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<document id="6E8A4271EF03076CF2325F2650ABE35A" ID-CLB-Dataset="29753" ID-DOI="10.11646/zootaxa.4442.3.4" ID-GBIF-Dataset="62cba2ff-77e1-42fa-b362-17a22409af07" ID-ISSN="1175-5326" ID-Zenodo-Dep="1303737" ID-ZooBank="C67563CF-9545-4AF0-A3AA-366338BB7F1A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530598579704" checkinUser="plazi" docAuthor="Cala-Riquelme, Franklyn, Quijano-Cuervo, Luis, Sabogal-González, Alexander &amp; Agnarsson, Ingi" docDate="2018" docId="5B6787B4FFC28D7ED0D7FE09FB6DC660" docLanguage="en" docName="zootaxa.4442.3.4.pdf" docOrigin="Zootaxa 4442 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Otiothops vaupes Cala-Riquelme &amp; Quijano-Cuervo &amp; Sabogal-González &amp; Agnarsson 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="420" masterDocId="A75EFFCCFFC48D79D040FF9DFFD7C400" masterDocTitle="New species of Otiothopinae (Araneae: Palpimanidae) from Colombia" masterLastPageNumber="426" masterPageNumber="413" pageNumber="419" updateTime="1698552413182" updateUser="ExternalLinkService">
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<mods:title id="5C984789638E6C79FDEA269FC0A391AD">New species of Otiothopinae (Araneae: Palpimanidae) from Colombia</mods:title>
</mods:titleInfo>
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<mods:namePart id="C643672A275BFF1A578589F2D2DB2E90">Cala-Riquelme, Franklyn</mods:namePart>
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<mods:namePart id="5AC5CB90556F9F7DC2B964CFB683BD05">Quijano-Cuervo, Luis</mods:namePart>
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<mods:namePart id="D10449836B0AA2B8A4BBF9DD65377D3A">Sabogal-González, Alexander</mods:namePart>
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<mods:namePart id="B9573C6FE1A7809CA85E96020CB4FC85">Agnarsson, Ingi</mods:namePart>
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<mods:title id="F4AAFF46B3373C693DDC756C5CAC172C">Zootaxa</mods:title>
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<mods:date id="7826E6C798C82007E6C3AB81A2257218">2018</mods:date>
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<mods:number id="08AE7E6183590E7AC7A1518EC9A24B3E">2018-07-03</mods:number>
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<treatment id="5B6787B4FFC28D7ED0D7FE09FB6DC660" ID-DOI="http://doi.org/10.5281/zenodo.5964805" ID-GBIF-Taxon="150767718" ID-Zenodo-Dep="5964805" LSID="urn:lsid:plazi:treatment:5B6787B4FFC28D7ED0D7FE09FB6DC660" httpUri="http://treatment.plazi.org/id/5B6787B4FFC28D7ED0D7FE09FB6DC660" lastPageId="7" lastPageNumber="420" pageId="6" pageNumber="419">
<subSubSection id="9BD46529FFC28D7FD0D7FE09FEF0C5CF" pageId="6" pageNumber="419" type="nomenclature">
<paragraph id="D37136A2FFC28D7FD0D7FE09FB91C5AE" blockId="6.[151,1094,404,463]" box="[151,1094,404,430]" pageId="6" pageNumber="419">
<heading id="883981CEFFC28D7FD0D7FE09FB91C5AE" bold="true" box="[151,1094,404,430]" fontSize="11" level="1" pageId="6" pageNumber="419" reason="1">
<emphasis id="E1BAEAB0FFC28D7FD0D7FE09FB91C5AE" bold="true" box="[151,1094,404,430]" pageId="6" pageNumber="419">
<taxonomicName id="14CE4D21FFC28D7FD0D7FE09FEB1C5AE" ID-CoL="4B6ZX" authority="Cala-Riquelme &amp; Quijano-Cuervo &amp; Sabogal-González &amp; Agnarsson, 2018" authorityName="Cala-Riquelme &amp; Quijano-Cuervo &amp; Sabogal-González &amp; Agnarsson" authorityYear="2018" box="[151,358,404,430]" class="Arachnida" family="Palpimanidae" genus="Otiothops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="419" phylum="Arthropoda" rank="species" species="vaupes" status="sp. nov.">
<emphasis id="E1BAEAB0FFC28D7FD0D7FE09FEB1C5AE" bold="true" box="[151,358,404,430]" italics="true" pageId="6" pageNumber="419">Otiothops vaupes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FA8957CBFFC28D7FD12EFE09FE67C5AE" box="[366,432,404,430]" pageId="6" pageNumber="419" rank="species">sp. n.</taxonomicNameLabel>
Cala-Riquelme, Quijano-Cuervo &amp; Agnarsson, 2018
</emphasis>
</heading>
</paragraph>
<paragraph id="D37136A2FFC28D7FD0D7FE2BFEF0C5CF" blockId="6.[151,1094,404,463]" box="[151,295,438,463]" pageId="6" pageNumber="419">
(
<figureCitation id="4BF52A27FFC28D7FD0DFFE2BFEC8C5CF" box="[159,287,438,463]" captionStart="FIGURES 1923" captionStartId="6.[151,264,1911,1934]" captionTargetBox="[171,1422,710,1838]" captionTargetId="figure@6.[128,1460,660,1891]" captionTargetPageId="6" captionText="FIGURES 1923. Otiothops vaupes sp. n. 19, Male holotype, habitus dorsal view. 20-23, Male holotype, left palp (E = embolus, Cy = cymbium, Bu = bulb): 2021, retrolateral view; 22, ventral view; 23, prolateral view." httpUri="https://zenodo.org/record/1303745/files/figure.png" pageId="6" pageNumber="419">Figs 1923</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9BD46529FFC28D7FD0D7FE63FE63C65F" pageId="6" pageNumber="419" type="materials_examined">
<paragraph id="D37136A2FFC28D7FD0D7FE63FE63C65F" blockId="6.[151,1436,510,607]" pageId="6" pageNumber="419">
<materialsCitation id="63A63CFFFFC28D7FD0D7FE63FC66C63B" ID-GBIF-Occurrence="1950907438" collectingDate="2004-04-01" collectionCode="MPUJ" collectorName="J. Pinzon" country="Colombia" county="Taraira" latitude="-0.564744" location="Male" longLatPrecision="1" longitude="-69.63411" municipality="Lago Taraira" pageId="6" pageNumber="419" specimenCode="ENT0058647" specimenCount="2" stateProvince="Vaupes" typeStatus="holotype">
<emphasis id="E1BAEAB0FFC28D7FD0D7FE63FE92C617" bold="true" box="[151,325,510,535]" pageId="6" pageNumber="419">
<typeStatus id="0C758800FFC28D7FD0D7FE63FF05C617" box="[151,210,510,535]" pageId="6" pageNumber="419">Type</typeStatus>
material.
</emphasis>
<location id="D6116079FFC28D7FD10BFE63FE52C617" LSID="urn:lsid:plazi:treatment:5B6787B4FFC28D7ED0D7FE09FB6DC660:D6116079FFC28D7FD10BFE63FE52C617" box="[331,389,510,535]" country="Colombia" county="Taraira" latitude="-0.564744" longLatPrecision="1" longitude="-69.63411" municipality="Lago Taraira" name="Male" pageId="6" pageNumber="419" stateProvince="Vaupes">Male</location>
<typeStatus id="0C758800FFC28D7FD1CCFE63FE39C617" box="[396,494,510,535]" pageId="6" pageNumber="419" type="holotype">holotype</typeStatus>
from
<collectingMunicipality id="3315ACD8FFC28D7FD272FE62FD13C617" box="[562,708,510,535]" pageId="6" pageNumber="419">Lago Taraira</collectingMunicipality>
,
<collectingCounty id="3A104E2EFFC28D7FD28FFE63FCF6C617" box="[719,801,510,535]" pageId="6" pageNumber="419">Taraira</collectingCounty>
,
<collectingRegion id="110AF840FFC28D7FD36CFE62FC57C617" box="[812,896,511,535]" country="Colombia" name="Vaupes" pageId="6" pageNumber="419">Vaupés</collectingRegion>
,
<collectingCountry id="ABD97632FFC28D7FD3CAFE63FC2CC617" box="[906,1019,510,535]" name="Colombia" pageId="6" pageNumber="419">Colombia</collectingCountry>
,
<geoCoordinate id="B6FA5065FFC28D7FD447FE63FB45C617" box="[1031,1170,510,535]" direction="south" orientation="latitude" pageId="6" pageNumber="419" precision="1" value="-0.564744">0.564744°S-</geoCoordinate>
<geoCoordinate id="B6FA5065FFC28D7FD4D8FE63FA95C617" box="[1176,1346,510,535]" direction="west" orientation="longitude" pageId="6" pageNumber="419" precision="1" value="-69.63411">69.6341099°W</geoCoordinate>
,
<collectingDate id="B734E98AFFC28D7FD50CFE62FF05C63B" pageId="6" pageNumber="419" value="2004-04-01">
<date id="A7701062FFC28D7FD50CFE62FF05C63B" pageId="6" pageNumber="419" value="2004-04-01">1 April 2004</date>
</collectingDate>
, tropical rainforest leaf litter,
<collectorName id="7E3B5374FFC28D7FD276FDBEFD76C63B" box="[566,673,546,571]" pageId="6" pageNumber="419">J. Pinzon</collectorName>
(
<collectionCode id="B5DFAE67FFC28D7FD2F4FDBFFD29C63B" box="[692,766,546,571]" pageId="6" pageNumber="419">MPUJ</collectionCode>
_
<specimenCode id="83689ED9FFC28D7FD34DFDBFFC7EC63B" box="[781,937,546,571]" collectionCode="ENT" country="Uganda" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="6" pageNumber="419">ENT0058647</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="63A63CFFFFC28D7FD380FDBEFE66C65F" ID-GBIF-Occurrence="1950907431" collectingDate="2004-04-01" collectionCode="MPUJ" collectorName="J. Pinzon" country="Colombia" county="Taraira" latitude="-0.564744" location="Male" longLatPrecision="1" longitude="-69.63411" municipality="Lago Taraira" pageId="6" pageNumber="419" specimenCode="ENT0058648.1" specimenCount="1" specimenCount-male="1" stateProvince="Vaupes" typeStatus="paratype">
<typeStatus id="0C758800FFC28D7FD380FDBEFBF1C63B" box="[960,1062,547,571]" pageId="6" pageNumber="419" type="paratype">Paratype</typeStatus>
:
<specimenCount id="C5C8FD2BFFC28D7FD475FDBEFB5FC63B" box="[1077,1160,546,571]" pageId="6" pageNumber="419" type="male">1 male</specimenCount>
, same data as holotype (
<collectionCode id="B5DFAE67FFC28D7FD0DEFDDBFF3FC65F" box="[158,232,582,607]" pageId="6" pageNumber="419">MPUJ</collectionCode>
_
<specimenCode id="83689ED9FFC28D7FD0B7FDDBFE7EC65F" box="[247,425,582,607]" collectionCode="ENT" country="Uganda" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="6" pageNumber="419">ENT0058648.1</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="87B1662AFFC28D7FD0D7F8EAFB59C3AC" httpUri="https://zenodo.org/record/1303745/files/figure.png" pageId="6" pageNumber="419" startId="6.[151,264,1911,1934]" targetBox="[171,1422,710,1838]" targetPageId="6">
<paragraph id="D37136A2FFC28D7FD0D7F8EAFB59C3AC" blockId="6.[151,1435,1911,1964]" pageId="6" pageNumber="419">
<emphasis id="E1BAEAB0FFC28D7FD0D7F8EAFE8FC38D" bold="true" box="[151,344,1911,1934]" pageId="6" pageNumber="419">FIGURES 1923.</emphasis>
<taxonomicName id="14CE4D21FFC28D7FD122F8E5FDC1C38E" authority="Cala-Riquelme &amp; Quijano-Cuervo &amp; Sabogal-González &amp; Agnarsson, 2018" authorityName="Cala-Riquelme &amp; Quijano-Cuervo &amp; Sabogal-González &amp; Agnarsson" authorityYear="2018" box="[354,534,1912,1934]" class="Arachnida" family="Palpimanidae" genus="Otiothops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="419" phylum="Arthropoda" rank="species" species="vaupes" status="sp. nov.">
<emphasis id="E1BAEAB0FFC28D7FD122F8E5FDC1C38E" box="[354,534,1912,1934]" italics="true" pageId="6" pageNumber="419">Otiothops vaupes</emphasis>
</taxonomicName>
<emphasis id="E1BAEAB0FFC28D7FD261F8E5FD56C38D" bold="true" box="[545,641,1912,1933]" pageId="6" pageNumber="419">
<taxonomicNameLabel id="FA8957CBFFC28D7FD261F8E5FD89C38D" box="[545,606,1912,1933]" pageId="6" pageNumber="419" rank="species">sp. n.</taxonomicNameLabel>
19
</emphasis>
, Male holotype, habitus dorsal view.
<emphasis id="E1BAEAB0FFC28D7FD456F8E5FB86C38D" bold="true" box="[1046,1105,1912,1933]" pageId="6" pageNumber="419">20-23</emphasis>
, Male holotype, left palp (E = embolus, Cy = cymbium, Bu = bulb):
<emphasis id="E1BAEAB0FFC28D7FD259F80AFD8FC3AC" bold="true" box="[537,600,1943,1964]" pageId="6" pageNumber="419">2021</emphasis>
, retrolateral view;
<emphasis id="E1BAEAB0FFC28D7FD359F80AFCE5C3AC" bold="true" box="[793,818,1943,1964]" pageId="6" pageNumber="419">22</emphasis>
, ventral view;
<emphasis id="E1BAEAB0FFC28D7FD38AF80AFC34C3AC" bold="true" box="[970,995,1943,1964]" pageId="6" pageNumber="419">23</emphasis>
, prolateral view.
</paragraph>
</caption>
<subSubSection id="9BD46529FFC38D7ED087FF0AFA85C4F8" pageId="7" pageNumber="420" type="diagnosis">
<paragraph id="D37136A2FFC38D7ED087FF0AFA85C4F8" blockId="7.[151,1436,151,608]" pageId="7" pageNumber="420">
<emphasis id="E1BAEAB0FFC38D7ED087FF0AFE96C4B0" bold="true" box="[199,321,151,176]" pageId="7" pageNumber="420">Diagnosis.</emphasis>
Males of
<taxonomicName id="14CE4D21FFC38D7ED1FBFF05FD55C4B0" authority="Cala-Riquelme &amp; Quijano-Cuervo &amp; Sabogal-González &amp; Agnarsson, 2018" authorityName="Cala-Riquelme &amp; Quijano-Cuervo &amp; Sabogal-González &amp; Agnarsson" authorityYear="2018" box="[443,642,152,176]" class="Arachnida" family="Palpimanidae" genus="Otiothops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="420" phylum="Arthropoda" rank="species" species="vaupes" status="sp. nov.">
<emphasis id="E1BAEAB0FFC38D7ED1FBFF05FD55C4B0" box="[443,642,152,176]" italics="true" pageId="7" pageNumber="420">Otiothops vaupes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FA8957CBFFC38D7ED2CCFF0AFD19C4B0" box="[652,718,151,176]" pageId="7" pageNumber="420" rank="species">
<emphasis id="E1BAEAB0FFC38D7ED2CCFF0AFD19C4B0" bold="true" box="[652,718,151,176]" pageId="7" pageNumber="420">sp. n.</emphasis>
</taxonomicNameLabel>
resemble those of species in the
<emphasis id="E1BAEAB0FFC38D7ED416FF05FB72C4AF" box="[1110,1189,152,175]" italics="true" pageId="7" pageNumber="420">typicus</emphasis>
group by the greatly inflated palpal bulb (
<figureCitation id="4BF52A27FFC38D7ED13FFF21FDD6C4D4" box="[383,513,188,213]" captionStart="FIGURES 1923" captionStartId="6.[151,264,1911,1934]" captionTargetBox="[171,1422,710,1838]" captionTargetId="figure@6.[128,1460,660,1891]" captionTargetPageId="6" captionText="FIGURES 1923. Otiothops vaupes sp. n. 19, Male holotype, habitus dorsal view. 20-23, Male holotype, left palp (E = embolus, Cy = cymbium, Bu = bulb): 2021, retrolateral view; 22, ventral view; 23, prolateral view." httpUri="https://zenodo.org/record/1303745/files/figure.png" pageId="7" pageNumber="420">Figs 2023</figureCitation>
; see also
<bibRefCitation id="B75F4B53FFC38D7ED22BFF21FCDFC4D5" box="[619,776,188,213]" pageId="7" pageNumber="420" refId="ref7069" refString="Platnick, N. I. (1975) A revision of the palpimanid spiders of the new subfamily Otiothopinae (Araneae: Palpimanidae). American Museum Novitates, 2562, 1 - 32" type="journal article">Platnick 1975</bibRefCitation>
;
<bibRefCitation id="B75F4B53FFC38D7ED354FF21FB81C4D4" box="[788,1110,188,213]" pageId="7" pageNumber="420" refId="ref6451" refString="Brescovit, A. D. &amp; Bonaldo, A. B. (1993) On the genus Otiothops in Brazil (Araneae, Palpimanidae). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie, 63, 47 - 50." type="journal article">Brescovit and Bonaldo 1993</bibRefCitation>
), but can be distinguished by the shape of the embolus, which is curved and widened close to the tip (
<figureCitation id="4BF52A27FFC38D7ED3FCFF42FB96C4F8" box="[956,1089,223,248]" captionStart="FIGURES 1923" captionStartId="6.[151,264,1911,1934]" captionTargetBox="[171,1422,710,1838]" captionTargetId="figure@6.[128,1460,660,1891]" captionTargetPageId="6" captionText="FIGURES 1923. Otiothops vaupes sp. n. 19, Male holotype, habitus dorsal view. 20-23, Male holotype, left palp (E = embolus, Cy = cymbium, Bu = bulb): 2021, retrolateral view; 22, ventral view; 23, prolateral view." httpUri="https://zenodo.org/record/1303745/files/figure.png" pageId="7" pageNumber="420">Figs. 20-23</figureCitation>
). Females are unknown.
</paragraph>
</subSubSection>
<subSubSection id="9BD46529FFC38D7ED087FE99FA40C51D" box="[199,1431,260,285]" pageId="7" pageNumber="420" type="etymology">
<paragraph id="D37136A2FFC38D7ED087FE99FA40C51D" blockId="7.[151,1436,151,608]" box="[199,1431,260,285]" pageId="7" pageNumber="420">
<emphasis id="E1BAEAB0FFC38D7ED087FE99FE99C51D" bold="true" box="[199,334,260,285]" pageId="7" pageNumber="420">Etymology.</emphasis>
This specific epithet is a noun in apposition referring to the
<typeStatus id="0C758800FFC38D7ED3B2FE98FBF5C51D" box="[1010,1058,261,285]" pageId="7" pageNumber="420">type</typeStatus>
locality from
<collectingRegion id="110AF840FFC38D7ED482FE99FAC1C51C" box="[1218,1302,260,284]" country="Colombia" name="Vaupes" pageId="7" pageNumber="420">Vaupés</collectingRegion>
,
<collectingCountry id="ABD97632FFC38D7ED560FE99FA45C51D" box="[1312,1426,260,285]" name="Colombia" pageId="7" pageNumber="420">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="9BD46529FFC38D7ED087FEBAFD9EC618" pageId="7" pageNumber="420" type="description">
<paragraph id="D37136A2FFC38D7ED087FEBAFD9EC618" blockId="7.[151,1436,151,608]" pageId="7" pageNumber="420">
<emphasis id="E1BAEAB0FFC38D7ED087FEBAFE8EC540" bold="true" box="[199,345,295,320]" pageId="7" pageNumber="420">Description.</emphasis>
<emphasis id="E1BAEAB0FFC38D7ED120FEB5FE4DC540" box="[352,410,296,320]" italics="true" pageId="7" pageNumber="420">Male</emphasis>
(
<typeStatus id="0C758800FFC38D7ED1E8FEBAFDD9C540" box="[424,526,295,320]" pageId="7" pageNumber="420" type="holotype">holotype</typeStatus>
, MPUJ_ENT0058647): Total length 5.8. Carapace: 2.7 long, 2.0 wide (at leg II), 1.2 high (at leg II). Leg I: femur 2.1 long, 1.0 wide. Abdomen: 2.8 long, 1.8 wide, scute 0.9 long. Posterior median eyes separated by less than their own diameter. Palp: femur 0.8 long, 0.3 wide; patella 0.25 long, 0.2 wide; tibia 0.37 long, 0.34 wide; cymbium 0.82 long; embolus 0.67 long. Sclerotized portions of body dark orange; abdominal scute and unsclerotized portion of dorsal abdomen orange-brown. Palpal femur not thickened; tibia longer than wide, considerably smaller than bulb; bulb inflated; embolus longer than bulb but shorter than cymbium, slightly curved with bifurcate tip (
<figureCitation id="4BF52A27FFC38D7ED1FAFE62FDEAC618" box="[442,573,511,536]" captionStart="FIGURES 1923" captionStartId="6.[151,264,1911,1934]" captionTargetBox="[171,1422,710,1838]" captionTargetId="figure@6.[128,1460,660,1891]" captionTargetPageId="6" captionText="FIGURES 1923. Otiothops vaupes sp. n. 19, Male holotype, habitus dorsal view. 20-23, Male holotype, left palp (E = embolus, Cy = cymbium, Bu = bulb): 2021, retrolateral view; 22, ventral view; 23, prolateral view." httpUri="https://zenodo.org/record/1303745/files/figure.png" pageId="7" pageNumber="420">Figs 2023</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9BD46529FFC38D7ED087FDB9FB6DC660" pageId="7" pageNumber="420" type="distribution">
<paragraph id="D37136A2FFC38D7ED087FDB9FBE8C63D" blockId="7.[151,1436,151,608]" box="[199,1087,548,573]" pageId="7" pageNumber="420">
<emphasis id="E1BAEAB0FFC38D7ED087FDB9FEB7C63D" bold="true" box="[199,352,548,573]" pageId="7" pageNumber="420">Distribution.</emphasis>
Known only from the
<typeStatus id="0C758800FFC38D7ED220FDB8FD47C63D" box="[608,656,549,573]" pageId="7" pageNumber="420">type</typeStatus>
locality at Taraira,
<collectingRegion id="110AF840FFC38D7ED32AFDB9FC69C63C" box="[874,958,548,572]" country="Colombia" name="Vaupes" pageId="7" pageNumber="420">Vaupés</collectingRegion>
,
<collectingCountry id="ABD97632FFC38D7ED389FDB9FBECC63D" box="[969,1083,548,573]" name="Colombia" pageId="7" pageNumber="420">Colombia</collectingCountry>
.
</paragraph>
<paragraph id="D37136A2FFC38D7ED087FDDAFB6DC660" blockId="7.[151,1436,151,608]" box="[199,1210,583,608]" pageId="7" pageNumber="420">
<emphasis id="E1BAEAB0FFC38D7ED087FDDAFE52C660" bold="true" box="[199,389,583,608]" pageId="7" pageNumber="420">Natural history.</emphasis>
The
<typeStatus id="0C758800FFC38D7ED1FEFDDAFDF7C660" box="[446,544,583,608]" pageId="7" pageNumber="420" type="holotype">holotype</typeStatus>
and
<typeStatus id="0C758800FFC38D7ED216FDD5FD61C660" box="[598,694,584,608]" pageId="7" pageNumber="420" type="paratype">paratype</typeStatus>
were found in leaf litter in tropical rainforest.
</paragraph>
</subSubSection>
</treatment>
</document>