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<mods:title id="81F0CD2C50BDD75E88C9A834815E7EC3">Revision of Phaenocora Ehrenberg, 1836 (Rhabditophora, Typhloplanidae, Phaenocorinae) with the description of two new species</mods:title>
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<mods:namePart id="6CCD37956BB1413AF3F2293B98140FBC">Houben, Albrecht M.</mods:namePart>
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<mods:namePart id="F00816DD7B9F216739534739C7AD91DD">Artois, Tom J.</mods:namePart>
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<paragraph id="47152B4EFFF9C50C17C40A71E72AFABF" blockId="29.[151,628,1367,1427]" box="[151,628,1367,1394]" pageId="29" pageNumber="330">
<heading id="1C5D9C22FFF9C50C17C40A71E72AFABF" bold="true" box="[151,628,1367,1394]" fontSize="11" level="1" pageId="29" pageNumber="330" reason="1">
<taxonomicName id="80AA50CDFFF9C50C17C40A71E72AFABF" ID-CoL="6V7QW" authority="Nasonov, 1919" authorityName="Nasonov" authorityYear="1919" box="[151,628,1367,1394]" class="Rhabditophora" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="29" pageNumber="330" phylum="Platyhelminthes" rank="species" species="achaeorum">
<emphasis id="75DEF75CFFF9C50C17C40A71E72AFABF" bold="true" box="[151,628,1367,1394]" pageId="29" pageNumber="330">
<emphasis id="75DEF75CFFF9C50C17C40A71E4E4FABC" bold="true" box="[151,442,1367,1393]" italics="true" pageId="29" pageNumber="330">Phaenocora achaeorum</emphasis>
<bibRefCitation id="233B56BFFFF9C50C16920A7EE72AFABF" author="Nasonov" box="[449,628,1368,1394]" pageId="29" pageNumber="330" refString="Nasonov, N. V. (1919) Contributions a la faune des Turbellaria de la Russie. Buletin de l'Academie de Sciences de Russie, Part I, 619 - 646. Part II + III, 1039 - 1053. Part IV, 1179 - 1197. [Tab. I - V] [In Russian]" type="book chapter" year="1919">Nasonov, 1919</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="47152B4EFFF9C50C17C40A5CE455FA5E" blockId="29.[151,628,1367,1427]" box="[151,267,1402,1427]" pageId="29" pageNumber="330">(Fig. 11C)</paragraph>
</subSubSection>
<subSubSection id="0FB078C5FFF9C50C17C40AE4E632F9F1" pageId="29" pageNumber="330" type="reference_group">
<paragraph id="47152B4EFFF9C50C17C40AE4E153FA3A" blockId="29.[151,1436,1474,1527]" pageId="29" pageNumber="330">
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<taxonomicName id="80AA50CDFFF9C50C17C40AE4E72FFA15" ID-CoL="6V7QW" authority="Nasonov 1919: 1043" authorityName="Nasonov" authorityPageNumber="1043" authorityYear="1919" box="[151,625,1474,1496]" class="Rhabditophora" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="29" pageNumber="330" phylum="Platyhelminthes" rank="species" species="achaeorum">
<emphasis id="75DEF75CFFF9C50C17C40AE4E4D3FA15" box="[151,397,1474,1496]" italics="true" pageId="29" pageNumber="330">Phaenocora achaeorum</emphasis>
<treatmentCitation id="C60B0D5FFFF9C50C16C40AE5E72FFA15" author="Nasonov" box="[407,625,1474,1496]" page="1043" pageId="29" pageNumber="330" year="1919">
<bibRefCitation id="233B56BFFFF9C50C16C40AE5E76FFA15" author="Nasonov" box="[407,561,1474,1496]" pageId="29" pageNumber="330" refString="Nasonov, N. V. (1919) Contributions a la faune des Turbellaria de la Russie. Buletin de l'Academie de Sciences de Russie, Part I, 619 - 646. Part II + III, 1039 - 1053. Part IV, 1179 - 1197. [Tab. I - V] [In Russian]" type="book chapter" year="1919">Nasonov 1919</bibRefCitation>
: 1043
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1044, plate 4:
<figureCitation id="DF9137CBFFF9C50C144B0AE4E62BFA15" box="[792,885,1474,1496]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="5.[151,250,1888,1910]" captionStartId-1="6.[151,250,1706,1728]" captionStartId-2="7.[151,250,1931,1953]" captionTargetBox-0="[216,1369,646,1866]" captionTargetId-0="figure@5.[743,1191,1176,1609]" captionTargetPageId-0="5" captionText-0="FIGURE 1. Photographs of live animals A, P. evelinae. B, P. foliacea from South Africa. C, P. foliacea from India. D, P. aglobulata sp. nov.. E H, P. gilberti sp. nov.: E F, pigmentation of the head. G, male copulatory organ. H, female genital canal." captionText-1="FIGURE 2. Phaenocora anomalocoela A, reproductive structures as seen on whole mount (G 211245). B, sagital reconstruction of the reproductive structures (caudal end at the left) (QM no. G 211246). C, partly evaginated male copulatory organ as seen on whole mount (QM no. G 211245). D, positioning of the reproductive structures on sagittal sections (caudal end at the left) (QM no. G 211246)." captionText-2="FIGURE 3. Phaenocora aglobulata sp. nov. A, habitus of a free swimming animal. B, reconstruction on sagittal sections of the male copulatory organ. (caudal end at the right) C, reconstruction on sagittal sections of female genital system. (caudal end at the right) D, schematic representation of the cirrus. (B D: SMNH Type- 8671)." httpUri-0="https://zenodo.org/record/224902/files/figure.png" pageId="29" pageNumber="330">Figs 13</figureCitation>
;
<treatmentCitation id="C60B0D5FFFF9C50C14D70AE4E019FA15" author="Stummer-Traunfels" box="[900,1351,1474,1496]" page="3396" pageId="29" pageNumber="330" year="1930">
<bibRefCitation id="233B56BFFFF9C50C14D70AE4E05AFA15" author="Stummer-Traunfels" box="[900,1284,1474,1496]" pageId="29" pageNumber="330" refString="Stummer-Traunfels, R. &amp; Meixner, J. (1930) Vierter Band: Vermes Abteilung Ic: Turbellaria 3.: Polycladida, Literatur V. In: Dr. H. G. Bronns Klassen und Ordnungen des Tier-Reichs, Leipzig, pp. 3371 - 3484." type="book chapter" year="1930">Stummer-Traunfels &amp; Meixner 1930</bibRefCitation>
: 3396
</treatmentCitation>
;
<treatmentCitation id="C60B0D5FFFF9C50C12070AE4E470FA3A" author="Gilbert" page="284" pageId="29" pageNumber="330" year="1935">
<bibRefCitation id="233B56BFFFF9C50C12070AE4E5A3FA3A" author="Gilbert" pageId="29" pageNumber="330" refString="Gilbert, C. M. (1935) A comparative study of three new American species of the genus Phaenocora with especial reference to their reproductive organs and their relationships with the other described forms of the genus. Acta zoologica, 16, 283 - 386. [Plate I - V]" type="journal article" year="1935">Gilbert 1935</bibRefCitation>
: 284
</treatmentCitation>
285, 293, 343, 352, 356,
<figureCitation id="DF9137CBFFF9C50C15650AC7E72FFA3A" box="[566,625,1505,1527]" captionStart="FIGURE 3" captionStartId="7.[151,250,1931,1953]" captionText="FIGURE 3. Phaenocora aglobulata sp. nov. A, habitus of a free swimming animal. B, reconstruction on sagittal sections of the male copulatory organ. (caudal end at the right) C, reconstruction on sagittal sections of female genital system. (caudal end at the right) D, schematic representation of the cirrus. (B D: SMNH Type- 8671)." pageId="29" pageNumber="330">Fig. 3</figureCitation>
E, tables 1, 2;
<treatmentCitation id="C60B0D5FFFF9C50C14510AC4E6EEFA3A" author="Marcus" box="[770,944,1505,1527]" page="72" pageId="29" pageNumber="330" year="1946">
<bibRefCitation id="233B56BFFFF9C50C14510AC4E6D5FA3A" author="Marcus" box="[770,907,1505,1527]" pageId="29" pageNumber="330" refString="Marcus, E. (1946) Sobre Turbellaria Brasilieros. Boletins da Faculdade de Filosofia, Ciencias e Letras, Zoologia, 11, 5 - 254. [Tab. I - XXXI]" type="journal article" year="1946">Marcus 1946</bibRefCitation>
: 72
</treatmentCitation>
, 81, 166.
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</paragraph>
<paragraph id="47152B4EFFF9C50C17C40903E632F9F1" blockId="29.[151,1437,1572,1885]" box="[151,876,1572,1597]" pageId="29" pageNumber="330">
<emphasis id="75DEF75CFFF9C50C17C40903E427F9F0" box="[151,377,1572,1597]" italics="true" pageId="29" pageNumber="330">Known distribution:</emphasis>
Tikhon (
<collectingCountry id="3FBD6BDEFFF9C50C168D0902E772F9F0" box="[478,556,1572,1597]" name="Russia" pageId="29" pageNumber="330">Russia</collectingCountry>
) in a creek (
<bibRefCitation id="233B56BFFFF9C50C15E50902E63EF9F1" author="Nasonov" box="[694,864,1572,1596]" pageId="29" pageNumber="330" refString="Nasonov, N. V. (1919) Contributions a la faune des Turbellaria de la Russie. Buletin de l'Academie de Sciences de Russie, Part I, 619 - 646. Part II + III, 1039 - 1053. Part IV, 1179 - 1197. [Tab. I - V] [In Russian]" type="book chapter" year="1919">Nasonov 1919</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0FB078C5FFF9C50C17940961E4AEF992" box="[199,496,1607,1632]" pageId="29" pageNumber="330" type="materials_examined">
<paragraph id="47152B4EFFF9C50C17940961E4AEF992" blockId="29.[151,1437,1572,1885]" box="[199,496,1607,1632]" pageId="29" pageNumber="330">
<emphasis id="75DEF75CFFF9C50C17940961E4FAF9AD" box="[199,420,1607,1632]" italics="true" pageId="29" pageNumber="330">Material examined:</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="0FB078C5FFF9C50C1794094BE591F8D8" pageId="29" pageNumber="330" type="diagnosis">
<paragraph id="47152B4EFFF9C50C1794094BE591F8D8" blockId="29.[151,1437,1572,1885]" pageId="29" pageNumber="330">
<emphasis id="75DEF75CFFF9C50C1794094BE41FF949" box="[199,321,1645,1668]" italics="true" pageId="29" pageNumber="330">Diagnosis:</emphasis>
Animals about
<quantity id="805286ABFFF9C50C16A4094AE767F949" box="[503,569,1644,1668]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="330" unit="mm" value="1.0">1 mm</quantity>
long, with black eyes that appear white under reflective light. Zoochlorellae not mentioned in literature.
<collectingCountry id="3FBD6BDEFFF9C50C16B109A9E742F965" box="[482,540,1679,1704]" name="Malta" pageId="29" pageNumber="330">Male</collectingCountry>
copulatory organ of the duplex-type IA. Prostate vesicle empties through a constriction at the distal end of the copulatory organ. Female genital structures only partly known. Female genital canal, genito-bursal duct and intestinal bursa present, hence either of the
<taxonomicName id="80AA50CDFFF9C50C14B209FAE129F922" box="[993,1143,1756,1775]" class="Rhabditophora" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="29" pageNumber="349" phylum="Platyhelminthes" rank="species" species="unipunctata">
<emphasis id="75DEF75CFFF9C50C14B209FAE129F922" box="[993,1143,1756,1775]" italics="true" pageId="29" pageNumber="330">UNIPUNCTATA</emphasis>
</taxonomicName>
-
<typeStatus id="981195ECFFF9C50C132D09FEE1EEF93D" box="[1150,1200,1752,1776]" pageId="29" pageNumber="330">type</typeStatus>
or the
<taxonomicName id="80AA50CDFFF9C50C125509FAE0CDF922" box="[1286,1427,1756,1775]" class="Rhabditophora" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="29" pageNumber="349" phylum="Platyhelminthes" rank="species" species="aglobulata" status="sp. nov.">
<emphasis id="75DEF75CFFF9C50C125509FAE0CDF922" box="[1286,1427,1756,1775]" italics="true" pageId="29" pageNumber="330">AGLOBULATA</emphasis>
</taxonomicName>
-
<typeStatus id="981195ECFFF9C50C17C409DBE595F8D8" box="[151,203,1789,1813]" pageId="29" pageNumber="330">type</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="0FB078C5FFF9C50C17940839E6B3F891" pageId="29" pageNumber="330" type="discussion">
<paragraph id="47152B4EFFF9C50C17940839E6B3F891" blockId="29.[151,1437,1572,1885]" pageId="29" pageNumber="330">
<emphasis id="75DEF75CFFF9C50C17940839E46CF8F5" box="[199,306,1823,1848]" italics="true" pageId="29" pageNumber="330">Remarks:</emphasis>
<bibRefCitation id="233B56BFFFF9C50C166A0839E734F8F5" author="Gilbert" box="[313,618,1823,1848]" pageId="29" pageNumber="330" refString="Gilbert, C. M. (1935) A comparative study of three new American species of the genus Phaenocora with especial reference to their reproductive organs and their relationships with the other described forms of the genus. Acta zoologica, 16, 283 - 386. [Plate I - V]" type="journal article" year="1935">Gilbert (1935; TextFig. 3E)</bibRefCitation>
reproduced the drawing of the male system of
<taxonomicName id="80AA50CDFFF9C50C13250807E051F8F5" box="[1142,1295,1823,1848]" class="Rhabditophora" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="29" pageNumber="330" phylum="Platyhelminthes" rank="species" species="achaeorum">
<emphasis id="75DEF75CFFF9C50C13250807E051F8F5" box="[1142,1295,1823,1848]" italics="true" pageId="29" pageNumber="330">P. achaeorum</emphasis>
</taxonomicName>
by
<bibRefCitation id="233B56BFFFF9C50C126B0806E77AF891" author="Nasonov" pageId="29" pageNumber="330" refString="Nasonov, N. V. (1919) Contributions a la faune des Turbellaria de la Russie. Buletin de l'Academie de Sciences de Russie, Part I, 619 - 646. Part II + III, 1039 - 1053. Part IV, 1179 - 1197. [Tab. I - V] [In Russian]" type="book chapter" year="1919">
Nasonov (1919; Plate 4
<figureCitation id="DF9137CBFFF9C50C16680862E421F891" box="[315,383,1860,1885]" captionStart="FIGURE 2" captionStartId="6.[151,250,1706,1728]" captionText="FIGURE 2. Phaenocora anomalocoela A, reproductive structures as seen on whole mount (G 211245). B, sagital reconstruction of the reproductive structures (caudal end at the left) (QM no. G 211246). C, partly evaginated male copulatory organ as seen on whole mount (QM no. G 211245). D, positioning of the reproductive structures on sagittal sections (caudal end at the left) (QM no. G 211246)." pageId="29" pageNumber="330">Fig. 2</figureCitation>
; our Fig. 11C)
</bibRefCitation>
, but erroneously labelled it
<taxonomicName id="80AA50CDFFF9C50C140F0863E6BBF890" box="[860,997,1860,1885]" class="Rhabditophora" family="Typhloplanidae" genus="Phaenocora" kingdom="Animalia" order="Rhabdocoela" pageId="29" pageNumber="330" phylum="Platyhelminthes" rank="species" species="vjatkensis">
<emphasis id="75DEF75CFFF9C50C140F0863E6BBF890" box="[860,997,1860,1885]" italics="true" pageId="29" pageNumber="330">P. vjatkensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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