357 lines
42 KiB
XML
357 lines
42 KiB
XML
<document id="B19A99ECD1FD8FC7E993C5E3696F0383" ID-DOI="10.11646/zootaxa.4028.3.2" ID-GBIF-Dataset="2c603243-ea11-4169-9f2d-244c653e1703" ID-ISSN="1175-5326" ID-Zenodo-Dep="232682" ID-ZooBank="1F5BEBE1-4BAE-4EB4-8BDE-ACA0FF2D1866" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461286835024" checkinUser="plazi" docAuthor="Garrouste, Romain & Nel, André" docDate="2015" docId="CF6C9266AA60525217AB62D9FD41FCFB" docLanguage="en" docName="zt04028p366.pdf" docOrigin="Zootaxa 4028 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Petrolestes hendersoni Cockerell 1927" docType="treatment" docVersion="8" lastPageNumber="359" masterDocId="3355EA1EAA615257173C6760FF91FFEF" masterDocTitle="New Eocene damselflies and first Cenozoic damsel-dragonfly of the isophlebiopteran lineage (Insecta: Odonata)" masterLastPageNumber="366" masterPageNumber="354" pageNumber="355" updateTime="1698615234362" updateUser="plazi">
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<mods:titleInfo id="EC36EF745B593AF4DCDDA1F93A5209C2">
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<mods:title id="5104A7C688903AE082F1ADF16ACB8319">New Eocene damselflies and first Cenozoic damsel-dragonfly of the isophlebiopteran lineage (Insecta: Odonata)</mods:title>
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</mods:titleInfo>
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<mods:name id="D97564B97005F49A1C64291377713D6E" type="personal">
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<mods:roleTerm id="08CA1FAF1A065381E559589B6494D80C">Author</mods:roleTerm>
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<mods:namePart id="3EC37C959EF202B78EB7F7F123F0665D">Garrouste, Romain</mods:namePart>
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</mods:name>
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<mods:name id="233431A99DE00E9E5CFFDE67EB43EC7B" type="personal">
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<mods:roleTerm id="82B8D3E25877171E646CFC9CE2FAF1C5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="19A21EB54DDA79B7015C379F1D992CCA">Nel, André</mods:namePart>
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<mods:typeOfResource id="DAD013E519077EEF187777BE02A8DEB1">text</mods:typeOfResource>
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<mods:title id="3B8712A6CCAD371D07073FE26238BB48">Zootaxa</mods:title>
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<mods:part id="D3272AD74CFADEC4C1B9C5FB65506C03">
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<mods:date id="561C028C914F59FED9FF30680A44EB0E">2015</mods:date>
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<mods:detail id="16CE44B3D13FDC6EE9D0AE5AB7A48F84" type="volume">
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<mods:number id="0F0F36E7F5363D9BF18C1AFC8D4DDC5D">4028</mods:number>
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<mods:detail id="31627A48E5B6B95FA997D14624F4DD5E" type="issue">
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<mods:number id="0A9AD979BFCAA12D0B3DCDE831102E60">3</mods:number>
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<mods:start id="1CBD1ADB4DE4720AAE2DAD91FF383003">354</mods:start>
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<mods:end id="35617B82273066FEA3D01A52D26BC4BE">366</mods:end>
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<mods:identifier id="A3042B9864054DC4DAB454453D5C8AA2" type="DOI">10.11646/zootaxa.4028.3.2</mods:identifier>
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<mods:identifier id="F5236EA7B8FF5321E86E02A706146C73" type="GBIF-Dataset">2c603243-ea11-4169-9f2d-244c653e1703</mods:identifier>
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<mods:identifier id="7F26E77789CC48D0F91BA49C27CBC63D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="EC06C33CB3F14B04E36E7D78D22A138A" type="Zenodo-Dep">232682</mods:identifier>
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<mods:identifier id="B5D3CF5338E22F0DEBDCE581EDE7CA0E" type="ZooBank">1F5BEBE1-4BAE-4EB4-8BDE-ACA0FF2D1866</mods:identifier>
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<treatment id="CF6C9266AA60525217AB62D9FD41FCFB" ID-DOI="http://doi.org/10.5281/zenodo.6105804" ID-GBIF-Taxon="119641570" ID-Zenodo-Dep="6105804" LSID="urn:lsid:plazi:treatment:CF6C9266AA60525217AB62D9FD41FCFB" httpUri="http://treatment.plazi.org/id/CF6C9266AA60525217AB62D9FD41FCFB" lastPageId="5" lastPageNumber="359" pageId="1" pageNumber="355">
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<subSubSection id="0FDF70FBAA60525617AB62D9FE83FA1A" pageId="1" pageNumber="355" type="nomenclature">
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<paragraph id="477A2370AA60525617AB62D9FDE1FA3C" blockId="1.[151,624,1465,1525]" box="[151,624,1465,1491]" pageId="1" pageNumber="355">
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<heading id="1C32941CAA60525617AB62D9FDE1FA3C" bold="true" box="[151,624,1465,1491]" fontSize="11" level="1" pageId="1" pageNumber="355" reason="1">
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<taxonomicName id="80C558F3AA60525617AB62D9FDE1FA3C" authority="Cockerell, 1927" authorityName="Cockerell" authorityYear="1927" box="[151,624,1465,1491]" class="Insecta" family="Dysagrionidae" genus="Petrolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="355" phylum="Arthropoda" rank="species" species="hendersoni">
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<emphasis id="75B1FF62AA60525617AB62D9FDE1FA3C" bold="true" box="[151,624,1465,1491]" pageId="1" pageNumber="355">
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<emphasis id="75B1FF62AA60525617AB62D9FE36FA3C" bold="true" box="[151,423,1465,1491]" italics="true" pageId="1" pageNumber="355">Petrolestes hendersoni</emphasis>
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Cockerell, 1927
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="477A2370AA60525617AB62BCFE83FA1A" blockId="1.[151,624,1465,1525]" box="[151,274,1500,1525]" pageId="1" pageNumber="355">
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(
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<figureCitation id="DFFE3FF5AA60525617A362BCFE99FA1A" box="[159,264,1500,1525]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,959,981]" captionStartId-1="2.[151,250,1344,1366]" captionStartId-2="2.[151,250,1952,1974]" captionStartId-3="3.[151,250,1061,1083]" captionStartId-4="4.[151,250,1933,1955]" captionTargetBox-0="[257,1329,267,935]" captionTargetBox-1="[256,1332,1066,1313]" captionTargetBox-2="[280,1310,1454,1923]" captionTargetBox-3="[340,1246,193,1039]" captionTargetBox-4="[267,1319,913,1911]" captionTargetId-0="figure@2.[257,1329,267,935]" captionTargetId-1="figure@2.[248,1339,1062,1323]" captionTargetId-2="figure@2.[272,1314,1447,1931]" captionTargetId-3="figure@3.[340,1247,193,1040]" captionTargetId-4="figure@4.[267,1320,913,1911]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="2" captionTargetPageId-3="3" captionTargetPageId-4="4" captionText-0="FIGURE 1. Petrolestes hendersoni Cockerell, 1927, USNM without number, photograph of habitus. Scale bar represents 5.0 mm." captionText-1="FIGURE 2. Petrolestes hendersoni Cockerell, 1927, USNM without number, drawing of forewing. Scale bar represents 3.0 mm." captionText-2="FIGURE 3. Petrolestes hendersoni Cockerell, 1927, USNM without number, drawing of hind wing. Scale bar represents 3.0 mm." captionText-3="FIGURE 4. Petrolestes hendersoni Cockerell, 1927, USNM without number, photograph of thorax, arrows correspond to the extremities of the interpleural suture. Scale bar represents 4.0 mm." captionText-4="FIGURE 5. Petrolestes hendersoni Cockerell, 1927, USNM without number, photograph of legs. Scale bar represents 4.0 mm." httpUri-0="https://zenodo.org/record/232683/files/figure.png" httpUri-1="https://zenodo.org/record/232684/files/figure.png" httpUri-2="https://zenodo.org/record/232685/files/figure.png" httpUri-3="https://zenodo.org/record/232686/files/figure.png" httpUri-4="https://zenodo.org/record/232687/files/figure.png" pageId="1" pageNumber="355">Figs. 1–5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="0FDF70FBAA60525617AB6144FC01F8FA" pageId="1" pageNumber="355" type="materials_examined">
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<paragraph id="477A2370AA60525617AB6144FEDCF923" blockId="1.[151,1437,1572,2029]" pageId="1" pageNumber="355">
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<emphasis id="75B1FF62AA60525617AB6144FE97F9D2" bold="true" box="[151,262,1572,1597]" pageId="1" pageNumber="355">Material.</emphasis>
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<typeStatus id="987E9DD2AA60525616336144FEE0F9D2" box="[271,369,1572,1597]" pageId="1" pageNumber="355" type="holotype">holotype</typeStatus>
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specimen
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<collectionCode id="21D4BBB5AA60525616D26144FDBFF9D3" LSID="urn:lsid:biocol.org:col:34854" box="[494,558,1572,1596]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="1" pageNumber="355">UCM</collectionCode>
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15619, University of Colorado, Boulder, Geology Section, Museum Annex,
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<collectingCountry id="3FD263E0AA60525617AB6128FF42F98F" box="[151,211,1608,1632]" name="United States of America" pageId="1" pageNumber="355">USA</collectingCountry>
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; new specimen
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<collectionCode id="21D4BBB5AA60525616BF6128FE45F98F" LSID="urn:lsid:biocol.org:col:34871" box="[387,468,1608,1632]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="355">USNM</collectionCode>
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without number, Smithsonian Institution, National Museum of Natural History,
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<collectingCountry id="3FD263E0AA605256126A6128FA03F98F" box="[1366,1426,1608,1632]" name="United States of America" pageId="1" pageNumber="355">USA</collectingCountry>
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. Age and outcrop. Eocene, Green River Formation; the
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<typeStatus id="987E9DD2AA60525615C1610DFCBCF96A" box="[765,813,1645,1669]" pageId="1" pageNumber="355">type</typeStatus>
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specimen
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<collectionCode id="21D4BBB5AA6052561499610CFC74F96B" LSID="urn:lsid:biocol.org:col:34854" box="[933,997,1644,1668]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="1" pageNumber="355">UCM</collectionCode>
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15619 comes from the Ute Trail, Roan Plateau, Garfield County; the new specimen
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<collectionCode id="21D4BBB5AA60525615B561F0FD4BF947" LSID="urn:lsid:biocol.org:col:34871" box="[649,730,1680,1704]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="355">USNM</collectionCode>
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without number comes from Wighton Garfield County; both in Colorado,
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<collectingCountry id="3FD263E0AA605256163161D4FED8F923" box="[269,329,1716,1740]" name="United States of America" pageId="1" pageNumber="355">USA</collectingCountry>
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.
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</paragraph>
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<paragraph id="477A2370AA60525617FB61B7FC01F8FA" blockId="1.[151,1437,1572,2029]" pageId="1" pageNumber="355">
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<emphasis id="75B1FF62AA60525617FB61B7FE28F91F" bold="true" box="[199,441,1751,1776]" pageId="1" pageNumber="355">Emended diagnosis.</emphasis>
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Wing characters only. Intercalaries between MA and MP present just distal of nodus level; 17 postnodals; wing rather large, 8.0 mm wide at nodus level.
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</paragraph>
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</subSubSection>
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<subSubSection id="0FDF70FBAA60525417FB607FFB59F8AA" lastPageId="3" lastPageNumber="357" pageId="1" pageNumber="355" type="description">
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<paragraph id="477A2370AA60525617FB607FFAF7F86F" blockId="1.[151,1437,1572,2029]" pageId="1" pageNumber="355">
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<emphasis id="75B1FF62AA60525617FB607FFDA4F8D7" bold="true" box="[199,565,1823,1848]" pageId="1" pageNumber="355">Description of new specimen.</emphasis>
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Body with head and five basal segment of abdomen attached to thorax, together with the four wings; only the left forewing is nearly complete, basal half of left hind wing and basal fourth of right wings preserved; no trace of secondary genital structures on abdominal segment 2 (female specimen).
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</paragraph>
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<paragraph id="477A2370AA60525517FB60ECFD2CFF3A" blockId="1.[151,1437,1572,2029]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="356" pageId="1" pageNumber="355">
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Head rather broad,
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<quantity id="803D8E95AA605256169D60ECFE69F84B" box="[417,504,1932,1957]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="1" pageNumber="355" unit="mm" value="3.6">3.6 mm</quantity>
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long,
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<quantity id="803D8E95AA605256157C60ECFD06F84B" box="[576,663,1932,1957]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="1" pageNumber="355" unit="mm" value="5.2">5.2 mm</quantity>
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wide, eyes 3.0 mm wide, widely separated,
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<quantity id="803D8E95AA60525613B160ECFB75F84B" box="[1165,1252,1932,1956]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="355" unit="mm" value="2.4">2.4 mm</quantity>
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apart; ocelli not discernable; thorax 6.0 mm high, moderately prognathous, thoracic skewness (sensu
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<bibRefCitation id="23545E81AA605256137060CFFA05F827" author="Needham" box="[1100,1428,1967,1992]" pageId="1" pageNumber="355" refString="Needham, J. G. & Anthony, M. H. (1903) The skewness of the thorax in the Odonata. Journal of the New York Entomological Society, 11, 117 - 123." type="journal article" year="1903">Needham and Anthony, 1903</bibRefCitation>
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) moderate, comparable to that of a modern
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<taxonomicName id="80C558F3AA60525615AC60B4FC98F802" box="[656,777,2004,2029]" class="Insecta" family="Aeshnidae" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="355" phylum="Arthropoda" rank="family">Aeshnidae</taxonomicName>
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, angle
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<date id="337B05B0AA605256145D60B5FC08F802" box="[865,921,2004,2029]" pageId="1" pageNumber="355" value="1935-10">x 35</date>
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°, angle z ca. 30°; interpleural suture (sensu
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<bibRefCitation id="23545E81AA63525517AB67F7FEA5FF5F" author="Asahina" box="[151,308,151,176]" pageId="2" pageNumber="356" refString="Asahina, S. (1957) On the pterothoracic interpleural suture of Mesozoic and Recent Odonata. J ournal of the Faculty of Sciences, Hokkaido University, Zoology, (6), 13, 1 - 7." type="journal article" year="1957">Asahina 1957</bibRefCitation>
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) incomplete but formed at dorsal 1/2; abdomen 2.0 mm wide. Legs robust, with tibia spines strong, slightly longer than distance between two spines.
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</paragraph>
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<caption id="13BA73F8AA63525517AB64DFFF57FC1B" httpUri="https://zenodo.org/record/232683/files/figure.png" pageId="2" pageNumber="356" targetBox="[257,1329,267,935]" targetPageId="2">
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<paragraph id="477A2370AA63525517AB64DFFF57FC1B" blockId="2.[151,1436,959,1012]" pageId="2" pageNumber="356">
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<emphasis id="75B1FF62AA63525517AB64DFFE85FC3B" bold="true" box="[151,276,959,981]" pageId="2" pageNumber="356">FIGURE 1.</emphasis>
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<taxonomicName id="80C558F3AA635255162764DFFD3EFC3A" authority="Cockerell, 1927" authorityName="Cockerell" authorityYear="1927" box="[283,687,959,981]" class="Insecta" family="Dysagrionidae" genus="Petrolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="356" phylum="Arthropoda" rank="species" species="hendersoni">
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<emphasis id="75B1FF62AA635255162764DFFD92FC3A" box="[283,515,959,981]" italics="true" pageId="2" pageNumber="356">Petrolestes hendersoni</emphasis>
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Cockerell, 1927
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</taxonomicName>
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, USNM without number, photograph of habitus. Scale bar represents 5.0 mm.
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</paragraph>
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</caption>
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<caption id="13BA73F8AA63525517AB6220FF57FA9A" httpUri="https://zenodo.org/record/232684/files/figure.png" pageId="2" pageNumber="356" targetBox="[256,1332,1066,1313]" targetPageId="2">
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<paragraph id="477A2370AA63525517AB6220FF57FA9A" blockId="2.[151,1436,1344,1397]" pageId="2" pageNumber="356">
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<emphasis id="75B1FF62AA63525517AB6220FE84FABA" bold="true" box="[151,277,1344,1366]" pageId="2" pageNumber="356">FIGURE 2.</emphasis>
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<taxonomicName id="80C558F3AA63525516216220FD25FAB9" authority="Cockerell, 1927" authorityName="Cockerell" authorityYear="1927" box="[285,692,1344,1366]" class="Insecta" family="Dysagrionidae" genus="Petrolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="356" phylum="Arthropoda" rank="species" species="hendersoni">
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<emphasis id="75B1FF62AA63525516216220FD94FAB9" box="[285,517,1344,1366]" italics="true" pageId="2" pageNumber="356">Petrolestes hendersoni</emphasis>
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Cockerell, 1927
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</taxonomicName>
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, USNM without number, drawing of forewing. Scale bar represents 3.0 mm.
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</paragraph>
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</caption>
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<caption id="13BA73F8AA63525517AB60C0FF57F83A" httpUri="https://zenodo.org/record/232685/files/figure.png" pageId="2" pageNumber="356" targetBox="[280,1310,1454,1923]" targetPageId="2">
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||
<paragraph id="477A2370AA63525517AB60C0FF57F83A" blockId="2.[151,1436,1952,2005]" pageId="2" pageNumber="356">
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||
<emphasis id="75B1FF62AA63525517AB60C0FE85F85A" bold="true" box="[151,276,1952,1974]" pageId="2" pageNumber="356">FIGURE 3.</emphasis>
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<taxonomicName id="80C558F3AA635255162760C0FD21F859" authority="Cockerell, 1927" authorityName="Cockerell" authorityYear="1927" box="[283,688,1952,1974]" class="Insecta" family="Dysagrionidae" genus="Petrolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="356" phylum="Arthropoda" rank="species" species="hendersoni">
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<emphasis id="75B1FF62AA635255162760C0FD93F859" box="[283,514,1952,1974]" italics="true" pageId="2" pageNumber="356">Petrolestes hendersoni</emphasis>
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||
Cockerell, 1927
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</taxonomicName>
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, USNM without number, drawing of hind wing. Scale bar represents 3.0 mm.
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</paragraph>
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</caption>
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<caption id="13BA73F8AA62525417AB6345FCA8FBB5" httpUri="https://zenodo.org/record/232686/files/figure.png" pageId="3" pageNumber="357" targetBox="[340,1246,193,1039]" targetPageId="3">
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||
<paragraph id="477A2370AA62525417AB6345FCA8FBB5" blockId="3.[151,1436,1061,1114]" pageId="3" pageNumber="357">
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||
<emphasis id="75B1FF62AA62525417AB6345FE82FBD5" bold="true" box="[151,275,1061,1083]" pageId="3" pageNumber="357">FIGURE 4.</emphasis>
|
||
<taxonomicName id="80C558F3AA62525416276345FD3EFBD4" authority="Cockerell, 1927" authorityName="Cockerell" authorityYear="1927" box="[283,687,1061,1083]" class="Insecta" family="Dysagrionidae" genus="Petrolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="357" phylum="Arthropoda" rank="species" species="hendersoni">
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<emphasis id="75B1FF62AA62525416276345FD92FBD4" box="[283,515,1061,1083]" italics="true" pageId="3" pageNumber="357">Petrolestes hendersoni</emphasis>
|
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Cockerell, 1927
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</taxonomicName>
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, USNM without number, photograph of thorax, arrows correspond to the extremities of the interpleural suture. Scale bar represents 4.0 mm.
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</paragraph>
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</caption>
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<paragraph id="477A2370AA62525417FB63E1FD83F911" blockId="3.[151,1437,1153,2006]" pageId="3" pageNumber="357">
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Wings hyaline, forewing ca. 40.0 mm long; max. width
|
||
<quantity id="803D8E95AA625254140A63E1FC1CFB76" box="[822,909,1153,1178]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="357" unit="mm" value="8.5">8.5 mm</quantity>
|
||
, wing 8.0 mm wide at nodus level; a very short petiole,
|
||
<quantity id="803D8E95AA62525417CF63C5FEDBFB52" box="[243,330,1189,1214]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="357" unit="mm" value="1.5">1.5 mm</quantity>
|
||
long; anal area with one row of cells between AA and AP; subdiscoidal cell broad elongate,
|
||
<quantity id="803D8E95AA625254124563C6FF52FB0E" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="357" unit="mm" value="1.8">1.8 mm</quantity>
|
||
long,
|
||
<quantity id="803D8E95AA625254163263AAFEF6FB0E" box="[270,359,1226,1250]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="357" unit="mm" value="0.8">0.8 mm</quantity>
|
||
wide; distance from base to arculus 5.0 mm, from arculus to nodus
|
||
<quantity id="803D8E95AA62525413B863AAFB7AFB0E" box="[1156,1259,1226,1250]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="3" pageNumber="357" unit="mm" value="11.2">11.2 mm</quantity>
|
||
, from nodus to pterostigma
|
||
<quantity id="803D8E95AA625254161F638DFE1BFAEA" box="[291,394,1261,1286]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="3" pageNumber="357" unit="mm" value="16.5">16.5 mm</quantity>
|
||
; nodus in a basal position; a significant increase of spine density at the apical costal margin; pterostigma long and broad, 4.0 mm long,
|
||
<quantity id="803D8E95AA625254154D6272FD56FAC6" box="[625,711,1298,1322]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="357" unit="mm" value="0.8">0.8 mm</quantity>
|
||
wide, covering six cells; pterostigmal brace perpendicular to RA and
|
||
<collectingCountry id="3FD263E0AA62525417FB6256FF61FAA1" box="[199,240,1334,1358]" name="Poland" pageId="3" pageNumber="357">RP</collectingCountry>
|
||
1; distance from wing base to Ax1
|
||
<quantity id="803D8E95AA62525415496256FD5CFAA2" box="[629,717,1334,1358]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="357" unit="mm" value="3.8">3.8 mm</quantity>
|
||
, from Ax1 to Ax2
|
||
<quantity id="803D8E95AA625254149F6255FC6DFAA2" box="[931,1020,1333,1358]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="357" unit="mm" value="1.5">1.5 mm</quantity>
|
||
, Ax2 nearly aligned with arculus; no supplementary antenodal crossvein; discoidal cell unicellular,
|
||
<quantity id="803D8E95AA625254146A623AFC3CFA9E" box="[854,941,1370,1394]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="357" unit="mm" value="1.4">1.4 mm</quantity>
|
||
long,
|
||
<quantity id="803D8E95AA62525414C9623AFBDDFA9E" box="[1013,1100,1370,1394]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="357" unit="mm" value="1.4">1.4 mm</quantity>
|
||
wide, distinctly broadened in its distal part, basal side
|
||
<quantity id="803D8E95AA625254168E621EFD9DFA7A" box="[434,524,1406,1430]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="357" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, anterior side
|
||
<quantity id="803D8E95AA6252541593621EFC99FA7A" box="[687,776,1406,1430]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="357" unit="mm" value="1.2">1.2 mm</quantity>
|
||
, posterior side
|
||
<quantity id="803D8E95AA625254148B621EFB80FA7A" box="[951,1041,1406,1430]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="357" unit="mm" value="1.4">1.4 mm</quantity>
|
||
, distal side MAb
|
||
<quantity id="803D8E95AA62525413E1621EFAA6FA7A" box="[1245,1335,1406,1430]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="357" unit="mm" value="1.4">1.4 mm</quantity>
|
||
; ScP not crossing through nodus as in
|
||
<emphasis id="75B1FF62AA62525416D462C1FDF5FA55" box="[488,612,1441,1466]" italics="true" pageId="3" pageNumber="357">Stenolestes</emphasis>
|
||
Scudder, 1895; nodal crossvein very oblique; subnodus slightly oblique with corresponding crossvein below (between
|
||
<collectingCountry id="3FD263E0AA625254159762A6FD40FA31" box="[683,721,1478,1502]" name="Poland" pageId="3" pageNumber="357">RP</collectingCountry>
|
||
1/2 and IR2) of inverted obliquity; ca. 17 postnodal crossveins preserved, not aligned with the 16 postsubnodal crossveins; bases of
|
||
<collectingCountry id="3FD263E0AA6252541492628AFC43F9ED" box="[942,978,1514,1538]" name="Poland" pageId="3" pageNumber="357">RP</collectingCountry>
|
||
3/4 and IR2 between arculus and nodus, closer to nodus than to arculus, base of
|
||
<collectingCountry id="3FD263E0AA6252541572616EFDE3F9C9" box="[590,626,1550,1574]" name="Poland" pageId="3" pageNumber="357">RP</collectingCountry>
|
||
3/4
|
||
<quantity id="803D8E95AA62525415A7616DFD61F9CA" box="[667,752,1549,1574]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="357" unit="mm" value="6.5">6.5 mm</quantity>
|
||
from arculus and
|
||
<quantity id="803D8E95AA6252541486616EFB81F9CA" box="[954,1040,1550,1574]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="3" pageNumber="357" unit="mm" value="4.7">4.7 mm</quantity>
|
||
from nodus; base of
|
||
<collectingCountry id="3FD263E0AA62525413C7616EFA8FF9C9" box="[1275,1310,1550,1574]" name="Poland" pageId="3" pageNumber="357">RP</collectingCountry>
|
||
2 two cells,
|
||
<quantity id="803D8E95AA62525417AB6152FF7FF9A6" box="[151,238,1586,1610]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="357" unit="mm" value="2.2">2.2 mm</quantity>
|
||
distal of subnodus; base of IR1 four cells and 4.0 mm distally; oblique crossvein ‘O’ absent; cubito-anal area with five rows of cells between CuA and posterior wing margin; CuA reaching posterior wing margin well distal of nodus level; CuA weakly curved; postdiscoidal area with one row of cells up to nodus level and two rows more distally; area between
|
||
<collectingCountry id="3FD263E0AA62525416EC61FEFE65F959" box="[464,500,1694,1718]" name="Poland" pageId="3" pageNumber="357">RP</collectingCountry>
|
||
3/4 and MA with at least two secondary longitudinal veins in between; three rows of cells between
|
||
<collectingCountry id="3FD263E0AA625254160861A2FEC9F935" box="[308,344,1730,1754]" name="Poland" pageId="3" pageNumber="357">RP</collectingCountry>
|
||
3/4 and IR2 and one row between
|
||
<collectingCountry id="3FD263E0AA62525415EF61A2FD6BF935" box="[723,762,1730,1754]" name="Poland" pageId="3" pageNumber="357">RP</collectingCountry>
|
||
1 and IR1; three rows of cells between IR1 and
|
||
<collectingCountry id="3FD263E0AA625254123B61A2FABBF935" box="[1287,1322,1730,1754]" name="Poland" pageId="3" pageNumber="357">RP</collectingCountry>
|
||
2 opposite pterostigma, but only one basally.
|
||
</paragraph>
|
||
<paragraph id="477A2370AA62525417FB6069FB59F8AA" blockId="3.[151,1437,1153,2006]" pageId="3" pageNumber="357">
|
||
Preserved part of hind wing nearly identical to forewing, even the shape of the discoidal cell nearly the same. Only the position of the nodus is slightly different, as distance from arculus to nodus is
|
||
<quantity id="803D8E95AA6252541357604EFB55F8AA" box="[1131,1220,1838,1862]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="3" pageNumber="357" unit="mm" value="8.7">8.7 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FDF70FBAA62525217FB6031FD41FCFB" lastPageId="5" lastPageNumber="359" pageId="3" pageNumber="357" type="discussion">
|
||
<paragraph id="477A2370AA62525317FB6031FB87FCB2" blockId="3.[151,1437,1153,2006]" lastBlockId="4.[151,1437,151,861]" lastPageId="4" lastPageNumber="358" pageId="3" pageNumber="357">
|
||
<emphasis id="75B1FF62AA62525417FB6031FEDDF885" bold="true" box="[199,332,1873,1898]" pageId="3" pageNumber="357">Discussion.</emphasis>
|
||
The comparison of the preserved parts (hind wing basal two-thirds) of the venation of the
|
||
<typeStatus id="987E9DD2AA62525412716032FAECF885" box="[1357,1405,1874,1898]" pageId="3" pageNumber="357">type</typeStatus>
|
||
of
|
||
<taxonomicName id="80C558F3AA62525417AB6015FE0BF861" box="[151,410,1909,1934]" class="Insecta" family="Dysagrionidae" genus="Petrolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="357" phylum="Arthropoda" rank="species" species="hendersoni">
|
||
<emphasis id="75B1FF62AA62525417AB6015FE0BF861" box="[151,410,1909,1934]" italics="true" pageId="3" pageNumber="357">Petrolestes hendersoni</emphasis>
|
||
</taxonomicName>
|
||
with that of this new fossil shows that they are nearly identical in every point (petiole length, nodus, subnodus, discoidal cell, antenodal and cubito-anal areas, relative positions of bases of branches of
|
||
<collectingCountry id="3FD263E0AA62525417AB60DEFF28F83A" box="[151,185,1982,2005]" name="Poland" pageId="3" pageNumber="357">RP</collectingCountry>
|
||
, number of rows of cells between main veins in basal part of wings). Also the dimensions of the wing structures are nearly identical in the two specimens.
|
||
<collectingCountry id="3FD263E0AA655253154467F8FD06FF5F" box="[632,663,152,176]" name="American Samoa" pageId="4" pageNumber="358">As</collectingCountry>
|
||
both specimens come from the Green River Formation in Colorado and even from outcrops in the same county, we consider that they belong to the same species. The new specimen allows more precision in several points concerning this taxon, viz. it has: no oblique vein ‘O’, a long pterostigma covering numerous cells, a long zigzagged IR1, a short petiole, and the discoidal cells of the fore and hind wings are nearly identical. These characters confirm the position of
|
||
<emphasis id="75B1FF62AA655253146F6647FC5EFEAF" box="[851,975,295,320]" italics="true" pageId="4" pageNumber="358">Petrolestes</emphasis>
|
||
close to
|
||
<emphasis id="75B1FF62AA655253130B6648FB2AFED0" box="[1079,1211,296,319]" italics="true" pageId="4" pageNumber="358">Congqingia</emphasis>
|
||
Zhang,
|
||
<quantity id="803D8E95AA65525312276648FAE3FEAF" box="[1307,1394,295,320]" metricMagnitude="1" metricUnit="m" metricValue="5.05968" pageId="4" pageNumber="358" unit="in" value="1992.0">1992 in</quantity>
|
||
the Dysagrioninae, rather than close to the
|
||
<taxonomicName id="80C558F3AA6552531569662CFD73FE8A" box="[597,738,332,357]" class="Insecta" family="Sieblosiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="358" phylum="Arthropoda" rank="family">Sieblosiidae</taxonomicName>
|
||
. In particular
|
||
<emphasis id="75B1FF62AA6552531443662CFC6AFE8A" box="[895,1019,332,357]" italics="true" pageId="4" pageNumber="358">Petrolestes</emphasis>
|
||
has not the subnodal veinlet vertical or anteriorly oriented and its nodus is not traversed by ScP, unlike in the
|
||
<taxonomicName id="80C558F3AA65525314EB660FFBF5FE67" box="[983,1124,367,392]" class="Insecta" family="Sieblosiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="358" phylum="Arthropoda" rank="family">Sieblosiidae</taxonomicName>
|
||
; it also has no oblique vein ‘O’.
|
||
<emphasis id="75B1FF62AA65525317F066F4FED9FE42" box="[204,328,404,429]" italics="true" pageId="4" pageNumber="358">Petrolestes</emphasis>
|
||
has also a significant increase of spine density at the apical costal margin, a character proposed by
|
||
<bibRefCitation id="23545E81AA65525317AB66D7FEACFE3F" author="Bechly" box="[151,317,439,464]" pageId="4" pageNumber="358" refString="Bechly, G. (1996) Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Special Volume 2, 402 pp." type="book" year="1996">Bechly (1996)</bibRefCitation>
|
||
as a potential synapomorphy of the Zygoptera, absent in the
|
||
<taxonomicName id="80C558F3AA655253131E66D7FA09FE3F" authority="Fleck et al. 2004" authorityName="Fleck et al." authorityYear="2004" box="[1058,1432,439,464]" class="Insecta" family="Sieblosiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="358" phylum="Arthropoda" rank="family">
|
||
Sieblosiidae (
|
||
<bibRefCitation id="23545E81AA655253138266D7FA00FE3F" author="Fleck" box="[1214,1425,439,464]" pageId="4" pageNumber="358" refString="Fleck, G., Bechly, G., Martinez-Delclos, X., Jarzembowski, E. A. & Nel, A. (2004) A revision of the Mesozoic dragonfly family Tarsophlebiidae, with a discussion on the phylogenetic positions of the Tarsophlebiidae and Sieblosiidae (Odonatoptera: Panodonata). Geodiversitas, 26, 33 - 60." type="journal article" year="2004">
|
||
Fleck
|
||
<emphasis id="75B1FF62AA655253123666D9FAD9FE3F" box="[1290,1352,439,464]" italics="true" pageId="4" pageNumber="358">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="75B1FF62AA65525317AB66BCFE82FE1A" box="[151,275,476,501]" italics="true" pageId="4" pageNumber="358">Petrolestes</emphasis>
|
||
and
|
||
<emphasis id="75B1FF62AA655253166166BDFE73FE1B" box="[349,482,477,500]" italics="true" pageId="4" pageNumber="358">Congqingia</emphasis>
|
||
share with
|
||
<emphasis id="75B1FF62AA65525315BD66BDFD66FE1B" box="[641,759,477,500]" italics="true" pageId="4" pageNumber="358">Dysagrion</emphasis>
|
||
a rather unique shape of the discoidal cell, potential synapomorphy even if it is also present in the
|
||
<taxonomicName id="80C558F3AA655253159B669FFCA2FDF7" box="[679,819,511,536]" class="Insecta" family="Sieblosiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="358" phylum="Arthropoda" rank="family">Sieblosiidae</taxonomicName>
|
||
. The presence of a ‘basal costal margin between wing base and nodus distinctly convex curved’ (a character absent in the
|
||
<taxonomicName id="80C558F3AA65525314936544FBAAFDD2" box="[943,1083,548,573]" class="Insecta" family="Sieblosiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="358" phylum="Arthropoda" rank="family">Sieblosiidae</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="23545E81AA655253134B6544FABBFDD2" author="Nel" box="[1143,1322,548,573]" pageId="4" pageNumber="358" refString="Nel, A., Petrulevicius, J. F., Gentilini, G. & Martinez-Delclos, X. (2005) Phylogenetic analysis of the Cenozoic family Sieblosiidae (Insecta: Odonata), with description of new taxa from Russia, Italy and France. Geobios, 38, 219 - 233. http: // dx. doi. org / 10.1016 / j. geobios. 2003.10.007" type="journal article" year="2005">
|
||
Nel
|
||
<emphasis id="75B1FF62AA65525313956545FB75FDD2" box="[1193,1252,548,573]" italics="true" pageId="4" pageNumber="358">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
), plus the ‘antesubnodal space without any crossveins’ (a character present in many Zygoptera and in the
|
||
<taxonomicName id="80C558F3AA65525313816527FAD9FD8F" box="[1213,1352,583,608]" class="Insecta" family="Sieblosiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="358" phylum="Arthropoda" rank="family">Sieblosiidae</taxonomicName>
|
||
) would be the unique putative synapomorphies of the Dysagrioninae with the modern
|
||
<taxonomicName id="80C558F3AA6552531317650CFA91FD6A" box="[1067,1280,620,645]" class="Insecta" family="Thaumatoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="358" phylum="Arthropoda" rank="family">Thaumatoneuridae</taxonomicName>
|
||
, after
|
||
<bibRefCitation id="23545E81AA6552531271650CFF74FD47" author="Bechly" pageId="4" pageNumber="358" refString="Bechly, G. (2014) Phylogenetic systematics of Odonata. Available from: http: // members. tripod. de / GBechly / phylosys. htm (Accessed 7 Oct. 2015)" type="journal article" year="2014">Bechly (2014)</bibRefCitation>
|
||
. If
|
||
<emphasis id="75B1FF62AA655253163665F1FE10FD47" box="[266,385,657,680]" italics="true" pageId="4" pageNumber="358">Dysagrion</emphasis>
|
||
shares with
|
||
<taxonomicName id="80C558F3AA655253153365EFFD53FD47" box="[527,706,655,680]" class="Insecta" family="Megapodagrionidae" genus="Thaumatoneura" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="358" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75B1FF62AA655253153365EFFD53FD47" box="[527,706,655,680]" italics="true" pageId="4" pageNumber="358">Thaumatoneura</emphasis>
|
||
</taxonomicName>
|
||
similar relative positions of the nodus and bases of the branches of
|
||
<collectingCountry id="3FD263E0AA655253178B65D5FF48FD23" box="[183,217,693,716]" name="Poland" pageId="4" pageNumber="358">RP</collectingCountry>
|
||
, it is not the case for
|
||
<emphasis id="75B1FF62AA65525316ED65D4FDDCFD22" box="[465,589,692,717]" italics="true" pageId="4" pageNumber="358">Petrolestes</emphasis>
|
||
and
|
||
<emphasis id="75B1FF62AA65525315BA65D5FC9BFD23" box="[646,778,693,716]" italics="true" pageId="4" pageNumber="358">Congqingia</emphasis>
|
||
, which have the bases of
|
||
<collectingCountry id="3FD263E0AA655253130F65D4FBC6FD23" box="[1075,1111,692,716]" name="Poland" pageId="4" pageNumber="358">RP</collectingCountry>
|
||
3/4 and IR2 basally recessed midway between the nodus and the arculus. Also the discoidal cell of
|
||
<taxonomicName id="80C558F3AA65525314DD65B7FB05FD1F" box="[993,1172,727,752]" class="Insecta" family="Megapodagrionidae" genus="Thaumatoneura" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="358" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75B1FF62AA65525314DD65B7FB05FD1F" box="[993,1172,727,752]" italics="true" pageId="4" pageNumber="358">Thaumatoneura</emphasis>
|
||
</taxonomicName>
|
||
is elongate and much narrower than in all the taxa currently included in the Dysagrioninae. Also
|
||
<taxonomicName id="80C558F3AA65525314D1659CFB31FCFA" box="[1005,1184,764,789]" class="Insecta" family="Megapodagrionidae" genus="Thaumatoneura" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="358" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75B1FF62AA65525314D1659CFB31FCFA" box="[1005,1184,764,789]" italics="true" pageId="4" pageNumber="358">Thaumatoneura</emphasis>
|
||
</taxonomicName>
|
||
has long spines on its legs, unlike
|
||
<emphasis id="75B1FF62AA655253161C647FFE0AFCD7" box="[288,411,799,824]" italics="true" pageId="4" pageNumber="358">Petrolestes</emphasis>
|
||
, and its thoracic skewness is much more pronounced. Thus the attribution of this group of damselflies as sister group of the modern
|
||
<taxonomicName id="80C558F3AA65525315566424FCADFCB2" box="[618,828,836,861]" class="Insecta" family="Thaumatoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="358" phylum="Arthropoda" rank="family">Thaumatoneuridae</taxonomicName>
|
||
remains debatable.
|
||
</paragraph>
|
||
<caption id="13BA73F8AA65525317AB60EDFA0CF84C" box="[151,1437,1933,1955]" httpUri="https://zenodo.org/record/232687/files/figure.png" pageId="4" pageNumber="358" targetBox="[267,1319,913,1911]" targetPageId="4">
|
||
<paragraph id="477A2370AA65525317AB60EDFA0CF84C" blockId="4.[151,1437,1933,1955]" box="[151,1437,1933,1955]" pageId="4" pageNumber="358">
|
||
<emphasis id="75B1FF62AA65525317AB60EDFE83F84D" bold="true" box="[151,274,1933,1955]" pageId="4" pageNumber="358">FIGURE 5.</emphasis>
|
||
<taxonomicName id="80C558F3AA655253162460EDFD39F84C" authority="Cockerell, 1927" authorityName="Cockerell" authorityYear="1927" box="[280,680,1933,1955]" class="Insecta" family="Dysagrionidae" genus="Petrolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="358" phylum="Arthropoda" rank="species" species="hendersoni">
|
||
<emphasis id="75B1FF62AA655253162460EDFE6EF84C" box="[280,511,1933,1955]" italics="true" pageId="4" pageNumber="358">Petrolestes hendersoni</emphasis>
|
||
Cockerell, 1927
|
||
</taxonomicName>
|
||
, USNM without number, photograph of legs. Scale bar represents 4.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="477A2370AA64525217FB67F7FEDCFDF7" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="359">
|
||
Several characters suggest that
|
||
<emphasis id="75B1FF62AA645252151B67F7FD33FF5F" box="[551,674,151,176]" italics="true" pageId="5" pageNumber="359">Petrolestes</emphasis>
|
||
(and consequently
|
||
<emphasis id="75B1FF62AA645252144167F8FB93FF40" box="[893,1026,152,175]" italics="true" pageId="5" pageNumber="359">Congqingia</emphasis>
|
||
and maybe
|
||
<emphasis id="75B1FF62AA64525213B667F9FA91FF5F" box="[1162,1280,153,176]" italics="true" pageId="5" pageNumber="359">Dysagrion</emphasis>
|
||
) could not be a Zygoptera: the head appears not as transverse as in a Zygoptera (but this could be due to the compression of the fossil that can strongly deform the head shape; similar situations occur for fossil
|
||
<taxonomicName id="80C558F3AA645252130B6781FBECFF17" box="[1079,1149,225,248]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="359" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75B1FF62AA645252130B6781FBECFF17" box="[1079,1149,225,248]" italics="true" pageId="5" pageNumber="359">Lestes</emphasis>
|
||
</taxonomicName>
|
||
in compression on rocks from the Oligocene of
|
||
<collectingCountry id="3FD263E0AA645252169A6665FE64FEF3" box="[422,501,261,284]" name="France" pageId="5" pageNumber="359">France</collectingCountry>
|
||
, so that the head seems to be less transverse than it was certainly in the living animal); the thorax is not very skewed, similarly to what happen in Epiproctophora (but this could be due to the robustness of this animal); the legs have spines shorter and stronger than in Zygoptera; this character is rather present in Epiproctophora. Also the position of the arculus near Ax2 instead of being midway between Ax2 and Ax1 or closer to Ax1 is a plesiomorphic character that would exclude a position within the Epiproctophora for
|
||
<emphasis id="75B1FF62AA64525217AB66D7FE82FE3F" box="[151,275,439,464]" italics="true" pageId="5" pageNumber="359">Petrolestes</emphasis>
|
||
. The presence of an incomplete interpleural suture but rather long and formed at dorsal ½ is a plesiomorphic character present in Zygoptera (the interpleural suture is complete in
|
||
<taxonomicName id="80C558F3AA645252135866BCFA86FE1A" box="[1124,1303,476,501]" class="Insecta" family="Megapodagrionidae" genus="Thaumatoneura" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="359" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="75B1FF62AA645252135866BCFA86FE1A" box="[1124,1303,476,501]" italics="true" pageId="5" pageNumber="359">Thaumatoneura</emphasis>
|
||
</taxonomicName>
|
||
) but not in Epiproctophora.
|
||
</paragraph>
|
||
<paragraph id="477A2370AA64525217FB6545FAC8FD47" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="359">
|
||
It remains that the thoracic skewness and relatively short leg spines of
|
||
<emphasis id="75B1FF62AA64525214CF6544FBFEFDD2" box="[1011,1135,548,573]" italics="true" pageId="5" pageNumber="359">Petrolestes</emphasis>
|
||
suggest affinities with the Epiproctophora rather than with Zygoptera.
|
||
<emphasis id="75B1FF62AA64525215A86527FC81FD8F" box="[660,784,583,608]" italics="true" pageId="5" pageNumber="359">Petrolestes</emphasis>
|
||
, and consequently
|
||
<emphasis id="75B1FF62AA64525214CD6528FBE4FDB0" box="[1009,1141,584,607]" italics="true" pageId="5" pageNumber="359">Congqingia</emphasis>
|
||
and
|
||
<emphasis id="75B1FF62AA645252138E6529FAB9FD8F" box="[1202,1320,585,608]" italics="true" pageId="5" pageNumber="359">Dysagrion</emphasis>
|
||
, could be taxa in the stem lineage of the Epiproctophora rather than Zygoptera. It would be more accurate to have information on the genital appendages of these damselflies before an attribution close to the Epiproctophora.
|
||
</paragraph>
|
||
<paragraph id="477A2370AA64525217FB65D4FD41FCFB" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="359">
|
||
<emphasis id="75B1FF62AA64525217FB65D4FED2FD22" box="[199,323,692,717]" italics="true" pageId="5" pageNumber="359">Petrolestes</emphasis>
|
||
differs from
|
||
<emphasis id="75B1FF62AA64525216E965D5FDC8FD23" box="[469,601,693,716]" italics="true" pageId="5" pageNumber="359">Congqingia</emphasis>
|
||
in the longer pterostigma, covering six cells, instead of two, in the broader area between IR1 and
|
||
<collectingCountry id="3FD263E0AA64525216A565B8FE2DFD1F" box="[409,444,728,752]" name="Poland" pageId="5" pageNumber="359">RP</collectingCountry>
|
||
2 with longer intercalary veins between them, and in the broader cubito-anal area with five-six rows of cells instead of two (Zhang 1992).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |