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<emphasis id="41E1EAF5F24FAC38FF67FD21FE7AA9CE" bold="true" box="[151,469,690,717]" italics="true" pageId="63" pageNumber="64">Onycholyda kumamotonis</emphasis>
(
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)
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(
<figureCitation id="EBAE2A62F24FAC38FF6FFD45FF59A9F2" box="[159,246,726,753]" captionStart="FIGURE 35" captionStartId="124.[151,250,1869,1893]" captionTargetBox="[151,1435,181,1845]" captionTargetId="figure-26@124.[151,1435,181,1845]" captionTargetPageId="124" captionText="FIGURE 35. Onycholyda kumamotonis, female, Japan (a) dorsally, (b) lateroventrally, (c) head &amp; thorax dorsally, (d) head &amp; thorax ventrolaterally, (e) face, (f) antenna, (g) sawsheath laterally, (h) claw." figureDoi="http://doi.org/10.5281/zenodo.6876576" httpUri="https://zenodo.org/record/6876576/files/figure.png" pageId="63" pageNumber="64">Figs 35</figureCitation>
,
<figureCitation id="EBAE2A62F24FAC38FEF2FD45FEB0A9F3" box="[258,287,726,752]" captionStart="FIGURE 36" captionStartId="125.[151,250,1869,1893]" captionTargetBox="[151,1435,181,1845]" captionTargetId="figure-19@125.[151,1435,181,1845]" captionTargetPageId="125" captionText="FIGURE 36. Onycholyda kumamotonis, male, Japan (a) dorsally, (b) lateroventrally, (c) head &amp; thorax dorsally, (d) head &amp; thorax ventrolaterally, (e) face, (f) antenna, (g) genitalia dorsally, (h) genitalia ventrally, (i) claw." figureDoi="http://doi.org/10.5281/zenodo.6876578" httpUri="https://zenodo.org/record/6876578/files/figure.png" pageId="63" pageNumber="64">36</figureCitation>
) (https://doi.org/10.6084/m9.figshare.11405067)
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<emphasis id="41E1EAF5F24FAC38FF67FC88FEF9A830" box="[151,342,795,819]" italics="true" pageId="63" pageNumber="64">Lyda kumamotonis</emphasis>
<bibRefCitation id="17044B16F24FAC38FEADFC88FD96A831" author="Matsumura, S." box="[349,569,794,819]" pageId="63" pageNumber="64" refId="ref74972" refString="Matsumura, S. (1912) Thousand Insects of Japan. Supplement IV. Keiseisha, Tokyo, 1 p. + 247 pp., 14 pls., 2 + 2 pp. [in Japanese and English]" type="book" year="1912">Matsumura, 1912: 75</bibRefCitation>
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.
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<treatmentCitationGroup id="538511C9F24FAC38FF67FCAAFA32A852" box="[151,1437,825,849]" pageId="63" pageNumber="64">
<treatmentCitation id="F23410F6F24FAC38FF67FCAAFDD5A852" author="Benes, K." box="[151,634,825,849]" page="387" pageId="63" pageNumber="64" year="1972">
<taxonomicName id="B4954D64F24FAC38FF67FCAAFDD5A852" authority=": Benes, 1972 b: 387" authorityName=": Benes" authorityPageNumber="387" authorityYear="1972" box="[151,634,825,849]" class="Insecta" family="Pamphiliidae" genus="Onycholyda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="kumamotonis">
<emphasis id="41E1EAF5F24FAC38FF67FCAAFE0DA852" box="[151,418,825,849]" italics="true" pageId="63" pageNumber="64">Onycholyda kumamotonis</emphasis>
:
<bibRefCitation id="17044B16F24FAC38FE43FCAAFDD5A852" author="Benes, K." box="[435,634,825,849]" pageId="63" pageNumber="64" pagination="378 - 395" refId="ref72824" refString="Benes, K. (1972 b) Generic classification of the tribe Pamphiliini (Hymenoptera, Pamphiliidae). Acta Entomologica Bohemoslovaca, 69, 378 - 395." type="journal article" year="1972">Beneš, 1972b: 387</bibRefCitation>
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;
<treatmentCitation id="F23410F6F24FAC38FD78FCAAFCD9A852" author="Shinohara, A." box="[648,886,825,849]" page="716" pageId="63" pageNumber="64" year="1985">
<bibRefCitation id="17044B16F24FAC38FD78FCAAFCD9A852" author="Shinohara, A." box="[648,886,825,849]" pageId="63" pageNumber="64" pagination="452 - 460" refId="ref76106" refString="Shinohara, A. (1985 c) Pamphilius itoi n. sp. from Japan and the southern Kuriles, with notes on Pamphilius balteatus (Fallen) (Hymenoptera, Pamphiliidae). Kontyu, Tokyo, 53, 452 - 460." type="journal article" year="1985">Shinohara, 1985c: 716</bibRefCitation>
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;
<treatmentCitation id="F23410F6F24FAC38FC74FCAAFBDDA852" author="Shinohara, A." box="[900,1138,825,849]" page="499" pageId="63" pageNumber="64" year="1987">
<bibRefCitation id="17044B16F24FAC38FC74FCAAFBDDA852" author="Shinohara, A." box="[900,1138,825,849]" pageId="63" pageNumber="64" pagination="486 - 501" refId="ref76282" refString="Shinohara, A. (1987 a) The sawfly genus Onycholyda (Hymenoptera, Pamphiliidae) of Japan V. Kontyu, Tokyo, 55, 486 - 501." type="journal article" year="1987">Shinohara, 1987a: 499</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="17044B16F24FAC38FB70FCAAFA32A852" author="Zhelochovtsev, A. N. &amp; Zinovjev, A. G." box="[1152,1437,825,849]" pageId="63" pageNumber="64" pagination="395 - 415" refId="ref81182" refString="Zhelochovtsev, A. N. &amp; Zinovjev, A. G. (1995) A list of the sawflies and horntails (Hymenoptera, Symphyta) of the fauna of Russia and adjacent territories. I. Entomologicheskoe Obozrenie, 74, 395 - 415. [in Russian]" type="journal article" year="1995">Zhelochovtsev &amp; Zinovjev,</bibRefCitation>
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1995: 398;
<bibRefCitation id="17044B16F24FAC38FEC9FCCBFD8EA873" author="Shinohara, A." box="[313,545,856,880]" pageId="63" pageNumber="64" pagination="359 - 438" refId="ref76809" refString="Shinohara, A. (2002 b) Systematics of the leaf-rolling or webspinning sawfly subfamily Pamphiliinae: a preliminary overview. In: Viitasaari, M. (Ed.), Sawflies (Hymenoptera, Symphyta) I. Tremex Press, Helsinki, pp. 359 - 438." type="book chapter" year="2002">Shinohara, 2002b: 422</bibRefCitation>
; Shinohara, 2004: 262;
<bibRefCitation id="17044B16F24FAC38FCE0FCCBFB91A873" author="Shinohara, A. &amp; Lelej, A. S." box="[784,1086,856,880]" pageId="63" pageNumber="64" pagination="922 - 942" refId="ref78590" refString="Shinohara, A. &amp; Lelej, A. S. (2007) 2. Sem. Pamphiliidae - Pautinnye Pilil'shchiki, ili Pilil'shchiki-tkachi. In: Lelej, A. S. (Ed.), Setchatokryloobraznye, Skorpionnitsy, Pereponchatokrylye. Ch. 5. Opredelitel' Nasekomykh Dal'nego Vostoka Rossii. Tom IV. Dal'nauka, Vladivostok, pp. 922 - 942." type="book chapter" year="2007">Shinohara &amp; Lelej, 2007: 929</bibRefCitation>
;
<bibRefCitation id="17044B16F24FAC38FBB8FCCBFA84A873" author="Taeger, A. &amp; Blank, S. M. &amp; Liston, A. D." box="[1096,1323,856,880]" pageId="63" pageNumber="64" pagination="1 - 1064" refId="ref80219" refString="Taeger, A., Blank, S. M. &amp; Liston, A. D. (2010) World catalog of Symphyta (Hymenoptera). Zootaxa, 2580 (1), 1 - 1064. https: // doi. org / 10.11646 / zootaxa. 2580.1.1" type="journal article" year="2010">
Taeger
<emphasis id="41E1EAF5F24FAC38FB62FCCAFB6AA873" box="[1170,1221,856,880]" italics="true" pageId="63" pageNumber="64">et al.</emphasis>
, 2010: 84
</bibRefCitation>
;
<bibRefCitation id="17044B16F24FAC38FAC5FCCBFA33A873" author="Sundukov, Yu. N. &amp; Lelej, A. S." box="[1333,1436,856,880]" pageId="63" pageNumber="64" pagination="62 - 119" refId="ref79983" refString="Sundukov, Yu. N. &amp; Lelej, A. S. (2012) Podotryad Symphyta - Sidyachebryukhie. In: Lelej, A. S. (Ed), Annotated Catalogue of the Insects of Russian Far East. Vol. I. Hymenoptera. Dalnauka, Vladivostok, pp. 62 - 119. [in Russian]" type="book chapter" year="2012">Sundukov</bibRefCitation>
&amp; Lelej, 2012: 108;
<bibRefCitation id="17044B16F24FAC38FE65FCE4FDDFA88C" author="Sundukov, Yu. N." box="[405,624,886,911]" pageId="63" pageNumber="64" pagination="241 - 258" refId="ref79811" refString="Sundukov, Yu. N. (2015) To the sawfly fauna (Hymenoptera, Symphyta) of southern Kuriles. A. I. Kurentsov's Annual Memorial Meetings, Issue XXVI, pp. 241 - 258. [in Russian with English abstract]" type="journal article" year="2015">Sundukov, 2015: 249</bibRefCitation>
;
<bibRefCitation id="17044B16F24FAC38FD8BFCE4FCFAA88C" author="Sundukov, Yu. N." box="[635,853,887,911]" pageId="63" pageNumber="64" pagination="20 - 117" refId="ref79858" refString="Sundukov, Yu. N. (2017) Suborder Symphyta - Sawflies and wood wasps. In: Lelej, A. S., Proshchalykin, M. Yu. &amp; Loktionov, V. M. (Eds.), Annotated catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute of the Russian Academy of Sciences, Supplement 6, pp. 20 - 117." type="journal article" year="2017">Sundukov, 2017: 103</bibRefCitation>
; Shinohara, 2019: 7;
<bibRefCitation id="17044B16F24FAC38FBDDFCE4FB53A88C" author="Shinohara, A." box="[1069,1276,887,911]" pageId="63" pageNumber="64" pagination="222 - 254" refId="ref77165" refString="Shinohara, A. (2020) Family Pamphiliidae. In: Naito, T., Shinohara, A., Hara, H. &amp; Ito, F., Sawflies and Woodwasps of Japan. Hokkaido University Press, Sapporo, pp. 3 - 23 + 222 - 254. [in Japanese]" type="book chapter" year="2020">Shinohara, 2020: 12</bibRefCitation>
, 236.
</paragraph>
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<taxonomicName id="B4954D64F24FAC38FF67FC06FE97A8AE" box="[151,312,917,941]" class="Insecta" family="Pamphiliidae" genus="Onycholyda" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="41E1EAF5F24FAC38FF67FC06FEBEA8AE" box="[151,273,917,941]" italics="true" pageId="63" pageNumber="64">Onycholyda</emphasis>
sp.
</taxonomicName>
:
<bibRefCitation id="17044B16F24FAC38FEB2FC06FD3EA8AE" author="Shinohara, A. &amp; Okutani, T." box="[322,657,917,941]" pageId="63" pageNumber="64" pagination="276 - 281" refId="ref78722" refString="Shinohara, A. &amp; Okutani, T. (1983) Host-plants of Japanese Pamphiliidae (Hymenoptera, Pamphiliidae). Kontyu, Tokyo, 51, 276 - 281." type="journal article" year="1983">Shinohara &amp; Okutani, 1983: 277</bibRefCitation>
.
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<typeStatus id="AC2E8845F24FAC38FF67FC4FFEA2A8F5" box="[151,269,988,1014]" pageId="63" pageNumber="64" type="lectotype">Lectotype</typeStatus>
designation.
</emphasis>
<bibRefCitation id="17044B16F24FAC38FE5FFC48FD2DA8F5" author="Shinohara, A." box="[431,642,987,1014]" pageId="63" pageNumber="64" pagination="711 - 720" refId="ref76151" refString="Shinohara, A. (1985 d) The sawfly genus Onycholyda (Hymenoptera, Pamphiliidae) of Japan II. Kontyu, Tokyo, 53, 711 - 720." type="journal article" year="1985">Shinohara (1985d)</bibRefCitation>
treated a specimen in Matsumuras collection in HU as the
<typeStatus id="AC2E8845F24FAC38FAC3FC48FA37A8F6" box="[1331,1432,987,1013]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
, stating “
<bibRefCitation id="17044B16F24FAC38FF02FB93FE69AF19" author="Matsumura, S." box="[242,454,1024,1050]" pageId="63" pageNumber="64" refId="ref74972" refString="Matsumura, S. (1912) Thousand Insects of Japan. Supplement IV. Keiseisha, Tokyo, 1 p. + 247 pp., 14 pls., 2 + 2 pp. [in Japanese and English]" type="book" year="1912">Matsumura (1912)</bibRefCitation>
described this species on the basis of a single female from Kiushu (
<collectingRegion id="B151F805F24FAC38FB4AFB93FAEEAF19" box="[1210,1345,1024,1050]" country="Japan" name="Kumamoto" pageId="63" pageNumber="64">Kumamoto</collectingRegion>
)”. However,
<bibRefCitation id="17044B16F24FAC38FF24FBB7FE0AAF3D" author="Matsumura, S." box="[212,421,1060,1086]" pageId="63" pageNumber="64" refId="ref74972" refString="Matsumura, S. (1912) Thousand Insects of Japan. Supplement IV. Keiseisha, Tokyo, 1 p. + 247 pp., 14 pls., 2 + 2 pp. [in Japanese and English]" type="book" year="1912">Matsumura (1912)</bibRefCitation>
did not give the actual number of the specimens examined and thus the specimen is a
<typeStatus id="AC2E8845F24FAC38FA9CFBB7FF64AF61" pageId="63" pageNumber="64" type="syntype">syntype</typeStatus>
. Here we designate this specimen as a
<typeStatus id="AC2E8845F24FAC38FD8DFBD4FD48AF62" box="[637,743,1095,1121]" pageId="63" pageNumber="64" type="lectotype">lectotype</typeStatus>
. It is labeled “
<collectingCountry id="0B827677F24FAC38FC76FBDBFC64AF61" box="[902,971,1096,1122]" name="Japan" pageId="63" pageNumber="64">Japan</collectingCountry>
,
<collectingRegion id="B151F805F24FAC38FC26FBDBFBF8AF61" box="[982,1111,1096,1122]" country="Japan" name="Kumamoto" pageId="63" pageNumber="64">Kumamoto</collectingRegion>
, Matsumura, 7/18” (HU).
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF24FAC38FF37FBF8FE08AFA9" pageId="63" pageNumber="64" type="materials_examined">
<paragraph id="732A36E7F24FAC38FF37FBF8FE08AFA9" blockId="63.[151,1437,987,1770]" pageId="63" pageNumber="64">
<emphasis id="41E1EAF5F24FAC38FF37FBF8FE1DAF86" bold="true" box="[199,434,1131,1157]" pageId="63" pageNumber="64">Material examined.</emphasis>
About
<specimenCount id="6593FD6EF24FAC38FDF6FBF8FD1FAF86" box="[518,688,1131,1158]" pageId="63" pageNumber="64" type="generic">355 specimens</specimenCount>
, including the
<typeStatus id="AC2E8845F24FAC38FCA8FBF8FC6DAF86" box="[856,962,1131,1157]" pageId="63" pageNumber="64" type="lectotype">lectotype</typeStatus>
. Two specimens are from the Russian Far East (
<bibRefCitation id="17044B16F24FAC38FF26FB1CFE34AFA9" author="Shinohara, A." box="[214,411,1167,1194]" pageId="63" pageNumber="64" pagination="711 - 720" refId="ref76151" refString="Shinohara, A. (1985 d) The sawfly genus Onycholyda (Hymenoptera, Pamphiliidae) of Japan II. Kontyu, Tokyo, 53, 711 - 720." type="journal article" year="1985">Shinohara 1985d</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF24FAC38FF37FB20FB2FAFCE" box="[199,1152,1203,1230]" pageId="63" pageNumber="64" type="distribution">
<paragraph id="732A36E7F24FAC38FF37FB20FB2FAFCE" blockId="63.[151,1437,987,1770]" box="[199,1152,1203,1230]" pageId="63" pageNumber="64">
<emphasis id="41E1EAF5F24FAC38FF37FB20FEF6AFCE" bold="true" box="[199,345,1203,1229]" pageId="63" pageNumber="64">Distribution</emphasis>
.
<collectingCountry id="0B827677F24FAC38FE97FB20FE1DAFCE" box="[359,434,1203,1229]" name="Russia" pageId="63" pageNumber="64">Russia</collectingCountry>
(
<collectingRegion id="B151F805F24FAC38FE31FB20FD86AFCE" box="[449,553,1203,1229]" country="Russia" name="Sakhalin" pageId="63" pageNumber="64">Sakhalin</collectingRegion>
),
<collectingCountry id="0B827677F24FAC38FDCCFB27FDD3AFCD" box="[572,636,1204,1230]" name="Japan" pageId="63" pageNumber="64">Japan</collectingCountry>
(
<collectingRegion id="B151F805F24FAC38FD7AFB20FD52AFCE" box="[650,765,1203,1229]" country="Japan" name="Hokkaido" pageId="63" pageNumber="64">Hokkaido</collectingRegion>
, Kunashiri Is., Honshu,?Kyushu).
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF24FAC38FF37FB4BFF43AE15" pageId="63" pageNumber="64" type="biology_ecology">
<paragraph id="732A36E7F24FAC38FF37FB4BFF43AE15" blockId="63.[151,1437,987,1770]" pageId="63" pageNumber="64">
<emphasis id="41E1EAF5F24FAC38FF37FB4BFEE4AFF1" bold="true" box="[199,331,1240,1266]" pageId="63" pageNumber="64">Host plant</emphasis>
.
<taxonomicName id="B4954D64F24FAC38FE90FB4BFE61AFF1" box="[352,462,1240,1266]" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
:
<taxonomicName id="B4954D64F24FAC38FE2FFB44FC13AFF2" authority="(Pall.) Maxim." authorityName="Maxim." baseAuthorityName="Pall." box="[479,956,1239,1265]" class="Magnoliopsida" family="Rosaceae" genus="Filipendula" kingdom="Plantae" order="Rosales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="camtschatica">
<emphasis id="41E1EAF5F24FAC38FE2FFB44FCACAFF2" box="[479,771,1239,1265]" italics="true" pageId="63" pageNumber="64">Filipendula camtschatica</emphasis>
(Pall.) Maxim.
</taxonomicName>
(
<bibRefCitation id="17044B16F24FAC38FC21FB44FAB7AFF1" author="Shinohara, A. &amp; Okutani, T." box="[977,1304,1239,1266]" pageId="63" pageNumber="64" pagination="276 - 281" refId="ref78722" refString="Shinohara, A. &amp; Okutani, T. (1983) Host-plants of Japanese Pamphiliidae (Hymenoptera, Pamphiliidae). Kontyu, Tokyo, 51, 276 - 281." type="journal article" year="1983">Shinohara &amp; Okutani 1983</bibRefCitation>
;
<bibRefCitation id="17044B16F24FAC38FADBFB44FF4FAE15" author="Shinohara, A." pageId="63" pageNumber="64" pagination="452 - 460" refId="ref76106" refString="Shinohara, A. (1985 c) Pamphilius itoi n. sp. from Japan and the southern Kuriles, with notes on Pamphilius balteatus (Fallen) (Hymenoptera, Pamphiliidae). Kontyu, Tokyo, 53, 452 - 460." type="journal article" year="1985">Shinohara 1985c</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF24FAC38FF37FAB3FD00ADEA" pageId="63" pageNumber="64" type="discussion">
<paragraph id="732A36E7F24FAC38FF37FAB3FD04AE81" blockId="63.[151,1437,987,1770]" pageId="63" pageNumber="64">
<emphasis id="41E1EAF5F24FAC38FF37FAB3FE9BAE39" bold="true" box="[199,308,1312,1338]" pageId="63" pageNumber="64">Remarks</emphasis>
. This species is not rare around
<taxonomicName id="B4954D64F24FAC38FD5EFA8CFC61AE3A" box="[686,974,1311,1337]" class="Magnoliopsida" family="Rosaceae" genus="Filipendula" kingdom="Plantae" order="Rosales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="camtschatica">
<emphasis id="41E1EAF5F24FAC38FD5EFA8CFC61AE3A" box="[686,974,1311,1337]" italics="true" pageId="63" pageNumber="64">Filipendula camtschatica</emphasis>
</taxonomicName>
bushes in
<collectingRegion id="B151F805F24FAC38FBBCFA8CFB12AE3A" box="[1100,1213,1311,1337]" country="Japan" name="Hokkaido" pageId="63" pageNumber="64">Hokkaido</collectingRegion>
, where two species of
<taxonomicName id="B4954D64F24FAC38FF47FAD0FE98AE5E" authorityName="Latreille" authorityYear="1803" box="[183,311,1347,1373]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="41E1EAF5F24FAC38FF47FAD0FE98AE5E" box="[183,311,1347,1373]" italics="true" pageId="63" pageNumber="64">Pamphilius</emphasis>
</taxonomicName>
,
<taxonomicName id="B4954D64F24FAC38FEB7FAD7FDC4AE5D" authority="(Smith, 1874)" authorityName="NSMT" baseAuthorityName="Smith" baseAuthorityYear="1874" box="[327,619,1347,1374]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="venustus">
<emphasis id="41E1EAF5F24FAC38FEB7FAD7FE6BAE5E" box="[327,452,1348,1373]" italics="true" pageId="63" pageNumber="64">P. venustus</emphasis>
(
<bibRefCitation id="17044B16F24FAC38FE25FAD0FDCDAE5D" author="Smith, F." box="[469,610,1347,1374]" pageId="63" pageNumber="64" pagination="373 - 409" refId="ref79508" refString="Smith, F. (1874) Descriptions of new species of Tenthredinidae, Ichneumonidae, Chrysididae, Formicidae &amp; c. of Japan. Transactions of the Entomological Society of London for the Year, 1874, 373 - 409. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00867. x" type="journal article" year="1874">Smith, 1874</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="B4954D64F24FAC38FD55FAD7FB85AE5D" authority="(Matsumura, 1912)" baseAuthorityName="Matsumura" baseAuthorityYear="1912" box="[677,1066,1348,1374]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="41E1EAF5F24FAC38FD55FAD7FCE8AE5E" box="[677,839,1348,1373]" italics="true" pageId="63" pageNumber="64">P. sapporensis</emphasis>
(
<bibRefCitation id="17044B16F24FAC38FCA7FAD7FB8EAE5D" author="Matsumura, S." box="[855,1057,1348,1374]" pageId="63" pageNumber="64" refId="ref74972" refString="Matsumura, S. (1912) Thousand Insects of Japan. Supplement IV. Keiseisha, Tokyo, 1 p. + 247 pp., 14 pls., 2 + 2 pp. [in Japanese and English]" type="book" year="1912">Matsumura, 1912</bibRefCitation>
)
</taxonomicName>
also occur together. These three pamphiliids are all associated with
<taxonomicName id="B4954D64F24FAC38FDD1FAF4FD0BAE82" baseAuthorityName="Kangas &amp; Syrjanen" baseAuthorityYear="1962" box="[545,676,1383,1409]" class="Magnoliopsida" family="Rosaceae" genus="Filipendula" kingdom="Plantae" order="Rosales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="genus">
<emphasis id="41E1EAF5F24FAC38FDD1FAF4FD0BAE82" box="[545,676,1383,1409]" italics="true" pageId="63" pageNumber="64">Filipendula</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="732A36E7F24FAC38FF37FA1FFD00ADEA" blockId="63.[151,1437,987,1770]" pageId="63" pageNumber="64">
In the COI analysis (
<figureCitation id="EBAE2A62F24FAC38FE43FA18FDBAAEA5" box="[435,533,1419,1446]" captionStart="FIGURE 139" captionStartId="228.[151,250,1667,1691]" captionTargetBox="[157,1426,190,1638]" captionTargetId="figure-26@228.[151,1436,181,1643]" captionTargetPageId="228" captionText="FIGURE 139. Condensed ML tree based on CO1 gene sequences, with details of Onycholyda, part 2." figureDoi="http://doi.org/10.5281/zenodo.6876841" httpUri="https://zenodo.org/record/6876841/files/figure.png" pageId="63" pageNumber="64">Fig. 139</figureCitation>
),
<specimenCount id="6593FD6EF24FAC38FDDAFA18FD70AEA6" box="[554,735,1419,1445]" pageId="63" pageNumber="64" type="generic">three specimens</specimenCount>
from
<collectingRegion id="B151F805F24FAC38FCD4FA18FC3DAEA6" box="[804,914,1419,1445]" country="Japan" name="Hokkaido" pageId="63" pageNumber="64">Hokkaido</collectingRegion>
(NSMT30554, 30788, 30789) and
<specimenCount id="6593FD6EF24FAC38FAD3FA18FF7DAEC9" pageId="63" pageNumber="64" type="generic">four specimens</specimenCount>
from Honshu (NSMT30730, 3078530787) each formed a different cluster and the two clusters had a sistergroup relationship supported by 100% UFBoot values. The maximum
<emphasis id="41E1EAF5F24FAC38FC5AFA47FC17AEEE" box="[938,952,1492,1517]" italics="true" pageId="63" pageNumber="64">p</emphasis>
-distance in the
<collectingRegion id="B151F805F24FAC38FB99FA40FB78AEEE" box="[1129,1239,1491,1517]" country="Japan" name="Hokkaido" pageId="63" pageNumber="64">Hokkaido</collectingRegion>
cluster was 0.2% and that in the Honshu cluster was 1%. The minimum
<emphasis id="41E1EAF5F24FAC38FCF3FA6BFCBEAD12" box="[771,785,1528,1553]" italics="true" pageId="63" pageNumber="64">p</emphasis>
-distance between the Honshu and
<collectingRegion id="B151F805F24FAC38FB6DFA64FAA4AD12" box="[1181,1291,1527,1553]" country="Japan" name="Hokkaido" pageId="63" pageNumber="64">Hokkaido</collectingRegion>
clusters was 2.2%. In the NaK tree (
<figureCitation id="EBAE2A62F24FAC38FE53F988FDAAAD35" box="[419,517,1563,1590]" captionStart="FIGURE 153" captionStartId="240.[151,250,1943,1967]" captionTargetBox="[178,1407,186,1910]" captionTargetId="figure-26@240.[172,1415,181,1919]" captionTargetPageId="240" captionText="FIGURE 153. Condensed ML tree based on NaK gene sequences, with details of Onycholyda." figureDoi="http://doi.org/10.5281/zenodo.6876872" httpUri="https://zenodo.org/record/6876872/files/figure.png" pageId="63" pageNumber="64">Fig. 153</figureCitation>
), the four Honshu specimens formed a cluster but the three
<collectingRegion id="B151F805F24FAC38FB41F988FAB0AD36" box="[1201,1311,1563,1589]" country="Japan" name="Hokkaido" pageId="63" pageNumber="64">Hokkaido</collectingRegion>
specimens did not and, with the Honshu cluster, formed a polytomous clade, which was sister to
<taxonomicName id="B4954D64F24FAC38FBAAF9D3FB62AD5A" authorityName=": Shinohara" authorityYear="1985" box="[1114,1229,1600,1626]" class="Insecta" family="Pamphiliidae" genus="Onycholyda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="armata">
<emphasis id="41E1EAF5F24FAC38FBAAF9D3FB62AD5A" box="[1114,1229,1600,1626]" italics="true" pageId="63" pageNumber="64">O. armata</emphasis>
</taxonomicName>
(this clade,
<taxonomicName id="B4954D64F24FAC38FAA6F9D3FEBDAD7E" authorityName=": Benes" authorityYear="1972" class="Insecta" family="Pamphiliidae" genus="Onycholyda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="kumamotonis">
<emphasis id="41E1EAF5F24FAC38FAA6F9D3FEBDAD7E" italics="true" pageId="63" pageNumber="64">O. kumamotonis</emphasis>
</taxonomicName>
+
<taxonomicName id="B4954D64F24FAC38FEDFF9F7FE0EAD7E" authorityName=": Shinohara" authorityYear="1985" box="[303,417,1636,1662]" class="Insecta" family="Pamphiliidae" genus="Onycholyda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="armata">
<emphasis id="41E1EAF5F24FAC38FEDFF9F7FE0EAD7E" box="[303,417,1636,1662]" italics="true" pageId="63" pageNumber="64">O. armata</emphasis>
</taxonomicName>
, was supported by 100% UFBoot value). The maximum
<emphasis id="41E1EAF5F24FAC38FBEEF9F7FB83AD7E" box="[1054,1068,1636,1661]" italics="true" pageId="63" pageNumber="64">p</emphasis>
-distance within
<taxonomicName id="B4954D64F24FAC38FB15F9F7FA33AD7E" authorityName=": Benes" authorityYear="1972" box="[1253,1436,1635,1662]" class="Insecta" family="Pamphiliidae" genus="Onycholyda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="kumamotonis">
<emphasis id="41E1EAF5F24FAC38FB15F9F7FA33AD7E" box="[1253,1436,1635,1662]" italics="true" pageId="63" pageNumber="64">O. kumamotonis</emphasis>
</taxonomicName>
was 0.4% and the nearest neighbour, diverging by a minimum of 0.2%, was
<emphasis id="41E1EAF5F24FAC38FC1DF91BFBC7ADA2" box="[1005,1128,1672,1698]" italics="true" pageId="63" pageNumber="64">
<taxonomicName id="B4954D64F24FAC38FC1DF91BFBCBADA2" authorityName=": Shinohara" authorityYear="1985" box="[1005,1124,1672,1698]" class="Insecta" family="Pamphiliidae" genus="Onycholyda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="armata">O. armata</taxonomicName>
.
</emphasis>
The
<collectingRegion id="B151F805F24FAC38FB50F914FAA1ADA2" box="[1184,1294,1671,1697]" country="Japan" name="Hokkaido" pageId="63" pageNumber="64">Hokkaido</collectingRegion>
and Honshu populations of
<taxonomicName id="B4954D64F24FAC38FECEF93FFE5AADC6" authorityName=": Benes" authorityYear="1972" box="[318,501,1707,1734]" class="Insecta" family="Pamphiliidae" genus="Onycholyda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="kumamotonis">
<emphasis id="41E1EAF5F24FAC38FECEF93FFE5AADC6" box="[318,501,1707,1734]" italics="true" pageId="63" pageNumber="64">O. kumamotonis</emphasis>
</taxonomicName>
certainly have some genetic divergence, but we will leave them as one species until more evidence of speciation becomes available.
</paragraph>
</subSubSection>
</treatment>
</document>