plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/40/2D/1B/402D1B3B533DFFDFC9FEFB1CFE67FFE0.xml
maintenance 9c643743c0 added 40
2024-06-21 12:34:30 +02:00

456 lines
54 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="08B432ECFFAAB36C9790766122D590FE" ID-DOI="10.11646/zootaxa.4790.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="3889527" ID-ZooBank="46EAA29B-7852-4374-B844-DA7E25A00ABD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591859566901" checkinUser="plazi" docAuthor="Nguyen, Tung T., Tran, Binh T. T., Lam, Dang H. &amp; Nguyen, Anh D." docDate="2020" docId="402D1B3B533DFFDFC9FEFB1CFE67FFE0" docLanguage="en" docName="zootaxa.4790.2.5.pdf" docOrigin="Zootaxa 4790 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amynthas dorsomorrioides Nguyen &amp; Tran &amp; Lam &amp; Nguyen 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="283" masterDocId="BC1463435339FFD9C969FF95FFE9FA24" masterDocTitle="Four new species of Amynthas earthworms in southeastern Vietnam (Annelida, Oligochaeta, Megascolecidae)" masterLastPageNumber="290" masterPageNumber="277" pageNumber="281" updateTime="1700276591635" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="BA496053407FA0403D3D89968046E242" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C736A30A766779E37820F191683844C2">
<mods:title id="4E6CD2B45E8D31F1D9463841D4CA72ED">Four new species of Amynthas earthworms in southeastern Vietnam (Annelida, Oligochaeta, Megascolecidae)</mods:title>
</mods:titleInfo>
<mods:name id="7B6271A58B171A8E667A6B51D68BBC3C" type="personal">
<mods:role id="42DEB590ABA75CA49B1EB6C654550011">
<mods:roleTerm id="0994DF89230B5A40677A23DAAEC95BB8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EAAB61A0F8505A26FB215C5BDD2067C0">Nguyen, Tung T.</mods:namePart>
</mods:name>
<mods:name id="4B6816951A7BA9E852673D41D6567516" type="personal">
<mods:role id="08931E6EBF8CFC13D206F561BEE3169A">
<mods:roleTerm id="D153C8E560AC2848B75A674D607A681B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="329A54F9768B9B4BC6E987D5E689EFF7">Tran, Binh T. T.</mods:namePart>
<mods:affiliation id="6D633325A264B5686E177CBEB77AACBD">Department of Zoology, Faculty of Biology, Hanoi National University of Education, Hanoi, Vietnam.</mods:affiliation>
</mods:name>
<mods:name id="7148E5F461290DAC8037519D87E7DC9A" type="personal">
<mods:role id="3C419305BFB910D5D25C32EC723D5076">
<mods:roleTerm id="6F99D4ACC844BB093C9D41C4F291A8EA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="215762CD8A283B9AF2AA3CD8238AAF9E">Lam, Dang H.</mods:namePart>
</mods:name>
<mods:name id="D0AEB793AEE62A73B8E66D967EBE1BA3" type="personal">
<mods:role id="EB3AEB1F8810970636728292AFD573C8">
<mods:roleTerm id="801E9F6B633FA0B787C74298BF466E1E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9539FBD4BA74793240CD4E0CF6A4201C">Nguyen, Anh D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="121166208879A92840A8767465A88F0A">text</mods:typeOfResource>
<mods:relatedItem id="5B2FDCFDA219304C148F8E54DC87B4BE" type="host">
<mods:titleInfo id="87FE96337658A1AD2A60F5BB8534D8B7">
<mods:title id="9479C91200F0ADA6BEE4922402A53237">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0F255BDE0BBAA31246DD528E223C23DD">
<mods:date id="8B30813814DE51850D52DAE7B8DB05DD">2020</mods:date>
<mods:detail id="8DBA30CB50DC5E9AE4C574343A5E5FE9" type="pubDate">
<mods:number id="C48BCCB548CF90BB1434E915F76DA847">2020-06-11</mods:number>
</mods:detail>
<mods:detail id="13C9337A8681C9D91C128886B2E519D1" type="volume">
<mods:number id="6F79F48099256D028AAD22FD6D52F6BB">4790</mods:number>
</mods:detail>
<mods:detail id="84C75B487EE3B1D1DA96784BDA369A68" type="issue">
<mods:number id="EE738BFE2873F919410936E7F00F10AB">2</mods:number>
</mods:detail>
<mods:extent id="0EDBFA135D7E8E5265775F71695C8538" unit="page">
<mods:start id="B17917DBC18F61A5369599A9DA23C22C">277</mods:start>
<mods:end id="44D93C8E1AAC2E9E74A6CA2A329B20DF">290</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D54D640A43FD33DC13E702922510C479">journal article</mods:classification>
<mods:identifier id="EFF69A33681418524D3FB7F5EA23B1F2" type="DOI">10.11646/zootaxa.4790.2.5</mods:identifier>
<mods:identifier id="3CDC99EA37A9DAEA0EA405C34BF15D76" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EE2B65D5D2F4B1957FE4A57EB5D0915F" type="Zenodo-Dep">3889527</mods:identifier>
<mods:identifier id="19DA678C04F18D277BAD2DE443A32734" type="ZooBank">46EAA29B-7852-4374-B844-DA7E25A00ABD</mods:identifier>
</mods:mods>
<treatment id="402D1B3B533DFFDFC9FEFB1CFE67FFE0" LSID="urn:lsid:plazi:treatment:402D1B3B533DFFDFC9FEFB1CFE67FFE0" httpUri="http://treatment.plazi.org/id/402D1B3B533DFFDFC9FEFB1CFE67FFE0" lastPageId="6" lastPageNumber="283" pageId="4" pageNumber="281">
<subSubSection id="809EF9A6533DFFDDC9FEFB1CFCD0FE87" box="[151,825,1161,1188]" pageId="4" pageNumber="281" type="nomenclature">
<paragraph id="C83BAA2D533DFFDDC9FEFB1CFCD0FE87" blockId="4.[151,825,1161,1223]" box="[151,825,1161,1188]" pageId="4" pageNumber="281">
<heading id="93731D41533DFFDDC9FEFB1CFCD0FE87" bold="true" box="[151,825,1161,1188]" fontSize="11" level="1" pageId="4" pageNumber="281" reason="1">
<emphasis id="FAF0763F533DFFDDC9FEFB1CFCD0FE87" bold="true" box="[151,825,1161,1188]" pageId="4" pageNumber="281">
<taxonomicName id="0F84D1AE533DFFDDC9FEFB1CFD26FE87" authority="Nguyen &amp; Nguyen" authorityName="Nguyen &amp; Tran &amp; Lam &amp; Nguyen" authorityYear="2020" box="[151,719,1161,1188]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="dorsomorrioides" status="sp. nov.">
<emphasis id="FAF0763F533DFFDDC9FEFB1CFE37FE80" bold="true" box="[151,478,1161,1188]" italics="true" pageId="4" pageNumber="281">Amynthas dorsomorrioides</emphasis>
Nguyen &amp; Nguyen
</taxonomicName>
,
<taxonomicNameLabel id="E1C3CB44533DFFDDCBB2FB1CFCD0FE87" box="[731,825,1161,1187]" pageId="4" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="809EF9A6533DFFDDC9FEFB38FEE1FEE3" box="[151,264,1197,1223]" pageId="4" pageNumber="281" type="description">
<paragraph id="C83BAA2D533DFFDDC9FEFB38FEE1FEE3" blockId="4.[151,825,1161,1223]" box="[151,264,1197,1223]" pageId="4" pageNumber="281">
(
<figureCitation id="50BFB6A8533DFFDDC9F6FB38FF16FEE3" box="[159,255,1197,1223]" captionStart="FIGURE 3" captionStartId="5.[151,250,1901,1926]" captionTargetBox="[162,1421,395,1864]" captionTargetId="figure@5.[154,1433,381,1876]" captionTargetPageId="5" captionText="FIGURE 3. Amynthas dorsomorrioides sp. nov. Holotype (CTU-EW.174.h01). A. Male region, ventral view. B. Testis sacs and seminal vesicle. C. Left prostate gland. D. Spermathecal region, dorsal view. E. Left intestinal caecum. F. Left spermatheca. G. Spermathecae (comp. F). H. Male region (comp. A). I. Prostate gland (comp. C). J. Intestinal caecum (comp. E). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889533" httpUri="https://zenodo.org/record/3889533/files/figure.png" pageId="4" pageNumber="281">Figure 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="809EF9A6533DFFDDC9FEFB61FD87FFBB" pageId="4" pageNumber="281" type="materials_examined">
<paragraph id="C83BAA2D533DFFDDC9FEFB61FD87FFBB" blockId="4.[151,1437,1268,2015]" pageId="4" pageNumber="281">
<emphasis id="FAF0763F533DFFDDC9FEFB61FE6AFF2B" bold="true" box="[151,387,1268,1295]" pageId="4" pageNumber="281">Material examined.</emphasis>
<materialsCitation id="78ECA070533DFFDDC8E2FB60FEE4FF73" collectingDate="2016-10-27" collectionCode="CTU-EW" collectorName="Nguyen Phuc Hau." county="Ba Ria - Vung Tau province" elevation="72" latitude="10.511111" location="Kim Dinh" longLatPrecision="21" longitude="107.126945" municipality="Ba Ria" pageId="4" pageNumber="281" specimenCount="1" typeStatus="holotype">
<emphasis id="FAF0763F533DFFDDC8E2FB60FE1BFF2B" box="[395,498,1269,1295]" italics="true" pageId="4" pageNumber="281">
<typeStatus id="173F148F533DFFDDC8E2FB60FE1BFF2B" box="[395,498,1269,1295]" pageId="4" pageNumber="281" type="holotype">Holotype</typeStatus>
</emphasis>
: 1 mature (complete body), (
<collectionCode id="AE9532E8533DFFDDCA2AFB60FC53FF34" box="[835,954,1269,1296]" pageId="4" pageNumber="281">CTU-EW</collectionCode>
.174.h01), shrubs and bushes (
<geoCoordinate id="ADB0CCEA533DFFDDCC62FB60FA7EFF2B" box="[1291,1431,1269,1295]" degrees="10" direction="north" minutes="30" orientation="latitude" pageId="4" pageNumber="281" precision="15" seconds="40" value="10.511111">10°30’40”N</geoCoordinate>
,
<geoCoordinate id="ADB0CCEA533DFFDDC9FEFA8CFEC6FF17" box="[151,303,1305,1331]" degrees="107" direction="east" minutes="07" orientation="longitude" pageId="4" pageNumber="281" precision="15" seconds="37" value="107.126945">107°07’37”E</geoCoordinate>
),
<quantity id="0F7C07C8533DFFDDC82BFA8CFE95FF17" box="[322,380,1305,1331]" metricMagnitude="1" metricUnit="m" metricValue="7.2" pageId="4" pageNumber="281" unit="m" value="72.0">
<elevation id="43A94D1E533DFFDDC82BFA8CFE95FF17" box="[322,380,1305,1331]" metricMagnitude="1" metricUnit="m" metricValue="7.2" pageId="4" pageNumber="281" unit="m" value="72.0">72 m</elevation>
</quantity>
,
<location id="CD5BFCF6533DFFDDC8EFFA8CFE1FFF17" LSID="urn:lsid:plazi:treatment:402D1B3B533DFFDFC9FEFB1CFE67FFE0:CD5BFCF6533DFFDDC8EFFA8CFE1FFF17" box="[390,502,1305,1331]" county="Ba Ria - Vung Tau province" latitude="10.511111" longLatPrecision="21" longitude="107.126945" municipality="Ba Ria" name="Kim Dinh" pageId="4" pageNumber="281">Kim Dinh</location>
commune,
<collectingMunicipality id="285F3057533DFFDDCB11FA8CFD2DFF17" box="[632,708,1305,1331]" pageId="4" pageNumber="281">Ba Ria</collectingMunicipality>
city,
<collectingCounty id="215AD2A1533DFFDDCB96FA8CFBD2FF17" box="[767,1083,1305,1331]" pageId="4" pageNumber="281">Ba Ria – Vung Tau province</collectingCounty>
,
<date id="BC3A8CED533DFFDDCD2CFA8CFAEAFF17" box="[1093,1283,1305,1331]" pageId="4" pageNumber="281" value="2016-10-27">
<collectingDate id="AC7E7505533DFFDDCD2CFA8CFAEAFF17" box="[1093,1283,1305,1331]" pageId="4" pageNumber="281" value="2016-10-27">27 October 2016</collectingDate>
</date>
, coll.
<collectorName id="6571CFFB533DFFDDCC2DFA8CFEE4FF73" pageId="4" pageNumber="281">Nguyen Phuc Hau.</collectorName>
</materialsCitation>
<materialsCitation id="78ECA070533DFFDDC87AFAABFD87FFBB" collectingDate="2016-10-25" collectionCode="CTU-EW, IEBR-EW" collectorName="Nguyen Phuc Hau." county="Ba Ria" elevation="33" latitude="10.406111" location="Long Hai" longLatPrecision="21" longitude="107.27167" municipality="Long Dien District" pageId="4" pageNumber="281" specimenCount="1" typeStatus="paratype">
<emphasis id="FAF0763F533DFFDDC87AFAABFE6CFF73" box="[275,389,1342,1367]" italics="true" pageId="4" pageNumber="281">
<typeStatus id="173F148F533DFFDDC87AFAABFE6CFF73" box="[275,389,1342,1367]" pageId="4" pageNumber="281" type="paratype">Paratypes</typeStatus>
</emphasis>
: 2 matures (complete body), (
<collectionCode id="AE9532E8533DFFDDCBBEFAA8FCA5FF73" box="[727,844,1341,1367]" pageId="4" pageNumber="281">CTU-EW</collectionCode>
.174.p02) and 2 matures (
<collectionCode id="AE9532E8533DFFDDCD08FAA8FB36FF73" box="[1121,1247,1341,1367]" pageId="4" pageNumber="281">IEBR-EW</collectionCode>
.174.p02), natural forests (
<geoCoordinate id="ADB0CCEA533DFFDDC99BFAF4FE96FF5F" box="[242,383,1377,1403]" degrees="10" direction="north" minutes="24" orientation="latitude" pageId="4" pageNumber="281" precision="15" seconds="22" value="10.406111">10°24’22”N</geoCoordinate>
;
<geoCoordinate id="ADB0CCEA533DFFDDC8E2FAF4FDCBFF58" box="[395,546,1377,1404]" degrees="107" direction="east" minutes="16" orientation="longitude" pageId="4" pageNumber="281" precision="15" seconds="18" value="107.27167">107°16’18”E</geoCoordinate>
),
<quantity id="0F7C07C8533DFFDDCB5EFAF4FD9AFF5F" box="[567,627,1377,1403]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="281" unit="m" value="33.0">
<elevation id="43A94D1E533DFFDDCB5EFAF4FD9AFF5F" box="[567,627,1377,1403]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="281" unit="m" value="33.0">33 m</elevation>
</quantity>
,
<location id="CD5BFCF6533DFFDDCB16FAF4FD03FF5F" LSID="urn:lsid:plazi:treatment:402D1B3B533DFFDFC9FEFB1CFE67FFE0:CD5BFCF6533DFFDDCB16FAF4FD03FF5F" box="[639,746,1377,1403]" county="Ba Ria" latitude="10.406111" longLatPrecision="21" longitude="107.27167" municipality="Long Dien District" name="Long Hai" pageId="4" pageNumber="281">Long Hai</location>
commune,
<collectingMunicipality id="285F3057533DFFDDCA06FAF4FBAEFF5F" box="[879,1095,1377,1403]" pageId="4" pageNumber="281">Long Dien District</collectingMunicipality>
,
<collectingCounty id="215AD2A1533DFFDDCD3AFAF4FB49FF5F" box="[1107,1184,1377,1403]" pageId="4" pageNumber="281">Ba Ria</collectingCounty>
– Vung Tau Province,
<date id="BC3A8CED533DFFDDC9FEFA11FEBFFFBB" box="[151,342,1412,1439]" pageId="4" pageNumber="281" value="2016-10-25">
<collectingDate id="AC7E7505533DFFDDC9FEFA11FEBFFFBB" box="[151,342,1412,1439]" pageId="4" pageNumber="281" value="2016-10-25">25 October 2016</collectingDate>
</date>
, coll.
<collectorName id="6571CFFB533DFFDDC8F0FA10FD87FFBB" box="[409,622,1413,1439]" pageId="4" pageNumber="281">Nguyen Phuc Hau.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="809EF9A6533DFFDDC9AEFA3CFF2DFFC3" pageId="4" pageNumber="281" type="etymology">
<paragraph id="C83BAA2D533DFFDDC9AEFA3CFF2DFFC3" blockId="4.[151,1437,1268,2015]" pageId="4" pageNumber="281">
<emphasis id="FAF0763F533DFFDDC9AEFA3CFEA7FFE7" bold="true" box="[199,334,1449,1475]" pageId="4" pageNumber="281">Etymology.</emphasis>
“
<taxonomicName id="0F84D1AE533DFFDDC80AFA3CFDF7FFE7" authorityName="Nguyen &amp; Tran &amp; Lam &amp; Nguyen" authorityYear="2020" box="[355,542,1449,1475]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="dorsomorrioides">
<emphasis id="FAF0763F533DFFDDC80AFA3CFDF7FFE7" box="[355,542,1449,1475]" italics="true" pageId="4" pageNumber="281">dorsomorrioides</emphasis>
</taxonomicName>
”, an adjective, to emphasize the similarity to the species
<taxonomicName id="0F84D1AE533DFFDDCDD9FA3CFF55FFC3" baseAuthorityName="Do &amp; Tran" baseAuthorityYear="1995" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="dorsomorrisi">
<emphasis id="FAF0763F533DFFDDCDD9FA3CFF55FFC3" italics="true" pageId="4" pageNumber="281">Amynthas dorsomorrisi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="809EF9A6533DFFDDC9AEFA65FD86FCBF" pageId="4" pageNumber="281" type="diagnosis">
<paragraph id="C83BAA2D533DFFDDC9AEFA65FD86FCBF" blockId="4.[151,1437,1268,2015]" pageId="4" pageNumber="281">
<emphasis id="FAF0763F533DFFDDC9AEFA65FEABFC2F" bold="true" box="[199,322,1520,1547]" pageId="4" pageNumber="281">Diagnosis.</emphasis>
Small size, length
<quantity id="0F7C07C8533DFFDDCB48FA65FD76FC2F" box="[545,671,1520,1547]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" metricValueMax="9.5" metricValueMin="5.9" pageId="4" pageNumber="281" unit="mm" value="77.0" valueMax="95.0" valueMin="59.0">59–95 mm</quantity>
, diameter
<quantity id="0F7C07C8533DFFDDCA71FA65FC4DFC2F" box="[792,932,1520,1547]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.1" metricValueMin="2.5" pageId="4" pageNumber="281" unit="mm" value="2.8" valueMax="3.1" valueMin="2.5">2.5–3.1 mm</quantity>
, segments 135–151. Prostomium epilobous. First dorsal pore in 12/13. Setal numbers:
<quantity id="0F7C07C8533DFFDDCB18F980FD3CFC0B" box="[625,725,1557,1583]" metricMagnitude="0" metricUnit="m" metricValue="1.6002" metricValueMax="1.6256" metricValueMin="1.5748" pageId="4" pageNumber="281" unit="in" value="63.0" valueMax="64.0" valueMin="62.0">62–64 in</quantity>
viii,
<quantity id="0F7C07C8533DFFDDCA79F981FC9AFC0B" box="[784,883,1556,1583]" metricMagnitude="0" metricUnit="m" metricValue="1.4097" metricValueMax="1.4478" metricValueMin="1.3716" pageId="4" pageNumber="281" unit="in" value="55.5" valueMax="57.0" valueMin="54.0">54–57 in</quantity>
xxx, 6–7 between male porophores in xviii. Two pairs of spermathecal pores located dorsally in intersegments 5/6/7. Ventral distance between two male pores about 0.3x body circumference. A pair of pre-setal genital markings present in xvii. Intestinal caeca simple. Holandric. Testis sacs separated. Septa 8/9/10 absent.
</paragraph>
</subSubSection>
<subSubSection id="809EF9A6533DFFDFC9AEF931FB81FB3B" lastPageId="6" lastPageNumber="283" pageId="4" pageNumber="281" type="description">
<paragraph id="C83BAA2D533DFFDDC9AEF931FBDAFD6B" blockId="4.[151,1437,1268,2015]" pageId="4" pageNumber="281">
<emphasis id="FAF0763F533DFFDDC9AEF931FEB0FC9B" bold="true" box="[199,345,1700,1727]" pageId="4" pageNumber="281">Description.</emphasis>
Cylindrical body, small size, length
<quantity id="0F7C07C8533DFFDDCB95F931FC90FC9B" box="[764,889,1700,1727]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" metricValueMax="9.5" metricValueMin="5.9" pageId="4" pageNumber="281" unit="mm" value="77.0" valueMax="95.0" valueMin="59.0">59–95 mm</quantity>
, diameter
<quantity id="0F7C07C8533DFFDDCA98F931FB92FC9B" box="[1009,1147,1700,1727]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.1" metricValueMin="2.5" pageId="4" pageNumber="281" unit="mm" value="2.8" valueMax="3.1" valueMin="2.5">2.5–3.1 mm</quantity>
, weight 0.2–0.42 gr, segments 135–151. Body transparent, clitellum light brown. Prostomium 1/2 epilobous. First dorsal pore in 12/13. Setae perichaetine, pre-clitellar setae stouter and denser than post-clitellar ones; setal number:
<quantity id="0F7C07C8533DFFDDCDD0F978FAF5FD23" box="[1209,1308,1773,1799]" metricMagnitude="0" metricUnit="m" metricValue="1.6002" metricValueMax="1.6256" metricValueMin="1.5748" pageId="4" pageNumber="281" unit="in" value="63.0" valueMax="64.0" valueMin="62.0">62–64 in</quantity>
viii,
<quantity id="0F7C07C8533DFFDDCC3FF979FF44FD0F" metricMagnitude="0" metricUnit="m" metricValue="1.4097" metricValueMax="1.4478" metricValueMin="1.3716" pageId="4" pageNumber="281" unit="in" value="55.5" valueMax="57.0" valueMin="54.0">54–57 in</quantity>
xxx, 6–7 between two male porophores in xviii; setal distance: aa=ab, zz&gt;zy. Clitellum within xiv–xvi, without dorsal pores, but with setae on ventrum xvi. Female pore single, in midventral xiv.
</paragraph>
<paragraph id="C83BAA2D533DFFDDC9AEF8CCFB38FDFB" blockId="4.[151,1437,1268,2015]" pageId="4" pageNumber="281">Two pairs of spermathecal pores located dorsally in intersegments 5/6/7. No genital markings in spermathecal region. Male porophores ventrally located in setal ring of xviii; ventral distance between male pores about 0.3x body circumference. A pair of large, round genital markings ventrally present in front of setal ring xvii. These markings slightly larger than the male porophores and shifted mediad with respect to the male porophores.</paragraph>
<paragraph id="C83BAA2D533CFFDCC9AEFF0CFCCEFADF" blockId="5.[151,1437,152,359]" pageId="5" pageNumber="282">Septa 5/6/7/8 thick, 8/9/10 absent, 10/11/12/13 thin. Oesophageal gizzard within viii–x. Intestinal origin at xiv; caeca simple, originating at xxvii and extending anteriorly to xxiv. Last hearts in xiii. Pharyngeal micronephridia developed 4/5/6. Typhlosole folded. Lymph glands absent.</paragraph>
<paragraph id="C83BAA2D533CFFDCC9AEFE90FE08FB43" blockId="5.[151,1437,152,359]" pageId="5" pageNumber="282">Two pairs of spermathecae in vi and vii. Ampulla oval-shaped; ducts about 1/3 ampulla length. Diverticula shorter than ampulla, slightly waved, and directly attached to base of ampulla duct; seminal chamber small ovalshaped. No accessory glands.</paragraph>
<caption id="9CFBFAA5533CFFDCC9FEF8F8FF04FDD5" ID-DOI="http://doi.org/10.5281/zenodo.3889533" ID-Zenodo-Dep="3889533" httpUri="https://zenodo.org/record/3889533/files/figure.png" pageId="5" pageNumber="282" startId="5.[151,250,1901,1926]" targetBox="[162,1421,395,1864]" targetPageId="5">
<paragraph id="C83BAA2D533CFFDCC9FEF8F8FF04FDD5" blockId="5.[151,1437,1901,2034]" pageId="5" pageNumber="282">
<emphasis id="FAF0763F533CFFDCC9FEF8F8FC1FFDA2" bold="true" box="[151,1014,1901,1926]" pageId="5" pageNumber="282">
FIGURE 3.
<taxonomicName id="0F84D1AE533CFFDCC872F8F8FDDFFDA2" authority="Nguyen &amp; Tran &amp; Lam &amp; Nguyen, 2020" authorityName="Nguyen &amp; Tran &amp; Lam &amp; Nguyen" authorityYear="2020" box="[283,566,1901,1926]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="282" phylum="Annelida" rank="species" species="dorsomorrioides" status="sp. nov.">
<emphasis id="FAF0763F533CFFDCC872F8F8FDDFFDA2" bold="true" box="[283,566,1901,1926]" italics="true" pageId="5" pageNumber="282">Amynthas dorsomorrioides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E1C3CB44533CFFDCCB57F8FBFD66FDA2" box="[574,655,1902,1926]" pageId="5" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (CTU-EW.174.h01). A.
</emphasis>
Male region, ventral view.
<emphasis id="FAF0763F533CFFDCCC7BF8FBFAC0FDA2" bold="true" box="[1298,1321,1902,1926]" pageId="5" pageNumber="282">B.</emphasis>
Testis sacs and seminal vesicle.
<emphasis id="FAF0763F533CFFDCC80EF804FE69FD8E" bold="true" box="[359,384,1937,1962]" pageId="5" pageNumber="282">C.</emphasis>
Left prostate gland.
<emphasis id="FAF0763F533CFFDCCB26F807FD8EFD8E" bold="true" box="[591,615,1938,1962]" pageId="5" pageNumber="282">D.</emphasis>
Spermathecal region, dorsal view.
<emphasis id="FAF0763F533CFFDCCAAEF807FC37FD8E" bold="true" box="[967,990,1938,1962]" pageId="5" pageNumber="282">E.</emphasis>
Left intestinal caecum.
<emphasis id="FAF0763F533CFFDCCDA6F807FB0BFD8E" bold="true" box="[1231,1250,1938,1962]" pageId="5" pageNumber="282">F.</emphasis>
Left spermatheca.
<emphasis id="FAF0763F533CFFDCC9FEF820FF58FDEA" bold="true" box="[151,177,1973,1998]" pageId="5" pageNumber="282">G.</emphasis>
Spermathecae (comp.
<emphasis id="FAF0763F533CFFDCC8F3F823FE40FDEA" bold="true" box="[410,425,1974,1998]" pageId="5" pageNumber="282">F</emphasis>
).
<emphasis id="FAF0763F533CFFDCC8D6F823FE30FDEA" bold="true" box="[447,473,1974,1998]" pageId="5" pageNumber="282">H.</emphasis>
Male region (comp.
<emphasis id="FAF0763F533CFFDCCBC7F820FD29FDEA" bold="true" box="[686,704,1973,1998]" pageId="5" pageNumber="282">A</emphasis>
).
<emphasis id="FAF0763F533CFFDCCBBFF823FD0FFDEA" bold="true" box="[726,742,1974,1998]" pageId="5" pageNumber="282">I.</emphasis>
Prostate gland (comp.
<emphasis id="FAF0763F533CFFDCCAB8F820FC0AFDEA" bold="true" box="[977,995,1973,1998]" pageId="5" pageNumber="282">C</emphasis>
).
<emphasis id="FAF0763F533CFFDCCA90F823FBE5FDEA" bold="true" box="[1017,1036,1974,1998]" pageId="5" pageNumber="282">J.</emphasis>
Intestinal caecum (comp.
<emphasis id="FAF0763F533CFFDCCC71F823FAC0FDEA" bold="true" box="[1304,1321,1974,1998]" pageId="5" pageNumber="282">E</emphasis>
). Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="C83BAA2D533FFFDFC9AEFF0CFC58FADF" blockId="6.[151,1437,153,1476]" pageId="6" pageNumber="283">Holandric. Testis sacs developed, separated in x–xi. Seminal vesicles well developed in xi–xii. Ovaries well developed in 12/13. Ovisacs invisible. Prostate glands deeply lobuled, paired in xviii–xix; ducts transparent, C-shaped. A pair of accessory glands in association with genital markings in xvii.</paragraph>
<paragraph id="C83BAA2D533FFFDFC9AEFE91FB81FB3B" blockId="6.[151,1437,153,1476]" box="[199,1128,260,287]" pageId="6" pageNumber="283">
<emphasis id="FAF0763F533FFFDFC9AEFE91FEC3FB3B" bold="true" box="[199,298,260,287]" pageId="6" pageNumber="283">Habitat.</emphasis>
The species was found in sandy or clay-sandy soils at a depth of
<quantity id="0F7C07C8533FFFDFCD6FFE90FB8DFB3B" box="[1030,1124,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="6" pageNumber="283" unit="cm" value="5.0" valueMax="10.0" valueMin="0.0">0–10cm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="809EF9A6533FFFDFC9AEFEBCFE67FFE0" pageId="6" pageNumber="283" type="discussion">
<paragraph id="C83BAA2D533FFFDFC9AEFEBCFD69FB94" blockId="6.[151,1437,153,1476]" pageId="6" pageNumber="283">
<emphasis id="FAF0763F533FFFDFC9AEFEBCFEDDFB67" bold="true" box="[199,308,297,323]" pageId="6" pageNumber="283">Remarks</emphasis>
. The new species can be assigned to the
<taxonomicName id="0F84D1AE533FFFDFCA6BFEBCFC23FB67" baseAuthorityName="Beddard" baseAuthorityYear="1892" box="[770,970,297,323]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="morrisi">
<emphasis id="FAF0763F533FFFDFCA6BFEBCFC23FB67" box="[770,970,297,323]" italics="true" pageId="6" pageNumber="283">Amynthas morrisi</emphasis>
</taxonomicName>
species group due to the presence of two pairs of spermathecal pores in 5/6/7 (
<bibRefCitation id="AC15D7DC533FFFDFCB50FED8FCC1FB43" author="Sims, R. W. &amp; Easton, E. G." box="[569,808,333,359]" pageId="6" pageNumber="283" pagination="169 - 268" refId="ref11103" refString="Sims, R. W. &amp; Easton, E. G. (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of Linnean Society, 4, 169 - 268. https: // doi. org / 10.1111 / j. 1095 - 8312.1972. tb 00694. x" type="journal article" year="1972">Sims &amp; Easton 1972</bibRefCitation>
). To date, the
<taxonomicName id="0F84D1AE533FFFDFCAAEFED8FBD1FB43" baseAuthorityName="Beddard" baseAuthorityYear="1892" box="[967,1080,333,359]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="morrisi">
<emphasis id="FAF0763F533FFFDFCAAEFED8FBD1FB43" box="[967,1080,333,359]" italics="true" pageId="6" pageNumber="283">A. morrisi</emphasis>
</taxonomicName>
group comprises 47 species, of which 13 species have been recorded in northern and central
<collectingCountry id="B093EABD533FFFDFCA21FEE4FC4EFBAF" box="[840,935,369,395]" name="Vietnam" pageId="6" pageNumber="283">Vietnam</collectingCountry>
(
<bibRefCitation id="AC15D7DC533FFFDFCADFFEE4FBC4FBAF" author="Thai, T. B." box="[950,1069,369,395]" pageId="6" pageNumber="283" pagination="817 - 830" refId="ref11223" refString="Thai, T. B. (1982) The fauna of earthworms of the Cucfuong forest reserve (Vietnam) and descriptions of new species of the genus Pheretima. Zoologicheskii Zhurnal, 61 (6), 817 - 830." type="journal article" year="1982">Thai 1982</bibRefCitation>
,
<bibRefCitation id="AC15D7DC533FFFDFCD50FEE4FB69FBAF" author="Thai, T. B." box="[1081,1152,369,395]" pageId="6" pageNumber="283" pagination="284 - 288" refId="ref11266" refString="Thai, T. B. (1984 a) New data on taxonomy of the genus Pheretima (Oligochaeta, Megascolecidae) of the fauna of Vietnam. Zoologicheskii Zhurnal, 63 (2), 284 - 288." type="journal article" year="1984">1984a</bibRefCitation>
, b;
<bibRefCitation id="AC15D7DC533FFFDFCDC0FEE4FAE0FBAF" author="Le, V. T." box="[1193,1289,368,395]" pageId="6" pageNumber="283" pagination="59 - 61" refId="ref10011" refString="Le, V. T. (1995) Two new species of Pheretima of earthworms was described from Lang Son and Cao Bang Provinces. TAP CHI SINH HOC, 17 (3), 59 - 61." type="journal article" year="1995">Le 1995</bibRefCitation>
;
<bibRefCitation id="AC15D7DC533FFFDFCC7CFEE4FA7EFBAF" author="Pham, T. H. H." box="[1301,1431,368,395]" pageId="6" pageNumber="283" pagination="22 - 24" refId="ref10895" refString="Pham, T. H. H. (1995) Three new species of earthworm of genus Pheretima Kinberg, 1867 (Megascolecidae: Oligochaeta) from Quang Nam - Da Nang Province. TAP CHI SINH HOC, 17 (1), 22 - 24." type="journal article" year="1995">Pham 1995</bibRefCitation>
;
<bibRefCitation id="AC15D7DC533FFFDFC9FEFE00FED3FB8B" author="Do, V. N. &amp; Le, V. T. &amp; Tran, M. K." box="[151,314,405,431]" pageId="6" pageNumber="283" pagination="22 - 28" refId="ref9595" refString="Do, V. N., Le, V. T. &amp; Tran, M. K. (1994) New species of earthworm genus Pheretima Kinberg, 1867 (Megascolecidae, Oligochaeta) from Lang Son and Nghe An Provinces. TAP CHI SINH HOC, 16 (1), 22 - 28." type="journal article" year="1994">
Do
<emphasis id="FAF0763F533FFFDFC9A9FE03FF1BFB8B" box="[192,242,405,431]" italics="true" pageId="6" pageNumber="283">et al</emphasis>
. 1994
</bibRefCitation>
,
<bibRefCitation id="AC15D7DC533FFFDFC82CFE01FE69FB8B" author="Do, V. N. &amp; Tran, M. K. &amp; Le, V. T." box="[325,384,404,431]" pageId="6" pageNumber="283" pagination="88 - 94" refId="ref9654" refString="Do, V. N., Tran, M. K. &amp; Le, V. T. (1995) New species of earthworm of genus Pheretima Kinberg, 1867 (Megascolecidae, Oligochaeta) from mountain of Nghe An Province. TAP CHI SINH HOC, 17 (3), 88 - 94." type="journal article" year="1995">1995</bibRefCitation>
;
<bibRefCitation id="AC15D7DC533FFFDFC8E5FE00FD9DFB94" author="Nguyen, T. T. &amp; Nguyen, A. D. &amp; Tran, T. T. B. &amp; Blakemore, R. J." box="[396,628,405,432]" pageId="6" pageNumber="283" pagination="1 - 92" refId="ref10413" refString="Nguyen, T. T., Nguyen, A. D., Tran, T. T. B. &amp; Blakemore, R. J. (2016 a) A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae). Zootaxa, 4140 (1), 1 - 92. https: // doi. org / 10.11646 / zootaxa. 4140.1.1" type="journal article" year="2016">
Nguyen
<emphasis id="FAF0763F533FFFDFC885FE03FDCDFB8B" box="[492,548,405,431]" italics="true" pageId="6" pageNumber="283">et al.</emphasis>
2016a
</bibRefCitation>
).
</paragraph>
<paragraph id="C83BAA2D533FFFDFC9AEFE2CFEF6F933" blockId="6.[151,1437,153,1476]" pageId="6" pageNumber="283">
Within the
<taxonomicName id="0F84D1AE533FFFDFC82DFE2CFE5CFBF7" baseAuthorityName="Beddard" baseAuthorityYear="1892" box="[324,437,441,467]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="morrisi">
<emphasis id="FAF0763F533FFFDFC82DFE2CFE5CFBF7" box="[324,437,441,467]" italics="true" pageId="6" pageNumber="283">A. morrisi</emphasis>
</taxonomicName>
group, the new species is similar to
<taxonomicName id="0F84D1AE533FFFDFCA27FE2CFB31FBF7" authority="(Do &amp; Tran, 1995)" baseAuthorityName="Do &amp; Tran" baseAuthorityYear="1995" box="[846,1240,440,467]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="dorsomorrisi">
<emphasis id="FAF0763F533FFFDFCA27FE2CFC16FBF7" box="[846,1023,441,467]" italics="true" pageId="6" pageNumber="283">A. dorsomorrisi</emphasis>
(
<bibRefCitation id="AC15D7DC533FFFDFCD67FE2CFB39FBF7" author="Do, V. N. &amp; Tran, M. K. &amp; Le, V. T." box="[1038,1232,440,467]" pageId="6" pageNumber="283" pagination="88 - 94" refId="ref9654" refString="Do, V. N., Tran, M. K. &amp; Le, V. T. (1995) New species of earthworm of genus Pheretima Kinberg, 1867 (Megascolecidae, Oligochaeta) from mountain of Nghe An Province. TAP CHI SINH HOC, 17 (3), 88 - 94." type="journal article" year="1995">Do &amp; Tran, 1995</bibRefCitation>
)
</taxonomicName>
in the absence of genital markings in the spermathecal region, simple intestinal caeca and a similar body size. Both species can be distinguished as follows: The new species has spermathecal pores located at some distance from the dorsal line (they are actually in dorsolateral position), a pre-setal pair of large round or oval genital markings in xvii, first dorsal pore in 12/13, about 6–7 setae between the male porophores, intestinal origin at xiv, and a folded typhlosole. In contrast,
<taxonomicName id="0F84D1AE533FFFDFC9FEFDF8FEA1F8A3" baseAuthorityName="Do &amp; Tran" baseAuthorityYear="1995" box="[151,328,621,647]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="dorsomorrisi">
<emphasis id="FAF0763F533FFFDFC9FEFDF8FEA1F8A3" box="[151,328,621,647]" italics="true" pageId="6" pageNumber="283">A. dorsomorrisi</emphasis>
</taxonomicName>
has spermathecal pores located closely to the dorsal line, one pair of small genital markings located behind the male pores in xviii, first dorsal pore in 11/12, no setae between two male porophores, intestinal origin at xv, and no typhlosole (
<bibRefCitation id="AC15D7DC533FFFDFC8F0FD20FDA8F8EB" author="Do, V. N. &amp; Tran, M. K. &amp; Le, V. T." box="[409,577,692,719]" pageId="6" pageNumber="283" pagination="88 - 94" refId="ref9654" refString="Do, V. N., Tran, M. K. &amp; Le, V. T. (1995) New species of earthworm of genus Pheretima Kinberg, 1867 (Megascolecidae, Oligochaeta) from mountain of Nghe An Province. TAP CHI SINH HOC, 17 (3), 88 - 94." type="journal article" year="1995">
Do
<emphasis id="FAF0763F533FFFDFC8ADFD23FE14F8EB" box="[452,509,693,719]" italics="true" pageId="6" pageNumber="283">et al.</emphasis>
1995
</bibRefCitation>
). In addition, the new species has 135–151 segments, transparent coloration and thinner body wall, whereas
<taxonomicName id="0F84D1AE533FFFDFCB6BFD4CFD5DF8D7" baseAuthorityName="Do &amp; Tran" baseAuthorityYear="1995" box="[514,692,729,755]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="dorsomorrisi">
<emphasis id="FAF0763F533FFFDFCB6BFD4CFD5DF8D7" box="[514,692,729,755]" italics="true" pageId="6" pageNumber="283">A. dorsomorrisi</emphasis>
</taxonomicName>
has 91 segments, light grey coloration and thicker body wall (
<bibRefCitation id="AC15D7DC533FFFDFCC11FD4CFEFBF933" author="Do, V. N. &amp; Tran, M. K. &amp; Le, V. T." pageId="6" pageNumber="283" pagination="88 - 94" refId="ref9654" refString="Do, V. N., Tran, M. K. &amp; Le, V. T. (1995) New species of earthworm of genus Pheretima Kinberg, 1867 (Megascolecidae, Oligochaeta) from mountain of Nghe An Province. TAP CHI SINH HOC, 17 (3), 88 - 94." type="journal article" year="1995">
Do
<emphasis id="FAF0763F533FFFDFC9FEFD6BFF26F933" box="[151,207,765,791]" italics="true" pageId="6" pageNumber="283">et al.</emphasis>
1995
</bibRefCitation>
).
</paragraph>
<paragraph id="C83BAA2D533FFFDFC9AEFCB4FD9FFEE3" blockId="6.[151,1437,153,1476]" pageId="6" pageNumber="283">
The new species shares the character state of dorsal spermathecal pores with several other
<taxonomicName id="0F84D1AE533FFFDFCDBDFCB4FAABF91F" authorityName="Kinberg" authorityYear="1867" box="[1236,1346,801,827]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="genus">
<emphasis id="FAF0763F533FFFDFCDBDFCB4FAABF91F" box="[1236,1346,801,827]" italics="true" pageId="6" pageNumber="283">Amynthas</emphasis>
</taxonomicName>
species outside the
<taxonomicName id="0F84D1AE533FFFDFC87EFCD0FE6EF97B" baseAuthorityName="Beddard" baseAuthorityYear="1892" box="[279,391,837,863]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="morrisi">
<emphasis id="FAF0763F533FFFDFC87EFCD0FE6EF97B" box="[279,391,837,863]" italics="true" pageId="6" pageNumber="283">A. morrisi</emphasis>
</taxonomicName>
-group, such as,
<taxonomicName id="0F84D1AE533FFFDFCB54FCD0FC9EF944" authority="(Chen, 1946)" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[573,887,837,864]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="acidophilus">
<emphasis id="FAF0763F533FFFDFCB54FCD0FD37F97B" box="[573,734,837,863]" italics="true" pageId="6" pageNumber="283">A. acidophilus</emphasis>
(
<bibRefCitation id="AC15D7DC533FFFDFCB85FCD0FC86F944" author="Chen, Y." box="[748,879,837,864]" pageId="6" pageNumber="283" pagination="83 - 141" refId="ref9559" refString="Chen, Y. (1946) On the terrestrial oligochaeta from Szechuan III. Contributions from the Biological Laboratory of the Science Society of China, Zoological Series, 16, 83 - 141." type="journal article" year="1946">Chen, 1946</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="0F84D1AE533FFFDFCAEDFCD0FB59F97B" authority="(Pham, 1995)" baseAuthorityName="Pham" baseAuthorityYear="1995" box="[900,1200,836,863]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="banaensis">
<emphasis id="FAF0763F533FFFDFCAEDFCD0FBFBF97B" box="[900,1042,837,863]" italics="true" pageId="6" pageNumber="283">A. banaensis</emphasis>
(
<bibRefCitation id="AC15D7DC533FFFDFCD49FCD0FB4EF97B" author="Pham, T. H. H." box="[1056,1191,836,863]" pageId="6" pageNumber="283" pagination="22 - 24" refId="ref10895" refString="Pham, T. H. H. (1995) Three new species of earthworm of genus Pheretima Kinberg, 1867 (Megascolecidae: Oligochaeta) from Quang Nam - Da Nang Province. TAP CHI SINH HOC, 17 (1), 22 - 24." type="journal article" year="1995">Pham, 1995</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="0F84D1AE533FFFDFCDD5FCD0FE5BF9A7" authority="(Le, 1994) (in Do et al. 1994)" authorityName="in Do et al." authorityYear="1994" baseAuthorityName="Le" baseAuthorityYear="1994" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="binhgiaensis">
<emphasis id="FAF0763F533FFFDFCDD5FCD0FA80F97B" box="[1212,1385,837,863]" italics="true" pageId="6" pageNumber="283">A. binhgiaensis</emphasis>
(Le, 1994) (in
<bibRefCitation id="AC15D7DC533FFFDFC86EFCFCFE40F9A7" author="Do, V. N. &amp; Le, V. T. &amp; Tran, M. K." box="[263,425,873,899]" pageId="6" pageNumber="283" pagination="22 - 28" refId="ref9595" refString="Do, V. N., Le, V. T. &amp; Tran, M. K. (1994) New species of earthworm genus Pheretima Kinberg, 1867 (Megascolecidae, Oligochaeta) from Lang Son and Nghe An Provinces. TAP CHI SINH HOC, 16 (1), 22 - 28." type="journal article" year="1994">
Do
<emphasis id="FAF0763F533FFFDFC859FCFFFE8BF9A7" box="[304,354,873,899]" italics="true" pageId="6" pageNumber="283">et al</emphasis>
. 1994
</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="0F84D1AE533FFFDFC8D6FCFCFCFAF9A7" authority="(Le, 1995)" baseAuthorityName="Le" baseAuthorityYear="1995" box="[447,787,872,899]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="multichaetiferus">
<emphasis id="FAF0763F533FFFDFC8D6FCFCFD7CF9A7" box="[447,661,873,899]" italics="true" pageId="6" pageNumber="283">A. multichaetiferus</emphasis>
(
<bibRefCitation id="AC15D7DC533FFFDFCBCCFCFCFCE2F9A7" author="Le, V. T." box="[677,779,872,899]" pageId="6" pageNumber="283" pagination="59 - 61" refId="ref10011" refString="Le, V. T. (1995) Two new species of Pheretima of earthworms was described from Lang Son and Cao Bang Provinces. TAP CHI SINH HOC, 17 (3), 59 - 61." type="journal article" year="1995">Le, 1995</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="0F84D1AE533FFFDFCA48FCFCFB55F9A7" authority="(Thai, 1982)" baseAuthorityName="Thai" baseAuthorityYear="1982" box="[801,1212,873,899]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="ophthalmopapillatus">
<emphasis id="FAF0763F533FFFDFCA48FCFCFBC0F9A7" box="[801,1065,873,899]" italics="true" pageId="6" pageNumber="283">A. ophthalmopapillatus</emphasis>
(
<bibRefCitation id="AC15D7DC533FFFDFCD51FCFCFB5DF9A7" author="Thai, T. B." box="[1080,1204,873,899]" pageId="6" pageNumber="283" pagination="817 - 830" refId="ref11223" refString="Thai, T. B. (1982) The fauna of earthworms of the Cucfuong forest reserve (Vietnam) and descriptions of new species of the genus Pheretima. Zoologicheskii Zhurnal, 61 (6), 817 - 830." type="journal article" year="1982">Thai, 1982</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="0F84D1AE533FFFDFCD9AFCFCFEDEF983" authority="(Grube, 1879)" baseAuthorityName="Grube" baseAuthorityYear="1879" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="rodericensis">
<emphasis id="FAF0763F533FFFDFCD9AFCFCFA75F9A7" box="[1267,1436,873,899]" italics="true" pageId="6" pageNumber="283">A. rodericensis</emphasis>
(
<bibRefCitation id="AC15D7DC533FFFDFC9F6FC18FEC6F983" author="Grube, E." box="[159,303,909,935]" pageId="6" pageNumber="283" pagination="554 - 556" refId="ref9889" refString="Grube, E. (1879) Annelida. Philosophical Transactions of the Royal Society, 168, 554 - 556. https: // doi. org / 10.1098 / rstl. 1879.0057." type="journal article" year="1879">Grube, 1879</bibRefCitation>
)
</taxonomicName>
. However, the new species is distinguished from these species by having two pair of spermathecal pores in 5/6/7 while
<taxonomicName id="0F84D1AE533FFFDFC8E9FC24FDF9F9EF" baseAuthorityName="Pham" baseAuthorityYear="1995" box="[384,528,945,971]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="banaensis">
<emphasis id="FAF0763F533FFFDFC8E9FC24FDF9F9EF" box="[384,528,945,971]" italics="true" pageId="6" pageNumber="283">A. banaensis</emphasis>
</taxonomicName>
and
<taxonomicName id="0F84D1AE533FFFDFCB21FC24FCA6F9EF" baseAuthorityName="Thai" baseAuthorityYear="1982" box="[584,847,945,971]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="ophthalmopapillatus">
<emphasis id="FAF0763F533FFFDFCB21FC24FCA6F9EF" box="[584,847,945,971]" italics="true" pageId="6" pageNumber="283">A. ophthalmopapillatus</emphasis>
</taxonomicName>
has one pair of pores in 5/6;
<taxonomicName id="0F84D1AE533FFFDFCDF2FC24FAD7F9EF" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[1179,1342,945,971]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="acidophilus">
<emphasis id="FAF0763F533FFFDFCDF2FC24FAD7F9EF" box="[1179,1342,945,971]" italics="true" pageId="6" pageNumber="283">A. acidophilus</emphasis>
</taxonomicName>
has two pairs in 6/7/8;
<taxonomicName id="0F84D1AE533FFFDFC85FFC40FE25F9CB" baseAuthorityName="Thai" baseAuthorityYear="1984" box="[310,460,981,1007]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="arrobustus">
<emphasis id="FAF0763F533FFFDFC85FFC40FE25F9CB" box="[310,460,981,1007]" italics="true" pageId="6" pageNumber="283">A. arrobustus</emphasis>
</taxonomicName>
and
<taxonomicName id="0F84D1AE533FFFDFCB69FC40FD3DF9CB" baseAuthorityName="Le" baseAuthorityYear="1995" box="[512,724,981,1007]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="multichaetiferus">
<emphasis id="FAF0763F533FFFDFCB69FC40FD3DF9CB" box="[512,724,981,1007]" italics="true" pageId="6" pageNumber="283">A. multichaetiferus</emphasis>
</taxonomicName>
has two pairs in 7/8/9;
<taxonomicName id="0F84D1AE533FFFDFCABDFC40FB69F9CB" authorityName="in Do et al." authorityYear="1994" baseAuthorityName="Le" baseAuthorityYear="1994" box="[980,1152,981,1007]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="binhgiaensis">
<emphasis id="FAF0763F533FFFDFCABDFC40FB69F9CB" box="[980,1152,981,1007]" italics="true" pageId="6" pageNumber="283">A. binhgiaensis</emphasis>
</taxonomicName>
has three pairs in 5/6/7/8, lastly
<taxonomicName id="0F84D1AE533FFFDFC9B3FC6CFE6BFE37" baseAuthorityName="Grube" baseAuthorityYear="1879" box="[218,386,1017,1043]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="rodericensis">
<emphasis id="FAF0763F533FFFDFC9B3FC6CFE6BFE37" box="[218,386,1017,1043]" italics="true" pageId="6" pageNumber="283">A. rodericensis</emphasis>
</taxonomicName>
four pairs in 5/6/7/8/9. In addition,
<taxonomicName id="0F84D1AE533FFFDFCA66FC6CFBD6FE37" authority="Nguyen &amp; Tran &amp; Lam &amp; Nguyen, 2020" authorityName="Nguyen &amp; Tran &amp; Lam &amp; Nguyen" authorityYear="2020" box="[783,1087,1017,1043]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="dorsomorrioides" status="sp. nov.">
<emphasis id="FAF0763F533FFFDFCA66FC6CFBD6FE37" box="[783,1087,1017,1043]" italics="true" pageId="6" pageNumber="283">Amynthas dorsomorrioides</emphasis>
</taxonomicName>
<emphasis id="FAF0763F533FFFDFCD2CFC6CFB77FE37" bold="true" box="[1093,1182,1017,1043]" pageId="6" pageNumber="283">
<taxonomicNameLabel id="E1C3CB44533FFFDFCD2CFC6CFB77FE37" box="[1093,1182,1017,1043]" pageId="6" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also differs from these species (but not from
<taxonomicName id="0F84D1AE533FFFDFC8E2FB88FDF3FE13" baseAuthorityName="Pham" baseAuthorityYear="1995" box="[395,538,1053,1079]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="banaensis">
<emphasis id="FAF0763F533FFFDFC8E2FB88FDF3FE13" box="[395,538,1053,1079]" italics="true" pageId="6" pageNumber="283">A. banaensis</emphasis>
</taxonomicName>
) in having a pre-setal pair of large round or oval genital markings in xvii. These species except
<taxonomicName id="0F84D1AE533FFFDFC82BFBD4FE3AFE7F" baseAuthorityName="Pham" baseAuthorityYear="1995" box="[322,467,1089,1115]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="banaensis">
<emphasis id="FAF0763F533FFFDFC82BFBD4FE3AFE7F" box="[322,467,1089,1115]" italics="true" pageId="6" pageNumber="283">A. banaensis</emphasis>
</taxonomicName>
have genital markings paired in xviii or paired in 17/18 and 18/19 (
<taxonomicName id="0F84D1AE533FFFDFCDB7FBD4FA65FE7F" authorityName="in Do et al." authorityYear="1994" baseAuthorityName="Le" baseAuthorityYear="1994" box="[1246,1420,1089,1115]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="binhgiaensis">
<emphasis id="FAF0763F533FFFDFCDB7FBD4FA65FE7F" box="[1246,1420,1089,1115]" italics="true" pageId="6" pageNumber="283">A. binhgiaensis</emphasis>
</taxonomicName>
). The new species shares the character states of one pair of genital markings in xvii and first dorsal pore in 12/13 but they are clearly distinguished by genital markings in the spermathecal region (absent in
<taxonomicName id="0F84D1AE533FFFDFCD0CFB1CFAD7FE87" authority="Nguyen &amp; Tran &amp; Lam &amp; Nguyen, 2020" authorityName="Nguyen &amp; Tran &amp; Lam &amp; Nguyen" authorityYear="2020" box="[1125,1342,1161,1187]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="dorsomorrioides" status="sp. nov.">
<emphasis id="FAF0763F533FFFDFCD0CFB1CFAD7FE87" box="[1125,1342,1161,1187]" italics="true" pageId="6" pageNumber="283">A. dorsomorrioides</emphasis>
</taxonomicName>
<emphasis id="FAF0763F533FFFDFCC2DFB1CFA75FE87" bold="true" box="[1348,1436,1161,1187]" pageId="6" pageNumber="283">
<taxonomicNameLabel id="E1C3CB44533FFFDFCC2DFB1CFA75FE87" box="[1348,1436,1161,1187]" pageId="6" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
vs. two pairs in vi and vii in
<taxonomicName id="0F84D1AE533FFFDFC8BEFB38FD8FFEE3" baseAuthorityName="Pham" baseAuthorityYear="1995" box="[471,614,1197,1223]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="banaensis">
<emphasis id="FAF0763F533FFFDFC8BEFB38FD8FFEE3" box="[471,614,1197,1223]" italics="true" pageId="6" pageNumber="283">A. banaensis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="C83BAA2D533FFFDFC9AEFB44FE67FFE0" blockId="6.[151,1437,153,1476]" pageId="6" pageNumber="283">
Regarding the arrangement of genital markings in the male region, the new species is fairly similar to
<taxonomicName id="0F84D1AE533FFFDFCC3AFB44FEC4FF2B" authority="(Chen, 1935)" baseAuthorityName="Chen" baseAuthorityYear="1935" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="wui">
<emphasis id="FAF0763F533FFFDFCC3AFB44FA75FECF" box="[1363,1436,1233,1259]" italics="true" pageId="6" pageNumber="283">A. wui</emphasis>
(
<bibRefCitation id="AC15D7DC533FFFDFC9F6FB60FECCFF2B" author="Chen, Y." box="[159,293,1268,1295]" pageId="6" pageNumber="283" pagination="109 - 122" refId="ref9482" refString="Chen, Y. (1935) On two new species of Oligochaeta from Amoy. Contributions from the Biological Laboratory of the Science Society of China, Zoological Series, 11 (4), 109 - 122." type="journal article" year="1935">Chen, 1935</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="0F84D1AE533FFFDFC801FB60FD91FF2B" authority="(Thai, 1984)" baseAuthorityName="Thai" baseAuthorityYear="1984" box="[360,632,1269,1295]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="adexilis">
<emphasis id="FAF0763F533FFFDFC801FB60FE09FF2B" box="[360,480,1269,1295]" italics="true" pageId="6" pageNumber="283">A. adexilis</emphasis>
(Thai, 1984)
</taxonomicName>
by having only one pair in xvii; further similarities include epilobous prostomium, holandry, simple intestinal caeca, and absence of septa 8/9/10. However, these species can be distinguished by position and number of spermathecal pores (laterodorsally 5/6/
<quantity id="0F7C07C8533FFFDFCD6BFAA9FBC7FF73" box="[1026,1070,1340,1367]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="6" pageNumber="283" unit="in" value="7.0">7 in</quantity>
<taxonomicName id="0F84D1AE533FFFDFCD5FFAA8FAF8FF73" authority="Nguyen &amp; Tran &amp; Lam &amp; Nguyen, 2020" authorityName="Nguyen &amp; Tran &amp; Lam &amp; Nguyen" authorityYear="2020" box="[1078,1297,1341,1367]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="dorsomorrioides" status="sp. nov.">
<emphasis id="FAF0763F533FFFDFCD5FFAA8FAF8FF73" box="[1078,1297,1341,1367]" italics="true" pageId="6" pageNumber="283">A. dorsomorrioides</emphasis>
</taxonomicName>
<emphasis id="FAF0763F533FFFDFCC70FAA8FA9DFF73" bold="true" box="[1305,1396,1341,1367]" pageId="6" pageNumber="283">
<taxonomicNameLabel id="E1C3CB44533FFFDFCC70FAA8FA9DFF73" box="[1305,1396,1341,1367]" pageId="6" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
vs. lateroventrally 5/
<quantity id="0F7C07C8533FFFDFC830FAF5FE6CFF5F" box="[345,389,1376,1403]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="283" unit="in" value="6.0">6 in</quantity>
<taxonomicName id="0F84D1AE533FFFDFC8E7FAF4FE3FFF5F" baseAuthorityName="Chen" baseAuthorityYear="1935" box="[398,470,1377,1403]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="wui">
<emphasis id="FAF0763F533FFFDFC8E7FAF4FE3FFF5F" box="[398,470,1377,1403]" italics="true" pageId="6" pageNumber="283">A. wui</emphasis>
</taxonomicName>
, lateroventrally 5/6/7/8 or 5/6/7/8/
<quantity id="0F7C07C8533FFFDFCA0BFAF5FC64FF5F" box="[866,909,1376,1403]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="6" pageNumber="283" unit="in" value="9.0">9 in</quantity>
<taxonomicName id="0F84D1AE533FFFDFCAFFFAF4FBE4FF5F" baseAuthorityName="Thai" baseAuthorityYear="1984" box="[918,1037,1377,1403]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="283" phylum="Annelida" rank="species" species="adexilis">
<emphasis id="FAF0763F533FFFDFCAFFFAF4FBE4FF5F" box="[918,1037,1377,1403]" italics="true" pageId="6" pageNumber="283">A. adexilis</emphasis>
</taxonomicName>
), status of male sexual system and intestinal origin (separated testis sacs and xiv in the new species vs. connected testis sacs and xv or xvi in these three species, respectively).
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink