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<mods:title id="F6BCFE859F37F70513307C7D3D9948E9">Brachyuran crabs of the family Macrophthalmidae Dana, 1851 (Decapoda: Brachyura: Macrophthalmidae) of the Persian Gulf</mods:title>
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<mods:date id="682BEA71ACDE5867F269DC503106399D">2011</mods:date>
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<treatment id="587287CE553FFFFAFF774F09FD8A7849" ID-DOI="http://doi.org/10.5281/zenodo.5417606" ID-GBIF-Taxon="119410909" ID-Zenodo-Dep="5417606" LSID="urn:lsid:plazi:treatment:587287CE553FFFFAFF774F09FD8A7849" httpUri="http://treatment.plazi.org/id/587287CE553FFFFAFF774F09FD8A7849" lastPageId="10" lastPageNumber="11" pageId="6" pageNumber="7">
<subSubSection id="98C16553553FFFF6FF774F09FE4F795C" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="D06436D8553FFFF6FF774F09FD8D793B" blockId="6.[151,621,793,852]" box="[151,621,793,819]" pageId="6" pageNumber="7">
<heading id="8B2C81B4553FFFF6FF774F09FD8D793B" bold="true" box="[151,621,793,819]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<taxonomicName id="17DB4D5B553FFFF6FF774F09FD8D793B" ID-CoL="84GMD" authority="Lucas, 1836" authorityName="Lucas" authorityYear="1836" box="[151,621,793,819]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="E2AFEACA553FFFF6FF774F09FD8D793B" bold="true" box="[151,621,793,819]" pageId="6" pageNumber="7">
<emphasis id="E2AFEACA553FFFF6FF774F09FE31793B" bold="true" box="[151,465,793,819]" italics="true" pageId="6" pageNumber="7">Macrophthalmus dentipes</emphasis>
<bibRefCitation id="B44A4B29553FFFF6FE394F09FD8D793B" author="Lucas" box="[473,621,793,819]" pageId="6" pageNumber="7" refString="Lucas, H. (1836) Macrophthalme, Macrophthalmus. (Crust.) In: Guerin-Meneville, F. E.: Dictionaire pittoresque d'Histoire naturelle, vol. 4, pp. 551, pl. 315, fig. 5." type="book chapter" year="1836">Lucas, 1836</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D06436D8553FFFF6FF774F2CFE4F795C" blockId="6.[151,621,793,852]" box="[151,431,828,852]" pageId="6" pageNumber="7">
(
<figureCitation id="48E02A5D553FFFF6FF7F4F2CFF0D795C" box="[159,237,828,852]" captionStart="FIGURE 4" captionStartId="7.[151,250,1906,1929]" captionTargetBox="[180,1393,655,1863]" captionTargetId="figure@7.[151,1436,601,1885]" captionTargetPageId="7" captionText="FIGURE 4. Macrophthalmus dentipes Lucas, 1836: a f, male (SMF 36863); f, male (SMF 26095). a, posterior surface of carapace, left side; b, lower orbital margin of male, right; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, merus of male cheliped, inner surface; f, cheliped of male outer surface." httpUri="https://zenodo.org/record/203102/files/figure.png" pageId="6" pageNumber="7">Figs. 4</figureCitation>
ae, 5af, 10ab)
</paragraph>
</subSubSection>
<subSubSection id="98C16553553FFFF6FF774F6FFAA87E07" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="D06436D8553FFFF6FF774F6FFCA3799C" blockId="6.[151,1437,895,1039]" box="[151,835,895,917]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="F0CB11F6553FFFF6FF774F6FFCA3799C" box="[151,835,895,917]" pageId="6" pageNumber="7">
<taxonomicName id="17DB4D5B553FFFF6FF774F6FFDB0799D" ID-CoL="84GMD" authority="Lucas 1836: 551" authorityName="Lucas" authorityPageNumber="551" authorityYear="1836" box="[151,592,895,917]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="E2AFEACA553FFFF6FF774F6FFE7D799D" box="[151,413,895,917]" italics="true" pageId="6" pageNumber="7">Macrophthalmus dentipes</emphasis>
<treatmentCitation id="517A10C9553FFFF6FE444F90FDB0799D" author="Lucas" box="[420,592,895,917]" page="551" pageId="6" pageNumber="7" year="1836">
<bibRefCitation id="B44A4B29553FFFF6FE444F90FDFD799D" author="Lucas" box="[420,541,895,917]" pageId="6" pageNumber="7" refString="Lucas, H. (1836) Macrophthalme, Macrophthalmus. (Crust.) In: Guerin-Meneville, F. E.: Dictionaire pittoresque d'Histoire naturelle, vol. 4, pp. 551, pl. 315, fig. 5." type="book chapter" year="1836">Lucas 1836</bibRefCitation>
: 551
</treatmentCitation>
</taxonomicName>
. —
<treatmentCitation id="517A10C9553FFFF6FD994F6FFCDF799C" author="Holthuis" box="[633,831,895,917]" page="401" pageId="6" pageNumber="7" year="1995">
<bibRefCitation id="B44A4B29553FFFF6FD994F6FFCEC799D" author="Holthuis" box="[633,780,895,917]" pageId="6" pageNumber="7" refString="Holthuis, L. B. (1995) The identities of Macrophthalmus rouxii Lucas, 1836, and M. dentipes Lucas, 1836, and the substitution of the latter name for M. pectinipes Guerin, 1838 (Decapoda, Brachyura, Ocypodidae). Crustaceana, 68 (3), 401 - 403." type="journal article" year="1995">Holthuis 1995</bibRefCitation>
: 401
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D06436D8553FFFF6FF774F8EFD2B79F8" blockId="6.[151,1437,895,1039]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="F0CB11F6553FFFF6FF774F8EFD2B79F8" pageId="6" pageNumber="7">
<taxonomicName id="17DB4D5B553FFFF6FF774F8EFE5179BC" ID-CoL="048dae79-c43f-4f0f-8339-a2d8d2b5a8cf" box="[151,433,926,948]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pectinipes">
<emphasis id="E2AFEACA553FFFF6FF774F8EFE5179BC" box="[151,433,926,948]" italics="true" pageId="6" pageNumber="7">Macrophthalmus pectinipes</emphasis>
</taxonomicName>
— Guerin-Méneville 1838: 1, pl. 23. —
<treatmentCitation id="517A10C9553FFFF6FC804F8DFBFB79BB" author="Alcock" box="[864,1051,925,947]" page="377" pageId="6" pageNumber="7" year="1900">
<bibRefCitation id="B44A4B29553FFFF6FC804F8DFC0779BB" author="Alcock" box="[864,999,925,947]" pageId="6" pageNumber="7" refString="Alcock, A. (1900) Material for a carcinological fauna of India. No. 6: The Brachyura Catometopa, or Grapsoidea. Journal of the Asiatic Society of Bengal, 69 (2), 279 - 456." type="journal article" year="1900">Alcock 1900</bibRefCitation>
: 377
</treatmentCitation>
. — Chhapgar 1957b: 512. —
<treatmentCitation id="517A10C9553FFFF6FAB64F8EFECF79D9" author="Barnes" page="237" pageId="6" pageNumber="7" year="1970">
<bibRefCitation id="B44A4B29553FFFF6FAB64F8EFF1D79D9" author="Barnes" pageId="6" pageNumber="7" refString="Barnes, R. S. K. (1970) The species of Macrophthalmus in the collections of the British Museum (Natural History). Bulletin of the British Museum of Natural History (Zoology), 20, 203 - 251." type="journal article" year="1970">Barnes 1970</bibRefCitation>
: 237
</treatmentCitation>
, fig. 10. —
<treatmentCitation id="517A10C9553FFFF6FE474FADFD9C79D9" author="Pretzmann" box="[423,636,956,978]" page="31" pageId="6" pageNumber="7" year="1971">
<bibRefCitation id="B44A4B29553FFFF6FE474FADFDB679D9" author="Pretzmann" box="[423,598,956,978]" pageId="6" pageNumber="7" refString="Pretzmann, G. (1971) Ergebnisse einiger Sammelreisen nach Vorderasien. 2. Teil: Marine Brachyura. Annalen des Naturhistorischen Museums Wien, 75, 477 - 487." type="journal article" year="1971">Pretzmann, 1971</bibRefCitation>
: 31
</treatmentCitation>
; 1974: 442. — Tirmizi 1981: 109. —
<treatmentCitation id="517A10C9553FFFF6FC1B4FACFB4879DA" author="Titgen" box="[1019,1192,956,978]" page="253" pageId="6" pageNumber="7" year="1982">
<bibRefCitation id="B44A4B29553FFFF6FC1B4FACFB9879D9" author="Titgen" box="[1019,1144,956,978]" pageId="6" pageNumber="7" refString="Titgen, R. H. (1982) The Systematics and Ecology of the Decapods of Dubai, and their Zoogeographic Relationships to the Arabian Gulf and the Western Indian Ocean. Unpublished D. Phil. Thesis, Texas A &amp; M University." type="book" year="1982">Titgen 1982</bibRefCitation>
: 253
</treatmentCitation>
(in list). —
<treatmentCitation id="517A10C9553FFFF6FAC44FADFF0F79F8" author="Jones" page="159" pageId="6" pageNumber="7" year="1986">
<bibRefCitation id="B44A4B29553FFFF6FAC44FADFA7879DA" author="Jones" box="[1316,1432,956,978]" pageId="6" pageNumber="7" refString="Jones, D. A. (1986) A Field Guide to the Sea Shores of Kuwait and the Arabian Gulf. University of Kuwait, Blandford Press, Poole, Kuwait, 192 pp." type="book" year="1986">Jones 1986</bibRefCitation>
: 159
</treatmentCitation>
, pl. 45. —
<treatmentCitation id="517A10C9553FFFF6FEBF4FCAFD9779F8" author="Tirmizi" box="[351,631,986,1008]" page="121" pageId="6" pageNumber="7" year="1996">
<bibRefCitation id="B44A4B29553FFFF6FEBF4FCAFDA579F8" author="Tirmizi" box="[351,581,986,1008]" pageId="6" pageNumber="7" refString="Tirmizi, N. M. &amp; Ghani, N. (1996) Marine Fauna of Pakistan: 5: Crustacea: Brachyura, Brachyrhyncha, Part 1: Xanthidae, Goneplacidae, Pinnotheridae, Ocypodidae, Grapsidae. Center of Excellence, University of Karachi, 1 - 188." type="journal article" year="1996">Tirmizi &amp; Ghani 1996</bibRefCitation>
: 121
</treatmentCitation>
, fig. 46.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D06436D8553FFFF6FF774FE9FAA87E07" blockId="6.[151,1437,895,1039]" box="[151,1352,1017,1039]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="F0CB11F6553FFFF6FF774FE9FAA87E07" box="[151,1352,1017,1039]" pageId="6" pageNumber="7">
<taxonomicName id="17DB4D5B553FFFF6FF774FE9FE1E7E07" ID-CoL="84GMD" box="[151,510,1017,1039]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="dentipes" subGenus="Venitus">
<emphasis id="E2AFEACA553FFFF6FF774FE9FEA37E07" box="[151,323,1017,1039]" italics="true" pageId="6" pageNumber="7">Macrophthalmus</emphasis>
(
<emphasis id="E2AFEACA553FFFF6FEB24FEAFE7B7E07" box="[338,411,1018,1039]" italics="true" pageId="6" pageNumber="7">Venitus</emphasis>
)
<emphasis id="E2AFEACA553FFFF6FE494FE9FE1E7E07" box="[425,510,1017,1039]" italics="true" pageId="6" pageNumber="7">dentipes</emphasis>
</taxonomicName>
<treatmentCitation id="517A10C9553FFFF6FDC44FE9FCCE7E07" author="Apel" box="[548,814,1017,1039]" page="135" pageId="6" pageNumber="7" year="1999">
<bibRefCitation id="B44A4B29553FFFF6FDC44FE9FD1B7E07" author="Apel" box="[548,763,1017,1039]" pageId="6" pageNumber="7" refString="Apel, M. &amp; Turkay, M. (1999) Taxonomic composition, distribution and zoogeographic relationships of the grapsid and ocypodid crab fauna of intertidal soft bottoms in the Arabian Gulf. Estuarine, Coastal and Shelf Science, 49 (suppl. A), 131 - 142." type="journal article" year="1999">Apel &amp; Türkay 1999</bibRefCitation>
: 135
</treatmentCitation>
. —
<treatmentCitation id="517A10C9553FFFF6FCB74FE9FC167E07" author="Apel" box="[855,1014,1017,1039]" page="110" pageId="6" pageNumber="7" year="2001">
<bibRefCitation id="B44A4B29553FFFF6FCB74FE9FC247E07" author="Apel" box="[855,964,1017,1039]" pageId="6" pageNumber="7" refString="Apel, M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguridea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes unpublished Ph. D. Thesis, Johann Wolfgang Goethe-Universitat, Frankfurt am Main 268 pp." type="book" year="2001">Apel 2001</bibRefCitation>
: 110
</treatmentCitation>
. —
<treatmentCitation id="517A10C9553FFFF6FBFF4FEAFB257E07" author="Barnes" box="[1055,1221,1018,1039]" page="33" pageId="6" pageNumber="7" year="2010">
<bibRefCitation id="B44A4B29553FFFF6FBFF4FEAFB427E07" author="Barnes" box="[1055,1186,1018,1039]" pageId="6" pageNumber="7" refString="Barnes, R. S. K. (2010) A review of the sentinel and allied crabs (Crustacea: Brachyura: Macrophthalmidae), with particular references to the genus Macrophthalmus. Raffles Bulletin of Zoology, 58 (1), 31 - 49." type="journal article" year="2010">Barnes 2010</bibRefCitation>
: 33
</treatmentCitation>
(in key), 43.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="98C16553553FFFF6FF774827FC4A7D14" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="D06436D8553FFFF6FF774827FD987E47" blockId="6.[151,1437,1078,2036]" box="[151,632,1078,1103]" pageId="6" pageNumber="7">
<emphasis id="E2AFEACA553FFFF6FF774827FED57E47" bold="true" box="[151,309,1078,1103]" pageId="6" pageNumber="7">
<typeStatus id="0F60887A553FFFF6FF774827FF327E47" box="[151,210,1079,1103]" pageId="6" pageNumber="7">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="A8CC7648553FFFF6FEDC4827FE7B7E47" box="[316,411,1079,1103]" name="India" pageId="6" pageNumber="7">Bombay</collectingCountry>
(=
<collectingCountry id="A8CC7648553FFFF6FE224827FDCC7E47" box="[450,556,1079,1103]" name="India" pageId="6" pageNumber="7">Mumbay</collectingCountry>
),
<collectingCountry id="A8CC7648553FFFF6FDDE4827FD987E47" box="[574,632,1079,1103]" name="India" pageId="6" pageNumber="7">India</collectingCountry>
</paragraph>
<paragraph id="D06436D8553FFFF6FF254847FD647C4C" blockId="6.[151,1437,1078,2036]" pageId="6" pageNumber="7">
<emphasis id="E2AFEACA553FFFF6FF254847FE4F7E78" bold="true" box="[197,431,1111,1136]" pageId="6" pageNumber="7">Material examined.</emphasis>
<emphasis id="E2AFEACA553FFFF6FE574849FDA87E78" box="[439,584,1112,1136]" italics="true" pageId="6" pageNumber="7">Persian Gulf</emphasis>
:
<emphasis id="E2AFEACA553FFFF6FDB84849FD687E78" box="[600,648,1113,1136]" italics="true" pageId="6" pageNumber="7">
<collectingCountry id="A8CC7648553FFFF6FDB84849FD687E78" box="[600,648,1113,1136]" name="Iran" pageId="6" pageNumber="7">Iran</collectingCountry>
</emphasis>
:
<specimenCount id="C6DDFD51553FFFF6FD784848FD0F7E78" box="[664,751,1112,1136]" pageId="6" pageNumber="7" type="male">3 males</specimenCount>
(
<collectionCode id="B6CAAE1D553FFFF6FD1F4848FCAA7E78" box="[767,842,1112,1136]" pageId="6" pageNumber="7">ZUTC</collectionCode>
Brach1163), Hormozgan, Bandar-Khamir,
<geoCoordinate id="B5EF501F553FFFF6FAD34848FA787E78" box="[1331,1432,1112,1136]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="28.8">28º 48'N</geoCoordinate>
,
<geoCoordinate id="B5EF501F553FFFF6FF77486CFF1C7E9C" box="[151,252,1148,1172]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="55.633335">55º 38'E</geoCoordinate>
, mangroves,
<date id="A4651018553FFFF6FE7A486CFDF97E9C" box="[410,537,1148,1172]" pageId="6" pageNumber="7" value="2005-11-31">31.11.2005</date>
, R. Naderloo;
<specimenCount id="C6DDFD51553FFFF6FD2A486CFCDB7E9C" box="[714,827,1148,1172]" pageId="6" pageNumber="7" type="female">3 females</specimenCount>
(
<collectionCode id="B6CAAE1D553FFFF6FCAE486CFC797E9C" box="[846,921,1148,1172]" pageId="6" pageNumber="7">ZUTC</collectionCode>
Brach1172), Hormozgan, Khor-Khalasi,
<geoCoordinate id="B5EF501F553FFFF6FA97486CFF2E7EB0" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="25.583334">25º 35'N</geoCoordinate>
,
<geoCoordinate id="B5EF501F553FFFF6FF3C48B0FEDE7EB0" box="[220,318,1184,1208]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="58.133335">58º 08'E</geoCoordinate>
, mud flat behind mangroves,
<date id="A4651018553FFFF6FD7648B0FCF77EB0" box="[662,791,1184,1208]" pageId="6" pageNumber="7" value="2006-10-26">26.10.2006</date>
, R. Naderloo;
<specimenCount id="C6DDFD51553FFFF6FC2048B0FBF87EB0" box="[960,1048,1184,1208]" pageId="6" pageNumber="7" type="male">4 males</specimenCount>
(
<collectionCode id="B6CAAE1D553FFFF6FBC948B0FB947EB0" box="[1065,1140,1184,1208]" pageId="6" pageNumber="7">ZUTC</collectionCode>
Brach1186), Hormozgan, Bandar-Khamir,
<geoCoordinate id="B5EF501F553FFFF6FEB448D4FE597ED4" box="[340,441,1220,1244]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="28.8">28º 48'N</geoCoordinate>
,
<geoCoordinate id="B5EF501F553FFFF6FE2648D4FDC77ED4" box="[454,551,1220,1244]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="55.633335">55º 38'E</geoCoordinate>
, muddy sandy shore, east of fishery jetty,
<date id="A4651018553FFFF6FBE748D4FB687ED4" box="[1031,1160,1220,1244]" pageId="6" pageNumber="7" value="2006-04-14">14.04.2006</date>
, R. Naderloo;
<specimenCount id="C6DDFD51553FFFF6FACF48D5FA7C7ED4" box="[1327,1436,1220,1244]" pageId="6" pageNumber="7" type="female">7 females</specimenCount>
(ovigerous) (
<collectionCode id="B6CAAE1D553FFFF6FEC748F8FE927F08" box="[295,370,1256,1280]" pageId="6" pageNumber="7">ZUTC</collectionCode>
Brach1210), Khuzestan, Mahshahr, Fishery jetty (Majidieh),
<geoCoordinate id="B5EF501F553FFFF6FBC848F8FB6C7F08" box="[1064,1164,1256,1280]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="30.466667">30º 28'N</geoCoordinate>
,
<geoCoordinate id="B5EF501F553FFFF6FB7848F8FB177F08" box="[1176,1271,1256,1280]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="49.183334">49º 11'E</geoCoordinate>
, muddy shore,
<date id="A4651018553FFFF6FF77491CFEF77F2C" box="[151,279,1292,1316]" pageId="6" pageNumber="7" value="2006-04-17">17.04.2006</date>
, R. Naderloo;
<specimenCount id="C6DDFD51553FFFF6FE2C491CFDCB7F2C" box="[460,555,1292,1316]" pageId="6" pageNumber="7" type="male">4 males</specimenCount>
,
<specimenCount id="C6DDFD51553FFFF6FDDC491CFD4E7F2C" box="[572,686,1292,1316]" pageId="6" pageNumber="7" type="female">2 females</specimenCount>
(
<collectionCode id="B6CAAE1D553FFFF6FD23491CFCEE7F2C" box="[707,782,1292,1316]" pageId="6" pageNumber="7">ZUTC</collectionCode>
Brach1211), Khuzestan, Shah Abdollah,
<geoCoordinate id="B5EF501F553FFFF6FB1D491CFA857F2C" box="[1277,1381,1292,1316]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="30.166666">30º 10'N</geoCoordinate>
,
<geoCoordinate id="B5EF501F553FFFF6FA97491CFF2B7F40" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="50.083332">50º 05'E</geoCoordinate>
, sandy mud-flat,
<date id="A4651018553FFFF6FE714920FDF17F40" box="[401,529,1328,1352]" pageId="6" pageNumber="7" value="2006-04-14">14.04.2006</date>
, R. Naderloo;
<specimenCount id="C6DDFD51553FFFF6FD594920FCF47F40" box="[697,788,1328,1352]" pageId="6" pageNumber="7" type="male">3 males</specimenCount>
,
<specimenCount id="C6DDFD51553FFFF6FCC14920FC6F7F40" box="[801,911,1328,1352]" pageId="6" pageNumber="7" type="female">2 females</specimenCount>
(
<collectionCode id="B6CAAE1D553FFFF6FC7F4920FC0A7F40" box="[927,1002,1328,1352]" pageId="6" pageNumber="7">ZUTC</collectionCode>
Brach1238), Khuzestan, Shah Abdollah,
<geoCoordinate id="B5EF501F553FFFF6FF274944FEC97F64" box="[199,297,1364,1388]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="30.166666">30º 10'N</geoCoordinate>
,
<geoCoordinate id="B5EF501F553FFFF6FED54944FE747F64" box="[309,404,1364,1388]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="50.083332">50º 05'E</geoCoordinate>
, sandy mud-flat,
<date id="A4651018553FFFF6FDB64944FD377F64" box="[598,727,1364,1388]" pageId="6" pageNumber="7" value="2006-04-14">14.04.2006</date>
, R. Naderloo;
<specimenCount id="C6DDFD51553FFFF6FC994944FC247F64" box="[889,964,1364,1388]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL = 28.35 CB = 28.15),
<specimenCount id="C6DDFD51553FFFF6FB134944FAB47F64" box="[1267,1364,1364,1388]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(
<collectionCode id="B6CAAE1D553FFFF6FA824944FA7C7F64" LSID="urn:lsid:biocol.org:col:34838" box="[1378,1436,1364,1388]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="7">SMF</collectionCode>
36863),
<collectingCountry id="A8CC7648553FFFF6FF174968FE7B7F98" box="[247,411,1400,1424]" name="Iran" pageId="6" pageNumber="7">Bandar-Abbas</collectingCountry>
, E. of city,
<geoCoordinate id="B5EF501F553FFFF6FDC64968FD6D7F98" box="[550,653,1400,1424]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="27.133333">27º 08'N</geoCoordinate>
,
<geoCoordinate id="B5EF501F553FFFF6FD7C4968FD1F7F98" box="[668,767,1400,1424]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="56.333332">56º 20'E</geoCoordinate>
, muddy sand flat with shell,
<date id="A4651018553FFFF6FBB54968FB357F98" box="[1109,1237,1400,1424]" pageId="6" pageNumber="7" value="2008-04-23">23.04.2008</date>
, R. Naderloo, A. Kazemi &amp; A. Keykhosravi;
<specimenCount id="C6DDFD51553FFFF6FE3C498CFDD77FBC" box="[476,567,1436,1460]" pageId="6" pageNumber="7" type="male">3 males</specimenCount>
,
<specimenCount id="C6DDFD51553FFFF6FDA5498CFD537FBC" box="[581,691,1436,1460]" pageId="6" pageNumber="7" type="female">3 females</specimenCount>
(
<collectionCode id="B6CAAE1D553FFFF6FD25498CFD1F7FBC" LSID="urn:lsid:biocol.org:col:34838" box="[709,767,1436,1460]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="7">SMF</collectionCode>
36866), Bandar-Khamir, E. of fishery Jetty,
<geoCoordinate id="B5EF501F553FFFF6FAE3498CFA8A7FBC" box="[1283,1386,1436,1460]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="26.933332">26º 56'N</geoCoordinate>
,
<geoCoordinate id="B5EF501F553FFFF6FA98498CFF2B7FD0" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="55.6">55º 36'E</geoCoordinate>
, Khor, muddy flat,
<date id="A4651018553FFFF6FE4349D0FDC37FD0" box="[419,547,1472,1496]" pageId="6" pageNumber="7" value="2008-04-24">24.04.2008</date>
, R. Naderloo, A. Kazemi &amp; A. Keykhosravi;
<specimenCount id="C6DDFD51553FFFF6FBFE49D0FB897FD0" box="[1054,1129,1472,1496]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(
<collectionCode id="B6CAAE1D553FFFF6FB9749D0FB517FD0" LSID="urn:lsid:biocol.org:col:34838" box="[1143,1201,1472,1496]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="7">SMF</collectionCode>
36864)
<collectingCountry id="A8CC7648553FFFF6FAED49D0FABC7FD0" box="[1293,1372,1472,1496]" name="Iran" pageId="6" pageNumber="7">Qeshm</collectingCountry>
I., N. coast,
<quantity id="17239B3D553FFFF6FF0249F4FEFE7FF4" box="[226,286,1508,1532]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="7" unit="km" value="3.0">3 km</quantity>
W. of Kuweii,
<geoCoordinate id="B5EF501F553FFFF6FE3849F4FDDF7FF4" box="[472,575,1508,1532]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="26.95">26º 57'N</geoCoordinate>
,
<geoCoordinate id="B5EF501F553FFFF6FDAD49F4FD517FF4" box="[589,689,1508,1532]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="56.0">56º 00'E</geoCoordinate>
, muddy flat,
<date id="A4651018553FFFF6FCAF49F4FC307FF4" box="[847,976,1508,1532]" pageId="6" pageNumber="7" value="2008-01-13">13.01.2008</date>
, R. Naderloo &amp; M. Türkay;
<specimenCount id="C6DDFD51553FFFF6FAC349F4FA9F7FF4" box="[1315,1407,1508,1532]" pageId="6" pageNumber="7" type="male">2 males</specimenCount>
,
<specimenCount id="C6DDFD51553FFFF6FA6E49F4FF067C28" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
,
<specimenCount id="C6DDFD51553FFFF6FF104A18FEDC7C28" box="[240,316,1544,1568]" pageId="6" pageNumber="7" type="generic">10 juv.</specimenCount>
(
<collectionCode id="B6CAAE1D553FFFF6FEAA4A18FE647C28" LSID="urn:lsid:biocol.org:col:34838" box="[330,388,1544,1568]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="7">SMF</collectionCode>
36865), Bandar-Khamir, E. of fishery Jetty,
<geoCoordinate id="B5EF501F553FFFF6FC924A18FC357C28" box="[882,981,1544,1568]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="26.933332">26º 56'N</geoCoordinate>
,
<geoCoordinate id="B5EF501F553FFFF6FC004A18FBDF7C28" box="[992,1087,1544,1568]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="55.6">55º 36'E</geoCoordinate>
, Khor, muddy flat,
<date id="A4651018553FFFF6FAF74A18FA777C28" box="[1303,1431,1544,1568]" pageId="6" pageNumber="7" value="2008-04-24">24.04.2008</date>
, R. Naderloo, A. Kazemi &amp; A. Keykhosravi.
</paragraph>
<paragraph id="D06436D8553FFFF6FF254A41FBB17C84" blockId="6.[151,1437,1078,2036]" pageId="6" pageNumber="7">
<emphasis id="E2AFEACA553FFFF6FF254A41FEF67C60" box="[197,278,1617,1640]" italics="true" pageId="6" pageNumber="7">
<collectingCountry id="A8CC7648553FFFF6FF254A41FEF67C60" box="[197,278,1617,1640]" name="Kuwait" pageId="6" pageNumber="7">Kuwait</collectingCountry>
</emphasis>
:
<specimenCount id="C6DDFD51553FFFF6FEC84A40FE977C60" box="[296,375,1616,1640]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(NHM 1978.194),
<collectingCountry id="A8CC7648553FFFF6FDBB4A40FD4D7C60" box="[603,685,1616,1640]" name="Kuwait" pageId="6" pageNumber="7">Kuwait</collectingCountry>
Bay, west coast of Sulaibikhat, near mangroves,
<date id="A4651018553FFFF6FB114A40FA917C60" box="[1265,1393,1616,1640]" pageId="6" pageNumber="7" value="1978-04-16">16.04.1978</date>
, D. Clayton;
<specimenCount id="C6DDFD51553FFFF6FF1E4A64FEBF7C84" box="[254,351,1652,1676]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(NHM 1978.195), mud flat creek at Al-Jahra, Clayton &amp; Vaughan.
</paragraph>
<paragraph id="D06436D8553FFFF6FF254A89FB7A7CB8" blockId="6.[151,1437,1078,2036]" box="[197,1178,1688,1712]" pageId="6" pageNumber="7">
<collectingCountry id="A8CC7648553FFFF6FF254A89FF157CB8" box="[197,245,1689,1712]" name="Iraq" pageId="6" pageNumber="7">
<emphasis id="E2AFEACA553FFFF6FF254A89FF157CB8" box="[197,245,1689,1712]" italics="true" pageId="6" pageNumber="7">Iraq</emphasis>
</collectingCountry>
:
<specimenCount id="C6DDFD51553FFFF6FEE44A88FEBA7CB8" box="[260,346,1688,1712]" pageId="6" pageNumber="7" type="male">4 males</specimenCount>
(NHM 1892.9.16.2-6), Fao, W.D. Cumming, examined by
<bibRefCitation id="B44A4B29553FFFF6FC144A89FB777CB8" author="Barnes" box="[1012,1175,1688,1712]" pageId="6" pageNumber="7" refString="Barnes, R. S. K. (1971) Biological results of the Snellius Expedition. XXIII. The Genus Macrophthalmus (Decapoda, Brachyura). Zoologische Verhandelingen, 115, 1 - 40." type="journal article" year="1971">Barnes (1971)</bibRefCitation>
.
</paragraph>
<paragraph id="D06436D8553FFFF6FF254AACFC4A7D14" blockId="6.[151,1437,1078,2036]" pageId="6" pageNumber="7">
<emphasis id="E2AFEACA553FFFF6FF254AACFE377CDD" bold="true" box="[197,471,1724,1749]" pageId="6" pageNumber="7">Comparative material.</emphasis>
<emphasis id="E2AFEACA553FFFF6FE004AACFD8D7CDC" box="[480,621,1724,1748]" italics="true" pageId="6" pageNumber="7">Arabian Sea</emphasis>
:
<specimenCount id="C6DDFD51553FFFF6FD9D4AACFD2D7CDC" box="[637,717,1724,1748]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
,
<specimenCount id="C6DDFD51553FFFF6FD3B4AACFCA87CDC" box="[731,840,1724,1748]" pageId="6" pageNumber="7" type="female">3 females</specimenCount>
(
<collectionCode id="B6CAAE1D553FFFF6FCB94AACFC737CDC" LSID="urn:lsid:biocol.org:col:34838" box="[857,915,1724,1748]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="7">SMF</collectionCode>
26095),
<collectingCountry id="A8CC7648553FFFF6FC1A4AACFBA17CDC" box="[1018,1089,1724,1748]" name="Oman" pageId="6" pageNumber="7">Oman</collectingCountry>
, Filim, W. of Barr al-Hikman,
<geoCoordinate id="B5EF501F553FFFF6FF774AF0FED47CF0" box="[151,308,1760,1784]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="1" value="20.609867">20º 36.592'N</geoCoordinate>
,
<geoCoordinate id="B5EF501F553FFFF6FEA84AF0FE017CF0" box="[328,481,1760,1784]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="58.18695">58º 11.217'E</geoCoordinate>
, muddy flat and mangroves,
<date id="A4651018553FFFF6FCB14AF0FC317CF0" box="[849,977,1760,1784]" pageId="6" pageNumber="7" value="2001-01-14">14.01.2001</date>
, M. Apel;
<specimenCount id="C6DDFD51553FFFF6FB824AF0FB247CF0" box="[1122,1220,1760,1784]" pageId="6" pageNumber="7" type="male">4 males</specimenCount>
,
<specimenCount id="C6DDFD51553FFFF6FB384AF0FAA17CF0" box="[1240,1345,1760,1784]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(NHM 1889:6:17:83-87),
<collectingCountry id="A8CC7648553FFFF6FE884B14FE2A7D14" box="[360,458,1796,1820]" name="Pakistan" pageId="6" pageNumber="7">Pakistan</collectingCountry>
, Sind, F. Day, examined by
<bibRefCitation id="B44A4B29553FFFF6FCE34B15FC477D14" author="Barnes" box="[771,935,1796,1820]" pageId="6" pageNumber="7" refString="Barnes, R. S. K. (1971) Biological results of the Snellius Expedition. XXIII. The Genus Macrophthalmus (Decapoda, Brachyura). Zoologische Verhandelingen, 115, 1 - 40." type="journal article" year="1971">Barnes (1971)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="98C16553553FFFF9FF254B38FC3F7F5C" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7" type="description">
<paragraph id="D06436D8553FFFF6FF254B38FBAF7DFC" blockId="6.[151,1437,1078,2036]" pageId="6" pageNumber="7">
<emphasis id="E2AFEACA553FFFF6FF254B38FE937D49" bold="true" box="[197,371,1832,1857]" pageId="6" pageNumber="7">Redescription.</emphasis>
Carapace (
<figureCitation id="48E02A5D553FFFF6FE104B38FDD37D48" box="[496,563,1832,1856]" captionStart="FIGURE 4" captionStartId="7.[151,250,1906,1929]" captionTargetBox="[180,1393,655,1863]" captionTargetId="figure@7.[151,1436,601,1885]" captionTargetPageId="7" captionText="FIGURE 4. Macrophthalmus dentipes Lucas, 1836: a f, male (SMF 36863); f, male (SMF 26095). a, posterior surface of carapace, left side; b, lower orbital margin of male, right; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, merus of male cheliped, inner surface; f, cheliped of male outer surface." httpUri="https://zenodo.org/record/203102/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
a) moderately wider than long (CB/CL = 1.61.7), slightly convex; large granules scattered on entire posterior surface except in narrow median, frontal regions, granules on branchial regions high, longitudinal row of 68 high granules on epibranchial region, granules on median regions low, round. Regions well defined; furrows wide, moderately deep, normally beset with dense setae. Frontal region remarkably deflexed downward, smooth; front very narrow, about 0.060.07 times as wide as carapace, strongly constricted medially; anterior edge remarkably concave, bi-lobed, frontal furrow markedly deep.
</paragraph>
<paragraph id="D06436D8553EFFF7FF254C87FEA17B37" blockId="7.[151,1436,151,572]" pageId="7" pageNumber="8">
Lateral margin of carapace (
<figureCitation id="48E02A5D553EFFF7FDE54C87FDAA7AA7" box="[517,586,151,175]" captionStart="FIGURE 4" captionStartId="7.[151,250,1906,1929]" captionTargetBox="[180,1393,655,1863]" captionTargetId="figure@7.[151,1436,601,1885]" captionTargetPageId="7" captionText="FIGURE 4. Macrophthalmus dentipes Lucas, 1836: a f, male (SMF 36863); f, male (SMF 26095). a, posterior surface of carapace, left side; b, lower orbital margin of male, right; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, merus of male cheliped, inner surface; f, cheliped of male outer surface." httpUri="https://zenodo.org/record/203102/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
a) with 3 distinct teeth (including exorbital tooth); first nearly subquadrate, with posterior margin smooth, curved forward; second triangular, with smooth margin, higher than first, greatest width of carapace between second lateral teeth; third very small, directed forwards; posterolateral margin nearly straight, slightly converging posteriorly, with small granules, beset with long setae; posterior margin with very small granules.
</paragraph>
<paragraph id="D06436D8553EFFF7FF254D5CFB8B7BA4" blockId="7.[151,1436,151,572]" pageId="7" pageNumber="8">
Eyestalks narrow (
<figureCitation id="48E02A5D553EFFF7FE774D5CFE3C7B6C" box="[407,476,332,356]" captionStart="FIGURE 4" captionStartId="7.[151,250,1906,1929]" captionTargetBox="[180,1393,655,1863]" captionTargetId="figure@7.[151,1436,601,1885]" captionTargetPageId="7" captionText="FIGURE 4. Macrophthalmus dentipes Lucas, 1836: a f, male (SMF 36863); f, male (SMF 26095). a, posterior surface of carapace, left side; b, lower orbital margin of male, right; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, merus of male cheliped, inner surface; f, cheliped of male outer surface." httpUri="https://zenodo.org/record/203102/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
a), long, but not reaching to exorbital angle; upper orbital margin fairly convex, regularly granular, granules large, high, pointed, directed laterally; lower orbital margin (
<figureCitation id="48E02A5D553EFFF7FBB74D7FFB7B7B8F" box="[1111,1179,367,391]" captionStart="FIGURE 4" captionStartId="7.[151,250,1906,1929]" captionTargetBox="[180,1393,655,1863]" captionTargetId="figure@7.[151,1436,601,1885]" captionTargetPageId="7" captionText="FIGURE 4. Macrophthalmus dentipes Lucas, 1836: a f, male (SMF 36863); f, male (SMF 26095). a, posterior surface of carapace, left side; b, lower orbital margin of male, right; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, merus of male cheliped, inner surface; f, cheliped of male outer surface." httpUri="https://zenodo.org/record/203102/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
b) with round granules on inner third, 56 long tubercles on outer portion of margin forming stridulating ridge.
</paragraph>
<paragraph id="D06436D8553EFFF7FF254DA7FDB37834" blockId="7.[151,1436,151,572]" pageId="7" pageNumber="8">Third maxillipeds leaving small gap between; ischium slightly more than twice the length of merus, inner margin of ischium, merus with long setae, outer margins with short setae, outer margin of ischium completely straight, outer margin of merus sharply sloping distally; outer surface of ischium, merus smooth, elliptical row of long setae proximally on outer surface of ischium.</paragraph>
<caption id="84A46650553EFFF7FF774B62FC907DCD" httpUri="https://zenodo.org/record/203102/files/figure.png" pageId="7" pageNumber="8" targetBox="[180,1393,655,1863]" targetPageId="7">
<paragraph id="D06436D8553EFFF7FF774B62FC907DCD" blockId="7.[151,1436,1906,1989]" pageId="7" pageNumber="8">
<emphasis id="E2AFEACA553EFFF7FF774B62FEF37D80" bold="true" box="[151,275,1906,1929]" pageId="7" pageNumber="8">FIGURE 4.</emphasis>
<taxonomicName id="17DB4D5B553EFFF7FEFA4B62FD477D80" authority="Lucas, 1836" authorityName="Lucas" authorityYear="1836" box="[282,679,1906,1928]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="E2AFEACA553EFFF7FEFA4B62FDC17D80" box="[282,545,1906,1928]" italics="true" pageId="7" pageNumber="8">Macrophthalmus dentipes</emphasis>
Lucas, 1836
</taxonomicName>
: af, male (SMF 36863); f, male (SMF 26095). a, posterior surface of carapace, left side; b, lower orbital margin of male, right; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, merus of male cheliped, inner surface; f, cheliped of male outer surface.
</paragraph>
</caption>
<caption id="84A466505531FFF8FF774B0CFC0D7D78" httpUri="https://zenodo.org/record/203103/files/figure.png" pageId="8" pageNumber="9" targetBox="[184,1375,215,1778]" targetPageId="8">
<paragraph id="D06436D85531FFF8FF774B0CFC0D7D78" blockId="8.[151,1436,1820,1904]" pageId="8" pageNumber="9">
<emphasis id="E2AFEACA5531FFF8FF774B0CFEF37D3A" bold="true" box="[151,275,1820,1843]" pageId="8" pageNumber="9">FIGURE 5.</emphasis>
<taxonomicName id="17DB4D5B5531FFF8FEFA4B0DFD477D3B" authority="Lucas, 1836" authorityName="Lucas" authorityYear="1836" box="[282,679,1821,1843]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="E2AFEACA5531FFF8FEFA4B0DFDC07D3B" box="[282,544,1821,1843]" italics="true" pageId="8" pageNumber="9">Macrophthalmus dentipes</emphasis>
Lucas, 1836
</taxonomicName>
: ad, male (SMF 36863); e, f, female (SMF 26095). a, G1, ventral surface (right); b, apical part of the same G1, dorsal surface (right); c, apical part of the same G1, ventral surface (right); d, male abdomen; e, female abdomen, penultimate segment and telson; f, female gonopore, right.
</paragraph>
</caption>
<paragraph id="D06436D85531FFF9FF254B8DFC8A7BC7" blockId="8.[151,1436,1949,2009]" lastBlockId="9.[151,1437,151,2012]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
Chelipeds nearly equal, sometimes subequal, remarkably long in large specimens (
<figureCitation id="48E02A5D5531FFF8FB9F4B8DFB257DBD" box="[1151,1221,1949,1973]" captionStart="FIGURE 4" captionStartId="7.[151,250,1906,1929]" captionTargetBox="[180,1393,655,1863]" captionTargetId="figure@7.[151,1436,601,1885]" captionTargetPageId="7" captionText="FIGURE 4. Macrophthalmus dentipes Lucas, 1836: a f, male (SMF 36863); f, male (SMF 26095). a, posterior surface of carapace, left side; b, lower orbital margin of male, right; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, merus of male cheliped, inner surface; f, cheliped of male outer surface." httpUri="https://zenodo.org/record/203102/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
f). Merus (
<figureCitation id="48E02A5D5531FFF8FAA14B8DFA677DBD" box="[1345,1415,1949,1973]" captionStart="FIGURE 4" captionStartId="7.[151,250,1906,1929]" captionTargetBox="[180,1393,655,1863]" captionTargetId="figure@7.[151,1436,601,1885]" captionTargetPageId="7" captionText="FIGURE 4. Macrophthalmus dentipes Lucas, 1836: a f, male (SMF 36863); f, male (SMF 26095). a, posterior surface of carapace, left side; b, lower orbital margin of male, right; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, merus of male cheliped, inner surface; f, cheliped of male outer surface." httpUri="https://zenodo.org/record/203102/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
e) long; inner upper margin with distinct stridulating ridge, large granules along it, larger distally, with scarce setae; inner lower margin with small granules, continuous with lower surface; upper margin with small granules on proximal portion, with scarce setae. Carpus smooth on outer surface, inner upper margin with denticles, distally larger, small granules along inner portion of these denticles. Palm long (
<figureCitation id="48E02A5D5530FFF9FC8E4CCFFC5D7AFF" box="[878,957,223,247]" captionStart="FIGURE 4" captionStartId="7.[151,250,1906,1929]" captionTargetBox="[180,1393,655,1863]" captionTargetId="figure@7.[151,1436,601,1885]" captionTargetPageId="7" captionText="FIGURE 4. Macrophthalmus dentipes Lucas, 1836: a f, male (SMF 36863); f, male (SMF 26095). a, posterior surface of carapace, left side; b, lower orbital margin of male, right; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, merus of male cheliped, inner surface; f, cheliped of male outer surface." httpUri="https://zenodo.org/record/203102/files/figure.png" pageId="9" pageNumber="10">Figs. 4</figureCitation>
c, d, f), about 1.82.3 times as long as high in proximal portion; outer surface smooth (
<figureCitation id="48E02A5D5530FFF9FD614D14FD317B14" box="[641,721,260,284]" captionStart="FIGURE 4" captionStartId="7.[151,250,1906,1929]" captionTargetBox="[180,1393,655,1863]" captionTargetId="figure@7.[151,1436,601,1885]" captionTargetPageId="7" captionText="FIGURE 4. Macrophthalmus dentipes Lucas, 1836: a f, male (SMF 36863); f, male (SMF 26095). a, posterior surface of carapace, left side; b, lower orbital margin of male, right; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, merus of male cheliped, inner surface; f, cheliped of male outer surface." httpUri="https://zenodo.org/record/203102/files/figure.png" pageId="9" pageNumber="10">Figs. 4</figureCitation>
c, f) without longitudinal ridge; inner surface (
<figureCitation id="48E02A5D5530FFF9FB3F4D14FAC47B14" box="[1247,1316,260,284]" captionStart="FIGURE 4" captionStartId="7.[151,250,1906,1929]" captionTargetBox="[180,1393,655,1863]" captionTargetId="figure@7.[151,1436,601,1885]" captionTargetPageId="7" captionText="FIGURE 4. Macrophthalmus dentipes Lucas, 1836: a f, male (SMF 36863); f, male (SMF 26095). a, posterior surface of carapace, left side; b, lower orbital margin of male, right; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, merus of male cheliped, inner surface; f, cheliped of male outer surface." httpUri="https://zenodo.org/record/203102/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
d) smooth, patch of dense setae on upper portion, parallel to upper margin; lower margin nearly smooth; upper margin with large granules on proximal portion, smaller distally. Fingers remarkably curved inward distally, movable finger with upper margin smooth, long setae densely along inner surface of upper margin, continuous on upper, outer surface, cutting edge with subproximal differentiated tooth, small teeth distally; immovable finger narrow, with relatively large teeth on cutting edge, long setae along inner surface.
</paragraph>
<paragraph id="D06436D85530FFF9FF254DCCFAE87857" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="10">Cheliped of females small; outer surface of palm smooth, longitudinal ridge short, extending from base of immovable finger to proximal half; upper margin with small granules, with long setae; lower margin smooth, with relatively short setae; fingers distinctly long, slightly less than 1.5 length of palm; cutting edge faintly tuberculate, chitinous distal margin long, more than half of length of finger, long setae on inner surface of fingers.</paragraph>
<paragraph id="D06436D85530FFF9FF254E7CFDCE7977" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="10">Walking legs moderately long; second, third longest, with margins of segments more granular than those of first, fourth. Merus long, that of third leg about 4 times as long as wide; merus of first, second, third walking legs with subdistal tooth on anterior margin, 34 large denticles distally on posterior margin, merus of females with large denticles on anterior margin. Carpus of first walking leg, with longitudinal ridge, that of second, third with two ridges of long granules on anterior margin, carpus of third with one distinct large denticle distally on posterior margin. Propodus of second with small denticles on anterior, posterior margins; propodus of third with small denticles on anterior margin, large, spine-shaped denticles on posterior margin; dense setae on upper portion of anterior surface of carpus, propodus of third.</paragraph>
<paragraph id="D06436D85530FFF9FF254F9CFA9B79AC" blockId="9.[151,1437,151,2012]" box="[197,1403,908,932]" pageId="9" pageNumber="10">Posteromedian margin of epistome nearly straight, short longitudinal ridge on anterior part of buccal cavity.</paragraph>
<paragraph id="D06436D85530FFF9FF254FBFFEBA7E3C" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="10">
Male abdomen (
<figureCitation id="48E02A5D5530FFF9FE614FBFFE2579CF" box="[385,453,943,967]" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1843]" captionTargetBox="[184,1375,215,1778]" captionTargetId="figure@8.[151,1436,193,1799]" captionTargetPageId="8" captionText="FIGURE 5. Macrophthalmus dentipes Lucas, 1836: a d, male (SMF 36863); e, f, female (SMF 26095). a, G 1, ventral surface (right); b, apical part of the same G 1, dorsal surface (right); c, apical part of the same G 1, ventral surface (right); d, male abdomen; e, female abdomen, penultimate segment and telson; f, female gonopore, right." httpUri="https://zenodo.org/record/203103/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
d) with segments 3, 4 of same length, slightly shorter than segment 5; segments 5, 6 nearly of same length, with lateral margins nearly straight; lateral margins of segment 6 with small depression at one third distal portion; telson very slightly longer than segment 6, lateral margins strongly converging distally, apically rounded.
</paragraph>
<paragraph id="D06436D85530FFF9FF25482FFEF07E74" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="10">
Female abdomen (
<figureCitation id="48E02A5D5530FFF9FE74482FFE367E5F" box="[404,470,1087,1111]" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1843]" captionTargetBox="[184,1375,215,1778]" captionTargetId="figure@8.[151,1436,193,1799]" captionTargetPageId="8" captionText="FIGURE 5. Macrophthalmus dentipes Lucas, 1836: a d, male (SMF 36863); e, f, female (SMF 26095). a, G 1, ventral surface (right); b, apical part of the same G 1, dorsal surface (right); c, apical part of the same G 1, ventral surface (right); d, male abdomen; e, female abdomen, penultimate segment and telson; f, female gonopore, right." httpUri="https://zenodo.org/record/203103/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
e) with large telson, slightly longer than segment 6, distinctly wider than half length of segment 6.
</paragraph>
<paragraph id="D06436D85530FFF9FF254898FD867F04" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="10">
Male G1 (
<figureCitation id="48E02A5D5530FFF9FEDC4898FE607EA8" box="[316,384,1160,1184]" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1843]" captionTargetBox="[184,1375,215,1778]" captionTargetId="figure@8.[151,1436,193,1799]" captionTargetPageId="8" captionText="FIGURE 5. Macrophthalmus dentipes Lucas, 1836: a d, male (SMF 36863); e, f, female (SMF 26095). a, G 1, ventral surface (right); b, apical part of the same G 1, dorsal surface (right); c, apical part of the same G 1, ventral surface (right); d, male abdomen; e, female abdomen, penultimate segment and telson; f, female gonopore, right." httpUri="https://zenodo.org/record/203103/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
a) moderately stout, slightly curved outward medially; distal half relatively narrowing, with apical chitinous process (
<figureCitation id="48E02A5D5530FFF9FE5748BCFDE67ECC" box="[439,518,1196,1220]" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1843]" captionTargetBox="[184,1375,215,1778]" captionTargetId="figure@8.[151,1436,193,1799]" captionTargetPageId="8" captionText="FIGURE 5. Macrophthalmus dentipes Lucas, 1836: a d, male (SMF 36863); e, f, female (SMF 26095). a, G 1, ventral surface (right); b, apical part of the same G 1, dorsal surface (right); c, apical part of the same G 1, ventral surface (right); d, male abdomen; e, female abdomen, penultimate segment and telson; f, female gonopore, right." httpUri="https://zenodo.org/record/203103/files/figure.png" pageId="9" pageNumber="10">Figs. 5</figureCitation>
b, c) remarkably long, narrow, curved outward at about 45°; distal opening (
<figureCitation id="48E02A5D5530FFF9FA8E48BCFF447EE0" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1843]" captionTargetBox="[184,1375,215,1778]" captionTargetId="figure@8.[151,1436,193,1799]" captionTargetPageId="8" captionText="FIGURE 5. Macrophthalmus dentipes Lucas, 1836: a d, male (SMF 36863); e, f, female (SMF 26095). a, G 1, ventral surface (right); b, apical part of the same G 1, dorsal surface (right); c, apical part of the same G 1, ventral surface (right); d, male abdomen; e, female abdomen, penultimate segment and telson; f, female gonopore, right." httpUri="https://zenodo.org/record/203103/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
b) large, distinct, subdistal on dorsal portion of apical process; long feather-shaped setae densely set along lateral margin, long setae around apical process.
</paragraph>
<paragraph id="D06436D85530FFF9FF254908FC3F7F5C" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="10">
Female gonopore (
<figureCitation id="48E02A5D5530FFF9FE7A4908FE3D7F38" box="[410,477,1304,1328]" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1843]" captionTargetBox="[184,1375,215,1778]" captionTargetId="figure@8.[151,1436,193,1799]" captionTargetPageId="8" captionText="FIGURE 5. Macrophthalmus dentipes Lucas, 1836: a d, male (SMF 36863); e, f, female (SMF 26095). a, G 1, ventral surface (right); b, apical part of the same G 1, dorsal surface (right); c, apical part of the same G 1, ventral surface (right); d, male abdomen; e, female abdomen, penultimate segment and telson; f, female gonopore, right." httpUri="https://zenodo.org/record/203103/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
f) on outer wall of depression, directed mesially, anterior margin distinctly rounded, elevated, outer margin slightly convex; operculum small, directed laterally.
</paragraph>
</subSubSection>
<subSubSection id="98C165535530FFF9FF254970FD607C7C" pageId="9" pageNumber="10" type="discussion">
<paragraph id="D06436D85530FFF9FF254970FD607C7C" blockId="9.[151,1437,151,2012]" pageId="9" pageNumber="10">
<emphasis id="E2AFEACA5530FFF9FF254970FED97F70" bold="true" box="[197,313,1376,1400]" pageId="9" pageNumber="10">Remarks.</emphasis>
<bibRefCitation id="B44A4B295530FFF9FEA44971FE097F70" author="Barnes" box="[324,489,1376,1400]" pageId="9" pageNumber="10" refString="Barnes, R. S. K. (1970) The species of Macrophthalmus in the collections of the British Museum (Natural History). Bulletin of the British Museum of Natural History (Zoology), 20, 203 - 251." type="journal article" year="1970">Barnes (1970)</bibRefCitation>
first recorded
<taxonomicName id="17DB4D5B5530FFF9FD7C4971FCC47F70" box="[668,804,1376,1400]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="E2AFEACA5530FFF9FD7C4971FCC47F70" box="[668,804,1376,1400]" italics="true" pageId="9" pageNumber="10">M. dentipes</emphasis>
</taxonomicName>
from the Persian Gulf at Al-Faw (Fao),
<collectingCountry id="A8CC76485530FFF9FAEE4971FADF7F70" box="[1294,1343,1377,1400]" name="Iraq" pageId="9" pageNumber="10">Iraq</collectingCountry>
.
<bibRefCitation id="B44A4B295530FFF9FAAE4971FF037F94" author="Barnes" pageId="9" pageNumber="10" refString="Barnes, R. S. K. (1970) The species of Macrophthalmus in the collections of the British Museum (Natural History). Bulletin of the British Museum of Natural History (Zoology), 20, 203 - 251." type="journal article" year="1970">Barnes (1970)</bibRefCitation>
, like many preceding authors, referred to this species using the junior synonym
<taxonomicName id="17DB4D5B5530FFF9FBBF4995FEEE7FC8" authority="Guerin-Meneville 1838" authorityName="Guerin-Meneville" authorityYear="1838" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pectinipes">
<emphasis id="E2AFEACA5530FFF9FBBF4995FB967F94" box="[1119,1142,1413,1436]" italics="true" pageId="9" pageNumber="10">M</emphasis>
.
<emphasis id="E2AFEACA5530FFF9FB644995FB167F94" box="[1156,1270,1413,1436]" italics="true" pageId="9" pageNumber="10">pectinipes</emphasis>
Guerin-Méneville 1838
</taxonomicName>
, and assigned it to the subgenus
<taxonomicName id="17DB4D5B5530FFF9FD7549B9FC6B7FC8" authority="Barnes, 1967" authorityName="Barnes" authorityYear="1967" box="[661,907,1448,1472]" class="Malacostraca" family="Macrophthalmidae" genus="Venitus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="E2AFEACA5530FFF9FD7549B9FD037FC8" box="[661,739,1449,1472]" italics="true" pageId="9" pageNumber="10">Venitus</emphasis>
<bibRefCitation id="B44A4B295530FFF9FD1049B9FC6B7FC8" author="Barnes" box="[752,907,1448,1472]" pageId="9" pageNumber="10" refString="Barnes, R. S. K. (1967) The Macrophthalminae of Australasia; with a review of the evolution and morphological diversity of the type genus Macrophthalmus. Transactions of the Zoological Society of London, 31, 195 - 262." type="journal article" year="1967">Barnes, 1967</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="B44A4B295530FFF9FC7949B9FB867FC8" author="Pretzmann" box="[921,1126,1448,1472]" pageId="9" pageNumber="10" refString="Pretzmann, G. (1971) Ergebnisse einiger Sammelreisen nach Vorderasien. 2. Teil: Marine Brachyura. Annalen des Naturhistorischen Museums Wien, 75, 477 - 487." type="journal article" year="1971">Pretzmann (1971)</bibRefCitation>
recorded the species from
<collectingCountry id="A8CC76485530FFF9FF7749DCFED87FEC" box="[151,312,1484,1508]" name="Iran" pageId="9" pageNumber="10">Bandar-Abbas</collectingCountry>
at the Iranian coast,
<bibRefCitation id="B44A4B295530FFF9FDCD49DDFD5F7FEC" author="Jones" box="[557,703,1484,1508]" pageId="9" pageNumber="10" refString="Jones, D. A. (1986) A Field Guide to the Sea Shores of Kuwait and the Arabian Gulf. University of Kuwait, Blandford Press, Poole, Kuwait, 192 pp." type="book" year="1986">Jones (1986)</bibRefCitation>
from
<collectingCountry id="A8CC76485530FFF9FCE849DCFCBD7FEC" box="[776,861,1484,1508]" name="Kuwait" pageId="9" pageNumber="10">Kuwait</collectingCountry>
, both following
<bibRefCitation id="B44A4B295530FFF9FBFC49DDFB5D7FEC" author="Barnes" box="[1052,1213,1484,1508]" pageId="9" pageNumber="10" refString="Barnes, R. S. K. (1970) The species of Macrophthalmus in the collections of the British Museum (Natural History). Bulletin of the British Museum of Natural History (Zoology), 20, 203 - 251." type="journal article" year="1970">Barnes (1970)</bibRefCitation>
using the name
<taxonomicName id="17DB4D5B5530FFF9FA9D49DDFEE87C00" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pectinipes">
<emphasis id="E2AFEACA5530FFF9FA9D49DDFA747FEC" box="[1405,1428,1485,1508]" italics="true" pageId="9" pageNumber="10">M</emphasis>
.
<emphasis id="E2AFEACA5530FFF9FF7749E1FEE87C00" box="[151,264,1521,1544]" italics="true" pageId="9" pageNumber="10">pectinipes</emphasis>
</taxonomicName>
. The older name
<taxonomicName id="17DB4D5B5530FFF9FE2D49E1FDB07C00" box="[461,592,1520,1544]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="E2AFEACA5530FFF9FE2D49E1FE047C00" box="[461,484,1521,1544]" italics="true" pageId="9" pageNumber="10">M</emphasis>
.
<emphasis id="E2AFEACA5530FFF9FE1349E0FDB07C00" box="[499,592,1520,1544]" italics="true" pageId="9" pageNumber="10">dentipes</emphasis>
</taxonomicName>
, however, was overlooked until
<bibRefCitation id="B44A4B295530FFF9FC5D49FFFB8F7C0F" author="Holthuis" box="[957,1135,1519,1543]" pageId="9" pageNumber="10" refString="Holthuis, L. B. (1995) The identities of Macrophthalmus rouxii Lucas, 1836, and M. dentipes Lucas, 1836, and the substitution of the latter name for M. pectinipes Guerin, 1838 (Decapoda, Brachyura, Ocypodidae). Crustaceana, 68 (3), 401 - 403." type="journal article" year="1995">Holthuis (1995)</bibRefCitation>
revived
<taxonomicName id="17DB4D5B5530FFF9FB3249E1FAB47C00" box="[1234,1364,1520,1544]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="E2AFEACA5530FFF9FB3249E1FB097C00" box="[1234,1257,1521,1544]" italics="true" pageId="9" pageNumber="10">M</emphasis>
.
<emphasis id="E2AFEACA5530FFF9FB1749E0FAB47C00" box="[1271,1364,1520,1544]" italics="true" pageId="9" pageNumber="10">dentipes</emphasis>
</taxonomicName>
as the valid name by showing that
<taxonomicName id="17DB4D5B5530FFF9FE374A05FD8E7C24" box="[471,622,1557,1580]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pectinipes">
<emphasis id="E2AFEACA5530FFF9FE374A05FE0E7C24" box="[471,494,1557,1580]" italics="true" pageId="9" pageNumber="10">M</emphasis>
.
<emphasis id="E2AFEACA5530FFF9FE1C4A05FD8E7C24" box="[508,622,1557,1580]" italics="true" pageId="9" pageNumber="10">pectinipes</emphasis>
</taxonomicName>
is an objective synonym of
<taxonomicName id="17DB4D5B5530FFF9FC4E4A05FBD17C24" box="[942,1073,1556,1580]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="E2AFEACA5530FFF9FC4E4A05FC257C24" box="[942,965,1557,1580]" italics="true" pageId="9" pageNumber="10">M</emphasis>
.
<emphasis id="E2AFEACA5530FFF9FC344A04FBD17C24" box="[980,1073,1556,1580]" italics="true" pageId="9" pageNumber="10">dentipes</emphasis>
</taxonomicName>
, and therefore must be replaced by the latter.
<bibRefCitation id="B44A4B295530FFF9FEC94A27FDC67C47" author="Apel" box="[297,550,1591,1615]" pageId="9" pageNumber="10" refString="Apel, M. &amp; Turkay, M. (1999) Taxonomic composition, distribution and zoogeographic relationships of the grapsid and ocypodid crab fauna of intertidal soft bottoms in the Arabian Gulf. Estuarine, Coastal and Shelf Science, 49 (suppl. A), 131 - 142." type="journal article" year="1999">Apel &amp; Türkay (1999)</bibRefCitation>
and
<bibRefCitation id="B44A4B295530FFF9FDBD4A27FD057C47" author="Apel" box="[605,741,1591,1615]" pageId="9" pageNumber="10" refString="Apel, M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguridea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes unpublished Ph. D. Thesis, Johann Wolfgang Goethe-Universitat, Frankfurt am Main 268 pp." type="book" year="2001">Apel (2001)</bibRefCitation>
listed
<taxonomicName id="17DB4D5B5530FFF9FCD04A29FC537C58" box="[816,947,1592,1616]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="E2AFEACA5530FFF9FCD04A29FCA77C58" box="[816,839,1593,1616]" italics="true" pageId="9" pageNumber="10">M</emphasis>
.
<emphasis id="E2AFEACA5530FFF9FCB54A28FC537C58" box="[853,947,1592,1616]" italics="true" pageId="9" pageNumber="10">dentipes</emphasis>
</taxonomicName>
from the Persian Gulf with reference to the records by
<bibRefCitation id="B44A4B295530FFF9FEF34A4DFE527C7C" author="Barnes" box="[275,434,1628,1652]" pageId="9" pageNumber="10" refString="Barnes, R. S. K. (1971) Biological results of the Snellius Expedition. XXIII. The Genus Macrophthalmus (Decapoda, Brachyura). Zoologische Verhandelingen, 115, 1 - 40." type="journal article" year="1971">Barnes (1971)</bibRefCitation>
and
<bibRefCitation id="B44A4B295530FFF9FE094A4DFD9C7C7C" author="Jones" box="[489,636,1628,1652]" pageId="9" pageNumber="10" refString="Jones, D. A. (1986) A Field Guide to the Sea Shores of Kuwait and the Arabian Gulf. University of Kuwait, Blandford Press, Poole, Kuwait, 192 pp." type="book" year="1986">Jones (1986)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="98C165535530FFFAFF254A6FFD8A7849" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="biology_ecology">
<paragraph id="D06436D85530FFFAFF254A6FFC5A7AF5" blockId="9.[151,1437,151,2012]" lastBlockId="10.[151,1436,151,577]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="E2AFEACA5530FFF9FF254A6FFEC47C90" bold="true" box="[197,292,1663,1688]" pageId="9" pageNumber="10">Biology.</emphasis>
<taxonomicName id="17DB4D5B5530FFF9FECE4A90FDB47C90" box="[302,596,1664,1688]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="E2AFEACA5530FFF9FECE4A90FDB47C90" box="[302,596,1664,1688]" italics="true" pageId="9" pageNumber="10">Macrophthalmus dentipes</emphasis>
</taxonomicName>
is the largest ocypodid crab (largest male CL = 40.11, CB =
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, largest female CL = 40.55, CB =
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) inhabiting the mid and low intertidal zones in muddy sand/sandy mud substrates (
<bibRefCitation id="B44A4B295530FFF9FEF54AD7FE787CD7" author="Jones" box="[277,408,1735,1759]" pageId="9" pageNumber="10" refString="Jones, D. A. (1986) A Field Guide to the Sea Shores of Kuwait and the Arabian Gulf. University of Kuwait, Blandford Press, Poole, Kuwait, 192 pp." type="book" year="1986">Jones 1986</bibRefCitation>
; Al-Zaidan
<emphasis id="E2AFEACA5530FFF9FDFF4AD9FDB17CE8" box="[543,593,1736,1760]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
. 2006) in the region. This burrowing crab occurs especially in soft muddy substrates at depths of more than
<quantity id="17239B3D5530FFF9FDF24AFCFDBB7D0C" box="[530,603,1772,1796]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="10" unit="cm" value="60.0">60 cm</quantity>
. The
<taxonomicName id="17DB4D5B5530FFF9FD794AFCFCB77D0C" box="[665,855,1772,1796]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="E2AFEACA5530FFF9FD794AFCFCB77D0C" box="[665,855,1772,1796]" italics="true" pageId="9" pageNumber="10">Macrophthalmus</emphasis>
</taxonomicName>
zone extends from the middle part of the mudskipper (
<taxonomicName id="17DB4D5B5530FFF9FF2C4B00FE3C7D20" box="[204,476,1808,1832]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="waltoni">
<emphasis id="E2AFEACA5530FFF9FF2C4B00FE3C7D20" box="[204,476,1808,1832]" italics="true" pageId="9" pageNumber="10">Periophthalmus waltoni</emphasis>
</taxonomicName>
) zone downwards to the low littoral (
<bibRefCitation id="B44A4B295530FFF9FC6D4B1FFBCB7D2F" author="Clayton" box="[909,1067,1807,1831]" pageId="9" pageNumber="10" refString="Clayton, D. A. (1986) Ecology of mudflats with particular reference to those of the northern Arabian Gulf. In: Halwagy, R., Clayton, D. &amp; Bbehbehani, M. (eds.): Marine Environment and Pollution. Proceedings of the 1 st Arabian Gulf Conference on Environment and Pollution, Kuwait, 7 - 9 February 1982. University of Kuwait, pp. 83 - 96." type="journal article" year="1986">Clayton 1986</bibRefCitation>
).
<taxonomicName id="17DB4D5B5530FFF9FBA04B00FA847D20" box="[1088,1380,1808,1832]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="E2AFEACA5530FFF9FBA04B00FA847D20" box="[1088,1380,1808,1832]" italics="true" pageId="9" pageNumber="10">Macrophthalmus dentipes</emphasis>
</taxonomicName>
digs large burrows with an opening of approximately
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in diameter, which sometimes can be confused with those of mudskippers, but in this crab the burrows entrance is marked by a sloppy, light-coloured mud pool with a mound of excavated mud Whereas, the species has not been recorded from the Arabian side on the western and southern parts of the Persian Gulf, it is a common species of muddy sand/sandy mud substrates along the northern and northeast coasts including
<collectingCountry id="A8CC76485530FFF9FE664BD4FE397DD4" box="[390,473,1988,2012]" name="Kuwait" pageId="9" pageNumber="10">Kuwait</collectingCountry>
(
<bibRefCitation id="B44A4B295530FFF9FE084BD4FD8A7DD4" author="Jones" box="[488,618,1988,2012]" pageId="9" pageNumber="10" refString="Jones, D. A. (1986) A Field Guide to the Sea Shores of Kuwait and the Arabian Gulf. University of Kuwait, Blandford Press, Poole, Kuwait, 192 pp." type="book" year="1986">Jones 1986</bibRefCitation>
;
<bibRefCitation id="B44A4B295530FFF9FD964BD4FCEF7DD4" author="Clayton" box="[630,783,1988,2012]" pageId="9" pageNumber="10" refString="Clayton, D. A. (1986) Ecology of mudflats with particular reference to those of the northern Arabian Gulf. In: Halwagy, R., Clayton, D. &amp; Bbehbehani, M. (eds.): Marine Environment and Pollution. Proceedings of the 1 st Arabian Gulf Conference on Environment and Pollution, Kuwait, 7 - 9 February 1982. University of Kuwait, pp. 83 - 96." type="journal article" year="1986">Clayton 1986</bibRefCitation>
) and the Iranian coast. The population density of this species was estimated based on burrow counts. In the muddy flat of Bandar-Khamir east to the fishery jetty (
<geoCoordinate id="B5EF501F5533FFFAFAD24C87FA777AA7" box="[1330,1431,151,175]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="925" value="26.933332">26º 56'N</geoCoordinate>
,
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) an average of 10 burrows were present per
<quantity id="17239B3D5533FFFAFD044CAFFCEE7ADF" box="[740,782,191,215]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="11" unit="m" value="5.0">5 m</quantity>
2, whereas near Mahtabi Police station (
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,
<geoCoordinate id="B5EF501F5533FFFAFAD54CAFFA737ADF" box="[1333,1427,191,215]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="55.333332">55º 20'E</geoCoordinate>
) in sandy mud substrate an average of 20 burrows per
<quantity id="17239B3D5533FFFAFD0D4CF5FCF87AF5" box="[749,792,229,253]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="11" unit="m" value="5.0">5 m</quantity>
2 was counted.
</paragraph>
<paragraph id="D06436D85533FFFAFF254D19FC477BDD" blockId="10.[151,1436,151,577]" pageId="10" pageNumber="11">
The pereopods of males of
<taxonomicName id="17DB4D5B5533FFFAFDEB4D1AFD737B29" box="[523,659,265,289]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis id="E2AFEACA5533FFFAFDEB4D1AFDC27B29" box="[523,546,266,289]" italics="true" pageId="10" pageNumber="11">M</emphasis>
.
<emphasis id="E2AFEACA5533FFFAFDD54D19FD737B29" box="[565,659,265,289]" italics="true" pageId="10" pageNumber="11">dentipes</emphasis>
</taxonomicName>
grow faster compared to the carapace, therefore large males have remarkably long legs, and long chelipeds in particular.
<bibRefCitation id="B44A4B295533FFFAFCE84D3EFC4B7B4D" author="Barnes" box="[776,939,301,325]" pageId="10" pageNumber="11" refString="Barnes, R. S. K. (1970) The species of Macrophthalmus in the collections of the British Museum (Natural History). Bulletin of the British Museum of Natural History (Zoology), 20, 203 - 251." type="journal article" year="1970">Barnes (1970)</bibRefCitation>
, in his detailed description, only mentioned the abnormal growth of the merus of the pereopods, but the rapid development of all segments was observed in the present study, which can be clearly detected in the remarkably long palm of the cheliped in large males (
<figureCitation id="48E02A5D5533FFFAFADE4D65FA637B85" box="[1342,1411,373,397]" captionStart="FIGURE 4" captionStartId="7.[151,250,1906,1929]" captionTargetBox="[180,1393,655,1863]" captionTargetId="figure@7.[151,1436,601,1885]" captionTargetPageId="7" captionText="FIGURE 4. Macrophthalmus dentipes Lucas, 1836: a f, male (SMF 36863); f, male (SMF 26095). a, posterior surface of carapace, left side; b, lower orbital margin of male, right; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, merus of male cheliped, inner surface; f, cheliped of male outer surface." httpUri="https://zenodo.org/record/203102/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
f). Furthermore, large males have more granules on the posterior surface of the carapace, which are markedly high in comparison to the low and round granules of carapace of small males.
</paragraph>
<paragraph id="D06436D85533FFFAFF254DF1FD8A7849" blockId="10.[151,1436,151,577]" pageId="10" pageNumber="11">
<emphasis id="E2AFEACA5533FFFAFF254DF1FDE37BF2" bold="true" box="[197,515,481,506]" pageId="10" pageNumber="11">Geographical distribution.</emphasis>
Northern Indian Ocean: northern and eastern Persian Gulf,
<collectingCountry id="A8CC76485533FFFAFB004DF1FAC47BF1" box="[1248,1316,481,505]" name="Oman" pageId="10" pageNumber="11">Oman</collectingCountry>
(Gulf of Masirah),
<collectingCountry id="A8CC76485533FFFAFEEB4E15FE8D7815" box="[267,365,517,541]" name="Pakistan" pageId="10" pageNumber="11">Pakistan</collectingCountry>
, west coast of
<collectingCountry id="A8CC76485533FFFAFDF74E15FDB47815" box="[535,596,517,541]" name="India" pageId="10" pageNumber="11">India</collectingCountry>
. A record from
<collectingCountry id="A8CC76485533FFFAFCEB4E15FC927815" box="[779,882,517,541]" name="Malaysia" pageId="10" pageNumber="11">Malaysia</collectingCountry>
by
<bibRefCitation id="B44A4B295533FFFAFC7E4E15FB8A7815" author="Henderson" box="[926,1130,517,541]" pageId="10" pageNumber="11" refString="Henderson, J. R. (1893) A contribution to Indian carcinology. Transactions of the Linnean Society of London, series 2, Zoology, 5 (10), 325 - 458, pls. 36 - 40." type="journal article" year="1893">Henderson (1893)</bibRefCitation>
is highly questionable and probably based on misidentified material.
</paragraph>
</subSubSection>
</treatment>
</document>