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Greenbaum, Dehling, Kusamba &amp; Portik
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Spiny Reed Frog 
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</taxonomicName>
(nec 
<taxonomicName id="C28DD7CEFFC5256CFEF3F89DFD7BF8D2" ID-CoL="72VR3" authority="Ahl 1930" authorityName="Ahl" authorityYear="1930" box="[368,659,1817,1841]" class="Amphibia" family="Hyperoliidae" genus="Megalixalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="17" pageNumber="218" phylum="Chordata" rank="species" species="laevis">
<emphasis id="37F9705FFFC5256CFEF3F89DFDC7F8D2" box="[368,559,1817,1841]" italics="true" pageId="17" pageNumber="218">Megalixalus laevis</emphasis>
<bibRefCitation id="611CD1BCFFC5256CFDB6F89DFD7BF8D2" author="Ahl, E." box="[565,659,1817,1841]" pageId="17" pageNumber="218" pagination="89 - 102" refId="ref17771" refString="Ahl, E. (1930) Ueber die afrikanischen Arten der Baumfroschgattung Megalixalus. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1930, 89 - 102." type="journal article" year="1930">Ahl 1930</bibRefCitation>
</taxonomicName>
)— 
<treatmentCitation id="842C8A5CFFC5256CFD37F89DFC80F8D2" author="Laurent, R." box="[692,872,1817,1841]" page="24" pageId="17" pageNumber="218" year="1950">
<bibRefCitation id="611CD1BCFFC5256CFD37F89DFC80F8D2" author="Laurent, R." box="[692,872,1817,1841]" pageId="17" pageNumber="218" pagination="5 - 120" refId="ref20371" refString="Laurent, R. (1950) Genres Afrixalus et Hyperolius (Amphibia Salientia). Exploration du Parc National Albert, Mission G. F. De Witte (1933 - 1935), 64, 5 - 120, 7 pls." type="journal article" year="1950">Laurent 1950: 24</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="842C8A5CFFC5256CFCF7F89EFBCBF8D2" author="Laurent, R. F." box="[884,1059,1817,1841]" page="59" pageId="17" pageNumber="218" year="1972">
<bibRefCitation id="611CD1BCFFC5256CFCF7F89EFBCBF8D2" author="Laurent, R. F." box="[884,1059,1817,1841]" pageId="17" pageNumber="218" pagination="1 - 125" refId="ref20584" refString="Laurent, R. F. (1972) Amphibiens. Exploration du Parc National des Virunga, Deuxieme Serie, 22, 1 - 125, 11 pls." type="journal article" year="1972">Laurent 1972: 59</bibRefCitation>
</treatmentCitation>
(partim); 
<treatmentCitation id="842C8A5CFFC5256CFB08F89EFAD3F8D2" author="Laurent, R. F." box="[1163,1339,1817,1841]" page="33" pageId="17" pageNumber="218" year="1982">
<bibRefCitation id="611CD1BCFFC5256CFB08F89EFAD3F8D2" author="Laurent, R. F." box="[1163,1339,1817,1841]" pageId="17" pageNumber="218" pagination="1 - 93" refId="ref20614" refString="Laurent, R. F. (1982) Le genre Afrixalus Laurent (Hyperoliidae) en Afrique Centrale. Annales du Musee Royal de l'Afrique Centrale, Serie in Octavo, Science Zoologique, 235, 1 - 93." type="journal article" year="1982">Laurent 1982: 33</bibRefCitation>
</treatmentCitation>
(partim); 
<treatmentCitation id="842C8A5CFFC5256CFF44F8BCFE9CF8B3" author="Schiotz, A." box="[199,372,1848,1872]" page="56" pageId="17" pageNumber="218" year="1999">
<bibRefCitation id="611CD1BCFFC5256CFF44F8BCFE9CF8B3" author="Schiotz, A." box="[199,372,1848,1872]" pageId="17" pageNumber="218" refId="ref22929" refString="Schiotz, A. (1999) Treefrogs of Africa. Edition Chimaira, Frankfurt am Main, 350 pp." type="book" year="1999">Schiøtz 1999: 56</bibRefCitation>
</treatmentCitation>
(partim); 
<treatmentCitation id="842C8A5CFFC5256CFE5EF8BCFCF1F8B3" author="Channing, A. &amp; Howell, K. M." box="[477,793,1848,1872]" page="137" pageId="17" pageNumber="218" year="2006">
<bibRefCitation id="611CD1BCFFC5256CFE5EF8BCFCF1F8B3" author="Channing, A. &amp; Howell, K. M." box="[477,793,1848,1872]" pageId="17" pageNumber="218" refId="ref18384" refString="Channing, A. &amp; Howell, K. M. (2006) Amphibians of East Africa. Cornell University Press, Ithaca, New York, New York, xi + 418 pp." type="book" year="2006">Channing &amp; Howell 2006: 137</bibRefCitation>
</treatmentCitation>
(partim); 
<treatmentCitation id="842C8A5CFFC5256CFC01F8BCFB9BF8B3" author="Spawls, S. &amp; Howell, K. M. &amp; Drewes, R. C." box="[898,1139,1848,1872]" page="183" pageId="17" pageNumber="218" year="2006">
<bibRefCitation id="611CD1BCFFC5256CFC01F8BCFB9BF8B3" author="Spawls, S. &amp; Howell, K. M. &amp; Drewes, R. C." box="[898,1139,1848,1872]" pageId="17" pageNumber="218" refId="ref22979" refString="Spawls, S., Howell, K. M. &amp; Drewes, R. C. (2006) Pocket Guide to the Reptiles and Amphibians of East Africa. A &amp; C Black Publishers, London, 240 pp." type="book" year="2006">
Spawls 
<emphasis id="37F9705FFFC5256CFC51F8BCFC17F8B3" box="[978,1023,1848,1872]" italics="true" pageId="17" pageNumber="218">et al</emphasis>
. 2006: 183
</bibRefCitation>
</treatmentCitation>
(partim); 
<treatmentCitation id="842C8A5CFFC5256CFB58F8BCFEC5F88D" author="Channing, A. &amp; Rodel, M. - O." page="158" pageId="17" pageNumber="218" year="2019">
<bibRefCitation id="611CD1BCFFC5256CFB58F8BCFEC5F88D" author="Channing, A. &amp; Rodel, M. - O." pageId="17" pageNumber="218" refId="ref18420" refString="Channing, A. &amp; Rodel, M. - O. (2019) Field Guide to the Frogs &amp; Other Amphibians of Africa. Struik Nature, Cape Town, 408 pp." type="book" year="2019">Channing &amp; Rödel 2019: 158</bibRefCitation>
</treatmentCitation>
(partim).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="4D97FFC6FFC5256EFF14F812FA97FBE8" lastPageId="19" lastPageNumber="220" pageId="17" pageNumber="218" type="materials_examined">
<paragraph id="0532AC4DFFC5256CFF14F812FE64F81B" blockId="17.[151,1436,1942,2040]" pageId="17" pageNumber="218">
<materialsCitation id="B5E5A610FFC5256CFF14F812FE64F81B" ID-GBIF-Occurrence="3887776325" collectingDate="2010-12-21" collectionCode="UTEP, ELI" collectorName="Chifundera Kusamba &amp; Felix I. Alonda" country="Democratic Republic of the Congo" elevation="1482" latitude="-3.15552" location="Kalundu" longLatPrecision="1" longitude="28.42108" municipality="South Kivu Province" pageId="17" pageNumber="218" specimenCode="UTEP 20805, ELI 605" specimenCount="1" typeStatus="holotype">
<emphasis id="37F9705FFFC5256CFF14F812FEE3F853" bold="true" box="[151,267,1942,1968]" pageId="17" pageNumber="218">
<typeStatus id="DA3612EFFFC5256CFF14F812FEEEF853" box="[151,262,1942,1968]" pageId="17" pageNumber="218" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="552B0436FFC5256CFE90F812FE4EF852" box="[275,422,1942,1969]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="17" pageNumber="218" type="Museum">UTEP 20805</specimenCode>
(field no. 
<specimenCode id="552B0436FFC5256CFD9EF812FD96F852" box="[541,638,1942,1969]" collectionCode="ELI" pageId="17" pageNumber="218">ELI 605</specimenCode>
), adult male, from the vicinity of 
<location id="0052FA96FFC5256CFC7FF812FBB4F853" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC5256CFC7FF812FBB4F853" box="[1020,1116,1942,1968]" country="Democratic Republic of the Congo" latitude="-3.15552" longLatPrecision="1" longitude="28.42108" municipality="South Kivu Province" name="Kalundu" pageId="17" pageNumber="218">Kalundu</location>
(
<geoCoordinate id="60B9CA8AFFC5256CFBEFF812FB13F853" box="[1132,1275,1942,1969]" degrees="03.15552" direction="south" orientation="latitude" pageId="17" pageNumber="218" precision="1" value="-3.15552">03.15552° S</geoCoordinate>
, 
<geoCoordinate id="60B9CA8AFFC5256CFA8BF812FA7FF853" box="[1288,1431,1942,1968]" degrees="28.42108" direction="east" orientation="longitude" pageId="17" pageNumber="218" precision="1" value="28.42108">28.42108° E</geoCoordinate>
, 
<quantity id="C27501A8FFC5256CFF14F83EFF07F837" box="[151,239,1978,2004]" metricMagnitude="3" metricUnit="m" metricValue="1.4819999999999998" pageId="17" pageNumber="218" unit="m" value="1482.0">
<elevation id="8EA04B7EFFC5256CFF14F83EFF07F837" box="[151,239,1978,2004]" metricMagnitude="3" metricUnit="m" metricValue="1.4819999999999998" pageId="17" pageNumber="218" unit="m" value="1482.0">1482 m</elevation>
</quantity>
), 
<collectingMunicipality id="E5563637FFC5256CFE80F83EFE18F837" box="[259,496,1978,2004]" pageId="17" pageNumber="218">South Kivu Province</collectingMunicipality>
, 
<collectingCountry id="7D9AECDDFFC5256CFE78F83EFDDFF837" box="[507,567,1978,2004]" name="Democratic Republic of the Congo" pageId="17" pageNumber="218">DRC</collectingCountry>
, collected on 
<date id="71338A8DFFC5256CFD52F83EFC4EF837" box="[721,934,1978,2004]" pageId="17" pageNumber="218" value="2010-12-21">
<collectingDate id="61777365FFC5256CFD52F83EFC4EF837" box="[721,934,1978,2004]" pageId="17" pageNumber="218" value="2010-12-21">21 December 2010</collectingDate>
</date>
by 
<collectorName id="A878C99BFFC5256CFC4CF83EFB56F837" box="[975,1214,1978,2004]" pageId="17" pageNumber="218">Chifundera Kusamba</collectorName>
and 
<collectorName id="A878C99BFFC5256CFB77F83EFA74F837" box="[1268,1436,1978,2004]" pageId="17" pageNumber="218">Félix I. Alonda</collectorName>
(
<figureCitation id="9DB6B0C8FFC5256CFF1CF85AFEF7F81B" box="[159,287,2014,2040]" captionStart="FIGURE 8" captionStartId="15.[151,250,1585,1609]" captionTargetBox="[151,1436,705,1562]" captionTargetId="figure-276@15.[151,1436,705,1562]" captionTargetPageId="15" captionText="FIGURE 8. Photos of the preserved holotype of Afrixalus phantasma sp. nov. (ZFMK 103454) in dorsal (A) and ventral (B) views (to scale), and plantar view of the left foot (C). Photos of the preserved holotype of A. lacustris sp. nov. (UTEP 20805) in dorsal (D) and ventral (E) views (to scale), and plantar view of the left foot (F)." figureDoi="http://doi.org/10.5281/zenodo.6986899" httpUri="https://zenodo.org/record/6986899/files/figure.png" pageId="17" pageNumber="218">Figs. 8D–F</figureCitation>
, 
<figureCitation id="9DB6B0C8FFC5256CFEA9F85AFE68F81B" box="[298,384,2014,2040]" captionStart="FIGURE 12" captionStartId="20.[151,250,1727,1751]" captionTargetBox="[151,1436,181,1704]" captionTargetId="figure-23@20.[151,1436,181,1704]" captionTargetPageId="20" captionText="FIGURE 12. Photos of Afrixalus lacustris sp. nov. in life. Adult male holotype (UTEP 20805), from the vicinity of Kalundu (03.15552° S, 28.42108° E, 1482 m), South Kivu Province, DRC in dorsal (A) and (B) ventral views, adult female (UTEP 20809) from Baraka, near shore of Lake Tanganyika, South Kivu Province, DRC (C), adult female (EPLU 395) from Epulu, Ituri Province, DRC (D), adult male (UTEP 22417) from Toyokana, Ituri Province, DRC (E), individual of unknown sex (DFH 1102) from Ngogo Research Center, Kibale Forest National Park, Western Region, Uganda (F). All photos taken the morning after capture." figureDoi="http://doi.org/10.5281/zenodo.6986240" httpUri="https://zenodo.org/record/6986240/files/figure.png" pageId="17" pageNumber="218">12A–B</figureCitation>
).
</materialsCitation>
</paragraph>
<caption id="51F2FCC5FFC6256FFF14F807FEB9F800" ID-DOI="http://doi.org/10.5281/zenodo.6986232" ID-Zenodo-Dep="6986232" httpUri="https://zenodo.org/record/6986232/files/figure.png" pageId="18" pageNumber="219" startId="18.[151,250,1923,1947]" targetBox="[179,1408,181,1898]" targetPageId="18">
<paragraph id="0532AC4DFFC6256FFF14F807FEB9F800" blockId="18.[151,1437,1923,2019]" pageId="18" pageNumber="219">
<emphasis id="37F9705FFFC6256FFF14F807FEC8F878" bold="true" box="[151,288,1923,1947]" pageId="18" pageNumber="219">FIGURE 11.</emphasis>
Photos of the habitat of (A) 
<emphasis id="37F9705FFFC6256FFDD2F807FC63F878" bold="true" box="[593,907,1923,1947]" pageId="18" pageNumber="219">
<taxonomicName id="C28DD7CEFFC6256FFDD2F807FCC7F878" authority="Greenbaum &amp; Portik &amp; Allen &amp; Vaughan &amp; Badjedjea &amp; Barej &amp; Behangana &amp; Conkey &amp; Dumbo &amp; Gonwouo &amp; Hirschfeld &amp; Hughes &amp; Igunzi &amp; Kusamba &amp; Lukwago &amp; Masudi &amp; Penner &amp; Reyes &amp; Rödel &amp; Roelke &amp; Romero &amp; Dehling, 2022" authorityName="Greenbaum &amp; Portik &amp; Allen &amp; Vaughan &amp; Badjedjea &amp; Barej &amp; Behangana &amp; Conkey &amp; Dumbo &amp; Gonwouo &amp; Hirschfeld &amp; Hughes &amp; Igunzi &amp; Kusamba &amp; Lukwago &amp; Masudi &amp; Penner &amp; Reyes &amp; Rödel &amp; Roelke &amp; Romero &amp; Dehling" authorityYear="2022" box="[593,815,1923,1947]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="18" pageNumber="219" phylum="Chordata" rank="species" species="phantasma" status="sp. nov.">
<emphasis id="37F9705FFFC6256FFDD2F807FCC7F878" bold="true" box="[593,815,1923,1947]" italics="true" pageId="18" pageNumber="219">Afrixalus phantasma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2CCACD24FFC6256FFCBBF807FC63F878" box="[824,907,1923,1947]" pageId="18" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Kamiranzovu Swamp, Nyungwe National Park, Rwanda, showing a forest edge in a montane marsh, and (B) 
<emphasis id="37F9705FFFC6256FFC81F823FC27F85C" bold="true" box="[770,975,1959,1983]" pageId="18" pageNumber="219">
<taxonomicName id="C28DD7CEFFC6256FFC81F823FC90F85C" authority="Greenbaum &amp; Portik &amp; Allen &amp; Vaughan &amp; Badjedjea &amp; Barej &amp; Behangana &amp; Conkey &amp; Dumbo &amp; Gonwouo &amp; Hirschfeld &amp; Hughes &amp; Igunzi &amp; Kusamba &amp; Lukwago &amp; Masudi &amp; Penner &amp; Reyes &amp; Rödel &amp; Roelke &amp; Romero &amp; Dehling, 2022" authorityName="Greenbaum &amp; Portik &amp; Allen &amp; Vaughan &amp; Badjedjea &amp; Barej &amp; Behangana &amp; Conkey &amp; Dumbo &amp; Gonwouo &amp; Hirschfeld &amp; Hughes &amp; Igunzi &amp; Kusamba &amp; Lukwago &amp; Masudi &amp; Penner &amp; Reyes &amp; Rödel &amp; Roelke &amp; Romero &amp; Dehling" authorityYear="2022" box="[770,888,1959,1983]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="18" pageNumber="219" phylum="Chordata" rank="species" species="lacustris" status="sp. nov.">
<emphasis id="37F9705FFFC6256FFC81F823FCFBF85C" bold="true" box="[770,787,1959,1983]" italics="true" pageId="18" pageNumber="219">A</emphasis>
. 
<emphasis id="37F9705FFFC6256FFC9CF823FC90F85C" bold="true" box="[799,888,1959,1983]" italics="true" pageId="18" pageNumber="219">lacustris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2CCACD24FFC6256FFCFDF823FC27F85C" box="[894,975,1959,1983]" pageId="18" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from the vicinity of Kalundu, DRC, showing transitional forest.
</paragraph>
</caption>
<paragraph id="0532AC4DFFC7256EFF44FF13FB45FF52" blockId="19.[151,1437,151,2037]" box="[199,1197,151,177]" pageId="19" pageNumber="220">
<emphasis id="37F9705FFFC7256EFF44FF13FE98FF52" bold="true" box="[199,368,151,177]" pageId="19" pageNumber="220">Paratopotype.</emphasis>
<materialsCitation id="B5E5A610FFC7256EFEF4FF13FD0EFF52" ID-GBIF-Occurrence="3887776323" box="[375,742,151,177]" collectionCode="UTEP" pageId="19" pageNumber="220" specimenCode="UTEP 20806, ELI 606" specimenCount="1">
<specimenCode id="552B0436FFC7256EFEF4FF13FDE1FF52" box="[375,521,151,177]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="19" pageNumber="220" type="Museum">UTEP 20806</specimenCode>
(field no. 
<specimenCode id="552B0436FFC7256EFDFDFF13FD36FF52" box="[638,734,151,177]" collectionCode="ELI" pageId="19" pageNumber="220">ELI 606</specimenCode>
)
</materialsCitation>
, adult male, collected with the 
<typeStatus id="DA3612EFFFC7256EFBC0FF13FB41FF52" box="[1091,1193,151,177]" pageId="19" pageNumber="220" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="0532AC4DFFC7256EFF44FF3EFA97FBE8" blockId="19.[151,1437,151,2037]" pageId="19" pageNumber="220">
<emphasis id="37F9705FFFC7256EFF44FF3EFE54FF37" bold="true" box="[199,444,186,212]" pageId="19" pageNumber="220">Referred specimens.</emphasis>
Specimens with an asterisk were not included in morphological analyses due to poor quality of preservation, lack of a voucher, or unavailability of specimen(s): 
<materialsCitation id="B5E5A610FFC7256EFC75FF5BFB40FEFE" ID-GBIF-Occurrence="3887776316" collectionCode="UTEP, ELI" country="Democratic Republic of the Congo" elevation="1482" latitude="-3.15552" location="Kalundu" longLatPrecision="1" longitude="28.42108" municipality="South Kivu Province" pageId="19" pageNumber="220" specimenCode="UTEP 20807-08, ELI 644, 649" specimenCount="1">
<specimenCode id="552B0436FFC7256EFC75FF5BFB5BFF1A" box="[1014,1203,223,249]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="19" pageNumber="220" type="Museum">UTEP 20807–08</specimenCode>
(field nos. 
<specimenCode id="552B0436FFC7256EFAB4FF5BFA70FF1A" box="[1335,1432,223,249]" collectionCode="ELI" pageId="19" pageNumber="220">ELI 644</specimenCode>
, 
<specimenCode id="552B0436FFC7256EFF14FE87FF2DFEFE" box="[151,197,259,285]" collectionCode="ELI" pageId="19" pageNumber="220">649</specimenCode>
), vicinity of 
<location id="0052FA96FFC7256EFED4FE87FE5FFEFE" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFED4FE87FE5FFEFE" box="[343,439,259,285]" country="Democratic Republic of the Congo" latitude="-3.15552" longLatPrecision="1" longitude="28.42108" municipality="South Kivu Province" name="Kalundu" pageId="19" pageNumber="220">Kalundu</location>
(
<geoCoordinate id="60B9CA8AFFC7256EFE44FE86FDBEFEFE" box="[455,598,258,285]" degrees="03.15552" direction="south" orientation="latitude" pageId="19" pageNumber="220" precision="1" value="-3.15552">03.15552° S</geoCoordinate>
, 
<geoCoordinate id="60B9CA8AFFC7256EFDE7FE87FD1BFEFE" box="[612,755,259,285]" degrees="28.42108" direction="east" orientation="longitude" pageId="19" pageNumber="220" precision="1" value="28.42108">28.42108° E</geoCoordinate>
, 
<quantity id="C27501A8FFC7256EFC83FE87FCB1FEFE" box="[768,857,259,285]" metricMagnitude="3" metricUnit="m" metricValue="1.4819999999999998" pageId="19" pageNumber="220" unit="m" value="1482.0">
<elevation id="8EA04B7EFFC7256EFC83FE87FCB1FEFE" box="[768,857,259,285]" metricMagnitude="3" metricUnit="m" metricValue="1.4819999999999998" pageId="19" pageNumber="220" unit="m" value="1482.0">1482 m</elevation>
</quantity>
), 
<collectingMunicipality id="E5563637FFC7256EFCEDFE87FBB6FEFE" box="[878,1118,259,285]" pageId="19" pageNumber="220">South Kivu Province</collectingMunicipality>
, 
<collectingCountry id="7D9AECDDFFC7256EFBE8FE87FB40FEFE" box="[1131,1192,259,285]" name="Democratic Republic of the Congo" pageId="19" pageNumber="220">DRC</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="B5E5A610FFC7256EFB37FE87FEAEFE87" ID-GBIF-Occurrence="3887776302" collectionCode="UTEP, ELI" country="Democratic Republic of the Congo" county="South Kivu Province" elevation="777" latitude="-4.10832" location="Baraka" longLatPrecision="1" longitude="29.09705" municipality="Lake Tanganyika" pageId="19" pageNumber="220" specimenCode="UTEP 20809, 22422, ELI 669-70" specimenCount="1">
<specimenCode id="552B0436FFC7256EFB37FE87FAA2FEFE" box="[1204,1354,259,285]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="19" pageNumber="220" type="Museum">UTEP 20809</specimenCode>
, 
<specimenCode id="552B0436FFC7256EFAD5FE87FA74FEFE" box="[1366,1436,259,285]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="19" pageNumber="220" type="Museum">22422</specimenCode>
(field nos. 
<specimenCode id="552B0436FFC7256EFE91FEA3FE76FEA2" box="[274,414,295,321]" collectionCode="ELI" pageId="19" pageNumber="220">ELI 669–70</specimenCode>
), 
<location id="0052FA96FFC7256EFE30FEA3FDEDFEA2" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFE30FEA3FDEDFEA2" box="[435,517,295,321]" country="Democratic Republic of the Congo" county="South Kivu Province" latitude="-4.10832" longLatPrecision="1" longitude="29.09705" municipality="Lake Tanganyika" name="Baraka" pageId="19" pageNumber="220">Baraka</location>
, near shore of 
<collectingMunicipality id="E5563637FFC7256EFD2DFEA3FC98FEA2" box="[686,880,295,321]" pageId="19" pageNumber="220">Lake Tanganyika</collectingMunicipality>
(
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, 
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, 
<quantity id="C27501A8FFC7256EFB39FEA3FAEDFEA2" box="[1210,1285,295,321]" metricMagnitude="2" metricUnit="m" metricValue="7.77" pageId="19" pageNumber="220" unit="m" value="777.0">
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), 
<collectingCounty id="EC53D4C1FFC7256EFA98FEA3FF15FE87" pageId="19" pageNumber="220">South Kivu Province</collectingCounty>
, 
<collectingCountry id="7D9AECDDFFC7256EFE8AFECEFEAEFE87" box="[265,326,330,356]" name="Democratic Republic of the Congo" pageId="19" pageNumber="220">DRC</collectingCountry>
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; 
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<specimenCode id="552B0436FFC7256EFED2FECFFDE4FE87" box="[337,524,330,357]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="19" pageNumber="220" type="Museum">UTEP 20810–11</specimenCode>
(field nos. 
<specimenCode id="552B0436FFC7256EFD0FFECEFCEEFE87" box="[652,774,330,356]" collectionCode="EBG" pageId="19" pageNumber="220">EBG 1316</specimenCode>
, 
<specimenCode id="552B0436FFC7256EFC92FECEFCA5FE87" box="[785,845,330,356]" collectionCode="EBG" pageId="19" pageNumber="220">1319</specimenCode>
), vicinity of 
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(
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, 
<geoCoordinate id="60B9CA8AFFC7256EFB3AFECEFAD1FE86" box="[1209,1337,330,357]" degrees="28.4495" direction="east" orientation="longitude" pageId="19" pageNumber="220" precision="5" value="28.4495">28.4495° E</geoCoordinate>
, 
<quantity id="C27501A8FFC7256EFAC6FECEFA67FE87" box="[1349,1423,330,356]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="19" pageNumber="220" unit="m" value="820.0">
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), 
<collectingMunicipality id="E5563637FFC7256EFF14FEEAFE68FE6B" box="[151,384,366,392]" pageId="19" pageNumber="220">South Kivu Province</collectingMunicipality>
, 
<collectingCountry id="7D9AECDDFFC7256EFE0AFEEAFE2EFE6B" box="[393,454,366,392]" name="Democratic Republic of the Congo" pageId="19" pageNumber="220">DRC</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="B5E5A610FFC7256EFE4CFEEBFCE2FE4F" ID-GBIF-Occurrence="3887776313" collectionCode="UTEP, MUSE" country="Democratic Republic of the Congo" county="South Kivu Province" elevation="1566" latitude="-2.362138" location="Kahuzi-Biega National Park" longLatPrecision="1" longitude="28.192589" municipality="Nanwa" pageId="19" pageNumber="220" specimenCode="UTEP 22423, MUSE 10192" specimenCount="1">
<specimenCode id="552B0436FFC7256EFE4CFEEBFDB6FE6B" box="[463,606,366,393]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="19" pageNumber="220" type="Museum">UTEP 22423</specimenCode>
(field no. 
<specimenCode id="552B0436FFC7256EFD4FFEEAFC80FE6B" box="[716,872,366,392]" collectionCode="MUSE" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35141" lsid="urn:lsid:biocol.org:col:35141" name="Museo delle Scienze" pageId="19" pageNumber="220" type="Museum">MUSE 10192</specimenCode>
), 
<location id="0052FA96FFC7256EFCFAFEEAFB5CFE6B" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFCFAFEEAFB5CFE6B" box="[889,1204,366,392]" country="Democratic Republic of the Congo" county="South Kivu Province" latitude="-2.362138" longLatPrecision="1" longitude="28.192589" municipality="Nanwa" name="Kahuzi-Biega National Park" pageId="19" pageNumber="220">Kahuzi-Biega National Park</location>
, 
<collectingMunicipality id="E5563637FFC7256EFB3EFEEBFAE4FE6A" box="[1213,1292,367,393]" pageId="19" pageNumber="220">Nanwa</collectingMunicipality>
(
<geoCoordinate id="60B9CA8AFFC7256EFA9BFEEAFF40FE4F" degrees="02.362138" direction="south" orientation="latitude" pageId="19" pageNumber="220" precision="1" value="-2.362138">02.362138° S</geoCoordinate>
, 
<geoCoordinate id="60B9CA8AFFC7256EFF35FE16FEBCFE4F" box="[182,340,402,429]" degrees="28.192589" direction="east" orientation="longitude" pageId="19" pageNumber="220" precision="1" value="28.192589">28.192589° E</geoCoordinate>
, 
<quantity id="C27501A8FFC7256EFEE2FE16FE52FE4F" box="[353,442,402,429]" metricMagnitude="3" metricUnit="m" metricValue="1.566" pageId="19" pageNumber="220" unit="m" value="1566.0">
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), 
<collectingCounty id="EC53D4C1FFC7256EFE53FE16FD28FE4F" box="[464,704,402,428]" pageId="19" pageNumber="220">South Kivu Province</collectingCounty>
, 
<collectingCountry id="7D9AECDDFFC7256EFD4EFE16FCE2FE4F" box="[717,778,402,428]" name="Democratic Republic of the Congo" pageId="19" pageNumber="220">DRC</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="B5E5A610FFC7256EFC95FE16FBFCFE33" ID-GBIF-Occurrence="3887776321" collectionCode="UTEP, MUSE" country="Democratic Republic of the Congo" county="South Kivu Province" elevation="1135" latitude="-3.47937" location="Itombwe Plateau" longLatPrecision="1" longitude="28.41305" municipality="Mbandakila" pageId="19" pageNumber="220" specimenCode="UTEP 22424, MUSE 10137" specimenCount="1">
<specimenCode id="552B0436FFC7256EFC95FE16FC42FE4F" box="[790,938,402,428]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="19" pageNumber="220" type="Museum">UTEP 22424</specimenCode>
(field no. 
<specimenCode id="552B0436FFC7256EFBA1FE16FB2AFE4F" box="[1058,1218,402,428]" collectionCode="MUSE" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35141" lsid="urn:lsid:biocol.org:col:35141" name="Museo delle Scienze" pageId="19" pageNumber="220" type="Museum">MUSE 10137</specimenCode>
), 
<location id="0052FA96FFC7256EFB54FE16FA70FE4F" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFB54FE16FA70FE4F" box="[1239,1432,402,428]" country="Democratic Republic of the Congo" county="South Kivu Province" latitude="-3.47937" longLatPrecision="1" longitude="28.41305" municipality="Mbandakila" name="Itombwe Plateau" pageId="19" pageNumber="220">Itombwe Plateau</location>
, 
<collectingMunicipality id="E5563637FFC7256EFF14FE31FEF5FE2C" box="[151,285,437,463]" pageId="19" pageNumber="220">Mbandakila</collectingMunicipality>
(
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, 
<geoCoordinate id="60B9CA8AFFC7256EFE44FE32FD8CFE33" box="[455,612,438,464]" degrees="28.413049" direction="east" orientation="longitude" pageId="19" pageNumber="220" precision="1" value="28.41305">28.413049° E</geoCoordinate>
, 
<quantity id="C27501A8FFC7256EFDF2FE31FD20FE33" box="[625,712,437,464]" metricMagnitude="3" metricUnit="m" metricValue="1.135" pageId="19" pageNumber="220" unit="m" value="1135.0">
<elevation id="8EA04B7EFFC7256EFDF2FE31FD20FE33" box="[625,712,437,464]" metricMagnitude="3" metricUnit="m" metricValue="1.135" pageId="19" pageNumber="220" unit="m" value="1135.0">1135 m</elevation>
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), 
<collectingCounty id="EC53D4C1FFC7256EFD5EFE31FC23FE2C" box="[733,971,437,463]" pageId="19" pageNumber="220">South Kivu Province</collectingCounty>
, 
<collectingCountry id="7D9AECDDFFC7256EFC54FE32FBFCFE33" box="[983,1044,438,464]" name="Democratic Republic of the Congo" pageId="19" pageNumber="220">DRC</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="B5E5A610FFC7256EFBA3FE32FBE0FE10" ID-GBIF-Occurrence="3887776335" collectionCode="RMCA" country="Democratic Republic of the Congo" county="South Kivu Province" elevation="2300" location="Itombwe Plateau" municipality="Tubutubu" pageId="19" pageNumber="220" specimenCode="RMCA 77-020" specimenCount="1">
<specimenCode id="552B0436FFC7256EFBA3FE32FB22FE2C" box="[1056,1226,437,464]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="19" pageNumber="220" type="Museum">RMCA 77-020</specimenCode>
-B-133–136* (
<specimenCount id="138B67C4FFC7256EFAEFFE31FEF8FE10" pageId="19" pageNumber="220" type="generic">four specimens</specimenCount>
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<location id="0052FA96FFC7256EFEA0FE5DFE0AFE10" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFEA0FE5DFE0AFE10" box="[291,482,473,499]" country="Democratic Republic of the Congo" county="South Kivu Province" municipality="Tubutubu" name="Itombwe Plateau" pageId="19" pageNumber="220">Itombwe Plateau</location>
, 
<collectingMunicipality id="E5563637FFC7256EFE6FFE5DFDB0FE10" box="[492,600,473,499]" pageId="19" pageNumber="220">Tubutubu</collectingMunicipality>
(
<quantity id="C27501A8FFC7256EFDE5FE5DFD57FE10" box="[614,703,473,499]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="19" pageNumber="220" unit="m" value="2300.0">
<elevation id="8EA04B7EFFC7256EFDE5FE5DFD57FE10" box="[614,703,473,499]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="19" pageNumber="220" unit="m" value="2300.0">2300 m</elevation>
</quantity>
), 
<collectingCounty id="EC53D4C1FFC7256EFD50FE5DFC28FE10" box="[723,960,473,499]" pageId="19" pageNumber="220">South Kivu Province</collectingCounty>
, 
<collectingCountry id="7D9AECDDFFC7256EFC48FE5DFBE0FE10" box="[971,1032,473,499]" name="Democratic Republic of the Congo" pageId="19" pageNumber="220">DRC</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="B5E5A610FFC7256EFB91FE5DFB37FDF4" ID-GBIF-Occurrence="3887776311" collectionCode="RMCA, MF" country="Democratic Republic of the Congo" elevation="700" latitude="-3.28" location="Kitutu" longLatPrecision="784" longitude="28.11" municipality="South Kivu Province" pageId="19" pageNumber="220" specimenCode="RMCA 77-020, MF 14-15" specimenCount="1">
<specimenCode id="552B0436FFC7256EFB91FE5DFB52FE10" box="[1042,1210,473,499]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="19" pageNumber="220" type="Museum">RMCA 77-020</specimenCode>
-B-89–92 (field nos. 
<specimenCode id="552B0436FFC7256EFF14FE7AFEE5FDFB" box="[151,269,509,536]" collectionCode="MF" pageId="19" pageNumber="220">MF 14–15</specimenCode>
[
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]), 
<location id="0052FA96FFC7256EFE65FE79FDC6FDF4" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFE65FE79FDC6FDF4" box="[486,558,509,535]" country="Democratic Republic of the Congo" latitude="-3.28" longLatPrecision="784" longitude="28.11" municipality="South Kivu Province" name="Kitutu" pageId="19" pageNumber="220">Kitutu</location>
(ca. 
<geoCoordinate id="60B9CA8AFFC7256EFDE5FE79FD21FDF4" box="[614,713,509,535]" degrees="03.28" direction="south" orientation="latitude" pageId="19" pageNumber="220" precision="555" value="-3.28">03.28° S</geoCoordinate>
, 
<geoCoordinate id="60B9CA8AFFC7256EFD54FE79FCD2FDFB" box="[727,826,509,536]" degrees="28.11" direction="east" orientation="longitude" pageId="19" pageNumber="220" precision="555" value="28.11">28.11° E</geoCoordinate>
, 
<quantity id="C27501A8FFC7256EFCC4FE79FC7AFDF4" box="[839,914,509,535]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="220" unit="m" value="700.0">
<elevation id="8EA04B7EFFC7256EFCC4FE79FC7AFDF4" box="[839,914,509,535]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="19" pageNumber="220" unit="m" value="700.0">700 m</elevation>
</quantity>
), 
<collectingMunicipality id="E5563637FFC7256EFC24FE79FB7EFDF4" box="[935,1174,509,535]" pageId="19" pageNumber="220">South Kivu Province</collectingMunicipality>
, 
<collectingCountry id="7D9AECDDFFC7256EFB21FE79FB37FDF4" box="[1186,1247,509,535]" name="Democratic Republic of the Congo" pageId="19" pageNumber="220">DRC</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="B5E5A610FFC7256EFB68FE79FB57FDD8" ID-GBIF-Occurrence="3887776332" collectionCode="RMCA" country="Democratic Republic of the Congo" elevation="650" latitude="-3.27" location="Itula" longLatPrecision="784" longitude="28.12" municipality="South Kivu Province" pageId="19" pageNumber="220" specimenCode="RMCA 77-020" specimenCount="1">
<specimenCode id="552B0436FFC7256EFB68FE79FA7DFDF4" box="[1259,1429,509,535]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="19" pageNumber="220" type="Museum">RMCA 77-020</specimenCode>
- B-0069–71 (
<specimenCount id="138B67C4FFC7256EFEA9FDA5FE46FDD8" box="[298,430,545,571]" pageId="19" pageNumber="220" type="generic">1 specimen</specimenCount>
), 
<location id="0052FA96FFC7256EFE47FDA5FE1FFDD8" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFE47FDA5FE1FFDD8" box="[452,503,545,571]" country="Democratic Republic of the Congo" latitude="-3.27" longLatPrecision="784" longitude="28.12" municipality="South Kivu Province" name="Itula" pageId="19" pageNumber="220">Itula</location>
(ca. 
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, 
<geoCoordinate id="60B9CA8AFFC7256EFD2BFDA5FCE7FDD8" box="[680,783,545,571]" degrees="28.12" direction="east" orientation="longitude" pageId="19" pageNumber="220" precision="555" value="28.12">28.12° E</geoCoordinate>
, 
<quantity id="C27501A8FFC7256EFC9EFDA5FC82FDD8" box="[797,874,545,572]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="19" pageNumber="220" unit="m" value="650.0">
<elevation id="8EA04B7EFFC7256EFC9EFDA5FC82FDD8" box="[797,874,545,572]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="19" pageNumber="220" unit="m" value="650.0">650 m</elevation>
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), 
<collectingMunicipality id="E5563637FFC7256EFC02FDA5FB9CFDD8" box="[897,1140,545,571]" pageId="19" pageNumber="220">South Kivu Province</collectingMunicipality>
, 
<collectingCountry id="7D9AECDDFFC7256EFB01FDA5FB57FDD8" box="[1154,1215,545,571]" name="Democratic Republic of the Congo" pageId="19" pageNumber="220">DRC</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="B5E5A610FFC7256EFB4EFDA5FB08FDBC" ID-GBIF-Occurrence="3887776319" collectionCode="RMCA" country="Democratic Republic of the Congo" elevation="1050" latitude="-3.54" location="Kamituga" longLatPrecision="784" longitude="27.69" municipality="South Kivu Province" pageId="19" pageNumber="220" specimenCode="RMCA 77-020" specimenCount="1">
<specimenCode id="552B0436FFC7256EFB4EFDA5FA91FDD8" box="[1229,1401,545,571]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="19" pageNumber="220" type="Museum">RMCA 77-020</specimenCode>
-B-0081–83, 77-20-B-0086, 
<location id="0052FA96FFC7256EFE36FDC0FDCBFDBD" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFE36FDC0FDCBFDBD" box="[437,547,580,606]" country="Democratic Republic of the Congo" latitude="-3.54" longLatPrecision="784" longitude="27.69" municipality="South Kivu Province" name="Kamituga" pageId="19" pageNumber="220">Kamituga</location>
(ca. 
<geoCoordinate id="60B9CA8AFFC7256EFDD8FDC0FD56FDBC" box="[603,702,580,607]" degrees="03.54" direction="south" orientation="latitude" pageId="19" pageNumber="220" precision="555" value="-3.54">03.54° S</geoCoordinate>
, 
<geoCoordinate id="60B9CA8AFFC7256EFD48FDC0FCC7FDBC" box="[715,815,580,607]" degrees="27.69" direction="east" orientation="longitude" pageId="19" pageNumber="220" precision="555" value="27.69">27.69° E</geoCoordinate>
, 
<quantity id="C27501A8FFC7256EFCBFFDC0FC7CFDBC" box="[828,916,580,607]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="19" pageNumber="220" unit="m" value="1050.0">
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</quantity>
), 
<collectingMunicipality id="E5563637FFC7256EFC2AFDC0FB7FFDBD" box="[937,1175,580,606]" pageId="19" pageNumber="220">South Kivu Province</collectingMunicipality>
, 
<collectingCountry id="7D9AECDDFFC7256EFB20FDC1FB08FDBC" box="[1187,1248,581,607]" name="Democratic Republic of the Congo" pageId="19" pageNumber="220">DRC</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="B5E5A610FFC7256EFB68FDC1FB56FD61" ID-GBIF-Occurrence="3887776320" collectionCode="RMCA" country="Democratic Republic of the Congo" elevation="1650" latitude="-3.15" location="Mwana" longLatPrecision="784" longitude="28.47" municipality="South Kivu Province" pageId="19" pageNumber="220" specimenCode="RMCA 77-020" specimenCount="1">
<specimenCode id="552B0436FFC7256EFB68FDC1FA7DFDBC" box="[1259,1429,581,607]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="19" pageNumber="220" type="Museum">RMCA 77-020</specimenCode>
- B-0072–73 (
<specimenCount id="138B67C4FFC7256EFEA4FDECFE4FFD61" box="[295,423,616,642]" pageId="19" pageNumber="220" type="generic">1 specimen</specimenCount>
), 
<location id="0052FA96FFC7256EFE3AFDEDFDE5FD60" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFE3AFDEDFDE5FD60" box="[441,525,617,643]" country="Democratic Republic of the Congo" latitude="-3.15" longLatPrecision="784" longitude="28.47" municipality="South Kivu Province" name="Mwana" pageId="19" pageNumber="220">Mwana</location>
(ca. 
<geoCoordinate id="60B9CA8AFFC7256EFDC0FDECFD4DFD61" box="[579,677,616,643]" degrees="03.15" direction="south" orientation="latitude" pageId="19" pageNumber="220" precision="555" value="-3.15">03.15° S</geoCoordinate>
, 
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, 
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), 
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, 
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; 
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(field no. 
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), 
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(
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, 
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, 
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), 
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, 
<collectingCountry id="7D9AECDDFFC7256EFBD1FD08FB67FD45" box="[1106,1167,652,678]" name="Democratic Republic of the Congo" pageId="19" pageNumber="220">DRC</collectingCountry>
; (field no. 
<specimenCode id="552B0436FFC7256EFA8DFD09FA63FD44" box="[1294,1419,652,679]" collectionCode="EPLU" pageId="19" pageNumber="220">EPLU 395</specimenCode>
*, photo and tissue only)
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, 
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ca. 
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<quantity id="C27501A8FFC7256EFE46FD2BFDFCFD29" box="[453,532,687,714]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="220" unit="km" value="0.5">0.5 km</quantity>
E of Epulu
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, near 
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(
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, 
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, 
<quantity id="C27501A8FFC7256EFB1FFD2BFB0EFD29" box="[1180,1254,687,714]" metricMagnitude="2" metricUnit="m" metricValue="7.55" pageId="19" pageNumber="220" unit="m" value="755.0">
<elevation id="8EA04B7EFFC7256EFB1FFD2BFB0EFD29" box="[1180,1254,687,714]" metricMagnitude="2" metricUnit="m" metricValue="7.55" pageId="19" pageNumber="220" unit="m" value="755.0">755 m</elevation>
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), 
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, 
<collectingCountry id="7D9AECDDFFC7256EFF14FD50FF3CFD0D" box="[151,212,724,750]" name="Democratic Republic of the Congo" pageId="19" pageNumber="220">DRC</collectingCountry>
</materialsCitation>
; 
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<specimenCode id="552B0436FFC7256EFF5CFD50FE99FD0E" box="[223,369,723,750]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="19" pageNumber="220" type="Museum">UTEP 22416</specimenCode>
(field no. 
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), 
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, 
<collectingMunicipality id="E5563637FFC7256EFB81FD57FB89FD0E" box="[1026,1121,723,749]" pageId="19" pageNumber="220">Buhoma</collectingMunicipality>
(
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, 
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, 
<quantity id="C27501A8FFC7256EFF14FD73FF06FCF2" box="[151,238,759,786]" metricMagnitude="3" metricUnit="m" metricValue="1.523" pageId="19" pageNumber="220" unit="m" value="1523.0">
<elevation id="8EA04B7EFFC7256EFF14FD73FF06FCF2" box="[151,238,759,786]" metricMagnitude="3" metricUnit="m" metricValue="1.523" pageId="19" pageNumber="220" unit="m" value="1523.0">1523 m</elevation>
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), 
<collectingCounty id="EC53D4C1FFC7256EFE83FD7CFE5CFCF2" box="[256,436,759,786]" pageId="19" pageNumber="220">Western Region</collectingCounty>
, 
<collectingCountry id="7D9AECDDFFC7256EFE3DFD73FDF1FCF2" box="[446,537,759,785]" name="Uganda" pageId="19" pageNumber="220">Uganda</collectingCountry>
; (field nos. 
<specimenCode id="552B0436FFC7256EFD1BFD7CFCD2FCF2" box="[664,826,759,786]" collectionCode="DFH" pageId="19" pageNumber="220">DFH 1102–03</specimenCode>
* photos and tissues only)
</materialsCitation>
, 
<materialsCitation id="B5E5A610FFC7256EFBE3FD73FB9DFCD6" ID-GBIF-Occurrence="3887776333" country="Uganda" county="Western Region" elevation="1363" latitude="0.49795" location="Forest National Park" longLatPrecision="1" longitude="30.42262" municipality="Ngogo Research Center" pageId="19" pageNumber="220" specimenCount="1" stateProvince="Kibale">
<collectingRegion id="C74962AFFFC7256EFBE3FD73FB43FCF2" box="[1120,1195,759,785]" country="Uganda" name="Kibale" pageId="19" pageNumber="220">Kibale</collectingRegion>
<location id="0052FA96FFC7256EFB33FD7CFA70FCF2" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFB33FD7CFA70FCF2" box="[1200,1432,759,786]" country="Uganda" county="Western Region" latitude="0.49795" longLatPrecision="1" longitude="30.42262" municipality="Ngogo Research Center" name="Forest National Park" pageId="19" pageNumber="220" stateProvince="Kibale">Forest National Park</location>
, 
<collectingMunicipality id="E5563637FFC7256EFF14FC9FFE4BFCD6" box="[151,419,795,821]" pageId="19" pageNumber="220">Ngogo Research Center</collectingMunicipality>
(
<geoCoordinate id="60B9CA8AFFC7256EFE31FC9FFDAEFCD6" box="[434,582,795,822]" degrees="00.49795" direction="north" orientation="latitude" pageId="19" pageNumber="220" precision="1" value="0.49795">00.49795° N</geoCoordinate>
, 
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, 
<quantity id="C27501A8FFC7256EFD6FFC9FFCADFCD6" box="[748,837,795,821]" metricMagnitude="3" metricUnit="m" metricValue="1.363" pageId="19" pageNumber="220" unit="m" value="1363.0">
<elevation id="8EA04B7EFFC7256EFD6FFC9FFCADFCD6" box="[748,837,795,821]" metricMagnitude="3" metricUnit="m" metricValue="1.363" pageId="19" pageNumber="220" unit="m" value="1363.0">1363 m</elevation>
</quantity>
), 
<collectingCounty id="EC53D4C1FFC7256EFCDBFC9FFBE7FCD6" box="[856,1039,795,821]" pageId="19" pageNumber="220">Western Region</collectingCounty>
, 
<collectingCountry id="7D9AECDDFFC7256EFB99FC9FFB9DFCD6" box="[1050,1141,795,821]" name="Uganda" pageId="19" pageNumber="220">Uganda</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="B5E5A610FFC7256EFB03FC9FFEBBFC9F" ID-GBIF-Occurrence="3887776317" collectionCode="CAS" country="Uganda" county="Western Region" latitude="-1.00975" location="Bwindi Impenetrable National Park" longLatPrecision="1" longitude="29.620693" municipality="Bizenga River" pageId="19" pageNumber="220" specimenCode="CAS 202032-36" specimenCount="1">
<specimenCode id="552B0436FFC7256EFB03FC9FFAD6FCD6" box="[1152,1342,795,821]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="19" pageNumber="220" type="Museum">CAS 202032–36</specimenCode>
, 
<location id="0052FA96FFC7256EFAC9FC9FFE38FCBA" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFAC9FC9FFE38FCBA" country="Uganda" county="Western Region" latitude="-1.00975" longLatPrecision="1" longitude="29.620693" municipality="Bizenga River" name="Bwindi Impenetrable National Park" pageId="19" pageNumber="220">Bwindi Impenetrable National Park</location>
, Buhoma rd., 
<quantity id="C27501A8FFC7256EFDF3FCBBFD41FCBA" box="[624,681,831,857]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="220" unit="km" value="2.0">2 km</quantity>
S (by rd.) of 
<collectingMunicipality id="E5563637FFC7256EFCC0FCBBFC0CFCBA" box="[835,996,831,857]" pageId="19" pageNumber="220">Bizenga River</collectingMunicipality>
(
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, 
<geoCoordinate id="60B9CA8AFFC7256EFB0CFCBBFAC5FCBA" box="[1167,1325,831,857]" degrees="29.620694" direction="east" orientation="longitude" pageId="19" pageNumber="220" precision="1" value="29.620693">29.620694° E</geoCoordinate>
), 
<collectingCounty id="EC53D4C1FFC7256EFAC2FCBBFF03FC9F" pageId="19" pageNumber="220">Western Region</collectingCounty>
, 
<collectingCountry id="7D9AECDDFFC7256EFF7BFCE6FEBBFC9F" box="[248,339,866,892]" name="Uganda" pageId="19" pageNumber="220">Uganda</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="B5E5A610FFC7256EFEE3FCE6FD36FC43" ID-GBIF-Occurrence="3887776330" collectionCode="CAS" country="Uganda" county="Western Region" elevation="1280" latitude="-0.905" location="Bwindi Impenetrable National Park" longLatPrecision="55" longitude="29.704971" municipality="Ishasha River" pageId="19" pageNumber="220" specimenCode="CAS 202133" specimenCount="1">
<specimenCode id="552B0436FFC7256EFEE3FCE6FE1FFC9F" box="[352,503,866,892]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="19" pageNumber="220" type="Museum">CAS 202133</specimenCode>
, 
<location id="0052FA96FFC7256EFD87FCE6FC73FC9F" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFD87FCE6FC73FC9F" box="[516,923,866,892]" country="Uganda" county="Western Region" latitude="-0.905" longLatPrecision="55" longitude="29.704971" municipality="Ishasha River" name="Bwindi Impenetrable National Park" pageId="19" pageNumber="220">Bwindi Impenetrable National Park</location>
, small tributary to 
<collectingMunicipality id="E5563637FFC7256EFBF4FCE6FAFAFC9F" box="[1143,1298,866,892]" pageId="19" pageNumber="220">Ishasha River</collectingMunicipality>
(
<geoCoordinate id="60B9CA8AFFC7256EFAA0FCE6FA7FFC9F" box="[1315,1431,866,893]" degrees="00.905" direction="south" orientation="latitude" pageId="19" pageNumber="220" precision="55" value="-0.905">00.905° S</geoCoordinate>
, 
<geoCoordinate id="60B9CA8AFFC7256EFF14FC02FEDFFC42" box="[151,311,902,929]" degrees="29.704972" direction="east" orientation="longitude" pageId="19" pageNumber="220" precision="1" value="29.704971">29.704972° E</geoCoordinate>
, 
<quantity id="C27501A8FFC7256EFEC4FC02FE4BFC43" box="[327,419,902,928]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="19" pageNumber="220" unit="m" value="1280.0">
<elevation id="8EA04B7EFFC7256EFEC4FC02FE4BFC43" box="[327,419,902,928]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="19" pageNumber="220" unit="m" value="1280.0">1280 m</elevation>
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), 
<collectingCounty id="EC53D4C1FFC7256EFE38FC03FD9CFC43" box="[443,628,902,929]" pageId="19" pageNumber="220">Western Region</collectingCounty>
, 
<collectingCountry id="7D9AECDDFFC7256EFD00FC02FD36FC43" box="[643,734,902,928]" name="Uganda" pageId="19" pageNumber="220">Uganda</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="B5E5A610FFC7256EFD6EFC02FC1DFC27" ID-GBIF-Occurrence="3887776324" collectionCode="CAS" country="Uganda" county="Western Region" latitude="-0.99275" location="Bwindi Impenetrable National Park" longLatPrecision="1" longitude="29.615723" municipality="Buhoma rd." pageId="19" pageNumber="220" specimenCode="CAS 202109" specimenCount="1">
<specimenCode id="552B0436FFC7256EFD6EFC02FC6DFC43" box="[749,901,902,928]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="19" pageNumber="220" type="Museum">CAS 202109</specimenCode>
, 
<location id="0052FA96FFC7256EFC17FC02FAD9FC43" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFC17FC02FAD9FC43" box="[916,1329,902,928]" country="Uganda" county="Western Region" latitude="-0.99275" longLatPrecision="1" longitude="29.615723" municipality="Buhoma rd." name="Bwindi Impenetrable National Park" pageId="19" pageNumber="220">Bwindi Impenetrable National Park</location>
, 
<location id="0052FA96FFC7256EFAC3FC02FF3DFC27" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFAC3FC02FF3DFC27" country="Uganda" county="Western Region" latitude="-0.99275" longLatPrecision="1" longitude="29.615723" municipality="Buhoma rd." name="Bizenga River" pageId="19" pageNumber="220">Bizenga River</location>
at 
<collectingMunicipality id="E5563637FFC7256EFF7BFC2EFE95FC27" box="[248,381,938,964]" pageId="19" pageNumber="220">Buhoma rd.</collectingMunicipality>
(
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, 
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), 
<collectingCounty id="EC53D4C1FFC7256EFD5AFC2EFC67FC27" box="[729,911,938,964]" pageId="19" pageNumber="220">Western Region</collectingCounty>
, 
<collectingCountry id="7D9AECDDFFC7256EFC19FC2EFC1DFC27" box="[922,1013,938,964]" name="Uganda" pageId="19" pageNumber="220">Uganda</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="B5E5A610FFC7256EFB82FC2EFCA4FC04" ID-GBIF-Occurrence="3887776336" collectionCode="CAS" country="Uganda" county="Western Region" location="Bwindi Impenetrable National Park" municipality="Ruhija" pageId="19" pageNumber="220" specimenCode="CAS 256035" specimenCount="1">
<specimenCode id="552B0436FFC7256EFB82FC2EFB70FC26" box="[1025,1176,938,965]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="19" pageNumber="220" type="Museum">CAS 256035</specimenCode>
*, 
<location id="0052FA96FFC7256EFB2CFC2EFEDFFC04" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFB2CFC2EFEDFFC04" country="Uganda" county="Western Region" municipality="Ruhija" name="Bwindi Impenetrable National Park" pageId="19" pageNumber="220">Bwindi Impenetrable National Park</location>
, rd. north of 
<collectingMunicipality id="E5563637FFC7256EFE52FC49FDF7FC04" box="[465,543,973,999]" pageId="19" pageNumber="220">Ruhija</collectingMunicipality>
, 
<collectingCounty id="EC53D4C1FFC7256EFDAFFC4AFD0CFC04" box="[556,740,973,1000]" pageId="19" pageNumber="220">Western Region</collectingCounty>
, 
<collectingCountry id="7D9AECDDFFC7256EFD72FC49FCA4FC04" box="[753,844,973,999]" name="Uganda" pageId="19" pageNumber="220">Uganda</collectingCountry>
</materialsCitation>
; 
<materialsCitation id="B5E5A610FFC7256EFCD9FC4AFA94FBE8" ID-GBIF-Occurrence="3887776309" collectionCode="CAS" country="Uganda" elevation="1121" latitude="0.4508" location="Mabira Forest Reserve" longLatPrecision="7" longitude="32.9493" municipality="Central Region" pageId="19" pageNumber="220" specimenCode="CAS 256128-31" specimenCount="1" stateProvince="Mukono">
<specimenCode id="552B0436FFC7256EFCD9FC4AFBF5FC0B" box="[858,1053,973,1000]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="19" pageNumber="220" type="Museum">CAS 256128–31</specimenCode>
*, 
<location id="0052FA96FFC7256EFBB5FC49FAD4FC0B" LSID="urn:lsid:plazi:treatment:8D241D5BFFC5256BFF14F917FAE2FB42:0052FA96FFC7256EFBB5FC49FAD4FC0B" box="[1078,1340,973,1000]" country="Uganda" latitude="0.4508" longLatPrecision="7" longitude="32.9493" municipality="Central Region" name="Mabira Forest Reserve" pageId="19" pageNumber="220" stateProvince="Mukono">Mabira Forest Reserve</location>
, marshy area adjacent to unpolluted stream (
<geoCoordinate id="60B9CA8AFFC7256EFDA5FC75FD76FBEF" box="[550,670,1009,1036]" degrees="0.4508" direction="north" orientation="latitude" pageId="19" pageNumber="220" precision="5" value="0.4508">0.4508° N</geoCoordinate>
, 
<geoCoordinate id="60B9CA8AFFC7256EFD2AFC75FCC1FBEF" box="[681,809,1009,1036]" degrees="32.9493" direction="east" orientation="longitude" pageId="19" pageNumber="220" precision="5" value="32.9493">32.9493° E</geoCoordinate>
, 
<quantity id="C27501A8FFC7256EFCB6FC75FC64FBE8" box="[821,908,1009,1035]" metricMagnitude="3" metricUnit="m" metricValue="1.121" pageId="19" pageNumber="220" unit="m" value="1121.0">
<elevation id="8EA04B7EFFC7256EFCB6FC75FC64FBE8" box="[821,908,1009,1035]" metricMagnitude="3" metricUnit="m" metricValue="1.121" pageId="19" pageNumber="220" unit="m" value="1121.0">1121 m</elevation>
</quantity>
), 
<collectingRegion id="C74962AFFFC7256EFC23FC75FBB5FBE8" box="[928,1117,1009,1035]" country="Uganda" name="Mukono" pageId="19" pageNumber="220">Mukono District</collectingRegion>
, 
<collectingMunicipality id="E5563637FFC7256EFBEBFC75FAFEFBE8" box="[1128,1302,1009,1035]" pageId="19" pageNumber="220">Central Region</collectingMunicipality>
, 
<collectingCountry id="7D9AECDDFFC7256EFAA2FC75FA94FBE8" box="[1313,1404,1009,1035]" name="Uganda" pageId="19" pageNumber="220">Uganda</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4D97FFC6FFC7256EFF44FB91FA8EF817" pageId="19" pageNumber="220" type="diagnosis">
<paragraph id="0532AC4DFFC7256EFF44FB91FA8EF817" blockId="19.[151,1437,151,2037]" pageId="19" pageNumber="220">
<emphasis id="37F9705FFFC7256EFF44FB91FEAAFBCC" bold="true" box="[199,322,1045,1071]" pageId="19" pageNumber="220">Diagnosis.</emphasis>
The species is referred to the genus 
<taxonomicName id="C28DD7CEFFC7256EFD4CFB91FCDEFBCC" authorityName="Laurent" authorityYear="1944" box="[719,822,1045,1071]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="genus">
<emphasis id="37F9705FFFC7256EFD4CFB91FCDEFBCC" box="[719,822,1045,1071]" italics="true" pageId="19" pageNumber="220">Afrixalus</emphasis>
</taxonomicName>
for exhibiting the following characteristics: fingers and toes webbed; tips of fingers and toes enlarged to discs; eye large; pupil vertically elliptical; tympanum indistinct; vomer ridges and teeth absent; dorsal surfaces finely shagreened with minute pointed tubercles; outer metatarsal tubercle distinct; gular gland in males. It is readily distinguished from most other members of the genus by its small size (SVL in males 
<quantity id="C27501A8FFC7256EFE3FFB20FD8AFB5C" box="[444,610,1188,1215]" metricMagnitude="-2" metricUnit="m" metricValue="2.0549999999999997" metricValueMax="2.2199999999999998" metricValueMin="1.89" pageId="19" pageNumber="220" unit="mm" value="20.549999999999997" valueMax="22.2" valueMin="18.9">18.9–22.2 mm</quantity>
, in females 
<quantity id="C27501A8FFC7256EFD6FFB20FC7CFB5D" box="[748,916,1188,1215]" metricMagnitude="-2" metricUnit="m" metricValue="2.325" metricValueMax="2.57" metricValueMin="2.08" pageId="19" pageNumber="220" unit="mm" value="23.25" valueMax="25.7" valueMin="20.8">20.8–25.7 mm</quantity>
), being smaller than 
<taxonomicName id="C28DD7CEFFC7256EFB02FB20FB13FB5D" baseAuthorityName="Peters" baseAuthorityYear="1875" box="[1153,1275,1188,1214]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="37F9705FFFC7256EFB02FB20FB13FB5D" box="[1153,1275,1188,1214]" italics="true" pageId="19" pageNumber="220">A. dorsalis</emphasis>
</taxonomicName>
(males 
<quantity id="C27501A8FFC7256EFAD5FB20FF2DFB01" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="19" pageNumber="220" unit="mm" value="26.5" valueMax="28.0" valueMin="25.0">25–28 mm</quantity>
, females 
<quantity id="C27501A8FFC7256EFEB3FB4CFE46FB01" box="[304,430,1224,1250]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="19" pageNumber="220" unit="mm" value="28.0" valueMax="30.0" valueMin="26.0">26–30 mm</quantity>
), 
<taxonomicName id="C28DD7CEFFC7256EFE42FB4CFDAEFB01" baseAuthorityName="Bianconi" baseAuthorityYear="1849" box="[449,582,1224,1250]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="fornasini">
<emphasis id="37F9705FFFC7256EFE42FB4CFDAEFB01" box="[449,582,1224,1250]" italics="true" pageId="19" pageNumber="220">A. fornasini</emphasis>
</taxonomicName>
(males 
<quantity id="C27501A8FFC7256EFD23FB4CFCF4FB01" box="[672,796,1224,1250]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="3.8" metricValueMin="3.0" pageId="19" pageNumber="220" unit="mm" value="34.0" valueMax="38.0" valueMin="30.0">30–38 mm</quantity>
, females 
<quantity id="C27501A8FFC7256EFC04FB43FBECFB01" box="[903,1028,1223,1250]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="19" pageNumber="220" unit="mm" value="37.5" valueMax="40.0" valueMin="35.0">35–40 mm</quantity>
), 
<taxonomicName id="C28DD7CEFFC7256EFB9BFB4CFB2AFB01" baseAuthorityName="Cope" baseAuthorityYear="1861" box="[1048,1218,1224,1250]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="fulvovittatus">
<emphasis id="37F9705FFFC7256EFB9BFB4CFB2AFB01" box="[1048,1218,1224,1250]" italics="true" pageId="19" pageNumber="220">A. fulvovittatus</emphasis>
</taxonomicName>
(males 
<quantity id="C27501A8FFC7256EFA9FFB4CFA70FB01" box="[1308,1432,1224,1250]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="19" pageNumber="220" unit="mm" value="25.0" valueMax="27.0" valueMin="23.0">23–27 mm</quantity>
, females 
<quantity id="C27501A8FFC7256EFF76FB6FFE9AFAE5" box="[245,370,1259,1286]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="19" pageNumber="220" unit="mm" value="26.5" valueMax="28.0" valueMin="25.0">25–28 mm</quantity>
), 
<taxonomicName id="C28DD7CEFFC7256EFE06FB68FE1CFAE6" box="[389,500,1259,1286]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="lacteus">
<emphasis id="37F9705FFFC7256EFE06FB68FE1CFAE6" box="[389,500,1259,1286]" italics="true" pageId="19" pageNumber="220">A. lacteus</emphasis>
</taxonomicName>
(males 
<quantity id="C27501A8FFC7256EFDCEFB68FD20FAE5" box="[589,712,1260,1286]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="19" pageNumber="220" unit="mm" value="24.5" valueMax="27.0" valueMin="22.0">22–27 mm</quantity>
, females 
<quantity id="C27501A8FFC7256EFCB1FB6FFC47FAE5" box="[818,943,1259,1286]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="19" pageNumber="220" unit="mm" value="27.0" valueMax="29.0" valueMin="25.0">25–29 mm</quantity>
), 
<taxonomicName id="C28DD7CEFFC7256EFC41FB68FB8DFAE6" authorityName="Laurent" authorityYear="1950" box="[962,1125,1259,1286]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="leucostictus">
<emphasis id="37F9705FFFC7256EFC41FB68FB8DFAE6" box="[962,1125,1259,1286]" italics="true" pageId="19" pageNumber="220">A. leucostictus</emphasis>
</taxonomicName>
(males 
<quantity id="C27501A8FFC7256EFB3DFB68FAD1FAE5" box="[1214,1337,1260,1286]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.2" metricValueMin="2.7" pageId="19" pageNumber="220" unit="mm" value="29.5" valueMax="32.0" valueMin="27.0">27–32 mm</quantity>
, females 
<quantity id="C27501A8FFC7256EFF14FA8BFEFCFACA" box="[151,276,1295,1322]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.6" metricValueMin="2.5" pageId="19" pageNumber="220" unit="mm" value="30.5" valueMax="36.0" valueMin="25.0">25–36 mm</quantity>
), 
<taxonomicName id="C28DD7CEFFC7256EFEA5FA94FE1AFACA" box="[294,498,1295,1322]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="manengubensis">
<emphasis id="37F9705FFFC7256EFEA5FA94FE1AFACA" box="[294,498,1295,1322]" italics="true" pageId="19" pageNumber="220">A. manengubensis</emphasis>
</taxonomicName>
(males to 
<quantity id="C27501A8FFC7256EFDE5FA8BFD50FACA" box="[614,696,1295,1321]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="19" pageNumber="220" unit="mm" value="32.0">32 mm</quantity>
), 
<taxonomicName id="C28DD7CEFFC7256EFD48FA94FC8CFACA" authorityName="Schiotz" authorityYear="1963" box="[715,868,1296,1322]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="nigeriensis">
<emphasis id="37F9705FFFC7256EFD48FA94FC8CFACA" box="[715,868,1296,1322]" italics="true" pageId="19" pageNumber="220">A. nigeriensis</emphasis>
</taxonomicName>
(males 
<quantity id="C27501A8FFC7256EFC3FFA8BFBDEFAC9" box="[956,1078,1295,1322]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.4" metricValueMin="2.8" pageId="19" pageNumber="220" unit="mm" value="31.0" valueMax="34.0" valueMin="28.0">28–34 mm</quantity>
, females 
<quantity id="C27501A8FFC7256EFB1CFA8BFAF4FACA" box="[1183,1308,1295,1322]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="19" pageNumber="220" unit="mm" value="33.5" valueMax="35.0" valueMin="32.0">32–35 mm</quantity>
), 
<taxonomicName id="C28DD7CEFFC7256EFAADFA94FA74FACA" baseAuthorityName="Ferreira" baseAuthorityYear="1906" box="[1326,1436,1296,1322]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="osorioi">
<emphasis id="37F9705FFFC7256EFAADFA94FA74FACA" box="[1326,1436,1296,1322]" italics="true" pageId="19" pageNumber="220">A. osorioi</emphasis>
</taxonomicName>
(males 27–31, females 
<quantity id="C27501A8FFC7256EFE26FAB7FDCEFAAE" box="[421,550,1331,1358]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="19" pageNumber="220" unit="mm" value="33.5" valueMax="35.0" valueMin="32.0">32–35 mm</quantity>
), 
<taxonomicName id="C28DD7CEFFC7256EFDBFFAB7FD06FAAE" box="[572,750,1331,1357]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="paradorsalis">
<emphasis id="37F9705FFFC7256EFDBFFAB7FD06FAAE" box="[572,750,1331,1357]" italics="true" pageId="19" pageNumber="220">A. paradorsalis</emphasis>
</taxonomicName>
(males 
<quantity id="C27501A8FFC7256EFCCEFAB7FC24FAAE" box="[845,972,1331,1357]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.4" metricValueMin="2.8" pageId="19" pageNumber="220" unit="mm" value="31.0" valueMax="34.0" valueMin="28.0">28–34 mm</quantity>
, females 
<quantity id="C27501A8FFC7256EFBBFFAB7FB55FAAE" box="[1084,1213,1331,1358]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="19" pageNumber="220" unit="mm" value="33.5" valueMax="35.0" valueMin="32.0">32–35 mm</quantity>
), 
<taxonomicName id="C28DD7CEFFC7256EFB50FAB7FA74FAAE" authorityName="Schiotz" authorityYear="1974" box="[1235,1436,1331,1357]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="37F9705FFFC7256EFB50FAB7FA74FAAE" box="[1235,1436,1331,1357]" italics="true" pageId="19" pageNumber="220">A. septentrionalis</emphasis>
</taxonomicName>
(males 
<quantity id="C27501A8FFC7256EFF68FAD3FE82FA92" box="[235,362,1367,1393]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="19" pageNumber="220" unit="mm" value="20.0" valueMax="21.0" valueMin="19.0">19–21 mm</quantity>
), 
<taxonomicName id="C28DD7CEFFC7256EFEFCFAD3FE19FA93" baseAuthorityName="Peters" baseAuthorityYear="1876" box="[383,497,1367,1393]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="vittiger">
<emphasis id="37F9705FFFC7256EFEFCFAD3FE19FA93" box="[383,497,1367,1393]" italics="true" pageId="19" pageNumber="220">A. vittiger</emphasis>
</taxonomicName>
(males 
<quantity id="C27501A8FFC7256EFDCDFAD2FD23FA92" box="[590,715,1366,1393]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="19" pageNumber="220" unit="mm" value="23.5" valueMax="25.0" valueMin="22.0">22–25 mm</quantity>
, females 
<quantity id="C27501A8FFC7256EFCBAFAD2FC50FA92" box="[825,952,1366,1393]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="19" pageNumber="220" unit="mm" value="26.5" valueMax="28.0" valueMin="25.0">25–28 mm</quantity>
), and 
<taxonomicName id="C28DD7CEFFC7256EFC7DFAD3FBB5FA93" baseAuthorityName="Laurent" baseAuthorityYear="1941" box="[1022,1117,1367,1393]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="wittei">
<emphasis id="37F9705FFFC7256EFC7DFAD3FBB5FA93" box="[1022,1117,1367,1393]" italics="true" pageId="19" pageNumber="220">A. wittei</emphasis>
</taxonomicName>
(males 
<quantity id="C27501A8FFC7256EFB39FAD3FADFFA92" box="[1210,1335,1367,1393]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.0" metricValueMin="2.7" pageId="19" pageNumber="220" unit="mm" value="28.5" valueMax="30.0" valueMin="27.0">27–30 mm</quantity>
, females 
<quantity id="C27501A8FFC7256EFF14FAFEFEFFFA77" box="[151,279,1402,1428]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.3" metricValueMin="2.9" pageId="19" pageNumber="220" unit="mm" value="31.0" valueMax="33.0" valueMin="29.0">29–33 mm</quantity>
); and larger size than 
<taxonomicName id="C28DD7CEFFC7256EFD9BFAFFFD77FA77" authorityName="Pickersgill" authorityYear="1984" box="[536,671,1402,1429]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="delicatus">
<emphasis id="37F9705FFFC7256EFD9BFAFFFD77FA77" box="[536,671,1402,1429]" italics="true" pageId="19" pageNumber="220">A. delicatus</emphasis>
</taxonomicName>
(males 
<quantity id="C27501A8FFC7256EFD7EFAFEFC93FA76" box="[765,891,1402,1429]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="19" pageNumber="220" unit="mm" value="17.0" valueMax="19.0" valueMin="15.0">15–19 mm</quantity>
, females 
<quantity id="C27501A8FFC7256EFC69FAFEFB8EFA77" box="[1002,1126,1402,1428]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="19" pageNumber="220" unit="mm" value="19.5" valueMax="22.0" valueMin="17.0">17–22 mm</quantity>
) and 
<taxonomicName id="C28DD7CEFFC7256EFB29FAFFFAA0FA77" baseAuthorityName="Pfeffer" baseAuthorityYear="1893" box="[1194,1352,1402,1429]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="stuhlmanni">
<emphasis id="37F9705FFFC7256EFB29FAFFFAA0FA77" box="[1194,1352,1402,1429]" italics="true" pageId="19" pageNumber="220">A. stuhlmanni</emphasis>
</taxonomicName>
(males 
<quantity id="C27501A8FFC7256EFF14FA1AFEF9FA5A" box="[151,273,1438,1465]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.1" metricValueMin="1.5" pageId="19" pageNumber="220" unit="mm" value="18.0" valueMax="21.0" valueMin="15.0">15–21 mm</quantity>
, females 
<quantity id="C27501A8FFC7256EFEFAFA1AFE1DFA5B" box="[377,501,1438,1465]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="19" pageNumber="220" unit="mm" value="21.0" valueMax="25.0" valueMin="17.0">17–25 mm</quantity>
). Its differs in dorsal coloration and pattern from all species with longitudinal stripes or bands on a yellowish-brown background (
<taxonomicName id="C28DD7CEFFC7256EFD10FA46FCA7FA3F" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[659,847,1474,1500]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="brachycnemis">
<emphasis id="37F9705FFFC7256EFD10FA46FCA7FA3F" box="[659,847,1474,1500]" italics="true" pageId="19" pageNumber="220">A. brachycnemis</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFCDEFA46FC30FA3F" authorityName="Pickersgill" authorityYear="1984" box="[861,984,1474,1500]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="crotalus">
<emphasis id="37F9705FFFC7256EFCDEFA46FC30FA3F" box="[861,984,1474,1500]" italics="true" pageId="19" pageNumber="220">A. crotalus</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFC64FA46FB84FA3F" authorityName="Pickersgill" authorityYear="1984" box="[999,1132,1474,1500]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="delicatus">
<emphasis id="37F9705FFFC7256EFC64FA46FB84FA3F" box="[999,1132,1474,1500]" italics="true" pageId="19" pageNumber="220">A. delicatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFBF9FA46FB1CFA3F" baseAuthorityName="Peters" baseAuthorityYear="1875" box="[1146,1268,1474,1500]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="dorsalis">
<emphasis id="37F9705FFFC7256EFBF9FA46FB1CFA3F" box="[1146,1268,1474,1500]" italics="true" pageId="19" pageNumber="220">A. dorsalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFA87FA46FA7EFA3F" authorityName="Largen" authorityYear="1974" box="[1284,1430,1474,1500]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="enseticola">
<emphasis id="37F9705FFFC7256EFA87FA46FA7EFA3F" box="[1284,1430,1474,1500]" italics="true" pageId="19" pageNumber="220">A. enseticola</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFF14FA62FEF5F9E3" baseAuthorityName="Bianconi" baseAuthorityYear="1849" box="[151,285,1510,1536]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="fornasini">
<emphasis id="37F9705FFFC7256EFF14FA62FEF5F9E3" box="[151,285,1510,1536]" italics="true" pageId="19" pageNumber="220">A. fornasini</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFEAEFA62FE3FFA1C" baseAuthorityName="Cope" baseAuthorityYear="1861" box="[301,471,1509,1536]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="fulvovittatus">
<emphasis id="37F9705FFFC7256EFEAEFA62FE3FFA1C" box="[301,471,1509,1536]" italics="true" pageId="19" pageNumber="220">A. fulvovittatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFE64FA62FD8AFA1C" baseAuthorityName="Loveridge" baseAuthorityYear="1954" box="[487,610,1509,1536]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="knysnae">
<emphasis id="37F9705FFFC7256EFE64FA62FD8AFA1C" box="[487,610,1509,1536]" italics="true" pageId="19" pageNumber="220">A. knysnae</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFDF1FA62FD02FA1C" authorityName="Dubois" authorityYear="1986" box="[626,746,1510,1536]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="morerei">
<emphasis id="37F9705FFFC7256EFDF1FA62FD02FA1C" box="[626,746,1510,1536]" italics="true" pageId="19" pageNumber="220">A. morerei</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFD7AFA62FC6CFA1C" box="[761,900,1509,1536]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="orophilus">
<emphasis id="37F9705FFFC7256EFD7AFA62FC6CFA1C" box="[761,900,1509,1536]" italics="true" pageId="19" pageNumber="220">A. orophilus</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFC17FA62FBB9FA1C" baseAuthorityName="Werner" baseAuthorityYear="1908" box="[916,1105,1509,1536]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="quadrivittatus">
<emphasis id="37F9705FFFC7256EFC17FA62FBB9FA1C" box="[916,1105,1509,1536]" italics="true" pageId="19" pageNumber="220">A. quadrivittatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFBE2FA62FB1FFA1C" baseAuthorityName="Boettger" baseAuthorityYear="1889" box="[1121,1271,1509,1536]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="schneideri">
<emphasis id="37F9705FFFC7256EFBE2FA62FB1FFA1C" box="[1121,1271,1509,1536]" italics="true" pageId="19" pageNumber="220">A. schneideri</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFA85FA62FA7DF9E3" baseAuthorityName="Cope" baseAuthorityYear="1862" box="[1286,1429,1510,1536]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="spinifrons">
<emphasis id="37F9705FFFC7256EFA85FA62FA7DF9E3" box="[1286,1429,1510,1536]" italics="true" pageId="19" pageNumber="220">A. spinifrons</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFF14F98EFEDAF9C0" baseAuthorityName="Pfeffer" baseAuthorityYear="1893" box="[151,306,1545,1572]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="stuhlmanni">
<emphasis id="37F9705FFFC7256EFF14F98EFEDAF9C0" box="[151,306,1545,1572]" italics="true" pageId="19" pageNumber="220">A. stuhlmanni</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFEC2F98EFE2DF9C0" baseAuthorityName="Laurent" baseAuthorityYear="1941" box="[321,453,1545,1572]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="upembae">
<emphasis id="37F9705FFFC7256EFEC2F98EFE2DF9C0" box="[321,453,1545,1572]" italics="true" pageId="19" pageNumber="220">A. upembae</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFE50F98EFD70F9C0" authorityName="Schiotz" authorityYear="1974" box="[467,664,1545,1572]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="septentrionalis">
<emphasis id="37F9705FFFC7256EFE50F98EFD70F9C0" box="[467,664,1545,1572]" italics="true" pageId="19" pageNumber="220">A. septentrionalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFD25F98EFCFEF9C0" baseAuthorityName="Peters" baseAuthorityYear="1876" box="[678,790,1546,1572]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="vittiger">
<emphasis id="37F9705FFFC7256EFD25F98EFCFEF9C0" box="[678,790,1546,1572]" italics="true" pageId="19" pageNumber="220">A. vittiger</emphasis>
</taxonomicName>
, and 
<taxonomicName id="C28DD7CEFFC7256EFCD0F98EFC59F9C0" baseAuthorityName="Laurent" baseAuthorityYear="1941" box="[851,945,1546,1572]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="wittei">
<emphasis id="37F9705FFFC7256EFCD0F98EFC59F9C0" box="[851,945,1546,1572]" italics="true" pageId="19" pageNumber="220">A. wittei</emphasis>
</taxonomicName>
); species showing a conspicuous large dark dorsal blotch on a uniform background on anterior dorsum in combination with large blotches on both sides of the hip (
<taxonomicName id="C28DD7CEFFC7256EFF4FF9D5FE98F988" authorityName="Laurent" authorityYear="1950" box="[204,368,1617,1643]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="leucostictus">
<emphasis id="37F9705FFFC7256EFF4FF9D5FE98F988" box="[204,368,1617,1643]" italics="true" pageId="19" pageNumber="220">A. leucostictus</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFEFCF9D5FDA5F988" box="[383,589,1617,1643]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="manengubensis">
<emphasis id="37F9705FFFC7256EFEFCF9D5FDA5F988" box="[383,589,1617,1643]" italics="true" pageId="19" pageNumber="220">A. manengubensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFDD8F9D5FD22F988" baseAuthorityName="Ferreira" baseAuthorityYear="1906" box="[603,714,1617,1643]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="osorioi">
<emphasis id="37F9705FFFC7256EFDD8F9D5FD22F988" box="[603,714,1617,1643]" italics="true" pageId="19" pageNumber="220">A. osorioi</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFD5AF9D5FC61F988" box="[729,905,1617,1643]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="paradorsalis">
<emphasis id="37F9705FFFC7256EFD5AF9D5FC61F988" box="[729,905,1617,1643]" italics="true" pageId="19" pageNumber="220">A. paradorsalis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="C28DD7CEFFC7256EFC4BF9D5FBB5F988" baseAuthorityName="Boettger" baseAuthorityYear="1889" box="[968,1117,1617,1643]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="schneideri">
<emphasis id="37F9705FFFC7256EFC4BF9D5FBB5F988" box="[968,1117,1617,1643]" italics="true" pageId="19" pageNumber="220">A. schneideri</emphasis>
</taxonomicName>
) or with longitudinal lateral dark bands (
<taxonomicName id="C28DD7CEFFC7256EFEAAF9F1FE3BF96D" baseAuthorityName="Laurent" baseAuthorityYear="1941" box="[297,467,1652,1679]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="equatorialis">
<emphasis id="37F9705FFFC7256EFEAAF9F1FE3BF96D" box="[297,467,1652,1679]" italics="true" pageId="19" pageNumber="220">A. equatorialis</emphasis>
</taxonomicName>
and 
<taxonomicName id="C28DD7CEFFC7256EFD92F9F1FD47F96D" authorityName="Schiotz" authorityYear="1963" box="[529,687,1653,1679]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="nigeriensis">
<emphasis id="37F9705FFFC7256EFD92F9F1FD47F96D" box="[529,687,1653,1679]" italics="true" pageId="19" pageNumber="220">A. nigeriensis</emphasis>
</taxonomicName>
); species with a uniformly colored dorsum without conspicuous pattern except a dark dorsolateral stripe (
<taxonomicName id="C28DD7CEFFC7256EFDE0F91DFD25F951" authorityName="Pickersgill" authorityYear="1984" box="[611,717,1689,1715]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="aureus">
<emphasis id="37F9705FFFC7256EFDE0F91DFD25F951" box="[611,717,1689,1715]" italics="true" pageId="19" pageNumber="220">A. aureus</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFD5FF91DFCA1F951" authorityName="Largen" authorityYear="1974" box="[732,841,1688,1715]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="clarkei">
<emphasis id="37F9705FFFC7256EFD5FF91DFCA1F951" box="[732,841,1688,1715]" italics="true" pageId="19" pageNumber="220">A. clarkei</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFCD4F91DFC33F951" authorityName="Pickersgill" authorityYear="1984" box="[855,987,1688,1715]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="delicatus">
<emphasis id="37F9705FFFC7256EFCD4F91DFC33F951" box="[855,987,1688,1715]" italics="true" pageId="19" pageNumber="220">A. delicatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFC6AF91DFB86F951" baseAuthorityName="Bianconi" baseAuthorityYear="1849" box="[1001,1134,1688,1715]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="fornasini">
<emphasis id="37F9705FFFC7256EFC6AF91DFB86F951" box="[1001,1134,1688,1715]" italics="true" pageId="19" pageNumber="220">A. fornasini</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFBFFF91DFB02F951" box="[1148,1258,1688,1715]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="lacteus">
<emphasis id="37F9705FFFC7256EFBFFF91DFB02F951" box="[1148,1258,1688,1715]" italics="true" pageId="19" pageNumber="220">A. lacteus</emphasis>
</taxonomicName>
, 
<taxonomicName id="C28DD7CEFFC7256EFB7AF91DFA74F951" authorityName="Laurent" authorityYear="1950" box="[1273,1436,1688,1715]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="leucostictus">
<emphasis id="37F9705FFFC7256EFB7AF91DFA74F951" box="[1273,1436,1688,1715]" italics="true" pageId="19" pageNumber="220">A. leucostictus</emphasis>
</taxonomicName>
[dorsum with small white tubercles], 
<taxonomicName id="C28DD7CEFFC7256EFDCFF938FCE8F935" baseAuthorityName="Barbour &amp; Loveridge" baseAuthorityYear="1928" box="[588,768,1724,1750]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="uluguruensis">
<emphasis id="37F9705FFFC7256EFDCFF938FCE8F935" box="[588,768,1724,1750]" italics="true" pageId="19" pageNumber="220">A. uluguruensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="C28DD7CEFFC7256EFCC6F938FC3AF935" baseAuthorityName="Mertens" baseAuthorityYear="1938" box="[837,978,1724,1750]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="weidholzi">
<emphasis id="37F9705FFFC7256EFCC6F938FC3AF935" box="[837,978,1724,1750]" italics="true" pageId="19" pageNumber="220">A. weidholzi</emphasis>
</taxonomicName>
[might have a black vertebral line and dark brown flanks]); and from all remaining species having either dark stripes running from tip of snout to back, crossing each other and continuing as dorsolateral stripes (
<taxonomicName id="C28DD7CEFFC7256EFCB8F880FC38F8FE" authorityName="Schiotz" authorityYear="1967" box="[827,976,1795,1822]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="vibekensis">
<emphasis id="37F9705FFFC7256EFCB8F880FC38F8FE" box="[827,976,1795,1822]" italics="true" pageId="19" pageNumber="220">A. vibekensis</emphasis>
</taxonomicName>
, also males without asperities); anterior part of dorsum yellowish with brown pattern, posterior part of dorsum translucent (
<taxonomicName id="C28DD7CEFFC7256EFBD7F8ACFB5EF8A2" authorityName="Ahl" authorityYear="1930" box="[1108,1206,1831,1858]" class="Amphibia" family="Hyperoliidae" genus="Megalixalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="laevis">
<emphasis id="37F9705FFFC7256EFBD7F8ACFB5EF8A2" box="[1108,1206,1831,1858]" italics="true" pageId="19" pageNumber="220">A. laevis</emphasis>
</taxonomicName>
, also males without asperities); posterior part of dorsum and dorsal sides of limbs semi-translucent with dense small dark white-edged speckles (
<taxonomicName id="C28DD7CEFFC7256EFE85F8EBFE39F86A" baseAuthorityName="Ahl" baseAuthorityYear="1930" box="[262,465,1903,1929]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="dorsimaculatus">
<emphasis id="37F9705FFFC7256EFE85F8EBFE39F86A" box="[262,465,1903,1929]" italics="true" pageId="19" pageNumber="220">A. dorsimaculatus</emphasis>
</taxonomicName>
); and dark blotches and transverse marks with white speckles (
<taxonomicName id="C28DD7CEFFC7256EFB12F8EBFAF6F86A" authorityName="Schiotz" authorityYear="1974" box="[1169,1310,1903,1929]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="sylvaticus">
<emphasis id="37F9705FFFC7256EFB12F8EBFAF6F86A" box="[1169,1310,1903,1929]" italics="true" pageId="19" pageNumber="220">A. sylvaticus</emphasis>
</taxonomicName>
). It is most similar to 
<taxonomicName id="C28DD7CEFFC7256EFE8EF817FE41F84F" authority="Greenbaum &amp; Portik &amp; Allen &amp; Vaughan &amp; Badjedjea &amp; Barej &amp; Behangana &amp; Conkey &amp; Dumbo &amp; Gonwouo &amp; Hirschfeld &amp; Hughes &amp; Igunzi &amp; Kusamba &amp; Lukwago &amp; Masudi &amp; Penner &amp; Reyes &amp; Rödel &amp; Roelke &amp; Romero &amp; Dehling, 2022" authorityName="Greenbaum &amp; Portik &amp; Allen &amp; Vaughan &amp; Badjedjea &amp; Barej &amp; Behangana &amp; Conkey &amp; Dumbo &amp; Gonwouo &amp; Hirschfeld &amp; Hughes &amp; Igunzi &amp; Kusamba &amp; Lukwago &amp; Masudi &amp; Penner &amp; Reyes &amp; Rödel &amp; Roelke &amp; Romero &amp; Dehling" authorityYear="2022" box="[269,425,1938,1965]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="phantasma" status="sp. nov.">
<emphasis id="37F9705FFFC7256EFE8EF817FE41F84F" box="[269,425,1938,1965]" italics="true" pageId="19" pageNumber="220">A. phantasma</emphasis>
</taxonomicName>
<emphasis id="37F9705FFFC7256EFE31F816FDE5F84F" bold="true" box="[434,525,1938,1964]" pageId="19" pageNumber="220">
<taxonomicNameLabel id="2CCACD24FFC7256EFE31F816FDE5F84F" box="[434,525,1938,1964]" pageId="19" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but differs from this species in being smaller (Table 1) and showing an overall darker coloration and more contrasting dorsal pattern that is obvious even in preserved specimens (
<figureCitation id="9DB6B0C8FFC7256EFB60F832FAA5F833" box="[1251,1357,1974,2000]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="13.[151,250,1222,1246]" captionStartId-1="15.[151,250,1585,1609]" captionTargetBox-0="[151,1436,345,1199]" captionTargetBox-1="[151,1436,705,1562]" captionTargetId-0="figure-95@13.[151,1436,345,1199]" captionTargetId-1="figure-276@15.[151,1436,705,1562]" captionTargetPageId-0="13" captionTargetPageId-1="15" captionText-0="FIGURE 7. Photos of Afrixalus phantasma sp. nov. in life. Adult male holotype, ZFMK 103454 (field no. JMD 723), from Gishwati-Mukura National Park, Rwanda, in the habitat showing nighttime coloration (A), adult male paratype, ZFMK 103460 (field no. JMD 679) from Kamiranzovu Swamp, Nyungwe National Park, showing daytime coloration in dorsal (B) and ventral views (C), adult male paratype UTEP 20802 (EBG 1198) from Kahuzi-Biega National Park, DRC (D), photographed the morning after capture." captionText-1="FIGURE 8. Photos of the preserved holotype of Afrixalus phantasma sp. nov. (ZFMK 103454) in dorsal (A) and ventral (B) views (to scale), and plantar view of the left foot (C). Photos of the preserved holotype of A. lacustris sp. nov. (UTEP 20805) in dorsal (D) and ventral (E) views (to scale), and plantar view of the left foot (F)." figureDoi-0="http://doi.org/10.5281/zenodo.6986895" figureDoi-1="http://doi.org/10.5281/zenodo.6986899" httpUri-0="https://zenodo.org/record/6986895/files/figure.png" httpUri-1="https://zenodo.org/record/6986899/files/figure.png" pageId="19" pageNumber="220">Figs. 7–8</figureCitation>
, 
<figureCitation id="9DB6B0C8FFC7256EFADAF832FA90F833" box="[1369,1400,1974,2000]" captionStart="FIGURE 12" captionStartId="20.[151,250,1727,1751]" captionTargetBox="[151,1436,181,1704]" captionTargetId="figure-23@20.[151,1436,181,1704]" captionTargetPageId="20" captionText="FIGURE 12. Photos of Afrixalus lacustris sp. nov. in life. Adult male holotype (UTEP 20805), from the vicinity of Kalundu (03.15552° S, 28.42108° E, 1482 m), South Kivu Province, DRC in dorsal (A) and (B) ventral views, adult female (UTEP 20809) from Baraka, near shore of Lake Tanganyika, South Kivu Province, DRC (C), adult female (EPLU 395) from Epulu, Ituri Province, DRC (D), adult male (UTEP 22417) from Toyokana, Ituri Province, DRC (E), individual of unknown sex (DFH 1102) from Ngogo Research Center, Kibale Forest National Park, Western Region, Uganda (F). All photos taken the morning after capture." figureDoi="http://doi.org/10.5281/zenodo.6986240" httpUri="https://zenodo.org/record/6986240/files/figure.png" pageId="19" pageNumber="220">12</figureCitation>
). It is readily distinguished from 
<taxonomicName id="C28DD7CEFFC7256EFE5CF85EFD91F817" authority="Greenbaum &amp; Portik &amp; Allen &amp; Vaughan &amp; Badjedjea &amp; Barej &amp; Behangana &amp; Conkey &amp; Dumbo &amp; Gonwouo &amp; Hirschfeld &amp; Hughes &amp; Igunzi &amp; Kusamba &amp; Lukwago &amp; Masudi &amp; Penner &amp; Reyes &amp; Rödel &amp; Roelke &amp; Romero &amp; Dehling, 2022" authorityName="Greenbaum &amp; Portik &amp; Allen &amp; Vaughan &amp; Badjedjea &amp; Barej &amp; Behangana &amp; Conkey &amp; Dumbo &amp; Gonwouo &amp; Hirschfeld &amp; Hughes &amp; Igunzi &amp; Kusamba &amp; Lukwago &amp; Masudi &amp; Penner &amp; Reyes &amp; Rödel &amp; Roelke &amp; Romero &amp; Dehling" authorityYear="2022" box="[479,633,2010,2036]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="19" pageNumber="220" phylum="Chordata" rank="species" species="phantasma" status="sp. nov.">
<emphasis id="37F9705FFFC7256EFE5CF85EFD91F817" box="[479,633,2010,2036]" italics="true" pageId="19" pageNumber="220">A. phantasma</emphasis>
</taxonomicName>
<emphasis id="37F9705FFFC7256EFD03F85EFD32F817" bold="true" box="[640,730,2010,2036]" pageId="19" pageNumber="220">
<taxonomicNameLabel id="2CCACD24FFC7256EFD03F85EFD32F817" box="[640,730,2010,2036]" pageId="19" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the more extensive hand and toe webbing (
<figureCitation id="9DB6B0C8FFC7256EFB67F85EFAB1F817" box="[1252,1369,2010,2036]" captionStart="FIGURE 8" captionStartId="15.[151,250,1585,1609]" captionTargetBox="[151,1436,705,1562]" captionTargetId="figure-276@15.[151,1436,705,1562]" captionTargetPageId="15" captionText="FIGURE 8. Photos of the preserved holotype of Afrixalus phantasma sp. nov. (ZFMK 103454) in dorsal (A) and ventral (B) views (to scale), and plantar view of the left foot (C). Photos of the preserved holotype of A. lacustris sp. nov. (UTEP 20805) in dorsal (D) and ventral (E) views (to scale), and plantar view of the left foot (F)." figureDoi="http://doi.org/10.5281/zenodo.6986899" httpUri="https://zenodo.org/record/6986899/files/figure.png" pageId="19" pageNumber="220">Fig. 8E–F</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="51F2FCC5FFC02569FF14F93BFEF6F868" ID-DOI="http://doi.org/10.5281/zenodo.6986240" ID-Zenodo-Dep="6986240" httpUri="https://zenodo.org/record/6986240/files/figure.png" pageId="20" pageNumber="221" startId="20.[151,250,1727,1751]" targetBox="[151,1436,181,1704]" targetPageId="20">
<paragraph id="0532AC4DFFC02569FF14F93BFEF6F868" blockId="20.[151,1436,1727,1932]" pageId="20" pageNumber="221">
<emphasis id="37F9705FFFC02569FF14F93BFEC8F934" bold="true" box="[151,288,1727,1751]" pageId="20" pageNumber="221">FIGURE 12.</emphasis>
Photos of 
<emphasis id="37F9705FFFC02569FE13F93BFD42F93B" bold="true" box="[400,682,1727,1752]" pageId="20" pageNumber="221">
<taxonomicName id="C28DD7CEFFC02569FE13F93BFDBAF934" authority="Greenbaum &amp; Portik &amp; Allen &amp; Vaughan &amp; Badjedjea &amp; Barej &amp; Behangana &amp; Conkey &amp; Dumbo &amp; Gonwouo &amp; Hirschfeld &amp; Hughes &amp; Igunzi &amp; Kusamba &amp; Lukwago &amp; Masudi &amp; Penner &amp; Reyes &amp; Rödel &amp; Roelke &amp; Romero &amp; Dehling, 2022" authorityName="Greenbaum &amp; Portik &amp; Allen &amp; Vaughan &amp; Badjedjea &amp; Barej &amp; Behangana &amp; Conkey &amp; Dumbo &amp; Gonwouo &amp; Hirschfeld &amp; Hughes &amp; Igunzi &amp; Kusamba &amp; Lukwago &amp; Masudi &amp; Penner &amp; Reyes &amp; Rödel &amp; Roelke &amp; Romero &amp; Dehling" authorityYear="2022" box="[400,594,1727,1751]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="20" pageNumber="221" phylum="Chordata" rank="species" species="lacustris" status="sp. nov.">
<emphasis id="37F9705FFFC02569FE13F93BFDBAF934" bold="true" box="[400,594,1727,1751]" italics="true" pageId="20" pageNumber="221">Afrixalus lacustris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2CCACD24FFC02569FDDAF944FD42F93B" box="[601,682,1728,1752]" pageId="20" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life. Adult male holotype (UTEP 20805), from the vicinity of Kalundu (03.15552° S, 28.42108° E, 1482 m), South Kivu Province, DRC in dorsal (A) and (B) ventral views, adult female (UTEP 20809) from Baraka, near shore of Lake Tanganyika, South Kivu Province, DRC (C), adult female (EPLU 395) from Epulu, Ituri Province, DRC (D), adult male (UTEP 22417) from Toyokana, Ituri Province, DRC (E), individual of unknown sex (DFH 1102) from Ngogo Research Center, Kibale Forest National Park, Western Region, Uganda (F). All photos taken the morning after capture.
</paragraph>
</caption>
<subSubSection id="4D97FFC6FFC12568FF44FF12FBFAF892" pageId="21" pageNumber="222" type="description">
<paragraph id="0532AC4DFFC12568FF44FF12FC49FD82" blockId="21.[151,1437,150,2013]" pageId="21" pageNumber="222">
<emphasis id="37F9705FFFC12568FF44FF12FE01FF52" bold="true" box="[199,489,150,177]" pageId="21" pageNumber="222">
Description of 
<typeStatus id="DA3612EFFFC12568FEF8FF13FE0CFF52" box="[379,484,151,177]" pageId="21" pageNumber="222" type="holotype">holotype</typeStatus>
.
</emphasis>
Measurements of the 
<typeStatus id="DA3612EFFFC12568FD68FF13FCA5FF52" box="[747,845,151,177]" pageId="21" pageNumber="222" type="holotype">holotype</typeStatus>
are provided in 
<tableCitation id="480F99F6FFC12568FB8EFF13FB8DFF52" box="[1037,1125,151,177]" captionStart="TABLE 2" captionStartId="14.[151,239,1628,1652]" captionTargetPageId="14" captionText="TABLE 2. Measurements (mm) of the adult male holotypes of Afrixalus phantasma sp. nov. and A. lacustris sp. nov." httpUri="http://table.plazi.org/id/51F2FCC5FFDA2573FF14F9D8FA63F996" pageId="21" pageNumber="222" tableUuid="51F2FCC5FFDA2573FF14F9D8FA63F996">Table 2</tableCitation>
. Body very slender, widest at temporal region, slightly tapering to groin (
<figureCitation id="9DB6B0C8FFC12568FD2DFF3FFCC1FF36" box="[686,809,187,213]" captionStart="FIGURE 8" captionStartId="15.[151,250,1585,1609]" captionTargetBox="[151,1436,705,1562]" captionTargetId="figure-276@15.[151,1436,705,1562]" captionTargetPageId="15" captionText="FIGURE 8. Photos of the preserved holotype of Afrixalus phantasma sp. nov. (ZFMK 103454) in dorsal (A) and ventral (B) views (to scale), and plantar view of the left foot (C). Photos of the preserved holotype of A. lacustris sp. nov. (UTEP 20805) in dorsal (D) and ventral (E) views (to scale), and plantar view of the left foot (F)." figureDoi="http://doi.org/10.5281/zenodo.6986899" httpUri="https://zenodo.org/record/6986899/files/figure.png" pageId="21" pageNumber="222">Fig. 8D–F</figureCitation>
); head small (HL/SVL 0.31, HW/SVL 0.31), about as long as wide (HW/HL 0.99); snout relatively long, rounded in dorsal view and in lateral profile, slightly wider than long; canthus rostralis hardly discernible, slightly concave between eye and nostril in dorsal view; loreal region oblique; nostrils rounded, directed anterolaterally and slightly dorsally; situated much closer to tip of snout and to eye, separated from each other by distance about equal to distance between eye and nostril (IN/EN 1.02); eye directed anterolaterally, strongly protruding, very large (ED/HL 0.39), its diameter shorter than snout (ED/SL 0.90); interorbital distance much wider than upper eyelid (IO/EW 1.61) and wider than internarial distance (IO/IN 1.31); tympanum covered by skin, not visible externally; upper jaw with dentition; teeth on premaxilla larger than teeth on maxilla; choanae small, located far anterolaterally at margins of roof of mouth, its anterior edge covered by maxillary bone, therefore appearing semicircular in ventral view; vomer ridges and teeth absent; tongue heartshaped, longer than broad, bilobed for about one-fourth of its length, free distally for about two-thirds its length; densely covered with minute papillae; median lingual process absent.
</paragraph>
<paragraph id="0532AC4DFFC12568FF44FDEFFC52FCF6" blockId="21.[151,1437,150,2013]" pageId="21" pageNumber="222">
Dorsal surfaces of head, trunk, and limbs and lateral surfaces of trunk finely shagreened; tiny pointed, widely scattered tubercles on dorsum, more numerous posteriorly than anteriorly; supratympanic fold absent; few tubercles at rear end of mandible without pointed tips; ventral side of head smooth; vocal sac present; gular gland of vocal sac smooth, large and wide, covering about 70 percent of throat width (
<figureCitation id="9DB6B0C8FFC12568FC4AFD53FBCAFD12" box="[969,1058,727,753]" captionStart="FIGURE 8" captionStartId="15.[151,250,1585,1609]" captionTargetBox="[151,1436,705,1562]" captionTargetId="figure-276@15.[151,1436,705,1562]" captionTargetPageId="15" captionText="FIGURE 8. Photos of the preserved holotype of Afrixalus phantasma sp. nov. (ZFMK 103454) in dorsal (A) and ventral (B) views (to scale), and plantar view of the left foot (C). Photos of the preserved holotype of A. lacustris sp. nov. (UTEP 20805) in dorsal (D) and ventral (E) views (to scale), and plantar view of the left foot (F)." figureDoi="http://doi.org/10.5281/zenodo.6986899" httpUri="https://zenodo.org/record/6986899/files/figure.png" pageId="21" pageNumber="222">Fig. 8E</figureCitation>
); chest smooth, abdomen weakly areolate; ventral side of limbs smooth; short transverse fold above vent.
</paragraph>
<paragraph id="0532AC4DFFC12568FF44FC9BFB29FC0E" blockId="21.[151,1437,150,2013]" pageId="21" pageNumber="222">
Forelimbs slender; hand large (HaL/SVL 0.28); tips of fingers enlarged into large disks, each with circummarginal groove; relative length of fingers: I &lt;II &lt;IV &lt;III; subarticular tubercles well developed, numbering one on Fingers I and II, two on Fingers III and IV, proximal tubercles on Fingers III and IV and tubercles on Fingers I and II singular, rounded; distal tubercles on Fingers III and IV indistinctly bipartite; webbing formula of the hand 
<emphasis id="37F9705FFFC12568FADFFC0FFA8FFC46" bold="true" box="[1372,1383,907,933]" pageId="21" pageNumber="222">I</emphasis>
2-/2 
<emphasis id="37F9705FFFC12568FF14FC2BFF45FC2A" bold="true" box="[151,173,943,969]" pageId="21" pageNumber="222">II</emphasis>
2-/3- 
<emphasis id="37F9705FFFC12568FF71FC2BFEFBFC2A" bold="true" box="[242,275,943,969]" pageId="21" pageNumber="222">III</emphasis>
2.25/2 
<emphasis id="37F9705FFFC12568FEEAFC2BFE60FC2A" bold="true" box="[361,392,943,969]" pageId="21" pageNumber="222">IV</emphasis>
; thenar tubercle oval, small and very low, about one-fourth length of metacarpal of Finger I; inner palmar tubercle and outer palmar tubercle very low, small, rounded, almost indiscernible.
</paragraph>
<paragraph id="0532AC4DFFC12568FF44FC73FBC1FB0A" blockId="21.[151,1437,150,2013]" pageId="21" pageNumber="222">
Hind limbs slender, moderately long; heel reaching to level of eye when legs adpressed forwardly to body; crus moderately long (TibL/SVL 0.44), slightly shorter than thigh; heels not touching each other when knees flexed and thighs held perpendicularly to median plane; foot subequal in length to crus (FoL/TibL 0.94); relative length of toes: I &lt;II &lt;III &lt;V &lt;IV; toe tips rounded, enlarged into large disks, each with circummarginal groove; subarticular tubercles singular, numbering one on Toes I and II, two on Toes III and V, and three on Toe IV; pedal webbing formula 
<emphasis id="37F9705FFFC12568FF76FB2FFEE8FB26" bold="true" box="[245,256,1195,1221]" pageId="21" pageNumber="222">I</emphasis>
1.5/2+ 
<emphasis id="37F9705FFFC12568FED7FB2FFE82FB26" bold="true" box="[340,362,1195,1221]" pageId="21" pageNumber="222">II</emphasis>
1+/2+ 
<emphasis id="37F9705FFFC12568FE3BFB2FFE31FB26" bold="true" box="[440,473,1195,1221]" pageId="21" pageNumber="222">III</emphasis>
1+/2+ 
<emphasis id="37F9705FFFC12568FDA4FB2FFDAEFB26" bold="true" box="[551,582,1195,1221]" pageId="21" pageNumber="222">IV</emphasis>
2/1+ 
<emphasis id="37F9705FFFC12568FD07FB2FFD70FB26" bold="true" box="[644,664,1195,1221]" pageId="21" pageNumber="222">V</emphasis>
(
<figureCitation id="9DB6B0C8FFC12568FD25FB2FFD13FB26" box="[678,763,1195,1221]" captionStart="FIGURE 8" captionStartId="15.[151,250,1585,1609]" captionTargetBox="[151,1436,705,1562]" captionTargetId="figure-276@15.[151,1436,705,1562]" captionTargetPageId="15" captionText="FIGURE 8. Photos of the preserved holotype of Afrixalus phantasma sp. nov. (ZFMK 103454) in dorsal (A) and ventral (B) views (to scale), and plantar view of the left foot (C). Photos of the preserved holotype of A. lacustris sp. nov. (UTEP 20805) in dorsal (D) and ventral (E) views (to scale), and plantar view of the left foot (F)." figureDoi="http://doi.org/10.5281/zenodo.6986899" httpUri="https://zenodo.org/record/6986899/files/figure.png" pageId="21" pageNumber="222">Fig. 8E</figureCitation>
); inner metatarsal tubercle moderately prominent, elongated, about half length of metatarsus of Toe I; outer metatarsal tubercle small, rounded.
</paragraph>
<paragraph id="0532AC4DFFC12568FF44FB76FC56F9B2" blockId="21.[151,1437,150,2013]" pageId="21" pageNumber="222">
<emphasis id="37F9705FFFC12568FF44FB76FE71FAEF" bold="true" box="[199,409,1266,1292]" pageId="21" pageNumber="222">Coloration in life.</emphasis>
During night, skin on dorsal side of head, trunk, forelimbs, crus and tarsus light brown with more or less regular, locally more densely clustered dark brown speckling; distinct dark brown transverse stripe between upper eyelids, another less distinct one in scapula region, and another narrow one at level of anterior end of pelvis; dorsal skin of thigh largely unpigmented with only a broad stripe of widely scattered brown pigmentation; distinct dark brown band along canthus, continuing behind eye on both sides of trunk to level of pelvis; tubercles below eye and in tympanic region at rear end of jaw white; dark brown stripe running diagonally on middle of tibia, forming interrupted, inverted U-shaped band together with pelvic spots when legs folded against body; skin of ventral side of head, trunk and limbs largely unpigmented and translucent; gular gland bright yellow; fingers and toes yellow (
<figureCitation id="9DB6B0C8FFC12568FEA5F997FE90F9CE" box="[294,376,1555,1581]" captionStart="FIGURE 12" captionStartId="20.[151,250,1727,1751]" captionTargetBox="[151,1436,181,1704]" captionTargetId="figure-23@20.[151,1436,181,1704]" captionTargetPageId="20" captionText="FIGURE 12. Photos of Afrixalus lacustris sp. nov. in life. Adult male holotype (UTEP 20805), from the vicinity of Kalundu (03.15552° S, 28.42108° E, 1482 m), South Kivu Province, DRC in dorsal (A) and (B) ventral views, adult female (UTEP 20809) from Baraka, near shore of Lake Tanganyika, South Kivu Province, DRC (C), adult female (EPLU 395) from Epulu, Ituri Province, DRC (D), adult male (UTEP 22417) from Toyokana, Ituri Province, DRC (E), individual of unknown sex (DFH 1102) from Ngogo Research Center, Kibale Forest National Park, Western Region, Uganda (F). All photos taken the morning after capture." figureDoi="http://doi.org/10.5281/zenodo.6986240" httpUri="https://zenodo.org/record/6986240/files/figure.png" pageId="21" pageNumber="222">Fig. 12</figureCitation>
). During day, basic dorsal coloration light brown; dark brown dorsal speckles, spots and stripes in more pronounced contrast to basic coloration, more distinctly visible.
</paragraph>
<paragraph id="0532AC4DFFC12568FF44F9DEFB42F97A" blockId="21.[151,1437,150,2013]" pageId="21" pageNumber="222">
<emphasis id="37F9705FFFC12568FF44F9DEFDEDF997" bold="true" box="[199,517,1626,1652]" pageId="21" pageNumber="222">Coloration in preservative.</emphasis>
Dorsal basic coloration largely faded to light brown; darker dorsal pattern elements dark brown, clearly visible; color of gular gland, fingers and toes faded to white (
<figureCitation id="9DB6B0C8FFC12568FBA5F9FBFB76F97A" box="[1062,1182,1663,1689]" captionStart="FIGURE 8" captionStartId="15.[151,250,1585,1609]" captionTargetBox="[151,1436,705,1562]" captionTargetId="figure-276@15.[151,1436,705,1562]" captionTargetPageId="15" captionText="FIGURE 8. Photos of the preserved holotype of Afrixalus phantasma sp. nov. (ZFMK 103454) in dorsal (A) and ventral (B) views (to scale), and plantar view of the left foot (C). Photos of the preserved holotype of A. lacustris sp. nov. (UTEP 20805) in dorsal (D) and ventral (E) views (to scale), and plantar view of the left foot (F)." figureDoi="http://doi.org/10.5281/zenodo.6986899" httpUri="https://zenodo.org/record/6986899/files/figure.png" pageId="21" pageNumber="222">Fig. 8D–F</figureCitation>
).
</paragraph>
<paragraph id="0532AC4DFFC12568FF44F926FC7CF8AE" blockId="21.[151,1437,150,2013]" pageId="21" pageNumber="222">
<emphasis id="37F9705FFFC12568FF44F926FED7F95F" bold="true" box="[199,319,1698,1724]" pageId="21" pageNumber="222">Variation.</emphasis>
The 
<typeStatus id="DA3612EFFFC12568FEF0F927FE36F95E" box="[371,478,1699,1725]" pageId="21" pageNumber="222" type="paratype">paratypes</typeStatus>
match the 
<typeStatus id="DA3612EFFFC12568FDD0F927FD5DF95E" box="[595,693,1699,1725]" pageId="21" pageNumber="222" type="holotype">holotype</typeStatus>
in general appearance and proportions. Coloration and color pattern varies from light-cream to yellowish-orange with a few relatively large and darker blotches and vermiculation, light speckling and indistinct dorsolateral stripe to an almost reticulated pattern of contrasting bright and dark elements with a broad and conspicuous brown dorsolateral band (
<figureCitation id="9DB6B0C8FFC12568FC8CF88BFC8BF8CA" box="[783,867,1807,1833]" captionStart="FIGURE 12" captionStartId="20.[151,250,1727,1751]" captionTargetBox="[151,1436,181,1704]" captionTargetId="figure-23@20.[151,1436,181,1704]" captionTargetPageId="20" captionText="FIGURE 12. Photos of Afrixalus lacustris sp. nov. in life. Adult male holotype (UTEP 20805), from the vicinity of Kalundu (03.15552° S, 28.42108° E, 1482 m), South Kivu Province, DRC in dorsal (A) and (B) ventral views, adult female (UTEP 20809) from Baraka, near shore of Lake Tanganyika, South Kivu Province, DRC (C), adult female (EPLU 395) from Epulu, Ituri Province, DRC (D), adult male (UTEP 22417) from Toyokana, Ituri Province, DRC (E), individual of unknown sex (DFH 1102) from Ngogo Research Center, Kibale Forest National Park, Western Region, Uganda (F). All photos taken the morning after capture." figureDoi="http://doi.org/10.5281/zenodo.6986240" httpUri="https://zenodo.org/record/6986240/files/figure.png" pageId="21" pageNumber="222">Fig. 12</figureCitation>
). Pedal webbing variation is 
<emphasis id="37F9705FFFC12568FB28F88BFB5EF8CA" bold="true" box="[1195,1206,1807,1833]" pageId="21" pageNumber="222">I</emphasis>
1.5[100]/2+[100] 
<emphasis id="37F9705FFFC12568FA05F88BFA74F8CA" bold="true" box="[1414,1436,1807,1833]" pageId="21" pageNumber="222">II</emphasis>
1+[100]/2+[100] 
<materialsCitation id="B5E5A610FFC12568FED8F8B7FE64F8AE" ID-GBIF-Occurrence="3887776314" box="[347,396,1842,1869]" collectionCode="III" pageId="21" pageNumber="222" specimenCode="III 1" specimenCount="1">
<specimenCode id="552B0436FFC12568FED8F8B7FE64F8AE" box="[347,396,1842,1869]" collectionCode="III" pageId="21" pageNumber="222">
<emphasis id="37F9705FFFC12568FED8F8B7FE94F8AE" bold="true" box="[347,380,1843,1869]" pageId="21" pageNumber="222">III</emphasis>
1
</specimenCode>
</materialsCitation>
+[50],1.25[50]/2+[100] 
<materialsCitation id="B5E5A610FFC12568FD1CF8B7FD3CF8AE" ID-GBIF-Occurrence="3887776307" box="[671,724,1843,1869]" collectionCode="IV" pageId="21" pageNumber="222" specimenCode="IV 2" specimenCount="1">
<specimenCode id="552B0436FFC12568FD1CF8B7FD3CF8AE" box="[671,724,1843,1869]" collectionCode="IV" pageId="21" pageNumber="222">
<emphasis id="37F9705FFFC12568FD1CF8B7FD56F8AE" bold="true" box="[671,702,1843,1869]" pageId="21" pageNumber="222">IV</emphasis>
2
</specimenCode>
</materialsCitation>
[100]/1+[100] 
<emphasis id="37F9705FFFC12568FCFAF8B7FC65F8AE" bold="true" box="[889,909,1843,1869]" pageId="21" pageNumber="222">V</emphasis>
.
</paragraph>
<paragraph id="0532AC4DFFC12568FF44F8D2FBFAF892" blockId="21.[151,1437,150,2013]" box="[199,1042,1878,1905]" pageId="21" pageNumber="222">
<emphasis id="37F9705FFFC12568FF44F8D2FE8AF893" bold="true" box="[199,354,1878,1904]" pageId="21" pageNumber="222">Bioacoustics.</emphasis>
The advertisement call and other vocalizations are unknown.
</paragraph>
</subSubSection>
<subSubSection id="4D97FFC6FFC1256BFF44F8FFFB25FDFA" lastPageId="22" lastPageNumber="223" pageId="21" pageNumber="222" type="biology_ecology">
<paragraph id="0532AC4DFFC1256BFF44F8FFFB25FDFA" blockId="21.[151,1437,150,2013]" lastBlockId="22.[151,1437,150,1185]" lastPageId="22" lastPageNumber="223" pageId="21" pageNumber="222">
<emphasis id="37F9705FFFC12568FF44F8FFFDF7F877" bold="true" box="[199,543,1914,1941]" pageId="21" pageNumber="222">Ecology and natural history.</emphasis>
We collected males and females in swamps in forest openings and near forest edges (
<figureCitation id="9DB6B0C8FFC12568FF6BF81BFEA5F85A" box="[232,333,1951,1977]" captionStart="FIGURE 11" captionStartId="18.[151,250,1923,1947]" captionTargetBox="[179,1408,181,1898]" captionTargetId="figure-23@18.[179,1408,181,1899]" captionTargetPageId="18" captionText="FIGURE 11. Photos of the habitat of (A) Afrixalus phantasma sp. nov. in Kamiranzovu Swamp, Nyungwe National Park, Rwanda, showing a forest edge in a montane marsh, and (B) A. lacustris sp. nov. from the vicinity of Kalundu, DRC, showing transitional forest." figureDoi="http://doi.org/10.5281/zenodo.6986232" httpUri="https://zenodo.org/record/6986232/files/figure.png" pageId="21" pageNumber="222">Fig. 11B</figureCitation>
), from vegetation at the edge of streams (or on vegetation overhanging them) in forest, and from non-forest vegetation near the shore of Lake Tanganyika. Egg deposition has not been observed and tadpoles are unknown. An adult male (CAS 256035) was found in Bwindi Impenetrable National Park (
<collectingCountry id="7D9AECDDFFC2256BFB15FF13FB18FF52" box="[1174,1264,151,177]" name="Uganda" pageId="22" pageNumber="223">Uganda</collectingCountry>
) ca. 
<quantity id="C27501A8FFC2256BFAA4FF13FABAFF52" box="[1319,1362,151,177]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="223" unit="m" value="2.0">2 m</quantity>
above ground in a tree fern “either calling or moving to a calling site (similar to ‘running’ of 
<taxonomicName id="C28DD7CEFFC2256BFBE2FF3FFB53FF37" authorityName="Girard" authorityYear="1853" box="[1121,1211,187,212]" class="Amphibia" family="Hyperoliidae" genus="Kassina" kingdom="Animalia" order="Anura" pageId="22" pageNumber="223" phylum="Chordata" rank="genus">
<emphasis id="37F9705FFFC2256BFBE2FF3FFB53FF37" box="[1121,1211,187,212]" italics="true" pageId="22" pageNumber="223">Kassina</emphasis>
</taxonomicName>
)” (D. Blackburn, in litt., 
<date id="71338A8DFFC2256BFF48FF5BFEB6FF1A" box="[203,350,223,249]" pageId="22" pageNumber="223" value="2021-07-22">22 July 2021</date>
). 
<bibRefCitation id="611CD1BCFFC2256BFEF2FF5BFDF1FF1A" author="Laurent, R. F." box="[369,537,222,249]" pageId="22" pageNumber="223" pagination="290 - 310" refId="ref20448" refString="Laurent, R. F. (1955) Apercu de la biogeographie des batraciens et des reptiles de la region des grand lacs. Bulletin de la Societe Zoologique de France, 79, 290 - 310." type="journal article" year="1955">Laurent (1955)</bibRefCitation>
noted the species from transitional forest. In his paper on amphibians of Virunga National Park, 
<bibRefCitation id="611CD1BCFFC2256BFEBCFE87FE00FEFE" author="Laurent, R. F." box="[319,488,259,285]" pageId="22" pageNumber="223" pagination="1 - 125" refId="ref20584" refString="Laurent, R. F. (1972) Amphibiens. Exploration du Parc National des Virunga, Deuxieme Serie, 22, 1 - 125, 11 pls." type="journal article" year="1972">Laurent (1972)</bibRefCitation>
noted the species is most commonly found in swamps with herbaceous vegetation in secondary forest, and more rarely, in primary forest. 
<bibRefCitation id="611CD1BCFFC2256BFD6EFEA3FC7EFEA2" author="Laurent, R. F." box="[749,918,295,321]" pageId="22" pageNumber="223" pagination="1 - 93" refId="ref20614" refString="Laurent, R. F. (1982) Le genre Afrixalus Laurent (Hyperoliidae) en Afrique Centrale. Annales du Musee Royal de l'Afrique Centrale, Serie in Octavo, Science Zoologique, 235, 1 - 93." type="journal article" year="1982">Laurent (1982)</bibRefCitation>
noted that, in general, the species is common in the foliage of shrubs in dense forests. He also recorded it from “une mare sombre” (i.e., a dark pond) in syntopy with 
<taxonomicName id="C28DD7CEFFC2256BFF52FEEBFEB4FE6A" authorityName="Peters" authorityYear="1854" box="[209,348,367,393]" class="Amphibia" family="Rhacophoridae" genus="Chiromantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="22" pageNumber="223" phylum="Chordata" rank="genus">
<emphasis id="37F9705FFFC2256BFF52FEEBFEB4FE6A" box="[209,348,367,393]" italics="true" pageId="22" pageNumber="223">Chiromantis</emphasis>
</taxonomicName>
at Kitutu (
<collectingCountry id="7D9AECDDFFC2256BFE59FEEBFDF1FE6A" box="[474,537,367,393]" name="Democratic Republic of the Congo" pageId="22" pageNumber="223">DRC</collectingCountry>
), marshes where it reproduces by sticking its eggs under the leaves of fountain grass (
<taxonomicName id="C28DD7CEFFC2256BFF60FE17FE0EFE4E" authority=", Shabunda" authorityName="Shabunda" box="[227,486,403,429]" class="Liliopsida" family="Poaceae" genus="Pennisetum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="22" pageNumber="223" phylum="Tracheophyta" rank="genus">
<emphasis id="37F9705FFFC2256BFF60FE17FE8EFE4F" box="[227,358,403,428]" italics="true" pageId="22" pageNumber="223">Pennisetum</emphasis>
, Shabunda
</taxonomicName>
, 
<collectingCountry id="7D9AECDDFFC2256BFE77FE17FDDAFE4E" box="[500,562,403,429]" name="Democratic Republic of the Congo" pageId="22" pageNumber="223">DRC</collectingCountry>
), a shaded bank of a pond in syntopy with 
<taxonomicName id="C28DD7CEFFC2256BFBA1FE17FB75FE4E" authorityName="Rapp" authorityYear="1842" box="[1058,1181,403,429]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="22" pageNumber="223" phylum="Chordata" rank="genus">
<emphasis id="37F9705FFFC2256BFBA1FE17FB75FE4E" box="[1058,1181,403,429]" italics="true" pageId="22" pageNumber="223">Hyperolius</emphasis>
</taxonomicName>
“ 
<emphasis id="37F9705FFFC2256BFB30FE17FAD6FE4E" box="[1203,1342,403,429]" italics="true" pageId="22" pageNumber="223">tuberculatus</emphasis>
” (likely 
<taxonomicName id="C28DD7CEFFC2256BFF14FE33FDD2FE32" authority="sensu Bell et al. 2017" authorityName="Bell" authorityYear="2017" box="[151,570,439,465]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="22" pageNumber="223" phylum="Chordata" rank="species" species="hutsebauti">
<emphasis id="37F9705FFFC2256BFF14FE33FF43FE33" box="[151,171,439,464]" italics="true" pageId="22" pageNumber="223">H</emphasis>
. 
<emphasis id="37F9705FFFC2256BFF3FFE33FE91FE33" box="[188,377,439,465]" italics="true" pageId="22" pageNumber="223">hutsebauti sensu</emphasis>
<bibRefCitation id="611CD1BCFFC2256BFE00FE33FDD2FE32" author="Bell, R. C. &amp; Parra, J. L. &amp; Badjedjea, G. &amp; Barej, M. F. &amp; Blackburn, D. C. &amp; Burger, M. &amp; Channing, A. &amp; Dehling, J. M. &amp; Greenbaum, E. &amp; Gvozdik, V. &amp; Kielgast, J. &amp; Kusamba, C. &amp; Lotters, S. &amp; McLaughlin, P. J. &amp; Nagy, Z. T. &amp; Rodel, M. - O. &amp; Portik, D. M. &amp; Stuart, B. L. &amp; VanDerWal, J. &amp; Zamudio, K. R." box="[387,570,439,465]" pageId="22" pageNumber="223" pagination="5223 - 5244" refId="ref18213" refString="Bell, R. C., Parra, J. L., Badjedjea, G., Barej, M. F., Blackburn, D. C., Burger, M., Channing, A., Dehling, J. M., Greenbaum, E., Gvozdik, V., Kielgast, J., Kusamba, C., Lotters, S., McLaughlin, P. J., Nagy, Z. T., Rodel, M. - O., Portik, D. M., Stuart, B. L., VanDerWal, J. &amp; Zamudio, K. R. (2017) Idiosyncratic responses to climate-driven forest fragmentation and marine incursions in reed frogs from Central Africa and the Gulf of Guinea Islands. Molecular Ecology, 26 (19), 5223 - 5244. https: // doi. org / 10.1111 / mec. 14260" type="journal article" year="2017">
Bell 
<emphasis id="37F9705FFFC2256BFE3FFE33FE18FE32" box="[444,496,439,465]" italics="true" pageId="22" pageNumber="223">et al</emphasis>
. 2017
</bibRefCitation>
</taxonomicName>
) and 
<taxonomicName id="C28DD7CEFFC2256BFDFDFE33FCE9FE32" authorityName="Laurent" authorityYear="1950" box="[638,769,439,465]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="22" pageNumber="223" phylum="Chordata" rank="species" species="frontalis">
<emphasis id="37F9705FFFC2256BFDFDFE33FD7AFE33" box="[638,658,439,464]" italics="true" pageId="22" pageNumber="223">H</emphasis>
. 
<emphasis id="37F9705FFFC2256BFD20FE33FCE9FE32" box="[675,769,439,465]" italics="true" pageId="22" pageNumber="223">frontalis</emphasis>
</taxonomicName>
; the center of this pond had reeds harboring 
<taxonomicName id="C28DD7CEFFC2256BFA93FE33FA75FE32" authorityName="Ahl" authorityYear="1931" box="[1296,1437,439,465]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="22" pageNumber="223" phylum="Chordata" rank="species" species="kivuensis">
<emphasis id="37F9705FFFC2256BFA93FE33FACCFE33" box="[1296,1316,439,464]" italics="true" pageId="22" pageNumber="223">H</emphasis>
. 
<emphasis id="37F9705FFFC2256BFAB5FE33FA75FE32" box="[1334,1437,439,465]" italics="true" pageId="22" pageNumber="223">kivuensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="C28DD7CEFFC2256BFF48FE5FFE4EFE16" box="[203,422,475,501]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="22" pageNumber="223" phylum="Chordata" rank="species" species="orophilus">
<emphasis id="37F9705FFFC2256BFF48FE5FFE4EFE16" box="[203,422,475,501]" italics="true" pageId="22" pageNumber="223">Afrixalus orophilus</emphasis>
</taxonomicName>
(Shabunda, 
<collectingCountry id="7D9AECDDFFC2256BFDB8FE5FFD92FE16" box="[571,634,475,501]" name="Democratic Republic of the Congo" pageId="22" pageNumber="223">DRC</collectingCountry>
). 
<bibRefCitation id="611CD1BCFFC2256BFD13FE5FFC87FE16" author="Laurent, R. F." box="[656,879,474,501]" pageId="22" pageNumber="223" pagination="350 - 358" refId="ref20655" refString="Laurent, R. F. (1983) About the herpetofauna of Central African montane forest. In: Rhodin, A. G. J. &amp; Miyata, K. (Eds.), Advances in Herpetology and Evolutionary Biology: Essays in Honor of Ernest E. Williams. Museum of Comparative Zoology, Cambridge, Massachusetts, pp. 350 - 358." type="book chapter" year="1983">Laurent (1983:352)</bibRefCitation>
listed the species (along with 
<taxonomicName id="C28DD7CEFFC2256BFB59FE5FFEA1FDFA" authorityName="Ahl" authorityYear="1924" class="Amphibia" family="Phrynobatrachidae" genus="Phrynobatrachus" kingdom="Animalia" order="Anura" pageId="22" pageNumber="223" phylum="Chordata" rank="species" species="petropedetoides">
<emphasis id="37F9705FFFC2256BFB59FE5FFEA1FDFA" italics="true" pageId="22" pageNumber="223">Phrynobatrachus petropedetoides</emphasis>
</taxonomicName>
and 
<taxonomicName id="C28DD7CEFFC2256BFEFCFE7BFD88FDFA" authorityName="Laurent" authorityYear="1950" box="[383,608,511,537]" class="Amphibia" family="Hyperoliidae" genus="Hyperolius" kingdom="Animalia" order="Anura" pageId="22" pageNumber="223" phylum="Chordata" rank="species" species="frontalis">
<emphasis id="37F9705FFFC2256BFEFCFE7BFD88FDFA" box="[383,608,511,537]" italics="true" pageId="22" pageNumber="223">Hyperolius frontalis</emphasis>
</taxonomicName>
) from “shadowy puddles of the thick transition forest.”
</paragraph>
</subSubSection>
<subSubSection id="4D97FFC6FFC2256BFF44FDA7FB89FD82" pageId="22" pageNumber="223" type="etymology">
<paragraph id="0532AC4DFFC2256BFF44FDA7FB89FD82" blockId="22.[151,1437,150,1185]" pageId="22" pageNumber="223">
<emphasis id="37F9705FFFC2256BFF44FDA7FEA6FDDE" bold="true" box="[199,334,547,573]" pageId="22" pageNumber="223">Etymology.</emphasis>
The species epithet is the Latin adjective “ 
<taxonomicName id="C28DD7CEFFC2256BFCA9FDA7FC62FDDE" authorityName="Greenbaum &amp; Portik &amp; Allen &amp; Vaughan &amp; Badjedjea &amp; Barej &amp; Behangana &amp; Conkey &amp; Dumbo &amp; Gonwouo &amp; Hirschfeld &amp; Hughes &amp; Igunzi &amp; Kusamba &amp; Lukwago &amp; Masudi &amp; Penner &amp; Reyes &amp; Rödel &amp; Roelke &amp; Romero &amp; Dehling" authorityYear="2022" box="[810,906,547,573]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="22" pageNumber="223" phylum="Chordata" rank="species" species="lacustris">
<emphasis id="37F9705FFFC2256BFCA9FDA7FC62FDDE" box="[810,906,547,573]" italics="true" pageId="22" pageNumber="223">lacustris</emphasis>
</taxonomicName>
,” meaning belonging to or dwelling in lakes; in allusion to the distribution of the new species in the region of the African Great 
<collectingRegion id="C74962AFFFC2256BFB95FDC3FBB4FD82" box="[1046,1116,583,609]" country="South Sudan" name="Lakes" pageId="22" pageNumber="223">Lakes</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="4D97FFC6FFC2256BFF44FDEEFAE2FB42" pageId="22" pageNumber="223" type="distribution">
<paragraph id="0532AC4DFFC2256BFF44FDEEFBF0FCBE" blockId="22.[151,1437,150,1185]" pageId="22" pageNumber="223">
<emphasis id="37F9705FFFC2256BFF44FDEEFDDDFD67" bold="true" box="[199,565,618,645]" pageId="22" pageNumber="223">Distribution and conservation.</emphasis>
The species is distributed from lowland rainforest of the 
<collectingCountry id="7D9AECDDFFC2256BFB3EFDEFFAE0FD66" box="[1213,1288,619,645]" name="Democratic Republic of the Congo" pageId="22" pageNumber="223">Congo</collectingCountry>
Basin at 
<quantity id="C27501A8FFC2256BFAF1FDEFFF45FD4A" metricMagnitude="2" metricUnit="m" metricValue="4.6" pageId="22" pageNumber="223" unit="m" value="460.0">460 m</quantity>
(Omaniundu, 
<collectingCountry id="7D9AECDDFFC2256BFECCFD0BFE63FD4A" box="[335,395,655,681]" name="Democratic Republic of the Congo" pageId="22" pageNumber="223">DRC</collectingCountry>
, 
<bibRefCitation id="611CD1BCFFC2256BFE15FD0BFDC5FD4A" author="Laurent, R. F." box="[406,557,655,681]" pageId="22" pageNumber="223" pagination="1 - 93" refId="ref20614" refString="Laurent, R. F. (1982) Le genre Afrixalus Laurent (Hyperoliidae) en Afrique Centrale. Annales du Musee Royal de l'Afrique Centrale, Serie in Octavo, Science Zoologique, 235, 1 - 93." type="journal article" year="1982">Laurent 1982</bibRefCitation>
) to transitional forests of the Albertine Rift at 
<quantity id="C27501A8FFC2256BFBB1FD0AFB6EFD4A" box="[1074,1158,654,681]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="22" pageNumber="223" unit="m" value="1650.0">1650 m</quantity>
(Mwana, 
<collectingCountry id="7D9AECDDFFC2256BFB7BFD0BFADBFD4A" box="[1272,1331,655,681]" name="Democratic Republic of the Congo" pageId="22" pageNumber="223">DRC</collectingCountry>
) and east as far as Mabira Forest, 
<collectingCountry id="7D9AECDDFFC2256BFE31FD37FDE1FD2E" box="[434,521,691,717]" name="Uganda" pageId="22" pageNumber="223">Uganda</collectingCountry>
(
<quantity id="C27501A8FFC2256BFD98FD37FD9EFD2E" box="[539,630,691,717]" metricMagnitude="3" metricUnit="m" metricValue="1.121" pageId="22" pageNumber="223" unit="m" value="1121.0">1121 m</quantity>
). As shown in 
<figureCitation id="9DB6B0C8FFC2256BFCA6FD37FC6FFD2E" box="[805,903,690,717]" captionStart="FIGURE 5" captionStartId="9.[151,250,1256,1280]" captionTargetBox="[155,1436,283,1233]" captionTargetId="figure-66@9.[151,1436,273,1233]" captionTargetPageId="9" captionText="FIGURE 5. Distribution map of Afrixalus phantasma sp. nov. (black symbols) and A. lacustris sp. nov. (white symbols). Star, triangle, and circle symbols represent holotype, examined specimens, and literature records (Drewes &amp; Vindum 1994; Laurent 1950, 1952, 1972, 1982; Vonesh 2001), respectively. Areas shaded in lighter gray are below 1700 meters, whereas areas shaded in darker gray are 1700 meters or higher. Lakes are shown in white. Gray triangle symbol represents A. lacustris specimens (RMCA 77-020-B-133–136) from a problematic locality at Tubutubu, Itombwe Plateau, DRC; see species account for details. B = Burundi, R = Rwanda." figureDoi="http://doi.org/10.5281/zenodo.6986889" httpUri="https://zenodo.org/record/6986889/files/figure.png" pageId="22" pageNumber="223">Figure 5</figureCitation>
, the species has a relatively large distribution, including several national parks and protected areas (e.g., Virunga National Park, Bwindi Impenetrable National Park), and thus is not likely to be threatened based on a limited distribution. Using GeoCAT, its extent of occurrence is estimated as 
<quantity id="C27501A8FFC2256BFEC2FC9BFE2FFCD8" box="[321,455,799,827]" metricMagnitude="8" metricUnit="m" metricValue="4.09238" pageId="22" pageNumber="223" unit="km" value="409238.0">409,238 km</quantity>
² and its area of occupancy as 
<quantity id="C27501A8FFC2256BFC95FC9BFC83FCD8" box="[790,875,799,827]" metricMagnitude="5" metricUnit="m" metricValue="1.6" pageId="22" pageNumber="223" unit="km" value="160.0">160 km</quantity>
² (
<bibRefCitation id="611CD1BCFFC2256BFC01FC9BFB87FCDA" author="Bachman, S. &amp; Moat, J. &amp; Hill, A. &amp; de la Torre, J. &amp; Scott, B." box="[898,1135,799,825]" pageId="22" pageNumber="223" pagination="117 - 126" refId="ref18010" refString="Bachman, S., Moat, J., Hill, A., de la Torre, J. &amp; Scott, B. (2011) Supporting Red List threat assessments with GeoCAT: Geospatial Conservation Assessment Tool. ZooKeys, 150, 117 - 126. https: // doi. org / 10.3897 / zookeys. 150.2109" type="journal article" year="2011">
Bachman 
<emphasis id="37F9705FFFC2256BFC77FC9BFBCDFCDA" box="[1012,1061,799,825]" italics="true" pageId="22" pageNumber="223">et al</emphasis>
. 2011
</bibRefCitation>
). We therefore classify this species as Least Concern according to the IUCN Red List criteria (
<bibRefCitation id="611CD1BCFFC2256BFC01FCC7FBE3FCBE" author="IUCN" box="[898,1035,835,861]" pageId="22" pageNumber="223" refId="ref19805" refString="IUCN (2021) IUCN Red List Categories and Criteria: Version 2021 - 3. Available from: https: // www. iucnredlist. org / (accessed 25 May 2022)" type="url" year="2021">IUCN 2021</bibRefCitation>
).
</paragraph>
<paragraph id="0532AC4DFFC2256BFF44FCE3FAE2FB42" blockId="22.[151,1437,150,1185]" pageId="22" pageNumber="223">
Four specimens (RMCA 77-020-B-133–136) from Tubutubu, Itombwe 
<collectingRegion id="C74962AFFFC2256BFB8CFCE3FB89FC62" box="[1039,1121,871,897]" country="Nigeria" name="Plateau" pageId="22" pageNumber="223">Plateau</collectingRegion>
(
<quantity id="C27501A8FFC2256BFBF5FCE3FB3BFC62" box="[1142,1235,871,897]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="22" pageNumber="223" unit="m" value="2300.0">2300 m</quantity>
) are problematic because they are morphologically consistent with 
<taxonomicName id="C28DD7CEFFC2256BFD4EFC0FFCA5FC46" authorityName="Greenbaum &amp; Portik &amp; Allen &amp; Vaughan &amp; Badjedjea &amp; Barej &amp; Behangana &amp; Conkey &amp; Dumbo &amp; Gonwouo &amp; Hirschfeld &amp; Hughes &amp; Igunzi &amp; Kusamba &amp; Lukwago &amp; Masudi &amp; Penner &amp; Reyes &amp; Rödel &amp; Roelke &amp; Romero &amp; Dehling" authorityYear="2022" box="[717,845,907,933]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="22" pageNumber="223" phylum="Chordata" rank="species" species="lacustris">
<emphasis id="37F9705FFFC2256BFD4EFC0FFD36FC46" box="[717,734,907,933]" italics="true" pageId="22" pageNumber="223">A</emphasis>
. 
<emphasis id="37F9705FFFC2256BFD6EFC0FFCA5FC46" box="[749,845,907,933]" italics="true" pageId="22" pageNumber="223">lacustris</emphasis>
</taxonomicName>
, but they were collected from a substantially higher elevation than all other known specimens. Coordinates for Tubutubu provided by RMCA (
<geoCoordinate id="60B9CA8AFFC2256BFB09FC2BFB54FC2A" box="[1162,1212,943,969]" degrees="4" direction="south" orientation="latitude" pageId="22" pageNumber="223" precision="55555" value="-4.0">4° S</geoCoordinate>
, 
<geoCoordinate id="60B9CA8AFFC2256BFB4BFC2BFAD1FC2A" box="[1224,1337,943,969]" degrees="28.933" direction="east" orientation="longitude" pageId="22" pageNumber="223" precision="55" value="28.933">28.933° E</geoCoordinate>
) suggest the locality is east of the highest point of the plateau at an elevation of about 
<quantity id="C27501A8FFC2256BFB89FC57FB89FC0E" box="[1034,1121,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="22" pageNumber="223" unit="m" value="1400.0">1400 m</quantity>
on Google Earth. Although it is possible that the specimens were collected at the latter elevation, 
<bibRefCitation id="611CD1BCFFC2256BFC34FC73FB8AFBF2" author="Laurent, R. F." box="[951,1122,1015,1041]" pageId="22" pageNumber="223" pagination="1 - 93" refId="ref20614" refString="Laurent, R. F. (1982) Le genre Afrixalus Laurent (Hyperoliidae) en Afrique Centrale. Annales du Musee Royal de l'Afrique Centrale, Serie in Octavo, Science Zoologique, 235, 1 - 93." type="journal article" year="1982">Laurent (1982)</bibRefCitation>
mentions bamboo forest as the habitat, which usually occurs at the highest elevations of the Itombwe 
<collectingRegion id="C74962AFFFC2256BFC58FB9FFBC5FBD6" box="[987,1069,1051,1077]" country="Nigeria" name="Plateau" pageId="22" pageNumber="223">Plateau</collectingRegion>
(
<bibRefCitation id="611CD1BCFFC2256BFBBEFB9FFB15FBD6" author="Doumenge, C." box="[1085,1277,1051,1077]" pageId="22" pageNumber="223" pagination="249 - 264" refId="ref18902" refString="Doumenge, C. (1998) Forest diversity, distribution, and dynamique in the Itombwe Mountains, South-Kivu, Congo Democratic Republic. Mountain Research and Development, 18 (3), 249 - 264. https: // doi. org / 10.2307 / 3674036" type="journal article" year="1998">Doumenge 1998</bibRefCitation>
). No clarity is offered by 
<bibRefCitation id="611CD1BCFFC2256BFE93FBBBFE52FBBA" author="Laurent, R. F." box="[272,442,1087,1113]" pageId="22" pageNumber="223" pagination="458 - 467" refId="ref20487" refString="Laurent, R. F. (1964) Adaptive modifications in frogs of an isolated highland fauna in Central Africa. Evolution, 18 (3), 458 - 467. https: // doi. org / 10.1111 / j. 1558 - 5646.1964. tb 01622. x" type="journal article" year="1964">Laurent (1964)</bibRefCitation>
, because he does not mention any 
<taxonomicName id="C28DD7CEFFC2256BFCBBFBBBFC77FBBA" authorityName="Laurent" authorityYear="1944" box="[824,927,1087,1113]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="22" pageNumber="223" phylum="Chordata" rank="genus">
<emphasis id="37F9705FFFC2256BFCBBFBBBFC77FBBA" box="[824,927,1087,1113]" italics="true" pageId="22" pageNumber="223">Afrixalus</emphasis>
</taxonomicName>
in his study of the ecology and distribution of amphibians of the Itombwe 
<collectingRegion id="C74962AFFFC2256BFE52FBE7FDCFFB9E" box="[465,551,1123,1149]" country="Nigeria" name="Plateau" pageId="22" pageNumber="223">Plateau</collectingRegion>
. Given this conflicting information, we do not include the Tubutubu specimens in our current understanding of the elevational distribution of 
<taxonomicName id="C28DD7CEFFC2256BFCCFFB03FC23FB42" authorityName="Greenbaum &amp; Portik &amp; Allen &amp; Vaughan &amp; Badjedjea &amp; Barej &amp; Behangana &amp; Conkey &amp; Dumbo &amp; Gonwouo &amp; Hirschfeld &amp; Hughes &amp; Igunzi &amp; Kusamba &amp; Lukwago &amp; Masudi &amp; Penner &amp; Reyes &amp; Rödel &amp; Roelke &amp; Romero &amp; Dehling" authorityYear="2022" box="[844,971,1159,1185]" class="Amphibia" family="Hyperoliidae" genus="Afrixalus" kingdom="Animalia" order="Anura" pageId="22" pageNumber="223" phylum="Chordata" rank="species" species="lacustris">
<emphasis id="37F9705FFFC2256BFCCFFB03FCB5FB42" box="[844,861,1159,1185]" italics="true" pageId="22" pageNumber="223">A</emphasis>
. 
<emphasis id="37F9705FFFC2256BFCE8FB03FC23FB42" box="[875,971,1159,1185]" italics="true" pageId="22" pageNumber="223">lacustris</emphasis>
</taxonomicName>
(see gray triangle in 
<figureCitation id="9DB6B0C8FFC2256BFB3AFB03FB14FB42" box="[1209,1276,1158,1185]" captionStart="FIGURE 5" captionStartId="9.[151,250,1256,1280]" captionTargetBox="[155,1436,283,1233]" captionTargetId="figure-66@9.[151,1436,273,1233]" captionTargetPageId="9" captionText="FIGURE 5. Distribution map of Afrixalus phantasma sp. nov. (black symbols) and A. lacustris sp. nov. (white symbols). Star, triangle, and circle symbols represent holotype, examined specimens, and literature records (Drewes &amp; Vindum 1994; Laurent 1950, 1952, 1972, 1982; Vonesh 2001), respectively. Areas shaded in lighter gray are below 1700 meters, whereas areas shaded in darker gray are 1700 meters or higher. Lakes are shown in white. Gray triangle symbol represents A. lacustris specimens (RMCA 77-020-B-133–136) from a problematic locality at Tubutubu, Itombwe Plateau, DRC; see species account for details. B = Burundi, R = Rwanda." figureDoi="http://doi.org/10.5281/zenodo.6986889" httpUri="https://zenodo.org/record/6986889/files/figure.png" pageId="22" pageNumber="223">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>