treatments-xml/data/10/28/87/102887B8837FFF86FF26F831BA5BF857.xml

<document id="7580680145CDD5177A4DB91A397327C4" ID-CLB-Dataset="54351" ID-DOI="10.11646/phytotaxa.595.1.10" ID-GBIF-Dataset="2ddc689e-e205-46b3-a9ed-4c8ade8630b1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7889423" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683097915035" checkinUser="plazi" docAuthor="Xu, Ke-Wang, Liu, Yu-Xuan, Chen, Yi-Fei &amp; Zhang, Li-Bing" docDate="2023" docId="102887B8837FFF86FF26F831BA5BF857" docLanguage="en" docName="phytotaxa.595.1.10.pdf" docOrigin="Phytotaxa 595 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.595.1.10" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Hymenasplenium hovenkampii K. W. Xu &amp; Li Bing Zhang 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="122" masterDocId="EC11FFC0837FFF84FFAEFF85B913FFA4" masterDocTitle="Hymenasplenium hovenkampii (Aspleniaceae), a new bipinnatisect fern species from Lesser Sunda Islands of Indonesia" masterLastPageNumber="124" masterPageNumber="120" pageNumber="120" updateTime="1699515196279" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="DB1EA69A66C9D6284321279078E63841" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="52C88AB82ADDAF9B8E2C0433B19794B6">
<mods:title id="42B7BDB73FA16F5905E005472EE60B1F">Hymenasplenium hovenkampii (Aspleniaceae), a new bipinnatisect fern species from Lesser Sunda Islands of Indonesia</mods:title>
</mods:titleInfo>
<mods:name id="9706036F35B4CAEE077CFDE042B11161" type="personal">
<mods:role id="92474242B994636E6B9BCB7977DA9B42">
<mods:roleTerm id="E50BAA98ECDA36D610EF295F98039F1C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC0C8822DB347D3E2A15810D71C39283">Xu, Ke-Wang</mods:namePart>
<mods:nameIdentifier id="2E96507B3FA380199288AAB40B12E5D8" type="ORCID">0000-0003-2239-0487</mods:nameIdentifier>
<mods:affiliation id="2D579C0C0EE9D4EC48B5FE57EFD1D4AE">Co-Innovation Center for Sustainable Forestry in Southern China, College of Biology and the Environment, Nanjing Forestry University, Nanjing, 510275, China. &amp; xukw 10 @ njfu. edu. cn; https: // orcid. org / 0000 - 0003 - 2239 - 0487</mods:affiliation>
<mods:nameIdentifier id="4FC3B73B4421681F0DADEB4B98AAD860" type="email">xukw10@njfu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="248D63533CDEEFD814FF89077AB2A95B" type="personal">
<mods:role id="F8082FAD8A5FB205A1B239842C8702E6">
<mods:roleTerm id="D9578D067DB30407676E82A731D74C76">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="05D05E07DCCD8D9B8948B9666A82E2CE">Liu, Yu-Xuan</mods:namePart>
<mods:nameIdentifier id="87F0516D1EF123FD19BBF6B2D6CBFA84" type="ORCID">0009-0006-1142-0205</mods:nameIdentifier>
<mods:affiliation id="45C0C35F46F9055A11D213535E45FCA7">Co-Innovation Center for Sustainable Forestry in Southern China, College of Biology and the Environment, Nanjing Forestry University, Nanjing, 510275, China. &amp; 13401244397 @ 163. com; https: // orcid. org / 0009 - 0006 - 1142 - 0205</mods:affiliation>
</mods:name>
<mods:name id="B09B3CD0F24B580FCF7D15378B7ECD96" type="personal">
<mods:role id="C69BE9047B93144F4D998DE5BA7EA558">
<mods:roleTerm id="1C3982972097EA247232E71B0CA6FFF0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="32CC4A9FF2A648F501EA7FC99097B385">Chen, Yi-Fei</mods:namePart>
<mods:nameIdentifier id="770695BBF99168EE01B0C8E371106964" type="ORCID">0009-0008-3680-7427</mods:nameIdentifier>
<mods:affiliation id="E0945C38591E1A0A921446C2A9F98E22">Co-Innovation Center for Sustainable Forestry in Southern China, College of Biology and the Environment, Nanjing Forestry University, Nanjing, 510275, China. &amp; 19852399199 @ 163. com; https: // orcid. org / 0009 - 0008 - 3680 - 7427</mods:affiliation>
</mods:name>
<mods:name id="527B03927965279299F3791AB70B4C68" type="personal">
<mods:role id="89DF49022A945251E84930AFEBCA4D3F">
<mods:roleTerm id="9F4DCDE27CF83ECDCF53C74BA8C1C7D4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DCB448028E57384E9D6803B36E31E209">Zhang, Li-Bing</mods:namePart>
<mods:nameIdentifier id="F09BC156DDF25481C21D3089E0A5825E" type="ORCID">0000-0002-4905-040X</mods:nameIdentifier>
<mods:affiliation id="5556A385D0BD56F2E0A6126630237AF6">Missouri Botanical Garden, P. O. Box 299, St. Louis, Missouri 63166 - 0299, U. S. A.; Chengdu Institute of Biology, Chinese Academy of Sciences, P. O. Box 416, Chengdu, Sichuan 610041, China. &amp; Libing. Zhang @ mobot. org; https: // orcid. org / 0000 - 0002 - 4905 - 040 X</mods:affiliation>
<mods:nameIdentifier id="19368540EF9AAEC18A8EF692F6DE64C5" type="email">hang@mobot.org</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="12FDD07111BB0C3CE020092490B09931">text</mods:typeOfResource>
<mods:relatedItem id="F41361AC1564574BDE397E8020B577B1" type="host">
<mods:titleInfo id="983C326201B842D0B34BE166F6D28C32">
<mods:title id="68DEF8CF6C4F54750FC70414CB073A4A">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="A80369A1C5AFFB7D084AA83450C5488C">
<mods:date id="FDB270C5C0023C94CA24EE8EF8C0067C">2023</mods:date>
<mods:detail id="B04C47EB5B8B8BBD152F04B153E1CEF1" type="pubDate">
<mods:number id="86F7CAA144B4A8D1280854206E5757F6">2023-05-03</mods:number>
</mods:detail>
<mods:detail id="7366C91217CFB6288E13ABABA5F49866" type="volume">
<mods:number id="C66063D7CECA4C7508EB78F2F3470ADA">595</mods:number>
</mods:detail>
<mods:detail id="E0EDAC66157C01D5A11E2D535927BCAE" type="issue">
<mods:number id="0A0EDEF78149B5A8C45A56B6329CE722">1</mods:number>
</mods:detail>
<mods:extent id="4F588C04B140F5B5396CC9F66364C7EC" unit="page">
<mods:start id="1CDDF0B82BFFFCA59CB079D903740DB7">120</mods:start>
<mods:end id="873B8EE568E51CE209D83ADAFA3B746D">124</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="18C9CE5ED27AE6609230C13D228A1C3D">
<mods:url id="511F12D1B806F8DE20AD9C21FC56BDB9">http://dx.doi.org/10.11646/phytotaxa.595.1.10</mods:url>
</mods:location>
<mods:classification id="EC863A7214BC3FA3778ACD93A5CC1C94">journal article</mods:classification>
<mods:identifier id="7708EB1FCB0646CA0534E4CFD16501C3" type="CLB-Dataset">54351</mods:identifier>
<mods:identifier id="A52C6CC972B7759C68BF2D3EB2E9950E" type="DOI">10.11646/phytotaxa.595.1.10</mods:identifier>
<mods:identifier id="6A971147214F3C8EA03399D26997255F" type="GBIF-Dataset">2ddc689e-e205-46b3-a9ed-4c8ade8630b1</mods:identifier>
<mods:identifier id="CD1652E6A389B0E1056991D141E74E55" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="00D9E44F77D3A2F87041A9314590F0E1" type="Zenodo-Dep">7889423</mods:identifier>
</mods:mods>
<treatment id="102887B8837FFF86FF26F831BA5BF857" ID-DOI="http://doi.org/10.5281/zenodo.7889425" ID-GBIF-Taxon="209580676" ID-Zenodo-Dep="7889425" LSID="urn:lsid:plazi:treatment:102887B8837FFF86FF26F831BA5BF857" httpUri="http://treatment.plazi.org/id/102887B8837FFF86FF26F831BA5BF857" lastPageId="2" lastPageNumber="122" pageId="0" pageNumber="120">
<subSubSection id="D09B6525837FFF84FF26F831BAEEF86A" box="[136,1021,1972,1999]" pageId="0" pageNumber="120" type="nomenclature">
<paragraph id="983E36AE837FFF84FF26F831BAEEF86A" blockId="0.[136,1021,1972,1999]" box="[136,1021,1972,1999]" pageId="0" pageNumber="120">
<heading id="C37681C2837FFF84FF26F831BAEEF86A" box="[136,1021,1972,1999]" fontSize="11" level="2" pageId="0" pageNumber="120" reason="3">
<taxonomicName id="5F814D2D837FFF84FF26F831BA0CF86A" ID-CoL="BLRVZ" authority="K.W.Xu &amp; Li Bing Zhang" authorityName="K. W. Xu &amp; Li Bing Zhang" authorityYear="2023" box="[136,799,1972,1999]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="0" pageNumber="120" phylum="Tracheophyta" rank="species" species="hovenkampii" status="sp. nov.">
<emphasis id="AAF5EABC837FFF84FF26F831B8E3F86B" bold="true" box="[136,496,1972,1999]" italics="true" pageId="0" pageNumber="120">Hymenasplenium hovenkampii</emphasis>
K.W.Xu &amp; Li Bing Zhang
</taxonomicName>
, 
<emphasis id="AAF5EABC837FFF84FC84F830BA6CF86A" box="[810,895,1973,1998]" italics="true" pageId="0" pageNumber="120">
<taxonomicNameLabel id="B1C657C7837FFF84FC84F830BA6CF86A" box="[810,895,1973,1998]" pageId="0" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="00BA2A2B837FFF84FC23F831BAFFF86A" box="[909,1004,1972,1998]" captionStart="FIGURE 1" captionStartId="1.[136,229,1996,2018]" captionTargetBox="[151,1436,423,1972]" captionTargetId="figure-128@1.[151,1436,423,1972]" captionTargetPageId="1" captionText="FIGURE 1. Hymenasplenium hovenkampii sp. nov.—A. Holotype.—B. Middle portion of lamina (Drawn by Ke-Wang Xu based on the type)." figureDoi="http://doi.org/10.5281/zenodo.7889427" httpUri="https://zenodo.org/record/7889427/files/figure.png" pageId="0" pageNumber="120">Figure 1</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D09B6525837FFF85FF26F859BAD3FE84" lastPageId="1" lastPageNumber="121" pageId="0" pageNumber="120" type="diagnosis">
<paragraph id="983E36AE837FFF85FF26F859BAD3FE84" blockId="0.[136,1451,2012,2070]" lastBlockId="1.[136,1452,159,361]" lastPageId="1" lastPageNumber="121" pageId="0" pageNumber="120">
Diagnosis:—This new species is morphologically similar to 
<taxonomicName id="5F814D2D837FFF84FD4DF859B837F7B1" authority="(Kunze ex Mettenius 1859: 133) Tagawa (1938: 84)" authorityName="Tagawa" authorityPageNumber="84" authorityYear="1938" baseAuthorityName="Kunze ex Mettenius" baseAuthorityPageNumber="133" baseAuthorityYear="1859" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="0" pageNumber="120" phylum="Tracheophyta" rank="species" species="cheilosorum">
<emphasis id="AAF5EABC837FFF84FD4DF859BD19F855" box="[739,1034,2012,2033]" italics="true" pageId="0" pageNumber="120">Hymenasplenium cheilosorum</emphasis>
(Kunze ex 
<bibRefCitation id="FC104B5F837FFF84FB2DF859BC42F855" author="Mettenius, G. H." box="[1155,1361,2012,2034]" pageId="0" pageNumber="120" refId="ref2405" refString="Mettenius, G. H. (1859) Abhandlungen herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft. 133 pp." type="book" year="1859">Mettenius 1859: 133</bibRefCitation>
) 
<bibRefCitation id="FC104B5F837FFF84FACFF859B837F7B1" author="Tagawa, M." pageId="0" pageNumber="120" pagination="72 - 87" refId="ref2715" refString="Tagawa, M. (1938) Spicilegium pteridographiae Asiae Orientalis. Acta Phytotaxonomica et Geobotanica 7 (2): 72 - 87." type="journal article" year="1938">Tagawa (1938: 84)</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="5F814D2D837FFF84FEFBF785BA43F7B2" authority="Li Bing Zhang, K. W. Xu &amp; C. W. Chen" authorityName="Li Bing Zhang, K. W. Xu &amp; C. W. Chen" box="[341,848,2048,2070]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="0" pageNumber="120" phylum="Tracheophyta" rank="species" species="sabahense">
<emphasis id="AAF5EABC837FFF84FEFBF785B8CBF7B1" box="[341,472,2048,2069]" italics="true" pageId="0" pageNumber="120">H. sabahense</emphasis>
Li Bing Zhang, K.W.Xu &amp; C.W.Chen
</taxonomicName>
in Xu 
<emphasis id="AAF5EABC837FFF84FC39F785BADAF7B1" box="[919,969,2048,2069]" italics="true" pageId="0" pageNumber="120">et al.</emphasis>
(2019: 38) in having sori terminal on subtending veins and situated in pinna lobes, but the former has laminae bipinnatisect, rachises reddish brown, and the acroscopic margins deeply lobed and usually with 4–5 lobes and each lobe deeply or shallowly crenate to dentate with two teeth, while both the latter two species have laminae 1-pinnate, rachises dark purplish to black, and the acroscopic margins relatively shallowly lobed with more than 5 lobes and each lobe shallowly crenate to dentate with two teeth (
<tableCitation id="D5030315837EFF85FCC4FE8EBAA2FE84" box="[874,945,267,289]" captionStart="TABLE 1" captionStartId="2.[137,229,160,184]" captionTargetPageId="2" captionText="TABLE 1. Morphological comparison of Hymenasplenium hovenkampii and the three morphologically similar species." httpUri="http://table.plazi.org/id/CCFE6626837DFF86FF27FF25BC94FF1C" pageId="1" pageNumber="121" tableUuid="CCFE6626837DFF86FF27FF25BC94FF1C">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D09B6525837EFF86FF26FEAABA5BF857" lastPageId="2" lastPageNumber="122" pageId="1" pageNumber="121" type="materials_examined">
<paragraph id="983E36AE837EFF85FF26FEAAB83BFECD" blockId="1.[136,1452,159,361]" pageId="1" pageNumber="121">
<emphasis id="AAF5EABC837EFF85FF26FEAAB9A9FEE0" bold="true" box="[136,186,303,324]" pageId="1" pageNumber="121">Type</emphasis>
:— 
<materialsCitation id="28E93CF3837EFF85FF78FEAAB837FECD" ID-GBIF-Occurrence="4098986301" collectingDate="1983-04-18" collectionCode="E" country="Indonesia" county="Manggarai" location="Schmutz" municipality="Ruteng" pageId="1" pageNumber="121" specimenCode="E 5646" specimenCount="1" stateProvince="Nusa Tenggara Timur" typeStatus="holotype">
<collectingCountry id="E096763E837EFF85FF78FEAAB84DFEE0" box="[214,350,303,324]" name="Indonesia" pageId="1" pageNumber="121">INDONESIA</collectingCountry>
. 
<collectingRegion id="5A45F84C837EFF85FEC4FEB5BB20FEE1" box="[362,563,304,325]" country="Indonesia" name="Nusa Tenggara Timur" pageId="1" pageNumber="121">East Nusa Tenggara</collectingRegion>
: Lesser Sunda Islands, 
<collectingCounty id="715F4E22837EFF85FCB3FEAABA94FEE1" box="[797,903,303,325]" pageId="1" pageNumber="121">Manggarai</collectingCounty>
, 
<collectingMunicipality id="785AACD4837EFF85FC3DFEB5BAC8FEE1" box="[915,987,304,325]" pageId="1" pageNumber="121">Ruteng</collectingMunicipality>
, 
<date id="EC3F106E837EFF85FC49FEAABD67FEE0" box="[999,1140,303,325]" pageId="1" pageNumber="121" value="1983-04-18">
<collectingDate id="FC7BE986837EFF85FC49FEAABD67FEE0" box="[999,1140,303,325]" pageId="1" pageNumber="121" value="1983-04-18">18 April 1983</collectingDate>
</date>
, 
<emphasis id="AAF5EABC837EFF85FB2FFEAABC32FEE0" box="[1153,1313,303,325]" italics="true" pageId="1" pageNumber="121">
<location id="9D5E6075837EFF85FB2FFEAABDC2FEE0" LSID="urn:lsid:plazi:treatment:102887B8837FFF86FF26F831BA5BF857:9D5E6075837EFF85FB2FFEAABDC2FEE0" box="[1153,1233,303,324]" country="Indonesia" county="Manggarai" municipality="Ruteng" name="Schmutz" pageId="1" pageNumber="121" stateProvince="Nusa Tenggara Timur">Schmutz</location>
<specimenCode id="C8279ED5837EFF85FB77FEB5BC32FEE0" box="[1241,1313,303,325]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="1" pageNumber="121" type="Herbarium">E 5646</specimenCode>
</emphasis>
(
<typeStatus id="473A880C837EFF85FA9EFEAABC9AFEE1" box="[1328,1417,303,325]" pageId="1" pageNumber="121" type="holotype">holotype</typeStatus>
: L-3507947!)
</materialsCitation>
.
</paragraph>
<caption id="CCFE6626837EFF85FF26F849B9D3F7A1" ID-DOI="http://doi.org/10.5281/zenodo.7889427" ID-Zenodo-Dep="7889427" httpUri="https://zenodo.org/record/7889427/files/figure.png" pageId="1" pageNumber="121" startId="1.[136,229,1996,2018]" targetBox="[151,1436,423,1972]" targetPageId="1" targetType="figure">
<paragraph id="983E36AE837EFF85FF26F849B9D3F7A1" blockId="1.[136,1451,1996,2053]" pageId="1" pageNumber="121">
<emphasis id="AAF5EABC837EFF85FF26F849B9EDF845" bold="true" box="[136,254,1996,2018]" pageId="1" pageNumber="121">FIGURE 1.</emphasis>
<emphasis id="AAF5EABC837EFF85FEAAF849BB6FF846" box="[260,636,1996,2018]" italics="true" pageId="1" pageNumber="121">
<taxonomicName id="5F814D2D837EFF85FEAAF849BB3FF845" authority="Xu &amp; Liu &amp; Chen &amp; Zhang, 2023" authorityName="Xu &amp; Liu &amp; Chen &amp; Zhang" authorityYear="2023" box="[260,556,1996,2017]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="1" pageNumber="121" phylum="Tracheophyta" rank="species" species="hovenkampii" status="sp. nov.">Hymenasplenium hovenkampii</taxonomicName>
<taxonomicNameLabel id="B1C657C7837EFF85FD9DF848BB6FF846" box="[563,636,1997,2018]" pageId="1" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
—A. Holotype.—B. Middle portion of lamina (Drawn by Ke-Wang Xu based on the type).
</paragraph>
</caption>
<caption id="CCFE6626837DFF86FF27FF25BC94FF1C" ID-Table-UUID="CCFE6626837DFF86FF27FF25BC94FF1C" box="[137,1415,160,184]" httpUri="http://table.plazi.org/id/CCFE6626837DFF86FF27FF25BC94FF1C" pageId="2" pageNumber="122" startId="2.[137,229,160,184]" targetBox="[144,1443,212,800]" targetIsTable="true" targetPageId="2" targetType="table">
<paragraph id="983E36AE837DFF86FF27FF25BC94FF1C" blockId="2.[137,1415,160,184]" box="[137,1415,160,184]" pageId="2" pageNumber="122">
<emphasis id="AAF5EABC837DFF86FF27FF25B813FF1C" bold="true" box="[137,256,160,184]" pageId="2" pageNumber="122">TABLE 1.</emphasis>
Morphological comparison of 
<taxonomicName id="5F814D2D837DFF86FDE2FF25BA80FF1C" authorityName="Xu &amp; Liu &amp; Chen &amp; Zhang" authorityYear="2023" box="[588,915,160,184]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="hovenkampii">
<emphasis id="AAF5EABC837DFF86FDE2FF25BA80FF1C" box="[588,915,160,184]" italics="true" pageId="2" pageNumber="122">Hymenasplenium hovenkampii</emphasis>
</taxonomicName>
and the three morphologically similar species.
</paragraph>
</caption>
<paragraph id="983E36AE837DFF86FF3EFF51BC28FC84" pageId="2" pageNumber="122">
<table id="EA81C40E837D007BFF3EFF51BCB0FC84" box="[144,1443,212,800]" gridcols="5" gridrows="9" pageId="2" pageNumber="122">
<tr id="26B134EC837D007BFF3EFF51BCB0FF43" box="[144,1443,212,231]" gridrow="0" pageId="2" pageNumber="122">
<th id="65605D90837D007BFF3EFF51B826FF43" box="[144,309,212,231]" gridcol="0" gridrow="0" pageId="2" pageNumber="122">
<emphasis id="AAF5EABC837DFF86FF3EFF51B9FDFF43" bold="true" box="[144,238,212,231]" pageId="2" pageNumber="122">Character</emphasis>
</th>
<th id="65605D90837D007BFEF1FF51BB7EFF43" box="[351,621,212,231]" gridcol="1" gridrow="0" pageId="2" pageNumber="122">
<taxonomicName id="5F814D2D837DFF86FEF1FF50B8F5FF43" authorityName="Xu &amp; Liu &amp; Chen &amp; Zhang" authorityYear="2023" box="[351,486,212,231]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="hovenkampii">
<emphasis id="AAF5EABC837DFF86FEF1FF50B8F5FF43" box="[351,486,212,231]" italics="true" pageId="2" pageNumber="122">H. hovenkampii</emphasis>
</taxonomicName>
</th>
<th id="65605D90837D007BFDD2FF51BA93FF43" box="[636,896,212,231]" gridcol="2" gridrow="0" pageId="2" pageNumber="122">
<taxonomicName id="5F814D2D837DFF86FDD2FF50BBECFF43" authorityName="Tagawa" authorityYear="1938" baseAuthorityName="Kunze ex Mettenius" baseAuthorityYear="1859" box="[636,767,212,231]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="cheilosorum">
<emphasis id="AAF5EABC837DFF86FDD2FF50BBECFF43" box="[636,767,212,231]" italics="true" pageId="2" pageNumber="122">H. cheilosorum</emphasis>
</taxonomicName>
</th>
<th id="65605D90837D007BFC3DFF51BD6DFF43" box="[915,1150,212,231]" gridcol="3" gridrow="0" pageId="2" pageNumber="122">
<taxonomicName id="5F814D2D837DFF86FC3DFF50BD0CFF43" box="[915,1055,212,231]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="inthanonense">
<emphasis id="AAF5EABC837DFF86FC3DFF50BD0CFF43" box="[915,1055,212,231]" italics="true" pageId="2" pageNumber="122">H. inthanonense</emphasis>
</taxonomicName>
</th>
<th id="65605D90837D007BFB02FF51BCB0FF43" box="[1196,1443,212,231]" gridcol="4" gridrow="0" pageId="2" pageNumber="122">
<emphasis id="AAF5EABC837DFF86FB02FF50BC0CFF43" box="[1196,1311,212,231]" italics="true" pageId="2" pageNumber="122">H. sabahense</emphasis>
</th>
</tr>
<tr id="26B134EC837D007BFF3EFE88BCB0FE84" box="[144,1443,269,288]" gridrow="1" pageId="2" pageNumber="122">
<th id="65605D90837D007BFF3EFE88B826FE84" box="[144,309,269,288]" gridcol="0" gridrow="1" pageId="2" pageNumber="122">Rhizomes</th>
<td id="65605D90837D007BFEF1FE88BB7EFE84" box="[351,621,269,288]" gridcol="1" gridrow="1" pageId="2" pageNumber="122">ca. 2 mm in diam.</td>
<td id="65605D90837D007BFDD2FE88BA93FE84" box="[636,896,269,288]" gridcol="2" gridrow="1" pageId="2" pageNumber="122">2.5–4 mm in diam.</td>
<td id="65605D90837D007BFC3DFE88BD6DFE84" box="[915,1150,269,288]" gridcol="3" gridrow="1" pageId="2" pageNumber="122">3–6.5 mm in diam.</td>
<td id="65605D90837D007BFB02FE88BCB0FE84" box="[1196,1443,269,288]" gridcol="4" gridrow="1" pageId="2" pageNumber="122">ca. 3 mm in diam.</td>
</tr>
<tr id="26B134EC837D007BFF3EFEC4BCB0FEF0" box="[144,1443,321,340]" gridrow="2" pageId="2" pageNumber="122">
<th id="65605D90837D007BFF3EFEC4B826FEF0" box="[144,309,321,340]" gridcol="0" gridrow="2" pageId="2" pageNumber="122">Laminae dissection</th>
<td id="65605D90837D007BFEF1FEC4BB7EFEF0" box="[351,621,321,340]" gridcol="1" gridrow="2" pageId="2" pageNumber="122">bipinnatisect</td>
<td id="65605D90837D007BFDD2FEC4BA93FEF0" box="[636,896,321,340]" gridcol="2" gridrow="2" pageId="2" pageNumber="122">1-pinnate</td>
<td id="65605D90837D007BFC3DFEC4BD6DFEF0" box="[915,1150,321,340]" gridcol="3" gridrow="2" pageId="2" pageNumber="122">1-pinnate</td>
<td id="65605D90837D007BFB02FEC4BCB0FEF0" box="[1196,1443,321,340]" gridcol="4" gridrow="2" pageId="2" pageNumber="122">1-pinnate</td>
</tr>
<tr id="26B134EC837D007BFF3EFEF3BCB0FE2D" box="[144,1443,374,393]" gridrow="3" pageId="2" pageNumber="122">
<th id="65605D90837D007BFF3EFEF3B826FE2D" box="[144,309,374,393]" gridcol="0" gridrow="3" pageId="2" pageNumber="122">Laminae size</th>
<td id="65605D90837D007BFEF1FEF3BB7EFE2D" box="[351,621,374,393]" gridcol="1" gridrow="3" pageId="2" pageNumber="122">14–20 × (1.5–)2–2.5(–3) cm</td>
<td id="65605D90837D007BFDD2FEF3BA93FE2D" box="[636,896,374,393]" gridcol="2" gridrow="3" pageId="2" pageNumber="122">15–35 × 3–5(–7) cm</td>
<td id="65605D90837D007BFC3DFEF3BD6DFE2D" box="[915,1150,374,393]" gridcol="3" gridrow="3" pageId="2" pageNumber="122">30–40 × 4–7 cm</td>
<td id="65605D90837D007BFB02FEF3BCB0FE2D" box="[1196,1443,374,393]" gridcol="4" gridrow="3" pageId="2" pageNumber="122">20–28 × 3–5 cm</td>
</tr>
<tr id="26B134EC837D007BFF3EFE2EBCB0FE1A" box="[144,1443,427,446]" gridrow="4" pageId="2" pageNumber="122">
<th id="65605D90837D007BFF3EFE2EB826FE1A" box="[144,309,427,446]" gridcol="0" gridrow="4" pageId="2" pageNumber="122">Rachises</th>
<td id="65605D90837D007BFEF1FE2EBB7EFE1A" box="[351,621,427,446]" gridcol="1" gridrow="4" pageId="2" pageNumber="122">reddish brown</td>
<td id="65605D90837D007BFDD2FE2EBA93FE1A" box="[636,896,427,446]" gridcol="2" gridrow="4" pageId="2" pageNumber="122">dark purplish to black</td>
<td id="65605D90837D007BFC3DFE2EBD6DFE1A" box="[915,1150,427,446]" gridcol="3" gridrow="4" pageId="2" pageNumber="122">dark purple</td>
<td id="65605D90837D007BFB02FE2EBCB0FE1A" box="[1196,1443,427,446]" gridcol="4" gridrow="4" pageId="2" pageNumber="122">dark purplish to black</td>
</tr>
<tr id="26B134EC837D007BFF3EFE5ABCB0FDB3" box="[144,1443,479,535]" gridrow="5" pageId="2" pageNumber="122">
<th id="65605D90837D007BFF3EFE5AB826FDB3" box="[144,309,479,535]" gridcol="0" gridrow="5" pageId="2" pageNumber="122">Pinnae</th>
<td id="65605D90837D007BFEF1FE5ABB7EFDB3" box="[351,621,479,535]" gridcol="1" gridrow="5" pageId="2" pageNumber="122">shortly stalked to almost sessile at apex, 30–40 pairs</td>
<td id="65605D90837D007BFDD2FE5ABA93FDB3" box="[636,896,479,535]" gridcol="2" gridrow="5" pageId="2" pageNumber="122">almost sessile, 25–40 pairs</td>
<td id="65605D90837D007BFC3DFE5ABD6DFDB3" box="[915,1150,479,535]" gridcol="3" gridrow="5" pageId="2" pageNumber="122">almost sessile, 30–40 pairs</td>
<td id="65605D90837D007BFB02FE5ABCB0FDB3" box="[1196,1443,479,535]" gridcol="4" gridrow="5" pageId="2" pageNumber="122">shortly stalked to almost sessile at apex, 25–40 pairs</td>
</tr>
<tr id="26B134EC837D007BFF3EFDBDBCB0FDEF" box="[144,1443,568,587]" gridrow="6" pageId="2" pageNumber="122">
<th id="65605D90837D007BFF3EFDBDB826FDEF" box="[144,309,568,587]" gridcol="0" gridrow="6" pageId="2" pageNumber="122">Pinna margin</th>
<td id="65605D90837D007BFEF1FDBDBB7EFDEF" box="[351,621,568,587]" gridcol="1" gridrow="6" pageId="2" pageNumber="122">usually with 4–5 lobes</td>
<td id="65605D90837D007BFDD2FDBDBA93FDEF" box="[636,896,568,587]" gridcol="2" gridrow="6" pageId="2" pageNumber="122">usually with 7–10 lobes</td>
<td id="65605D90837D007BFC3DFDBDBD6DFDEF" box="[915,1150,568,587]" gridcol="3" gridrow="6" pageId="2" pageNumber="122">usually with 7–9 lobes</td>
<td id="65605D90837D007BFB02FDBDBCB0FDEF" box="[1196,1443,568,587]" gridcol="4" gridrow="6" pageId="2" pageNumber="122">usually with 5–7 lobes</td>
</tr>
<tr id="26B134EC837D007BFF3EFDE9BCB0FD07" box="[144,1443,620,675]" gridrow="7" pageId="2" pageNumber="122">
<th id="65605D90837D007BFF3EFDE9B826FD07" box="[144,309,620,675]" gridcol="0" gridrow="7" pageId="2" pageNumber="122">Veins</th>
<td id="65605D90837D007BFEF1FDE9BB7EFD07" box="[351,621,620,675]" gridcol="1" gridrow="7" pageId="2" pageNumber="122">usually 3–4 or rarely all basal basiscopic veins lacking</td>
<td id="65605D90837D007BFDD2FDE9BA93FD07" box="[636,896,620,675]" gridcol="2" gridrow="7" pageId="2" pageNumber="122">usually 6 or 7 basal basiscopic veins lacking</td>
<td id="65605D90837D007BFC3DFDE9BD6DFD07" box="[915,1150,620,675]" gridcol="3" gridrow="7" pageId="2" pageNumber="122">6–8 basal basiscopic veins lacking</td>
<td id="65605D90837D007BFB02FDE9BCB0FD07" box="[1196,1443,620,675]" gridcol="4" gridrow="7" pageId="2" pageNumber="122">5–6 basal basiscopic veins lacking</td>
</tr>
<tr id="26B134EC837D007BFF3EFD40BCB0FC84" box="[144,1443,709,800]" gridrow="8" pageId="2" pageNumber="122">
<th id="65605D90837D007BFF3EFD40B826FC84" box="[144,309,709,800]" gridcol="0" gridrow="8" pageId="2" pageNumber="122">Sori</th>
<td id="65605D90837D007BFEF1FD40BB7EFC84" box="[351,621,709,800]" gridcol="1" gridrow="8" pageId="2" pageNumber="122">1.5–2 mm long, at tip of subtending veins and located in or below marginal teeth</td>
<td id="65605D90837D007BFDD2FD40BA93FC84" box="[636,896,709,800]" gridcol="2" gridrow="8" pageId="2" pageNumber="122">1–3 mm long, at tip of subtending veins and located in marginal teeth</td>
<td id="65605D90837D007BFC3DFD40BD6DFC84" box="[915,1150,709,800]" gridcol="3" gridrow="8" pageId="2" pageNumber="122">2–4 mm long, supramedial, not confined to lobes</td>
<td id="65605D90837D007BFB02FD40BCB0FC84" box="[1196,1443,709,800]" gridcol="4" gridrow="8" pageId="2" pageNumber="122">2–4 mm long, at tip of subtending veins and located in marginal teeth</td>
</tr>
</table>
</paragraph>
<paragraph id="983E36AE837DFF86FF26FCDDBC94FAE2" blockId="2.[136,1452,856,2035]" pageId="2" pageNumber="122">
Plants perennial, evergreen, 
<quantity id="5F799B4B837DFF86FE66FCDDBB24FCD7" box="[456,567,856,883]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="2" pageNumber="122" unit="cm" value="30.0" valueMax="35.0" valueMin="25.0">25–35 cm</quantity>
tall. Rhizomes long creeping, ca. 
<quantity id="5F799B4B837DFF86FC17FCDCBAE8FCD7" box="[953,1019,857,883]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="122" unit="mm" value="2.0">2 mm</quantity>
in diameter, apex scaly, scales brown, those narrowly triangular or lanceolate with entire margins; root yellowish brown when dried, slender, rhizome and root usually covered with yellow brown floccose indument. Fronds remote, membranous-herbaceous; petioles slender, shiny, reddish brown, glabrous, 
<quantity id="5F799B4B837DFF86FE40FC40BB5CFC7B" box="[494,591,965,991]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="2" pageNumber="122" unit="cm" value="7.0" valueMax="10.0" valueMin="4.0">4–10 cm</quantity>
long, ca. 
<quantity id="5F799B4B837DFF86FD13FC40BBEDFC7B" box="[701,766,965,991]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="122" unit="mm" value="1.0">1 mm</quantity>
in diameter. Laminae narrowly lanceolate, bipinnatisect, 14– 20 × (1.5–)2–2.5(–3) cm, base reduced, widest near base, gradually narrowing toward apex, apex acuminate; rachises 
<quantity id="5F799B4B837DFF86FF26FB89B9E8FB83" box="[136,251,1036,1063]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="2" pageNumber="122" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
in diameter, glabrous, shiny, reddish brown, adaxial side grooved and with 2 green narrow wings. Pinnae shortly stalked to almost sessile at apex, 30–40 pairs, base nearly opposite, middle and apex alternate, rectangular to trapeziform, (middle pairs) (1–)1.2–1.8(–2.5) × 
<quantity id="5F799B4B837DFF86FD34FBD1BA0BFBCB" box="[666,792,1108,1135]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="122" unit="cm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 cm</quantity>
, 
<quantity id="5F799B4B837DFF86FC8CFBD0BA6EFBCB" box="[802,893,1109,1135]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="2" pageNumber="122" unit="mm" value="2.5" valueMax="4.0" valueMin="1.0">1–4 mm</quantity>
apart, base asymmetrical, acroscopic side truncate and subparallel with rachis, basiscopic side cuneate, acroscopic margins deeply lobed, usually with 4–5 lobes, each lobe usually deeply or shallowly crenate to dentate with two teeth, teeth obtuse or sometimes acute, entire, basiscopic margins entire at base, lobed at the apex; pinna apex obtuse to acute, lower pinnae deflexed or spreading, upper pinnae ascending. Veins visible on both side of pinnae, green, forking and terminating in marginal teeth, 1 vein per marginal tooth and ending below tooth, usually 3–4 or rarely all basal basiscopic veins lacking. Sori elliptic to linear, 
<quantity id="5F799B4B837DFF86FA94FA8DBCBFFA87" box="[1338,1452,1288,1315]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="122" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
long, at tip of subtending veins and located in or below marginal teeth; indusia brown, linear, membranous, entire.
</paragraph>
<paragraph id="983E36AE837DFF86FF13FAD5BB46FA16" blockId="2.[136,1452,856,2035]" pageId="2" pageNumber="122">
<emphasis id="AAF5EABC837DFF86FF13FAD5B8F3FACE" bold="true" box="[189,480,1360,1387]" pageId="2" pageNumber="122">Habitat and distribution</emphasis>
:— 
<taxonomicName id="5F814D2D837DFF86FDADFAD5BA49FACE" authorityName="Xu &amp; Liu &amp; Chen &amp; Zhang" authorityYear="2023" box="[515,858,1360,1386]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="hovenkampii">
<emphasis id="AAF5EABC837DFF86FDADFAD5BA49FACE" box="[515,858,1360,1386]" italics="true" pageId="2" pageNumber="122">Hymenasplenium hovenkampii</emphasis>
</taxonomicName>
<materialsCitation id="28E93CF3837DFF86FCCEFAD4BB41FA16" ID-GBIF-Occurrence="4098986302" country="Indonesia" county="It" elevation="1300" location="This" municipality="Ruteng" pageId="2" pageNumber="122" specimenCount="1" stateProvince="Nusa Tenggara Timur">
was observed to grow in shaded and wet places along streams in forests at an elevation of ca. 
<quantity id="5F799B4B837DFF86FDE3FAF0BBB5FA2B" box="[589,678,1397,1423]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="122" unit="m" value="1300.0">
<elevation id="13ACD19D837DFF86FDE3FAF0BBB5FA2B" box="[589,678,1397,1423]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="122" unit="m" value="1300.0">1300 m</elevation>
</quantity>
. 
<location id="9D5E6075837DFF86FD1CFAF1BBF7FA2A" LSID="urn:lsid:plazi:treatment:102887B8837FFF86FF26F831BA5BF857:9D5E6075837DFF86FD1CFAF1BBF7FA2A" box="[690,740,1396,1422]" country="Indonesia" county="It" municipality="Ruteng" name="This" pageId="2" pageNumber="122" stateProvince="Nusa Tenggara Timur">This</location>
species is currently only known from 
<collectingMunicipality id="785AACD4837DFF86FB35FAF0BDFFFA2B" box="[1179,1260,1397,1423]" pageId="2" pageNumber="122">Ruteng</collectingMunicipality>
of 
<collectingRegion id="5A45F84C837DFF86FABAFAF0B9C8FA16" country="Indonesia" name="Nusa Tenggara Timur" pageId="2" pageNumber="122">Lesser Sunda Islands</collectingRegion>
. 
<collectingCounty id="715F4E22837DFF86FF4BFA1CB9E5FA17" box="[229,246,1433,1459]" pageId="2" pageNumber="122">It</collectingCounty>
is likely endemic to 
<collectingCountry id="E096763E837DFF86FE4FFA1DBB41FA16" box="[481,594,1432,1458]" name="Indonesia" pageId="2" pageNumber="122">Indonesia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="983E36AE837DFF86FF13FA39B8C0FA5E" blockId="2.[136,1452,856,2035]" pageId="2" pageNumber="122">
Etymology:—In honor of the late Professor Peter Hovenkamp based at 
<collectionCode id="FE90AE6B837DFF86FC73FA38BAFEFA73" box="[989,1005,1469,1495]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="2" pageNumber="122" type="Herbarium">L</collectionCode>
for his contributions to pteridology and Flora Malesiana in particular.
</paragraph>
<paragraph id="983E36AE837DFF86FF13F980B9B5F80F" blockId="2.[136,1452,856,2035]" pageId="2" pageNumber="122">
<emphasis id="AAF5EABC837DFF86FF13F980B813F9BB" bold="true" box="[189,256,1541,1567]" pageId="2" pageNumber="122">Notes</emphasis>
:— 
<taxonomicName id="5F814D2D837DFF86FE8AF981BB90F9BA" authorityName="Xu &amp; Liu &amp; Chen &amp; Zhang" authorityYear="2023" box="[292,643,1540,1566]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="hovenkampii">
<emphasis id="AAF5EABC837DFF86FE8AF981BB90F9BA" box="[292,643,1540,1566]" italics="true" pageId="2" pageNumber="122">Hymenasplenium hovenkampii</emphasis>
</taxonomicName>
is obviously a member of the 
<taxonomicName id="5F814D2D837DFF86FBA6F980BDA8F9BA" authorityName="Tagawa" authorityYear="1938" baseAuthorityName="Kunze ex Mettenius" baseAuthorityYear="1859" box="[1032,1211,1540,1566]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="cheilosorum">
<emphasis id="AAF5EABC837DFF86FBA6F980BDA8F9BA" box="[1032,1211,1540,1566]" italics="true" pageId="2" pageNumber="122">H. cheilosorum</emphasis>
</taxonomicName>
group based on its sori terminal on subtending veins and situated in or below ultimate segments of pinnae (
<figureCitation id="00BA2A2B837DFF86FB26F9ADBDFEF9E7" box="[1160,1261,1576,1603]" captionStart="FIGURE 2" captionStartId="3.[136,229,1291,1313]" captionTargetBox="[151,1436,190,1267]" captionTargetId="figure-32@3.[151,1436,190,1267]" captionTargetPageId="3" captionText="FIGURE 2. Hymenasplenium hovenkampii sp. nov.—A. Enlarged portion of laminae (some sori distributed in the pinnae are indicated by white arrows).—B. Fronds showing that the pinnae are lobed to the costa." figureDoi="http://doi.org/10.5281/zenodo.7889429" httpUri="https://zenodo.org/record/7889429/files/figure.png" pageId="2" pageNumber="122">Figure 2</figureCitation>
, Xu 
<emphasis id="AAF5EABC837DFF86FA86F9ACBC70F9E6" box="[1320,1379,1576,1602]" italics="true" pageId="2" pageNumber="122">et al.</emphasis>
2018, 2019a). This species is the only species with bipinnatisect laminae in 
<taxonomicName id="5F814D2D837DFF86FC35F9C9BD4EF9C2" authorityName="Hayata" authorityYear="1927" box="[923,1117,1612,1638]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="genus">
<emphasis id="AAF5EABC837DFF86FC35F9C9BD4EF9C2" box="[923,1117,1612,1638]" italics="true" pageId="2" pageNumber="122">Hymenasplenium</emphasis>
</taxonomicName>
. Up to now, three species, 
<taxonomicName id="5F814D2D837DFF86FA3EF9C8B801F92E" authorityName="Tagawa" authorityYear="1938" baseAuthorityName="Kunze ex Mettenius" baseAuthorityYear="1859" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="cheilosorum">
<emphasis id="AAF5EABC837DFF86FA3EF9C8B801F92E" italics="true" pageId="2" pageNumber="122">H. cheilosorum</emphasis>
</taxonomicName>
, 
<taxonomicName id="5F814D2D837DFF86FE8FF9F4BA0EF92E" authority="N. Murakami &amp; J. Yokoyama" authorityName="N. Murakami &amp; J. Yokoyama" box="[289,797,1648,1675]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="inthanonense">
<emphasis id="AAF5EABC837DFF86FE8FF9F4B8CBF92E" box="[289,472,1648,1674]" italics="true" pageId="2" pageNumber="122">H. inthanonense</emphasis>
N.Murakami &amp; J.Yokoyama
</taxonomicName>
in 
<bibRefCitation id="FC104B5F837DFF86FCECF9F5BD72F92F" author="Iwatsuki, K. &amp; Yokoyama, J. &amp; Murakami, N." box="[834,1121,1648,1675]" pageId="2" pageNumber="122" pagination="40 - 52" refId="ref2320" refString="Iwatsuki, K., Yokoyama, J. &amp; Murakami, N. (1998) Hymenasplenium inthanonense (Aspleniaceae), a new fern species from Doi Inthanon, and its phylogenetic status. Thai Forest Bulletin (Botany) 26: 40 - 52." type="journal article" year="1998">
Iwatsuki 
<emphasis id="AAF5EABC837DFF86FC07F9F4BAF1F92E" box="[937,994,1648,1674]" italics="true" pageId="2" pageNumber="122">et al.</emphasis>
(1998: 43)
</bibRefCitation>
, and 
<taxonomicName id="5F814D2D837DFF86FB32F9F4BC27F92E" box="[1180,1332,1648,1674]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="sabahense">
<emphasis id="AAF5EABC837DFF86FB32F9F4BC27F92E" box="[1180,1332,1648,1674]" italics="true" pageId="2" pageNumber="122">H. sabahense</emphasis>
</taxonomicName>
have been recorded in the 
<taxonomicName id="5F814D2D837DFF86FE9BF910B8F2F90A" authorityName="Tagawa" authorityYear="1938" baseAuthorityName="Kunze ex Mettenius" baseAuthorityYear="1859" box="[309,481,1684,1710]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="cheilosorum">
<emphasis id="AAF5EABC837DFF86FE9BF910B8F2F90A" box="[309,481,1684,1710]" italics="true" pageId="2" pageNumber="122">H. cheilosorum</emphasis>
</taxonomicName>
group. The pinnae of the three species are deeply or shallowly lobed. Among them, the pinnae of 
<taxonomicName id="5F814D2D837DFF86FF59F93CB89DF976" box="[247,398,1720,1746]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="sabahense">
<emphasis id="AAF5EABC837DFF86FF59F93CB89DF976" box="[247,398,1720,1746]" italics="true" pageId="2" pageNumber="122">H. sabahense</emphasis>
</taxonomicName>
are deeply lobed to half of the pinna width, 
<taxonomicName id="5F814D2D837DFF86FCD7F93CBD36F976" authorityName="Tagawa" authorityYear="1938" baseAuthorityName="Kunze ex Mettenius" baseAuthorityYear="1859" box="[889,1061,1720,1746]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="cheilosorum">
<emphasis id="AAF5EABC837DFF86FCD7F93CBD36F976" box="[889,1061,1720,1746]" italics="true" pageId="2" pageNumber="122">H. cheilosorum</emphasis>
</taxonomicName>
lobed up to 2/5 of the pinna width, and 
<taxonomicName id="5F814D2D837DFF86FF16F958B863F952" box="[184,368,1756,1782]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="inthanonense">
<emphasis id="AAF5EABC837DFF86FF16F958B863F952" box="[184,368,1756,1782]" italics="true" pageId="2" pageNumber="122">H. inthanonense</emphasis>
</taxonomicName>
only shallowly lobed. However, the pinnae of 
<taxonomicName id="5F814D2D837DFF86FC2DF958BD26F952" authorityName="Xu &amp; Liu &amp; Chen &amp; Zhang" authorityYear="2023" box="[899,1077,1756,1782]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="hovenkampii">
<emphasis id="AAF5EABC837DFF86FC2DF958BD26F952" box="[899,1077,1756,1782]" italics="true" pageId="2" pageNumber="122">H. hovenkampii</emphasis>
</taxonomicName>
are lobed to the costa, forming a bipinnatisect laminae. This is very different from other members in the genus. In addition, the acroscopic margins of the new species are deeply lobed and usually with 4–5 lobes and each lobe is deeply or shallowly crenate to dentate with two teeth, while the acroscopic margins of the other three species in the 
<taxonomicName id="5F814D2D837DFF86FBBFF8CCBDD3F8C6" authorityName="Tagawa" authorityYear="1938" baseAuthorityName="Kunze ex Mettenius" baseAuthorityYear="1859" box="[1041,1216,1864,1890]" class="Polypodiopsida" family="Aspleniaceae" genus="Hymenasplenium" kingdom="Plantae" order="Polypodiales" pageId="2" pageNumber="122" phylum="Tracheophyta" rank="species" species="cheilosorum">
<emphasis id="AAF5EABC837DFF86FBBFF8CCBDD3F8C6" box="[1041,1216,1864,1890]" italics="true" pageId="2" pageNumber="122">H. cheilosorum</emphasis>
</taxonomicName>
group are relatively shallowly lobed and with more than 5 lobes and each lobe is usually shallowly crenate to dentate with two teeth (
<tableCitation id="D5030315837DFF86FAC2F8E9B986F80F" captionStart="TABLE 1" captionStartId="2.[137,229,160,184]" captionTargetPageId="2" captionText="TABLE 1. Morphological comparison of Hymenasplenium hovenkampii and the three morphologically similar species." httpUri="http://table.plazi.org/id/CCFE6626837DFF86FF27FF25BC94FF1C" pageId="2" pageNumber="122" tableUuid="CCFE6626837DFF86FF27FF25BC94FF1C">Table 1</tableCitation>
).
</paragraph>
<paragraph id="983E36AE837DFF86FF13F831BA5BF857" blockId="2.[136,1452,856,2035]" pageId="2" pageNumber="122">
<materialsCitation id="28E93CF3837DFF86FF13F831BA57F857" ID-GBIF-Occurrence="4098986303" collectingDate="1982-05-20" collectionCode="E" country="Indonesia" county="Selected" elevation="1300" location="Ruteng" municipality="Manggarai" pageId="2" pageNumber="122" specimenCode="E 5352" specimenCount="1" stateProvince="Nusa Tenggara Timur" typeStatus="paratype">
<collectingCounty id="715F4E22837DFF86FF13F831B80FF86A" box="[189,284,1972,1998]" pageId="2" pageNumber="122">Selected</collectingCounty>
specimen examined (
<typeStatus id="473A880C837DFF86FDA0F830BB67F86B" box="[526,628,1973,1999]" pageId="2" pageNumber="122" type="paratype">Paratype</typeStatus>
):— 
<collectingCountry id="E096763E837DFF86FD33F830BA2FF86B" box="[669,828,1973,1999]" name="Indonesia" pageId="2" pageNumber="122">INDONESIA</collectingCountry>
. 
<collectingRegion id="5A45F84C837DFF86FCEBF830BD3BF86B" box="[837,1064,1973,1999]" country="Indonesia" name="Nusa Tenggara Timur" pageId="2" pageNumber="122">East Nusa Tenggara</collectingRegion>
: Lesser Sunda Islands, 
<collectingMunicipality id="785AACD4837DFF86FA85F831BCB4F86A" box="[1323,1447,1972,1998]" pageId="2" pageNumber="122">Manggarai</collectingMunicipality>
, 
<location id="9D5E6075837DFF86FF26F85CB9CFF857" LSID="urn:lsid:plazi:treatment:102887B8837FFF86FF26F831BA5BF857:9D5E6075837DFF86FF26F85CB9CFF857" box="[136,220,2009,2035]" country="Indonesia" county="Selected" municipality="Manggarai" name="Ruteng" pageId="2" pageNumber="122" stateProvince="Nusa Tenggara Timur">Ruteng</location>
, 
<quantity id="5F799B4B837DFF86FF49F85CB82DF857" box="[231,318,2009,2035]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="122" unit="m" value="1300.0">
<elevation id="13ACD19D837DFF86FF49F85CB82DF857" box="[231,318,2009,2035]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="122" unit="m" value="1300.0">1300 m</elevation>
</quantity>
, 
<date id="EC3F106E837DFF86FEE4F85CB8F1F857" box="[330,482,2009,2035]" pageId="2" pageNumber="122" value="1982-05-20">
<collectingDate id="FC7BE986837DFF86FEE4F85CB8F1F857" box="[330,482,2009,2035]" pageId="2" pageNumber="122" value="1982-05-20">20 May 1982</collectingDate>
</date>
, 
<emphasis id="AAF5EABC837DFF86FE43F85DBBB2F857" box="[493,673,2008,2035]" italics="true" pageId="2" pageNumber="122">
Schmutz 
<specimenCode id="C8279ED5837DFF86FDFFF85CBBB2F857" box="[593,673,2009,2035]" collectionCode="E" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="2" pageNumber="122" type="Herbarium">E 5352</specimenCode>
</emphasis>
(L-3507945!)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>