat. rev.">Udugama splendens</taxonomicName>
(Germar, 1830) by 
<bibRefCitation author="Distant, WL" journalOrPublisher="Taylor &amp; Francis, London" pageId="12" pageNumber="381" refId="B6" refString="Distant, WL, 1906. The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera). Taylor &amp; Francis, London" title="The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera)." year="1906">Distant 1906</bibRefCitation>
: 249.
</paragraph>
<paragraph pageId="3" pageNumber="372">
<taxonomicName authority="Melichar, 1903" authorityName="Song &amp; Malenovský &amp; Chen &amp; Deckert &amp; Liang" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Udugama exoleta" order="Hemiptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="exoleta" status="stat. rev.">Udugama exoleta</taxonomicName>
: 
<bibRefCitation author="Kirkaldy, GW" journalOrPublisher="Annales de la Societe entomologique de Belgique Bruxelles" pageId="12" pageNumber="381" pagination="9 - 14" refId="B12" refString="Kirkaldy, GW, 1908. Notes on some Sinhalese Rhynchota. Annales de la Societe entomologique de Belgique Bruxelles 52: 9 - 14" title="Notes on some Sinhalese Rhynchota." volume="52" year="1908">Kirkaldy 1908</bibRefCitation>
: 14.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="372" type="materials_examined">
<paragraph pageId="3" pageNumber="372">Material examined.</paragraph>
<paragraph pageId="3" pageNumber="372">
<materialsCitation collectingDate="2018-01-01" collectingDateMax="2018-12-31" collectingDateMin="2018-01-01" collectorName="T. R. Bell" country="INDIA" location="INDIA" specimenCount="2" specimenCount-female="2">
<emphasis bold="true" pageId="3" pageNumber="372">
<collectingCountry name="India">INDIA</collectingCountry>
</emphasis>
: 
<specimenCount type="female">2 females</specimenCount>
, [no state indicated], 1934-394, 
<collectorName>T. R. Bell</collectorName>
leg. (BMNH)
</materialsCitation>
; 
<materialsCitation collectingDate="1907-10-03" collectorName="Distant" country="India" location="Calcutta" specimenCount="1" specimenCount-female="1" stateProvince="West Bengal">
<collectingRegion country="India" name="West Bengal">West Bengal</collectingRegion>
: 
<specimenCount type="female">1 female</specimenCount>
, 
<location LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:3DB4C01D641E74FF1BA04872391CA973" country="India" name="Calcutta" stateProvince="West Bengal">Calcutta</location>
[= Kolkata], 
<collectingDate value="1907-10-03">3.x.1907</collectingDate>
(coll. 
<collectorName>Distant</collectorName>
, BMNH)
</materialsCitation>
; 
<materialsCitation collectingDate="2005-09-28" collectingDateMax="2005-10-02" collectingDateMin="2005-09-28" collectorName="C. R. Bartlett" country="India" county="Sindhudurg district" elevation="730" latitude="18.443611" location="Tamini village" longLatPrecision="21" longitude="73.42944" municipality="Pune district" specimenCount="2" specimenCount-male="2" stateProvince="Maharashtra">
<collectingRegion country="India" name="Maharashtra">Maharashtra</collectingRegion>
: 
<specimenCount type="male">1 male</specimenCount>
, 
<collectingCounty>Sindhudurg district</collectingCounty>
, roadside on ridge 
<locationDeviation location="Amboli">
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
W Amboli
</locationDeviation>
, 
<geoCoordinate degrees="15" direction="north" minutes="58" orientation="latitude" precision="15" seconds="04" value="15.967777">15°58'04&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="73" direction="east" minutes="59" orientation="longitude" precision="15" seconds="23" value="73.989716">73°59'23&quot;E</geoCoordinate>
, alt. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.296912" unit="ft" value="2394.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.296912" unit="ft" value="2394.0">2394 ft</elevation>
</quantity>
, pasture and successional, moist deciduous forest, 
<collectingDate value="2005-09-28">28.ix.2005</collectingDate>
, 
<collectorName>C. R. Bartlett</collectorName>
leg. (UDCC); 
<specimenCount type="male">1 male</specimenCount>
, 
<collectingMunicipality>Pune district</collectingMunicipality>
, 
<locationDeviation location="Mulshi Lake">
<quantity metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
E Mulshi Lake
</locationDeviation>
near 
<location LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:0E0AF540AA13EB925D876EA27FD74657" country="India" county="Sindhudurg district" latitude="18.443611" longLatPrecision="21" longitude="73.42944" municipality="Pune district" name="Tamini village" stateProvince="Maharashtra">Tamini village</location>
, 
<geoCoordinate degrees="18" direction="north" minutes="26" orientation="latitude" precision="15" seconds="37" value="18.443611">18°26'37&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="73" direction="east" minutes="25" orientation="longitude" precision="15" seconds="46" value="73.42944">73°25'46&quot;E</geoCoordinate>
, alt. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.239256" unit="ft" value="2047.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.239256" unit="ft" value="2047.0">2047 ft</elevation>
</quantity>
, deciduous forest and open areas, 
<collectingDate value="2005-10-02">2.x.2005</collectingDate>
, 
<collectorName>C. R. Bartlett</collectorName>
(UDCC)
</materialsCitation>
; 
<materialsCitation collectingDate="2005-09-24" collectingDateMax="2005-09-25" collectingDateMin="2005-09-24" collectorName="C. R. Bartlett" country="India" elevation="104" latitude="15.378612" location="Bhagwan Mahaveer Sanctuary" longLatPrecision="21" longitude="74.23111" municipality="Sanguem district" specimenCount="2" specimenCount-female="2" stateProvince="Goa">
<collectingRegion country="India" name="Goa">Goa</collectingRegion>
: 
<specimenCount type="female">2 females</specimenCount>
, 
<collectingMunicipality>Sanguem district</collectingMunicipality>
, near 
<location LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:B9F6385924AEDFABD5552A72E9594C4D" country="India" latitude="15.378612" longLatPrecision="21" longitude="74.23111" municipality="Sanguem district" name="Bhagwan Mahaveer Sanctuary" stateProvince="Goa">Bhagwan Mahaveer Sanctuary</location>
<locationDeviation location="Molem">
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</quantity>
E Molem
</locationDeviation>
, 
<geoCoordinate degrees="15" direction="north" minutes="22" orientation="latitude" precision="15" seconds="43" value="15.378612">15°22'43&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="74" direction="east" minutes="13" orientation="longitude" precision="15" seconds="52" value="74.23111">74°13'52&quot;E</geoCoordinate>
, alt. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0424159999999998" unit="ft" value="342.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.0424159999999998" unit="ft" value="342.0">342 ft</elevation>
</quantity>
, moist deciduous forest, 
<collectingDate value="2005-09-24" valueMax="2005-09-25" valueMin="2005-09-24">24-25.ix.2005</collectingDate>
, 
<collectorName>C. R. Bartlett</collectorName>
leg. (UDCC)
</materialsCitation>
; 
<materialsCitation collectingDate="1914-09-16" collectingDateMax="1938-09-22" collectingDateMin="1914-09-16" collectorName="F. H. Gravely, Distant, Tenmalai, Thenmala" country="India" county="Malabar" elevation="61" location="Parambikulam" municipality="Nadungayam" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Kerala">
<collectingRegion country="India" name="Kerala">Kerala</collectingRegion>
: 
<specimenCount type="male">2 males</specimenCount>
, 
<collectingCounty>Malabar</collectingCounty>
, 
<collectingMunicipality>Nadungayam</collectingMunicipality>
[forest near Nilambur], 
<quantity metricMagnitude="1" metricUnit="m" metricValue="6.096" unit="ft" value="200.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="6.096" unit="ft" value="200.0">200 ft</elevation>
</quantity>
, 
<collectingDate value="1938-09-16" valueMax="1938-09-22" valueMin="1938-09-16">16.-22.ix.1938</collectingDate>
(BMNH); 
<specimenCount type="female">1 female</specimenCount>
, 
<location LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:EA752546781A4B03AAAE4974749660C5" country="India" county="Malabar" municipality="Nadungayam" name="Parambikulam" stateProvince="Kerala">Parambikulam</location>
, alt. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.4676" metricValueMax="9.7536" metricValueMin="5.1816" unit="ft" value="2450.0" valueMax="3200.0" valueMin="1700.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.4676" metricValueMax="9.7536" metricValueMin="5.1816" unit="ft" value="2450.0" valueMax="3200.0" valueMin="1700.0">1700-3200 ft</elevation>
</quantity>
, 
<collectingDate value="1914-09-16" valueMax="1914-09-24" valueMin="1914-09-16">16.-24.ix.1914</collectingDate>
, 
<collectorName>F. H. Gravely</collectorName>
leg. (coll. 
<collectorName>Distant</collectorName>
, BMNH); 
<specimenCount type="female">2 females</specimenCount>
, 
<collectorName>Tenmalai</collectorName>
[= 
<collectorName>Thenmala</collectorName>
], 
<collectingDate value="1937-05-12" valueMax="1937-05-15" valueMin="1937-05-12">12.-15.v.1937</collectingDate>
(BMNH)
</materialsCitation>
; 
<materialsCitation collectingDate="2002-05-03" collectingDateMax="2002-05-15" collectingDateMin="2002-05-03" collectorName="L. Dembicky" country="India" elevation="1100" location="Kunchappanai" municipality="Nilgiri Hills" specimenCount="1" specimenCount-female="1" stateProvince="Tamil Nadu">
<collectingRegion country="India" name="Tamil Nadu">Tamil Nadu</collectingRegion>
: 
<specimenCount type="female">1 female</specimenCount>
, 
<collectingMunicipality>Nilgiri Hills</collectingMunicipality>
, 
<locationDeviation location="Kotagiri">
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.1" unit="km" value="11.0">11 km</quantity>
SE Kotagiri
</locationDeviation>
, 
<location LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:83B62F08653406187A3E8D1770BB5334" country="India" municipality="Nilgiri Hills" name="Kunchappanai" stateProvince="Tamil Nadu">Kunchappanai</location>
, 11°24ʹN, 76°56ʹE, alt. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">
1100 
<normalizedToken originalValue="±">+/-</normalizedToken>
100 m
</elevation>
</quantity>
, 
<collectingDate value="2002-05-03" valueMax="2002-05-15" valueMin="2002-05-03">3.-15.v.2002</collectingDate>
, 
<collectorName>
L. 
<normalizedToken originalValue="Dembický">Dembicky</normalizedToken>
</collectorName>
leg. (MMBC)
</materialsCitation>
. 
<materialsCitation collectingDate="2018-01-01" collectingDateMax="2018-12-31" collectingDateMin="2018-01-01" collectorName="Melichar" country="SRI LANKA" location="Puttalam" municipality="Puttalam district" specimenCount="1" specimenCount-female="1" stateProvince="North Western Province">
<emphasis bold="true" pageId="3" pageNumber="372">
<collectingCountry name="Sri Lanka">SRI LANKA</collectingCountry>
</emphasis>
: 
<collectingRegion country="Sri Lanka" name="North Western">North Western Province</collectingRegion>
: 
<specimenCount type="female">1 female</specimenCount>
, 
<collectingMunicipality>Puttalam district</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:98C618000FDF276A3CE5108D2A7408FB" country="SRI LANKA" municipality="Puttalam district" name="Puttalam" stateProvince="North Western Province">Puttalam</location>
, 
<normalizedToken originalValue="“12.”">&quot;12.&quot;</normalizedToken>
[=?1912] (coll. 
<collectorName>Melichar</collectorName>
, MMBC)
</materialsCitation>
; 
<materialsCitation collectingDate="1970-03-10" collectingDateMax="1970-03-19" collectingDateMin="1970-03-10" collectorName="D. Davis, B. Rowe" country="Sri Lanka" county="Anuradnapura district" elevation="61" location="Hunuwilagama" municipality="Wilpattu National Wildlife Park" specimenCount="1" specimenCount-male="1" stateProvince="North Central Province">
<collectingRegion country="Sri Lanka" name="North Central">North Central Province</collectingRegion>
: 
<specimenCount type="male">1 male</specimenCount>
, 
<collectingCounty>Anuradnapura district</collectingCounty>
, 
<collectingMunicipality>Wilpattu National Wildlife Park</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:A31F36E8D5B7B903881939635780450A" country="Sri Lanka" county="Anuradnapura district" municipality="Wilpattu National Wildlife Park" name="Hunuwilagama" stateProvince="North Central Province">Hunuwilagama</location>
, 
<location LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:29112F908E0B495AC2435F947F7EB513" country="Sri Lanka" county="Anuradnapura district" municipality="Wilpattu National Wildlife Park" name="Wildlife Soc. Bungalow" stateProvince="North Central Province">Wildlife Soc. Bungalow</location>
, 
<quantity metricMagnitude="1" metricUnit="m" metricValue="6.096" unit="ft" value="200.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="6.096" unit="ft" value="200.0">200 ft</elevation>
</quantity>
, 
<collectingDate value="1970-03-10" valueMax="1970-03-19" valueMin="1970-03-10">10-19.iii.1970</collectingDate>
, 
<collectorName>D. Davis</collectorName>
&amp; 
<collectorName>B. Rowe</collectorName>
leg. (USNM)
</materialsCitation>
; 
<materialsCitation collectingDate="1970-01-31" collectingDateMax="1995-11-29" collectingDateMin="1970-01-31" collectorName="M. Schaffer, Samaragamuwa, D. Davis, B. Rowe" country="Sri Lanka" county="Kandy district" elevation="107" location="Ratnapura district" municipality="Talwatte" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Central Province">
<collectingRegion country="Sri Lanka" name="Central">Central Province</collectingRegion>
: 
<specimenCount type="female">1 female</specimenCount>
, 
<collectingCounty>Kandy district</collectingCounty>
, 
<collectingMunicipality>Talwatte</collectingMunicipality>
, 
<collectingDate value="1995-11-29">29.xi.1995</collectingDate>
, 
<collectorName>M. Schaffer</collectorName>
leg. (BMNH); 
<collectorName>Samaragamuwa</collectorName>
: 
<specimenCount type="male">1 male</specimenCount>
, 
<location LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:2E9E28A70AF361FD3F83CE34AA26F573" country="Sri Lanka" county="Kandy district" municipality="Talwatte" name="Ratnapura district" stateProvince="Central Province">Ratnapura district</location>
, 
<location LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:B6A2690BE3D4973D3EFB97AAF6D8FD37" country="Sri Lanka" county="Kandy district" municipality="Talwatte" name="Uggalkaltota" stateProvince="Central Province">Uggalkaltota</location>
, 
<location LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:30E27F8C6BDAC6C4F6620CC968DAF417" country="Sri Lanka" county="Kandy district" municipality="Talwatte" name="Irrigation Bungalow" stateProvince="Central Province">Irrigation Bungalow</location>
, alt. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.0668" unit="ft" value="350.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.0668" unit="ft" value="350.0">350 ft</elevation>
</quantity>
, 
<collectingDate value="1970-01-31" valueMax="1970-02-08" valueMin="1970-01-31">31.i-8.ii.1970</collectingDate>
, 
<collectorName>D. Davis</collectorName>
&amp; 
<collectorName>B. Rowe</collectorName>
leg. (USNM)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="372" type="redescription">
<paragraph pageId="3" pageNumber="372">Redescription.</paragraph>
<paragraph pageId="3" pageNumber="372">Measurements (2 males, 8 females). Body length (from apex of head to tip of forewings): male 11.1 mm, female 12.3-14.7 mm; head length (from apex of cephalic process to base of eyes): male 1.28-1.30 mm, female 1.28-1.40 mm; head width (including eyes): male 1.40-1.45 mm, female 1.48-1.60 mm; forewing length: male 9.0-9.4 mm, female 10.2-12.2 mm.</paragraph>
<paragraph pageId="3" pageNumber="372">
Coloration. General coloration as in generic description (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Orthopagus species. (A, B) O. bartletti sp. n., male, holotype, Karnataka, India; (C, D) O. exoletus (Melichar), male and female, Sri Lanka; (E, F) O. hainanensis sp. n., male and female, paratypes, Hainan, China; (G, H) O. lunulifer Uhler, male and female, China; (I, J) O. philippinus Melichar, male and female, Philippines; (K, L) O. splendens (Germar), male and female, China." figureDoi="10.3897/zse.94.26859.figure2" httpUri="https://binary.pensoft.net/fig/221398" pageId="3" pageNumber="372">2A</figureCitation>
). Vertex predominantly light ochraceous, with 3-4 pairs of small dark brown markings: an elongate patch on each side of midline apically, a small spot at each lateral keel medially, a roundish spot on each side of midline at basal third, and a small spot in each postero-lateral corner (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5A</figureCitation>
). Frons light ochraceous with small dark brown spots along intermediate and lateral carinae, frons base slightly infuscated (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5C</figureCitation>
). Forewing membrane pattern as in Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Forewing of Orthopagus species. (A) O. bartletti sp. n.; (B) O. exoletus (Melichar); (C) O. hainanensis sp. n.; (D) O. lunulifer Uhler; (E) O. philippinus Melichar; (F) O. splendens (Germar)." figureDoi="10.3897/zse.94.26859.figure3" httpUri="https://binary.pensoft.net/fig/221399" pageId="3" pageNumber="372">3B</figureCitation>
. Hind wing membrane with a relatively narrow dark brown streak along the apical portion of CuA1 vein, extending along hind wing apical margin in some specimens.
</paragraph>
<paragraph pageId="3" pageNumber="372">
Structure. Head with cephalic process very short, not inflated (Figs 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5A-C</figureCitation>
). Vertex (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5A</figureCitation>
) with ratio of length at midline to width between eyes 1.2-1.5. Transition of vertex to frons narrowly rounded in lateral view (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5B</figureCitation>
). Frons relatively broad, with ratio of length at midline to maximum width 2.3-2.7.
</paragraph>
<paragraph pageId="3" pageNumber="372">
Male genitalia. Pygofer, in lateral view, with dorso-posterior margin produced into a relatively large but narrow, tooth-like, apically obtuse process (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5E</figureCitation>
); in ventral view (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5F</figureCitation>
) much longer than in dorsal view (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5D</figureCitation>
) with ratio of ventral to dorsal width about 2.6. Gonostyles (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5E</figureCitation>
) large and broad medially, with dorsal margin weakly sinuate medially. Aedeagus (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5G-I</figureCitation>
) relatively small and slender, endosomal processes curved laterad and slightly ventro- or dorsoanteriad, membranous, without distinct minute superficial spines; phallobase with three pairs of relatively small (not conspicuously inflated) membranous lobes: a pair of elongate lateral lobes with their apices gradually convergent and tapering posteriad (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5H-I</figureCitation>
), a pair of elongate thumb-like ventral lobes, directed ventroposteriad (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5H</figureCitation>
), and a pair of small thumb-like dorsal lobes, directed dorsolaterad (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5G</figureCitation>
). Segment X, in lateral view, elongate, basal half narrow, gradually widening to apex beyond middle (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5E</figureCitation>
); in dorsal view, widest medially, with ratio of length to maximum width 1.1 (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5D</figureCitation>
).
</paragraph>
<paragraph pageId="3" pageNumber="372">Female genitalia as in generic description.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="372" type="distribution">
<paragraph pageId="3" pageNumber="372">Distribution.</paragraph>
<paragraph pageId="3" pageNumber="372">
India (southwestern part and West Bengal), Sri Lanka (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Geographical distribution of the Orthopagus species." figureDoi="10.3897/zse.94.26859.figure11" httpUri="https://binary.pensoft.net/fig/221408" pageId="3" pageNumber="372">11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="373" type="remarks">
<paragraph pageId="4" pageNumber="373">
<pageBreakToken pageId="4" pageNumber="373" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph pageId="4" pageNumber="373">
<taxonomicName authority="Melichar, 1903" authorityName="Song &amp; Malenovský &amp; Chen &amp; Deckert &amp; Liang" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Udugama exoleta" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="exoleta">
<emphasis italics="true" pageId="4" pageNumber="373">Udugama exoleta</emphasis>
</taxonomicName>
was described from Sri Lanka as the type species of 
<taxonomicName authority="Melichar, 1903" authorityName="Melichar" authorityYear="1903" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Udugama" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="373">Udugama</emphasis>
</taxonomicName>
(
<bibRefCitation author="Melichar, L" journalOrPublisher="Transactions of the Sapporo Natural History Society" pageId="12" pageNumber="381" refId="B20" refString="Melichar, L, 1903. Homopteren Fauna von Ceylon. Berlin: Verlag von Felix L. Dames, 248 pp." title="Homopteren Fauna von Ceylon. Berlin: Verlag von Felix L. Dames, 248 pp." year="1903">Melichar 1903</bibRefCitation>
). 
<bibRefCitation author="Distant, WL" journalOrPublisher="Taylor &amp; Francis, London" pageId="12" pageNumber="381" refId="B6" refString="Distant, WL, 1906. The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera). Taylor &amp; Francis, London" title="The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera)." year="1906">Distant (1906)</bibRefCitation>
synonymized this species name with 
<taxonomicName authority="Melichar, 1903" authorityName="Song &amp; Malenovský &amp; Chen &amp; Deckert &amp; Liang" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Udugama splendens" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="splendens">
<emphasis italics="true" pageId="4" pageNumber="373">Udugama splendens</emphasis>
</taxonomicName>
described from Java, Indonesia. However, 
<bibRefCitation author="Kirkaldy, GW" journalOrPublisher="Annales de la Societe entomologique de Belgique Bruxelles" pageId="12" pageNumber="381" pagination="9 - 14" refId="B12" refString="Kirkaldy, GW, 1908. Notes on some Sinhalese Rhynchota. Annales de la Societe entomologique de Belgique Bruxelles 52: 9 - 14" title="Notes on some Sinhalese Rhynchota." volume="52" year="1908">Kirkaldy (1908)</bibRefCitation>
, probably based on comparisons of figures in 
<bibRefCitation author="Melichar, L" journalOrPublisher="Transactions of the Sapporo Natural History Society" pageId="12" pageNumber="381" refId="B20" refString="Melichar, L, 1903. Homopteren Fauna von Ceylon. Berlin: Verlag von Felix L. Dames, 248 pp." title="Homopteren Fauna von Ceylon. Berlin: Verlag von Felix L. Dames, 248 pp." year="1903">Melichar (1903)</bibRefCitation>
and 
<bibRefCitation author="Distant, WL" journalOrPublisher="Taylor &amp; Francis, London" pageId="12" pageNumber="381" refId="B6" refString="Distant, WL, 1906. The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera). Taylor &amp; Francis, London" title="The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera)." year="1906">Distant (1906)</bibRefCitation>
, commented that 
<taxonomicName genus="U." lsidName="U. exoleta" pageId="4" pageNumber="373" rank="species" species="exoleta">
<emphasis italics="true" pageId="4" pageNumber="373">U. exoleta</emphasis>
</taxonomicName>
was &quot;very different&quot; from 
<taxonomicName genus="U." lsidName="U. splendens" pageId="4" pageNumber="373" rank="species" species="splendens">
<emphasis italics="true" pageId="4" pageNumber="373">U. splendens</emphasis>
</taxonomicName>
in having a much longer face. Nevertheless, the synonymy of 
<taxonomicName genus="U." lsidName="U. exoleta" pageId="4" pageNumber="373" rank="species" species="exoleta">
<emphasis italics="true" pageId="4" pageNumber="373">U. exoleta</emphasis>
</taxonomicName>
with 
<taxonomicName genus="U." lsidName="U. splendens" pageId="4" pageNumber="373" rank="species" species="splendens">
<emphasis italics="true" pageId="4" pageNumber="373">U. splendens</emphasis>
</taxonomicName>
was accepted by 
<bibRefCitation author="Melichar, L" journalOrPublisher="Zoologisch-Botanischen Gesellschaft in Wien" pageId="12" pageNumber="381" pagination="1 - 221" refId="B21" refString="Melichar, L, 1912. Monographie der Dictyophorinen (Homoptera). Abhandlungen der K. K. Zoologisch-Botanischen Gesellschaft in Wien 7 (1): 1 - 221" title="Monographie der Dictyophorinen (Homoptera). Abhandlungen der K. K." volume="7" year="1912">Melichar (1912)</bibRefCitation>
, and later included in 
<bibRefCitation DOI="https://doi.org/10.1093/aesa/40.3.544" author="Metcalf, ZP" journalOrPublisher="Smith College, Northampton, USA" pageId="12" pageNumber="381" refId="B23" refString="Metcalf, ZP, 1946. General catalogue of the Hemiptera, Fasci. IV. Fulgoroidea, Part 8 Dictyopharidae. Smith College, Northampton, USA, DOI: https://doi.org/10.1093/aesa/40.3.544" title="General catalogue of the Hemiptera, Fasci. IV. Fulgoroidea, Part 8 Dictyopharidae." url="https://doi.org/10.1093/aesa/40.3.544" year="1946">
<normalizedToken originalValue="Metcalf’s">Metcalf's</normalizedToken>
(1946)
</bibRefCitation>
catalogue of world 
<taxonomicName authorityName="Spinola" authorityYear="1839" class="Insecta" family="Dictyopharidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="family">Dictyopharidae</taxonomicName>
.
</paragraph>
<paragraph pageId="4" pageNumber="373">
Based on our critical review of the published information and examination of 
<taxonomicName authorityName="Uhler" authorityYear="1897" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="373">Orthopagus</emphasis>
</taxonomicName>
material from Sri Lanka which agrees with the original description of 
<taxonomicName genus="U." lsidName="U. exoleta" pageId="4" pageNumber="373" rank="species" species="exoleta">
<emphasis italics="true" pageId="4" pageNumber="373">U. exoleta</emphasis>
</taxonomicName>
, we propose here to resurrect 
<taxonomicName authorityName=", comb. n." authorityYear="2018" baseAuthorityName="Melichar" baseAuthorityYear="1903" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus exoletus" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="exoletus">
<emphasis italics="true" pageId="4" pageNumber="373">Orthopagus exoletus</emphasis>
</taxonomicName>
comb. n. from the synonymy with 
<taxonomicName genus="O." lsidName="O. splendens" pageId="4" pageNumber="373" rank="species" species="splendens">
<emphasis italics="true" pageId="4" pageNumber="373">O. splendens</emphasis>
</taxonomicName>
and to restore it as a valid species. 
<taxonomicName authorityName=", comb. n." authorityYear="2018" baseAuthorityName="Melichar" baseAuthorityYear="1903" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus exoletus" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="exoletus">
<emphasis italics="true" pageId="4" pageNumber="373">Orthopagus exoletus</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName genus="O." lsidName="O. splendens" pageId="4" pageNumber="373" rank="species" species="splendens">
<emphasis italics="true" pageId="4" pageNumber="373">O. splendens</emphasis>
</taxonomicName>
by the coloration of the vertex and the structure of the male genitalia, particularly the structure of the endosomal processes of aedeagus, lobes of the phallobase and the shape of the segment X. The relative length of frons mentioned by 
<bibRefCitation author="Kirkaldy, GW" journalOrPublisher="Annales de la Societe entomologique de Belgique Bruxelles" pageId="12" pageNumber="381" pagination="9 - 14" refId="B12" refString="Kirkaldy, GW, 1908. Notes on some Sinhalese Rhynchota. Annales de la Societe entomologique de Belgique Bruxelles 52: 9 - 14" title="Notes on some Sinhalese Rhynchota." volume="52" year="1908">Kirkaldy (1908)</bibRefCitation>
is probably not a relevant diagnostic character because the length of the head in 
<taxonomicName authorityName="Uhler" authorityYear="1897" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="373">Orthopagus</emphasis>
</taxonomicName>
species varies within a certain range.
</paragraph>
<paragraph pageId="4" pageNumber="373">
According to 
<bibRefCitation author="Melichar, L" journalOrPublisher="Transactions of the Sapporo Natural History Society" pageId="12" pageNumber="381" refId="B20" refString="Melichar, L, 1903. Homopteren Fauna von Ceylon. Berlin: Verlag von Felix L. Dames, 248 pp." title="Homopteren Fauna von Ceylon. Berlin: Verlag von Felix L. Dames, 248 pp." year="1903">Melichar (1903)</bibRefCitation>
, 
<taxonomicName genus="U." lsidName="U. exoleta" pageId="4" pageNumber="373" rank="species" species="exoleta">
<emphasis italics="true" pageId="4" pageNumber="373">U. exoleta</emphasis>
</taxonomicName>
was described based on five female specimens from 
<normalizedToken originalValue="“Moruwale”">&quot;Moruwale&quot;</normalizedToken>
, deposited in the collection of the museum in Colombo, Sri Lanka. This material was not available to our study. Nevertheless, we have studied one female from Sri Lanka, 
<normalizedToken originalValue="“Puttalam”">&quot;Puttalam&quot;</normalizedToken>
preserved in 
<normalizedToken originalValue="Melichar’s">Melichar's</normalizedToken>
personal collection in MMBC. Even though this specimen bears original identification labels handwritten by Melichar as &quot; 
<taxonomicName authority="Melichar, 1903" authorityName="Melichar" authorityYear="1903" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Udugama" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="373">Udugama</emphasis>
</taxonomicName>
&quot; and &quot; 
<taxonomicName authority="Melichar, 1903" authorityName="Song &amp; Malenovský &amp; Chen &amp; Deckert &amp; Liang" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Udugama exoleta" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="exoleta">
<emphasis italics="true" pageId="4" pageNumber="373">Udugama exoleta</emphasis>
</taxonomicName>
det. Melichar&quot; and a dark red label 
<normalizedToken originalValue="“Typus”">&quot;Typus&quot;</normalizedToken>
originally also attached to the specimen by Melichar, it probably cannot be considered as a syntype because it differs in its locality and deposition from the information published in the original description and probably it was also collected later than the original species description had been published. Melichar did not use type labels in the modern sense. He had rather adopted the practice of placing a 
<normalizedToken originalValue="ʻtypeʼ">ʻtypeʼ</normalizedToken>
label on one or more specimens of the most taxa (even on species previously described by other authors and identified by Melichar), specimens presumably which he himself used for comparison (
<bibRefCitation author="Young, DA" journalOrPublisher="Budapest" pageId="13" pageNumber="382" pagination="465 - 467" refId="B52" refString="Young, DA, Soos, A, 1964. Types of Cicadellinae (Homoptera, Cicadellidae) in the Hungarian Natural History Museum. Annales historico-naturales Musei nationalis hungarici. Budapest 56: 465 - 467" title="Types of Cicadellinae (Homoptera, Cicadellidae) in the Hungarian Natural History Museum. Annales historico-naturales Musei nationalis hungarici." volume="56" year="1964">
Young and 
<normalizedToken originalValue="Soós">Soos</normalizedToken>
1964
</bibRefCitation>
, 
<bibRefCitation author="Wilson, MR" journalOrPublisher="Brno" pageId="13" pageNumber="382" pagination="137 - 153" refId="B48" refString="Wilson, MR, Malenovsky, I, 2007. Tropiduchidae described by Leopold Melichar (Hemiptera, Fulgoromorpha). Acta Musei Moraviae, Scientiae biologicae. Brno 92: 137 - 153" title="Tropiduchidae described by Leopold Melichar (Hemiptera, Fulgoromorpha). Acta Musei Moraviae, Scientiae biologicae." volume="92" year="2007">
Wilson and 
<normalizedToken originalValue="Malenovský">Malenovsky</normalizedToken>
2007
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>