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<document id="09AC77B37CF07D543FD3F876C54BFD60" ID-DOI="10.11646/zootaxa.4790.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="3889527" ID-ZooBank="46EAA29B-7852-4374-B844-DA7E25A00ABD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591859566901" checkinUser="plazi" docAuthor="Nguyen, Tung T., Tran, Binh T. T., Lam, Dang H. &amp; Nguyen, Anh D." docDate="2020" docId="402D1B3B533BFFDDC9FEFD8FFC60FE13" docLanguage="en" docName="zootaxa.4790.2.5.pdf" docOrigin="Zootaxa 4790 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amynthas longiprostaticus Nguyen &amp; Tran &amp; Lam &amp; Nguyen 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="281" masterDocId="BC1463435339FFD9C969FF95FFE9FA24" masterDocTitle="Four new species of Amynthas earthworms in southeastern Vietnam (Annelida, Oligochaeta, Megascolecidae)" masterLastPageNumber="290" masterPageNumber="277" pageNumber="279" updateTime="1700276591635" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="252D73A21E8B14D306217ECD13381273" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5EB19DB915A78E2C474176E0DFC3613A">
<mods:title id="944C5487A48DFE53626E786FDE7613CC">Four new species of Amynthas earthworms in southeastern Vietnam (Annelida, Oligochaeta, Megascolecidae)</mods:title>
</mods:titleInfo>
<mods:name id="E5CF9CB7D3878D1A7B9B1F028CB3DD98" type="personal">
<mods:role id="661BC6132EFFE47BA931F0E90372ADEB">
<mods:roleTerm id="356EDEF67EF8F80A2E805DDD130F9805">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4DB3A059DC3C8BDF8DEB15B69952DD51">Nguyen, Tung T.</mods:namePart>
</mods:name>
<mods:name id="CB337000FFA6295857CF6AE619C11B2B" type="personal">
<mods:role id="BB138491191FE82C8E3A8689E55D99E3">
<mods:roleTerm id="60822D8ED521F2BE90EA4E3ACC50FEF8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7097BF7A78E0F4A6C83EFE06694E30AB">Tran, Binh T. T.</mods:namePart>
<mods:affiliation id="FC28AAA32679B2D6B0F6A32793930D76">Department of Zoology, Faculty of Biology, Hanoi National University of Education, Hanoi, Vietnam.</mods:affiliation>
</mods:name>
<mods:name id="9E23D1646BB2E4E87DFBD65E69BDB606" type="personal">
<mods:role id="0C4048C73734075C40944F1CB5088CA6">
<mods:roleTerm id="56FC4CE3314A46EA6AA92ACF1993B969">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FD4958673E93D5318A0F0DFBA5B5F65E">Lam, Dang H.</mods:namePart>
</mods:name>
<mods:name id="7759FD1BA429EF4A4EF79704D79BA3A1" type="personal">
<mods:role id="036F5159DE2D9F2C4AF5F4C71DA0A72D">
<mods:roleTerm id="4EA784BABAC3C8D55945D5602B246898">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B0C7484B4DF64A34C314BC738D94D45">Nguyen, Anh D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A384E68FDB79E05B4ED3BD246CB6759E">text</mods:typeOfResource>
<mods:relatedItem id="055767ADCB0C80C0658D60FA331E2064" type="host">
<mods:titleInfo id="E667A8CC606545C19AE4618447DC0536">
<mods:title id="064E9C66D5374397B1B233FE8B0B2BC5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A80583E6E7F89B5EA8CD52968303416B">
<mods:date id="E8CB932389EC3E02F7931C5299E53312">2020</mods:date>
<mods:detail id="A860CE194F746C902E044128BA68DDA1" type="pubDate">
<mods:number id="6DD84B7087FF793B8E54CB02ECF2FB5D">2020-06-11</mods:number>
</mods:detail>
<mods:detail id="139514947432A4A5491C8680FB2410F4" type="volume">
<mods:number id="0C2F72518A201346409D48AFED6B011D">4790</mods:number>
</mods:detail>
<mods:detail id="C154934A18BCBEFA9AB350F86B3E1A0E" type="issue">
<mods:number id="E4EE1FBDE4C1F911F46AD107A403E428">2</mods:number>
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<mods:start id="928734478B4EB7611C49AC94A4F712BC">277</mods:start>
<mods:end id="14EE403FE6E891E4C17A2E04E29A2FF7">290</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="C6D7A7E119C19B81370969F871C17C87">journal article</mods:classification>
<mods:identifier id="8947475C17409887920230C096AAFBFC" type="DOI">10.11646/zootaxa.4790.2.5</mods:identifier>
<mods:identifier id="ED7E5ACE9F839CB4E2E91789DD5456AC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3F0ACE87E9DB9EAB67490464CF76F68E" type="Zenodo-Dep">3889527</mods:identifier>
<mods:identifier id="476539AC28A0C76C492C5E533FF239DB" type="ZooBank">46EAA29B-7852-4374-B844-DA7E25A00ABD</mods:identifier>
</mods:mods>
<treatment id="402D1B3B533BFFDDC9FEFD8FFC60FE13" LSID="urn:lsid:plazi:treatment:402D1B3B533BFFDDC9FEFD8FFC60FE13" httpUri="http://treatment.plazi.org/id/402D1B3B533BFFDDC9FEFD8FFC60FE13" lastPageId="4" lastPageNumber="281" pageId="2" pageNumber="279">
<subSubSection id="809EF9A6533BFFDBC9FEFD8FFCF9F811" box="[151,784,538,565]" pageId="2" pageNumber="279" type="nomenclature">
<paragraph id="C83BAA2D533BFFDBC9FEFD8FFCF9F811" blockId="2.[151,784,538,600]" box="[151,784,538,565]" pageId="2" pageNumber="279">
<heading id="93731D41533BFFDBC9FEFD8FFCF9F811" bold="true" box="[151,784,538,565]" fontSize="11" level="1" pageId="2" pageNumber="279" reason="1">
<emphasis id="FAF0763F533BFFDBC9FEFD8FFCF9F811" bold="true" box="[151,784,538,565]" pageId="2" pageNumber="279">
<taxonomicName id="0F84D1AE533BFFDBC9FEFD8FFD4CF811" authority="Nguyen &amp; Lam" authorityName="Nguyen &amp; Tran &amp; Lam &amp; Nguyen" authorityYear="2020" box="[151,677,538,565]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="279" phylum="Annelida" rank="species" species="longiprostaticus" status="sp. nov.">
<emphasis id="FAF0763F533BFFDBC9FEFD8FFE32F811" bold="true" box="[151,475,538,565]" italics="true" pageId="2" pageNumber="279">Amynthas longiprostaticus</emphasis>
Nguyen &amp; Lam
</taxonomicName>
,
<taxonomicNameLabel id="E1C3CB44533BFFDBCBDAFD8EFCF9F811" box="[691,784,539,565]" pageId="2" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="809EF9A6533BFFDBC9FEFDABFEE1F87C" box="[151,264,574,600]" pageId="2" pageNumber="279" type="description">
<paragraph id="C83BAA2D533BFFDBC9FEFDABFEE1F87C" blockId="2.[151,784,538,600]" box="[151,264,574,600]" pageId="2" pageNumber="279">
(
<figureCitation id="50BFB6A8533BFFDBC9F6FDABFEE9F87C" box="[159,256,574,600]" captionStart="FIGURE 2" captionStartId="3.[151,250,1785,1810]" captionTargetBox="[164,1423,200,1747]" captionTargetId="figure@3.[154,1433,181,1760]" captionTargetPageId="3" captionText="FIGURE 2. Amynthas longiprostaticus sp. nov. Holotype (CTU-EW.088.h01).A. Male region, ventral view. B. Left intestinal caecum. C. Left prostate gland. D. Spermathecal region, ventral view. E. Testis sacs and seminal vesicles. F. Right spermatheca. G. Male region (comp. A). H. Spermatheca (comp. F). I. Prostate gland (comp. C). J. Genital marking in x. K, L. Transverse section of male porophore, different section planes (Non-type: CTU-EW.088.03). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3889531" httpUri="https://zenodo.org/record/3889531/files/figure.png" pageId="2" pageNumber="279">Figure 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="809EF9A6533BFFDBC9FEFD10FEE8F970" pageId="2" pageNumber="279" type="materials_examined">
<paragraph id="C83BAA2D533BFFDBC9FEFD10FEE8F970" blockId="2.[151,1437,645,2005]" pageId="2" pageNumber="279">
<emphasis id="FAF0763F533BFFDBC9FEFD10FE6EF884" bold="true" box="[151,391,645,672]" pageId="2" pageNumber="279">Material examined.</emphasis>
<materialsCitation id="78ECA070533BFFDBC8FCFD13FA74F8E0" collectingDate="2013-10-13" collectionCode="CTU-EW" collectorName="Le Van Nhan." latitude="11.425278" location="Tan Phu District" longLatPrecision="21" longitude="107.42833" municipality="Cat Tien National Park" pageId="2" pageNumber="279" specimenCount="1" stateProvince="Dong Nai" typeStatus="holotype">
<emphasis id="FAF0763F533BFFDBC8FCFD13FDEDF884" box="[405,516,646,672]" italics="true" pageId="2" pageNumber="279">
<typeStatus id="173F148F533BFFDBC8FCFD13FE16F884" box="[405,511,646,672]" pageId="2" pageNumber="279" type="holotype">Holotype</typeStatus>
:
</emphasis>
1 mature (complete body), (
<collectionCode id="AE9532E8533BFFDBCA0DFD13FC33F885" box="[868,986,646,673]" pageId="2" pageNumber="279">CTU-EW</collectionCode>
.088.h01), natural forests (
<geoCoordinate id="ADB0CCEA533BFFDBCC65FD10FA7EF884" box="[1292,1431,645,672]" degrees="11" direction="north" minutes="25" orientation="latitude" pageId="2" pageNumber="279" precision="15" seconds="31" value="11.425278">11°25’31”N</geoCoordinate>
;
<geoCoordinate id="ADB0CCEA533BFFDBC9FEFD3CFEC7F8E0" box="[151,302,681,708]" degrees="107" direction="east" minutes="25" orientation="longitude" pageId="2" pageNumber="279" precision="15" seconds="42" value="107.42833">107°25’42”E</geoCoordinate>
),
<collectingMunicipality id="285F3057533BFFDBC82AFD3FFDAEF8E0" box="[323,583,682,708]" pageId="2" pageNumber="279">Cat Tien National Park</collectingMunicipality>
,
<location id="CD5BFCF6533BFFDBCB3BFD3EFCE4F8E0" LSID="urn:lsid:plazi:treatment:402D1B3B533BFFDDC9FEFD8FFC60FE13:CD5BFCF6533BFFDBCB3BFD3EFCE4F8E0" box="[594,781,682,709]" latitude="11.425278" longLatPrecision="21" longitude="107.42833" municipality="Cat Tien National Park" name="Tan Phu District" pageId="2" pageNumber="279" stateProvince="Dong Nai">Tan Phu District</location>
,
<collectingRegion id="0A4064CF533BFFDBCA71FD3EFC1AF8E0" box="[792,1011,682,709]" country="Vietnam" name="Dong Nai" pageId="2" pageNumber="279">Dong Nai Province</collectingRegion>
,
<date id="BC3A8CED533BFFDBCA96FD3FFB57F8E0" box="[1023,1214,682,708]" pageId="2" pageNumber="279" value="2013-10-13">
<collectingDate id="AC7E7505533BFFDBCA96FD3FFB57F8E0" box="[1023,1214,682,708]" pageId="2" pageNumber="279" value="2013-10-13">13 October 2013</collectingDate>
</date>
, coll.
<collectorName id="6571CFFB533BFFDBCC6BFD3EFA74F8E0" box="[1282,1437,682,709]" pageId="2" pageNumber="279">Le Van Nhan.</collectorName>
</materialsCitation>
<materialsCitation id="78ECA070533BFFDBC9FEFD5AFF08F928" collectingDate="2013-10-13" collectionCode="CTU-EW, IEBR-EW" collectorName="Le Van Nhan." latitude="11.425278" location="Tan Phu District" longLatPrecision="21" longitude="107.42833" municipality="Cat Tien National Park" pageId="2" pageNumber="279" specimenCount="1" stateProvince="Dong Nai" typeStatus="paratype">
<emphasis id="FAF0763F533BFFDBC9FEFD5AFEFBF8CC" box="[151,274,719,744]" italics="true" pageId="2" pageNumber="279">
<typeStatus id="173F148F533BFFDBC9FEFD5AFEE5F8CC" box="[151,268,719,744]" pageId="2" pageNumber="279" type="paratype">Paratypes</typeStatus>
:
</emphasis>
5 matures (complete body) (
<collectionCode id="AE9532E8533BFFDBCB37FD5BFD3AF8CC" box="[606,723,718,744]" pageId="2" pageNumber="279">CTU-EW</collectionCode>
.088.p02), 4 matures (
<collectionCode id="AE9532E8533BFFDBCAACFD5BFBABF8CC" box="[965,1090,718,744]" pageId="2" pageNumber="279">IEBR-EW</collectionCode>
.088.p02), same data as for holotype
</materialsCitation>
.
<materialsCitation id="78ECA070533BFFDBC986FD66FD77F914" collectingDate="2013-10-13" collectionCode="CTU-EW" collectorName="Le Van Nhan." latitude="11.425278" location="Tan Phu District" longLatPrecision="21" longitude="107.42833" municipality="Cat Tien National Park" pageId="2" pageNumber="279" specimenCount="4" specimenCount-immature="4" stateProvince="Dong Nai" typeStatus="paratype">
<emphasis id="FAF0763F533BFFDBC986FD66FE80F928" box="[239,361,755,780]" italics="true" pageId="2" pageNumber="279">Non-types:</emphasis>
30 matures (complete body), 1 mature (two fragments and two section slides),
<specimenCount id="DE8261A4533BFFDBCC67FD67FA75F928" box="[1294,1436,754,780]" pageId="2" pageNumber="279" type="immature">4 immatures</specimenCount>
(
<collectionCode id="AE9532E8533BFFDBC9F6FC83FEF9F914" box="[159,272,790,816]" pageId="2" pageNumber="279">CTU-EW</collectionCode>
.088.03) same data as for holotype
</materialsCitation>
;
<materialsCitation id="78ECA070533BFFDBCBC2FC83FF14F970" collectingDate="2013-10-13" collectionCode="IEBR-EW" collectorName="Le Van Nhan." latitude="11.425278" location="Tan Phu District" longLatPrecision="21" longitude="107.42833" municipality="Cat Tien National Park" pageId="2" pageNumber="279" specimenCount="1" stateProvince="Dong Nai" typeStatus="paratype">
17 matures (complete body) (
<collectionCode id="AE9532E8533BFFDBCA97FC83FB91F914" box="[1022,1144,790,816]" pageId="2" pageNumber="279">IEBR-EW</collectionCode>
.088.03), same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="809EF9A6533BFFDBC9AEFCCBFB8FF95C" box="[199,1126,862,889]" pageId="2" pageNumber="279" type="etymology">
<paragraph id="C83BAA2D533BFFDBC9AEFCCBFB8FF95C" blockId="2.[151,1437,645,2005]" box="[199,1126,862,889]" pageId="2" pageNumber="279">
<emphasis id="FAF0763F533BFFDBC9AEFCCBFEA7F95C" bold="true" box="[199,334,862,888]" pageId="2" pageNumber="279">Etymology.</emphasis>
“
<taxonomicName id="0F84D1AE533BFFDBC808FCCBFDFFF95C" authorityName="Nguyen &amp; Tran &amp; Lam &amp; Nguyen" authorityYear="2020" box="[353,534,862,888]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="279" phylum="Annelida" rank="species" species="longiprostaticus">
<emphasis id="FAF0763F533BFFDBC808FCCBFDFFF95C" box="[353,534,862,888]" italics="true" pageId="2" pageNumber="279">longiprostaticus</emphasis>
</taxonomicName>
”, an adjective, to emphasize the long prostate gland.
</paragraph>
</subSubSection>
<subSubSection id="809EF9A6533BFFDBC9AEFC14FB26FE08" pageId="2" pageNumber="279" type="diagnosis">
<paragraph id="C83BAA2D533BFFDBC9AEFC14FB26FE08" blockId="2.[151,1437,645,2005]" pageId="2" pageNumber="279">
<emphasis id="FAF0763F533BFFDBC9AEFC14FEABF9B8" bold="true" box="[199,322,897,924]" pageId="2" pageNumber="279">Diagnosis.</emphasis>
Small size, length
<quantity id="0F7C07C8533BFFDBCB7AFC14FD64F9B9" box="[531,653,897,925]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" metricValueMax="7.1" metricValueMin="5.1" pageId="2" pageNumber="279" unit="mm" value="61.0" valueMax="71.0" valueMin="51.0">51–71 mm</quantity>
, diameter
<quantity id="0F7C07C8533BFFDBCB96FC17FC6EF9B9" box="[767,903,898,925]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="2" pageNumber="279" unit="mm" value="2.55" valueMax="2.7" valueMin="2.4">2.4–2.7 mm</quantity>
, segments 81–112. Prostomium epilobous. First dorsal pore in 11/12. Setal numbers:
<quantity id="0F7C07C8533BFFDBCB5EFC30FD73F9E4" box="[567,666,933,960]" metricMagnitude="0" metricUnit="m" metricValue="1.2573" metricValueMax="1.4478" metricValueMin="1.0668" pageId="2" pageNumber="279" unit="in" value="49.5" valueMax="57.0" valueMin="42.0">42–57 in</quantity>
viii,
<quantity id="0F7C07C8533BFFDBCBBFFC33FCD0F9E4" box="[726,825,934,960]" metricMagnitude="0" metricUnit="m" metricValue="1.4605000000000001" metricValueMax="1.6764000000000003" metricValueMin="1.2446000000000002" pageId="2" pageNumber="279" unit="in" value="57.5" valueMax="66.0" valueMin="49.0">49–66 in</quantity>
xxx, 9–12 between male porophores in xviii. A single pair of spermathecal pores, lateroventral, in intersegment 5/6. One pair of genital markings present ventrally in xviii. Septa 8/9/10 absent. Intestinal caeca simple. Holandric. Testis sacs separated. Prostate glands well developed, moderately lobuled, paired in xviii–xxviii; ducts long, waved and directly attached to body wall.
</paragraph>
</subSubSection>
<subSubSection id="809EF9A6533BFFDBC9AEFBA0FB31FD60" pageId="2" pageNumber="279" type="description">
<paragraph id="C83BAA2D533BFFDBC9AEFBA0FBBDFEC4" blockId="2.[151,1437,645,2005]" pageId="2" pageNumber="279">
<emphasis id="FAF0763F533BFFDBC9AEFBA0FEB0FE74" bold="true" box="[199,345,1077,1104]" pageId="2" pageNumber="279">Description.</emphasis>
Body cylindrical, small size; length
<quantity id="0F7C07C8533BFFDBCA6AFBA0FC68FE75" box="[771,897,1077,1105]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" metricValueMax="7.1" metricValueMin="5.1" pageId="2" pageNumber="279" unit="mm" value="61.0" valueMax="71.0" valueMin="51.0">51–71 mm</quantity>
, diameter
<quantity id="0F7C07C8533BFFDBCA93FBA3FB6FFE75" box="[1018,1158,1078,1105]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="2" pageNumber="279" unit="mm" value="2.55" valueMax="2.7" valueMin="2.4">2.4–2.7 mm</quantity>
, weight 0.2–0.3 gr, segments 81–112. Body light grey, ventrum paler than dorsum, clitellum darkish brown. Prostomium 1/3 epilobous. First dorsal pore in 11/12. Setae perichaetine, pre-clitellar setae stouter and sparser than post-clitellar ones; setal numbers:
<quantity id="0F7C07C8533BFFDBC86FFB34FE81FE98" box="[262,360,1185,1212]" metricMagnitude="0" metricUnit="m" metricValue="1.2573" metricValueMax="1.4478" metricValueMin="1.0668" pageId="2" pageNumber="279" unit="in" value="49.5" valueMax="57.0" valueMin="42.0">42–57 in</quantity>
viii,
<quantity id="0F7C07C8533BFFDBC8C9FB37FDEBFE98" box="[416,514,1186,1212]" metricMagnitude="0" metricUnit="m" metricValue="1.4605000000000001" metricValueMax="1.6764000000000003" metricValueMin="1.2446000000000002" pageId="2" pageNumber="279" unit="in" value="57.5" valueMax="66.0" valueMin="49.0">49–66 in</quantity>
xxx, 6–8 between male porophores in xviii; setal distance: aa&gt;ab, zz&gt;zy. Clitellum within xiv–xvi, without dorsal pores and setae. Female pore single, in midventral xiv.
</paragraph>
<paragraph id="C83BAA2D533BFFDBC9AEFB7FFEC2FF55" blockId="2.[151,1437,645,2005]" pageId="2" pageNumber="279">A single pair of spermathecal pores located lateroventrally in intersegments 5/6. Genital markings single, located ventrally in front of setal rings of x or xi. Male porophores located on setal ring of xviii; ventral distance between two male pores about 0.35X body circumference. A single conial genital marking located ventrally, next to male pore.</paragraph>
<paragraph id="C83BAA2D533BFFDBC9AEFAEFFC55FFF8" blockId="2.[151,1437,645,2005]" pageId="2" pageNumber="279">Septa 5/6/7/8 thick, 8/9/10 absent, 10/11/12/13 thin. Oesophageal gizzard within viii–x. Intestinal origin at xv; caeca simple, originating at xxvii and extending anteriorly to xxii. Last hearts in xiii. Pharyngeal micronephridia developed 5/6/7. Typhlosole simple, lamelliform. Lymph glands absent.</paragraph>
<paragraph id="C83BAA2D533BFFDBC9AEFA73FE00FCB4" blockId="2.[151,1437,645,2005]" pageId="2" pageNumber="279">A pair of spermathecae in vi. Ampulla oval-shaped; ducts about 1/2 ampulla, robust, but getting thinner basally. Diverticula thin, shorter than ampulla, strongly twisted, directly attached to the middle of ampullar duct; seminal chamber small, oval or bean-shaped. Spermathecal ducts without nephridia. A large accessory gland located next to each ampullar duct, sometimes covering the duct. Additional accessory gland with duct in x or xi, in association with external genital marking.</paragraph>
<paragraph id="C83BAA2D533BFFDBC9AEF90FFAB9FCD8" blockId="2.[151,1437,645,2005]" pageId="2" pageNumber="279">Holandric. Testis sacs developed, separated in x–xi. Seminal vesicles well developed in xi–xii. Ovaries well developed in 12/13. Ovisacs invisible. Prostate glands deeply lobuled, paired in xviii–xxviii; ducts long, waved and directly attached to body wall. A pair of large accessory glands in association with genital markings in xviii.</paragraph>
<paragraph id="C83BAA2D533BFFDBC9AEF890FB31FD60" blockId="2.[151,1437,645,2005]" pageId="2" pageNumber="279">
<emphasis id="FAF0763F533BFFDBC9AEF890FEC3FD04" bold="true" box="[199,298,1797,1824]" pageId="2" pageNumber="279">Habitat.</emphasis>
The species was found in natural forests of Cat Tien National Park, where the forests are among the best-conserved in
<collectingCountry id="B093EABD533BFFDBC80BF8BFFE2CFD60" box="[354,453,1834,1860]" name="Vietnam" pageId="2" pageNumber="279">Vietnam</collectingCountry>
. It was collected in the
<quantity id="0F7C07C8533BFFDBCBA5F8BFFCC4FD61" box="[716,813,1834,1861]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="279" unit="cm" value="5.0" valueMax="10.0" valueMin="0.0">0–10 cm</quantity>
mineral soil below a thick litter layer.
</paragraph>
</subSubSection>
<subSubSection id="809EF9A6533BFFDDC9AEF8DBFC60FE13" lastPageId="4" lastPageNumber="281" pageId="2" pageNumber="279" type="discussion">
<paragraph id="C83BAA2D533BFFDBC9AEF8DBFD95FDF1" blockId="2.[151,1437,645,2005]" pageId="2" pageNumber="279">
<emphasis id="FAF0763F533BFFDBC9AEF8DBFED2FD4C" bold="true" box="[199,315,1870,1896]" pageId="2" pageNumber="279">Remarks.</emphasis>
The new species can be classified into the
<taxonomicName id="0F84D1AE533BFFDBCA58F8DBFBEEFD4C" baseAuthorityName="Horst" baseAuthorityYear="1893" box="[817,1031,1870,1896]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="279" phylum="Annelida" rank="species" species="minimus">
<emphasis id="FAF0763F533BFFDBCA58F8DBFBEEFD4C" box="[817,1031,1870,1896]" italics="true" pageId="2" pageNumber="279">Amynthas minimus</emphasis>
</taxonomicName>
group characterized by one pair of spermathecal pores in 5/6 (
<bibRefCitation id="AC15D7DC533BFFDBC8A8F8E7FD43FDA8" author="Sims, R. W. &amp; Easton, E. G." box="[449,682,1906,1933]" pageId="2" pageNumber="279" pagination="169 - 268" refId="ref11103" refString="Sims, R. W. &amp; Easton, E. G. (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of Linnean Society, 4, 169 - 268. https: // doi. org / 10.1111 / j. 1095 - 8312.1972. tb 00694. x" type="journal article" year="1972">Sims &amp; Easton 1972</bibRefCitation>
). This group is known to comprise 25 species (
<bibRefCitation id="AC15D7DC533BFFDBCDC6F8E7FA7EFDA8" author="Sims, R. W. &amp; Easton, E. G." box="[1199,1431,1906,1933]" pageId="2" pageNumber="279" pagination="169 - 268" refId="ref11103" refString="Sims, R. W. &amp; Easton, E. G. (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of Linnean Society, 4, 169 - 268. https: // doi. org / 10.1111 / j. 1095 - 8312.1972. tb 00694. x" type="journal article" year="1972">Sims &amp; Easton 1972</bibRefCitation>
;
<bibRefCitation id="AC15D7DC533BFFDBC9FEF803FEE4FD94" author="Thai, T. B." box="[151,269,1942,1968]" pageId="2" pageNumber="279" pagination="817 - 830" refId="ref11223" refString="Thai, T. B. (1982) The fauna of earthworms of the Cucfuong forest reserve (Vietnam) and descriptions of new species of the genus Pheretima. Zoologicheskii Zhurnal, 61 (6), 817 - 830." type="journal article" year="1982">Thai 1982</bibRefCitation>
;
<bibRefCitation id="AC15D7DC533BFFDBC870F803FE75FD94" author="Thai, T. B." box="[281,412,1942,1968]" pageId="2" pageNumber="279" pagination="284 - 288" refId="ref11266" refString="Thai, T. B. (1984 a) New data on taxonomy of the genus Pheretima (Oligochaeta, Megascolecidae) of the fauna of Vietnam. Zoologicheskii Zhurnal, 63 (2), 284 - 288." type="journal article" year="1984">Thai 1984a</bibRefCitation>
;
<bibRefCitation id="AC15D7DC533BFFDBC8C1F802FDA4FD94" author="Do, V. N. &amp; Tran, M. K. &amp; Le, V. T." box="[424,589,1941,1969]" pageId="2" pageNumber="279" pagination="88 - 94" refId="ref9654" refString="Do, V. N., Tran, M. K. &amp; Le, V. T. (1995) New species of earthworm of genus Pheretima Kinberg, 1867 (Megascolecidae, Oligochaeta) from mountain of Nghe An Province. TAP CHI SINH HOC, 17 (3), 88 - 94." type="journal article" year="1995">
Do
<emphasis id="FAF0763F533BFFDBC8BBF802FDE2FD94" box="[466,523,1942,1968]" italics="true" pageId="2" pageNumber="279">et al.</emphasis>
1995
</bibRefCitation>
;
<bibRefCitation id="AC15D7DC533BFFDBCB30F803FD35FD94" author="Pham, T. H. H." box="[601,732,1941,1968]" pageId="2" pageNumber="279" pagination="22 - 24" refId="ref10895" refString="Pham, T. H. H. (1995) Three new species of earthworm of genus Pheretima Kinberg, 1867 (Megascolecidae: Oligochaeta) from Quang Nam - Da Nang Province. TAP CHI SINH HOC, 17 (1), 22 - 24." type="journal article" year="1995">Pham 1995</bibRefCitation>
;
<bibRefCitation id="AC15D7DC533BFFDBCB81F802FC10FD94" author="Bantaowong, U. &amp; James, S. W. &amp; Panha, S." box="[744,1017,1941,1969]" pageId="2" pageNumber="279" pagination="167 - 178" refId="ref9302" refString="Bantaowong, U., James, S. W. &amp; Panha, S. (2015) Three new earthworm species of the genus Amynthas Kinberg, 1867 from Thailand (Clitellata: Megascolecidae). Tropical Natural History, 15 (2), 167 - 178." type="journal article" year="2015">
Bantaowong
<emphasis id="FAF0763F533BFFDBCA14F802FC5FFD94" box="[893,950,1942,1968]" italics="true" pageId="2" pageNumber="279">et al.</emphasis>
2015
</bibRefCitation>
), ten of which are known in northern and central
<collectingCountry id="B093EABD533BFFDBC870F82FFE91FDF0" box="[281,376,1978,2004]" name="Vietnam" pageId="2" pageNumber="279">Vietnam</collectingCountry>
(
<bibRefCitation id="AC15D7DC533BFFDBC8EEF82FFD98FDF1" author="Nguyen, T. T. &amp; Nguyen, A. D. &amp; Tran, T. T. B. &amp; Blakemore, R. J." box="[391,625,1978,2005]" pageId="2" pageNumber="279" pagination="1 - 92" refId="ref10413" refString="Nguyen, T. T., Nguyen, A. D., Tran, T. T. B. &amp; Blakemore, R. J. (2016 a) A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae). Zootaxa, 4140 (1), 1 - 92. https: // doi. org / 10.11646 / zootaxa. 4140.1.1" type="journal article" year="2016">
Nguyen
<emphasis id="FAF0763F533BFFDBC881F82EFDC9FDF0" box="[488,544,1978,2004]" italics="true" pageId="2" pageNumber="279">et al.</emphasis>
2016a
</bibRefCitation>
).
</paragraph>
<caption id="9CFBFAA5533AFFDAC9FEF96CFB49FD59" ID-DOI="http://doi.org/10.5281/zenodo.3889531" ID-Zenodo-Dep="3889531" httpUri="https://zenodo.org/record/3889531/files/figure.png" pageId="3" pageNumber="280" startId="3.[151,250,1785,1810]" targetBox="[164,1423,200,1747]" targetPageId="3">
<paragraph id="C83BAA2D533AFFDAC9FEF96CFB49FD59" blockId="3.[151,1437,1785,1918]" pageId="3" pageNumber="280">
<emphasis id="FAF0763F533AFFDAC9FEF96CFC0AFD36" bold="true" box="[151,995,1785,1810]" pageId="3" pageNumber="280">
FIGURE 2.
<taxonomicName id="0F84D1AE533AFFDAC87EF96CFDC7FD36" authority="Nguyen &amp; Tran &amp; Lam &amp; Nguyen, 2020" authorityName="Nguyen &amp; Tran &amp; Lam &amp; Nguyen" authorityYear="2020" box="[279,558,1785,1810]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="280" phylum="Annelida" rank="species" species="longiprostaticus" status="sp. nov.">
<emphasis id="FAF0763F533AFFDAC87EF96CFDC7FD36" bold="true" box="[279,558,1785,1810]" italics="true" pageId="3" pageNumber="280">Amynthas longiprostaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E1C3CB44533AFFDACB5BF96FFD6BFD36" box="[562,642,1786,1810]" pageId="3" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
Holotype (CTU-EW.088.h01).A.
</emphasis>
Male region, ventral view.
<emphasis id="FAF0763F533AFFDACD9DF96FFAE2FD36" bold="true" box="[1268,1291,1786,1810]" pageId="3" pageNumber="280">B.</emphasis>
Left intestinal caecum.
<emphasis id="FAF0763F533AFFDAC999F888FEE0FD12" bold="true" box="[240,265,1821,1846]" pageId="3" pageNumber="280">C.</emphasis>
Left prostate gland.
<emphasis id="FAF0763F533AFFDAC8B1F88BFE18FD12" bold="true" box="[472,497,1822,1846]" pageId="3" pageNumber="280">D.</emphasis>
Spermathecal region, ventral view.
<emphasis id="FAF0763F533AFFDACA33F88BFC98FD12" bold="true" box="[858,881,1822,1846]" pageId="3" pageNumber="280">E.</emphasis>
Testis sacs and seminal vesicles.
<emphasis id="FAF0763F533AFFDACDA9F88BFB3AFD12" bold="true" box="[1216,1235,1822,1846]" pageId="3" pageNumber="280">F.</emphasis>
Right spermatheca.
<emphasis id="FAF0763F533AFFDAC9FEF8D4FF58FD7E" bold="true" box="[151,177,1857,1882]" pageId="3" pageNumber="280">G.</emphasis>
Male region (comp.
<emphasis id="FAF0763F533AFFDAC8E0F8D4FE72FD7E" bold="true" box="[393,411,1857,1882]" pageId="3" pageNumber="280">A</emphasis>
).
<emphasis id="FAF0763F533AFFDAC8D8F8D7FE22FD7E" bold="true" box="[433,459,1858,1882]" pageId="3" pageNumber="280">H.</emphasis>
Spermatheca (comp.
<emphasis id="FAF0763F533AFFDACBC3F8D7FD50FD7E" bold="true" box="[682,697,1858,1882]" pageId="3" pageNumber="280">F</emphasis>
).
<emphasis id="FAF0763F533AFFDACBA6F8D7FD36FD7E" bold="true" box="[719,735,1858,1882]" pageId="3" pageNumber="280">I.</emphasis>
Prostate gland (comp.
<emphasis id="FAF0763F533AFFDACAA5F8D4FC37FD7E" bold="true" box="[972,990,1857,1882]" pageId="3" pageNumber="280">C</emphasis>
).
<emphasis id="FAF0763F533AFFDACA9DF8D7FBEEFD7E" bold="true" box="[1012,1031,1858,1882]" pageId="3" pageNumber="280">J.</emphasis>
Genital marking in x.
<emphasis id="FAF0763F533AFFDACD86F8D7FAEAFD7E" bold="true" box="[1263,1283,1858,1882]" pageId="3" pageNumber="280">K</emphasis>
,
<emphasis id="FAF0763F533AFFDACC79F8D7FAC1FD7E" bold="true" box="[1296,1320,1858,1882]" pageId="3" pageNumber="280">L.</emphasis>
Transverse section of male porophore, different section planes (Non-type:
<emphasis id="FAF0763F533AFFDACA7CF8F0FC2EFD5A" bold="true" box="[789,967,1893,1918]" pageId="3" pageNumber="280">CTU-EW.088.03</emphasis>
). Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="C83BAA2D533DFFDDC9AEFF0CFE96F8A3" blockId="4.[151,1437,153,1079]" pageId="4" pageNumber="281">
Within the
<taxonomicName id="0F84D1AE533DFFDDC82CFF0CFE2AFA97" baseAuthorityName="Horst" baseAuthorityYear="1893" box="[325,451,153,179]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="minimus">
<emphasis id="FAF0763F533DFFDDC82CFF0CFE2AFA97" box="[325,451,153,179]" italics="true" pageId="4" pageNumber="281">A. minimus</emphasis>
</taxonomicName>
group, it is the only species with prostate glands that extend over up to 11 segments; in all other species the glands are shorter. Apart from that character, the new species is somewhat similar to
<taxonomicName id="0F84D1AE533DFFDDCC47FF28FED9FADF" authority="(Gates, 1930)" baseAuthorityName="Gates" baseAuthorityYear="1930" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="papilio">
<emphasis id="FAF0763F533DFFDDCC47FF28FA75FAF3" box="[1326,1436,189,215]" italics="true" pageId="4" pageNumber="281">A. papilio</emphasis>
(
<bibRefCitation id="AC15D7DC533DFFDDC9F6FF74FECEFADF" author="Gates, G. E." box="[159,295,225,251]" pageId="4" pageNumber="281" pagination="257 - 356" refId="ref9793" refString="Gates, G. E. (1930) The earthworms of Burma. I. Records of the Indian Museum, 32, 257 - 356." type="journal article" year="1930">Gates, 1930</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="0F84D1AE533DFFDDC80CFF74FCA8FADF" authority="Bantaowong &amp; Panha, 2015" authorityName="Bantaowong &amp; Panha" authorityYear="2015" box="[357,833,224,251]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="khaohayod">
<emphasis id="FAF0763F533DFFDDC80CFF74FE16FADF" box="[357,511,225,251]" italics="true" pageId="4" pageNumber="281">A. khaohayod</emphasis>
<bibRefCitation id="AC15D7DC533DFFDDCB6FFF74FCA8FADF" author="Bantaowong, U. &amp; James, S. W. &amp; Panha, S." box="[518,833,224,251]" pageId="4" pageNumber="281" pagination="167 - 178" refId="ref9302" refString="Bantaowong, U., James, S. W. &amp; Panha, S. (2015) Three new earthworm species of the genus Amynthas Kinberg, 1867 from Thailand (Clitellata: Megascolecidae). Tropical Natural History, 15 (2), 167 - 178." type="journal article" year="2015">Bantaowong &amp; Panha, 2015</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation id="AC15D7DC533DFFDDCA07FF74FB94FADF" author="Bantaowong, U. &amp; James, S. W. &amp; Panha, S." box="[878,1149,224,251]" pageId="4" pageNumber="281" pagination="167 - 178" refId="ref9302" refString="Bantaowong, U., James, S. W. &amp; Panha, S. (2015) Three new earthworm species of the genus Amynthas Kinberg, 1867 from Thailand (Clitellata: Megascolecidae). Tropical Natural History, 15 (2), 167 - 178." type="journal article" year="2015">
Bantaowong et
<emphasis id="FAF0763F533DFFDDCD74FF74FBD3FADF" box="[1053,1082,225,251]" italics="true" pageId="4" pageNumber="281">al.</emphasis>
2015
</bibRefCitation>
), both also in the
<taxonomicName id="0F84D1AE533DFFDDCC2BFF74FF2DFB3B" baseAuthorityName="Horst" baseAuthorityYear="1893" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="minimus">
<emphasis id="FAF0763F533DFFDDCC2BFF74FF2DFB3B" italics="true" pageId="4" pageNumber="281">A. minimus</emphasis>
</taxonomicName>
group, by presence of a pair of genital markings in xviii and simple intestinal caeca.
<taxonomicName id="0F84D1AE533DFFDDCDF5FE90FA9BFB3B" authority="Nguyen &amp; Tran &amp; Lam &amp; Nguyen, 2020" authorityName="Nguyen &amp; Tran &amp; Lam &amp; Nguyen" authorityYear="2020" box="[1180,1394,261,287]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="longiprostaticus" status="sp. nov.">
<emphasis id="FAF0763F533DFFDDCDF5FE90FA9BFB3B" box="[1180,1394,261,287]" italics="true" pageId="4" pageNumber="281">A. longiprostaticus</emphasis>
</taxonomicName>
<emphasis id="FAF0763F533DFFDDCC12FE90FF21FB67" bold="true" pageId="4" pageNumber="281">
<taxonomicNameLabel id="E1C3CB44533DFFDDCC12FE90FF21FB67" pageId="4" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="0F84D1AE533DFFDDC833FEBCFE1AFB67" authorityName="Bantaowong &amp; Panha" authorityYear="2015" box="[346,499,297,323]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="khaohayod">
<emphasis id="FAF0763F533DFFDDC833FEBCFE1AFB67" box="[346,499,297,323]" italics="true" pageId="4" pageNumber="281">A. khaohayod</emphasis>
</taxonomicName>
in ventral distance between the two male porophores (0.35X vs. 0.18X), presence of genital markings in the spermathecal and male regions (vs. absence of genital markings), first dorsal pore in 11/12 (vs. in 12/13), and body size (length
<quantity id="0F7C07C8533DFFDDCB2DFEE5FD51FBAF" box="[580,696,368,395]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" metricValueMax="7.1" metricValueMin="5.1" pageId="4" pageNumber="281" unit="mm" value="61.0" valueMax="71.0" valueMin="51.0">51–71mm</quantity>
, diameter
<quantity id="0F7C07C8533DFFDDCA5BFEE4FC5CFBAF" box="[818,949,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="4" pageNumber="281" unit="mm" value="2.55" valueMax="2.7" valueMin="2.4">2.4–2.7mm</quantity>
, vs. length
<quantity id="0F7C07C8533DFFDDCD52FEE4FB7DFBAF" box="[1083,1172,369,395]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="281" unit="mm" value="230.0">230mm</quantity>
, diameter
<quantity id="0F7C07C8533DFFDDCC65FEE5FA79FBAF" box="[1292,1424,368,395]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="7.5" metricValueMin="6.8" pageId="4" pageNumber="281" unit="mm" value="7.15" valueMax="7.5" valueMin="6.8">6.8–7.5mm</quantity>
), the shape of the male genital markings (conical-shaped vs. glandular-shaped) and the shape of spermathecae (ovalshaped ampulla small, oval or bean-shaped seminal chamber vs. spatulate-shaped ampulla, roughly elliptic seminal chamber). Similarly, the new species is also differentiated from
<taxonomicName id="0F84D1AE533DFFDDCA1AFE48FC0AFBD3" baseAuthorityName="Gates" baseAuthorityYear="1930" box="[883,995,477,503]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="papilio">
<emphasis id="FAF0763F533DFFDDCA1AFE48FC0AFBD3" box="[883,995,477,503]" italics="true" pageId="4" pageNumber="281">A. papilio</emphasis>
</taxonomicName>
in the presence of genital markings in the spermathecal region (vs. absence of genital markings), first dorsal pore in 11/12 (vs. in 5/6), and size of genital markings in the male region (same size as male porophore vs. larger than male porophore, extending beyond xviii). In both species compared, the male genital markings are much larger than the ones in the new species, and the prostates are shorter.
</paragraph>
<paragraph id="C83BAA2D533DFFDDC9AEFD04FE7CF9A0" blockId="4.[151,1437,153,1079]" pageId="4" pageNumber="281">
The character state of exceptionally long prostate glands is, within
<taxonomicName id="0F84D1AE533DFFDDCABAFD04FBA8F88F" authorityName="Kinberg" authorityYear="1867" box="[979,1089,657,683]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="genus">
<emphasis id="FAF0763F533DFFDDCABAFD04FBA8F88F" box="[979,1089,657,683]" italics="true" pageId="4" pageNumber="281">Amynthas</emphasis>
</taxonomicName>
, shared by
<taxonomicName id="0F84D1AE533DFFDDCDA2FD04FEA0F8EB" authority="(Michaelsen, 1907)" baseAuthorityName="Michaelsen" baseAuthorityYear="1907" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="osmastoni">
<emphasis id="FAF0763F533DFFDDCDA2FD04FAB6F88F" box="[1227,1375,657,683]" italics="true" pageId="4" pageNumber="281">A. osmastoni</emphasis>
(
<bibRefCitation id="AC15D7DC533DFFDDCC18FD04FEA9F8EB" author="Michaelsen, W." pageId="4" pageNumber="281" pagination="143 - 93" refId="ref10051" refString="Michaelsen, W. (1907) Neue Oligochaten von Vorder-Indien, Ceylon, Birma und den Andaman-Inseln. Mitteilungen aus dem Naturhistorischen Museum in Hamburg, 24, 143 - 93." type="journal article" year="1907">Michaelsen, 1907</bibRefCitation>
)
</taxonomicName>
(xv–xxiv) and
<taxonomicName id="0F84D1AE533DFFDDC893FD20FBBCF8EB" authority="Shen, Chang, Li, Chih &amp; Chen, 2013" authorityName="Shen, Chang, Li, Chih &amp; Chen" authorityYear="2013" box="[506,1109,693,719]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="taiwumontis">
<emphasis id="FAF0763F533DFFDDC893FD20FD4AF8EB" box="[506,675,693,719]" italics="true" pageId="4" pageNumber="281">A. taiwumontis</emphasis>
<bibRefCitation id="AC15D7DC533DFFDDCBC5FD20FBBCF8EB" author="Shen, H. &amp; Chang, C. &amp; Li, C. &amp; Chih, W. &amp; Chen, J." box="[684,1109,693,719]" pageId="4" pageNumber="281" pagination="471 - 482" refId="ref11035" refString="Shen, H., Chang, C., Li, C., Chih, W. &amp; Chen, J. (2013) Four new earthworm species of the genus Amynthas (Oligochaeta: Megascolecidae) from Kinmen, Taiwan. Zootaxa, 3599 (5), 471 - 482. https: // doi. org / 10.11646 / zootaxa. 3599.5.4" type="journal article" year="2013">Shen, Chang, Li, Chih &amp; Chen, 2013</bibRefCitation>
</taxonomicName>
(xvi–xxiv). However,
<taxonomicName id="0F84D1AE533DFFDDCC30FD20FF19F8D7" baseAuthorityName="Michaelsen" baseAuthorityYear="1907" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="osmastoni">
<emphasis id="FAF0763F533DFFDDCC30FD20FF19F8D7" italics="true" pageId="4" pageNumber="281">A. osmastoni</emphasis>
</taxonomicName>
is longer than
<quantity id="0F7C07C8533DFFDDC8F3FD4DFE11F8D7" box="[410,504,728,755]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="281" unit="mm" value="250.0">250 mm</quantity>
and has three pairs of spermathecal pores in 6/7/8/9, first dorsal pore in 12/13, no genital markings in the male region.
<taxonomicName id="0F84D1AE533DFFDDCB45FD68FD3AF933" authorityName="Shen, Chang, Li, Chih &amp; Chen" authorityYear="2013" box="[556,723,765,791]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="taiwumontis">
<emphasis id="FAF0763F533DFFDDCB45FD68FD3AF933" box="[556,723,765,791]" italics="true" pageId="4" pageNumber="281">A. taiwumontis</emphasis>
</taxonomicName>
is about
<quantity id="0F7C07C8533DFFDDCA5CFD69FC20F933" box="[821,969,764,791]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="4" pageNumber="281" unit="mm" value="140.0" valueMax="150.0" valueMin="130.0">130–150 mm</quantity>
long, has four pairs of spermathecal pores in 5/6/7/8/9, first dorsal pore in 11/12, and no markings in the male region. The new species differs from these two species in one pair of spermathecal pores in 5/6 and presence of genital markings in the male region, and has smaller size (
<quantity id="0F7C07C8533DFFDDC9BBFCFDFEA5F9A7" box="[210,332,872,899]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" metricValueMax="7.1" metricValueMin="5.5" pageId="4" pageNumber="281" unit="mm" value="63.0" valueMax="71.0" valueMin="55.0">55–71 mm</quantity>
long).
</paragraph>
<paragraph id="C83BAA2D533DFFDDC9AEFC18FC60FE13" blockId="4.[151,1437,153,1079]" pageId="4" pageNumber="281">
Regarding the morphology of the male region, the new species is fairly similar to
<taxonomicName id="0F84D1AE533DFFDDCDE5FC18FF32F9EF" authority="(Thai, 1984)" baseAuthorityName="Thai" baseAuthorityYear="1984" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="brevicapitatus">
<emphasis id="FAF0763F533DFFDDCDE5FC18FAA7F983" box="[1164,1358,909,935]" italics="true" pageId="4" pageNumber="281">A. brevicapitatus</emphasis>
(Thai, 1984)
</taxonomicName>
,
<taxonomicName id="0F84D1AE533DFFDDC98FFC24FDD5F9EF" authority="(Thai, 1984)" baseAuthorityName="Thai" baseAuthorityYear="1984" box="[230,572,945,971]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="deonganganus">
<emphasis id="FAF0763F533DFFDDC98FFC24FE4EF9EF" box="[230,423,945,971]" italics="true" pageId="4" pageNumber="281">A. deonganganus</emphasis>
(Thai, 1984)
</taxonomicName>
,
<taxonomicName id="0F84D1AE533DFFDDCB2EFC24FC9AF9E8" authority="(Kinberg, 1867)" baseAuthorityName="Kinberg" baseAuthorityYear="1867" box="[583,883,945,972]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="corticis">
<emphasis id="FAF0763F533DFFDDCB2EFC24FD50F9EF" box="[583,697,945,971]" italics="true" pageId="4" pageNumber="281">A. corticis</emphasis>
(
<bibRefCitation id="AC15D7DC533DFFDDCBA1FC24FC82F9E8" author="Kinberg, J. G. H." box="[712,875,945,972]" pageId="4" pageNumber="281" pagination="97 - 103" refId="ref9924" refString="Kinberg, J. G. H. (1867) Annulata Nova. Ofversigt af Kongliga Vetenskaps-akademiens forhandlingar, 23, 97 - 103." type="journal article" year="1867">Kinberg, 1867</bibRefCitation>
)
</taxonomicName>
and to
<taxonomicName id="0F84D1AE533DFFDDCAACFC24FAA9F9EF" authority="(Goto &amp; Hatai, 1899)" baseAuthorityName="Goto &amp; Hatai" baseAuthorityYear="1899" box="[965,1344,945,971]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="281" phylum="Annelida" rank="species" species="carnosus">
<emphasis id="FAF0763F533DFFDDCAACFC24FBA1F9EF" box="[965,1096,945,971]" italics="true" pageId="4" pageNumber="281">A. carnosus</emphasis>
(
<bibRefCitation id="AC15D7DC533DFFDDCD3FFC24FAD0F9EF" author="Goto, S. &amp; Hatai, S." box="[1110,1337,945,971]" pageId="4" pageNumber="281" pagination="13 - 24" refId="ref9855" refString="Goto, S. &amp; Hatai, S. (1899) New or imperpectly known species of earthworm, No. 2. Annotationes Zoologicae Japonenses, 3, 13 - 24." type="journal article" year="1899">Goto &amp; Hatai, 1899</bibRefCitation>
)
</taxonomicName>
, species with one pair of genital markings in immediate vicinity to the male pores. These three species differ from the new one in three or four pairs of spermathecae, connected testis sacs, and in a more anterior location of genital markings in the spermathecal region (when present), among other characters.
</paragraph>
</subSubSection>
</treatment>
</document>
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