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<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende &amp; Pinto-da-Rocha, Ricardo" docDate="2022" docId="40D7F7A9EF4F57D094D22FA682E04E4C" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Ayacucho spielbergi Benedetti &amp; Pinto-da-Rocha 2022, sp. nov." docType="treatment" docUuid="81B944C8-2A70-44BE-AA34-DA96925262B0" docUuidSource="ZooBank" docVersion="1" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1660711556436" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Benedetti, Alipio Rezende</mods:namePart>
<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil &amp; Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pinto-da-Rocha, Ricardo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>309</mods:start>
<mods:end>388</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/asp.80.e73829</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e73829</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-80-309</mods:identifier>
<mods:identifier type="ZooBank">D5C0468B99A14EF39237D9BC51A8BDA3</mods:identifier>
<mods:identifier type="Pensoft-UUID">11907D1B6F665D96B5FD2F9751431366</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:81B944C8-2A70-44BE-AA34-DA96925262B0" httpUri="http://treatment.plazi.org/id/40D7F7A9EF4F57D094D22FA682E04E4C" lastPageNumber="309" pageId="0" pageNumber="309">
<subSubSection pageId="0" pageNumber="309" type="nomenclature">
<paragraph pageId="0" pageNumber="309">
3.20.
<taxonomicName LSID="http://zoobank.org/81B944C8-2A70-44BE-AA34-DA96925262B0" authority="Benedetti &amp; Pinto-da-Rocha, 2022" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho spielbergi" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="spielbergi" status="sp. nov.">Ayacucho spielbergi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="309">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="description">
<paragraph pageId="0" pageNumber="309">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">Figs 3A</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">, 7O, P</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D Ayacucho inermis (Roewer, 1957) comb. nov.; E - F Ayacucho pasolinii sp. nov.; G - H Ayacucho spielbergi sp. nov.; I - J Ayacucho titschacki Roewer, 1949; K - L Ayacucho triarmatus nom. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure12" httpUri="https://binary.pensoft.net/fig/729886" pageId="0" pageNumber="309">, 12G, H</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Live specimens of Ayacucho. A: A. glauberrochai sp. nov., male. B: female. C: A. pomacocha sp. nov., male. D, female. E: A. silvae sp. nov., male. F: female. G: A. pasolinii sp. nov., male. H: A. spielbergi sp. nov., male." figureDoi="10.3897/asp.80.e73829.figure22" httpUri="https://binary.pensoft.net/fig/729896" pageId="0" pageNumber="309">, 22H</figureCitation>
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">, 28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="description">
<paragraph pageId="0" pageNumber="309">Description.</paragraph>
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">
MALE:
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) DSW: 4.0-4.9 (4.9); DSL: 4.6-5.0 (5.0); CL: 1.6-2.0 (2.0). FIVL: 7.6-8.6 (8.6). ChL: 2.1-3.9 (3.9).
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
: (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Live specimens of Ayacucho. A: A. glauberrochai sp. nov., male. B: female. C: A. pomacocha sp. nov., male. D, female. E: A. silvae sp. nov., male. F: female. G: A. pasolinii sp. nov., male. H: A. spielbergi sp. nov., male." figureDoi="10.3897/asp.80.e73829.figure22" httpUri="https://binary.pensoft.net/fig/729896" pageId="0" pageNumber="309">22H</figureCitation>
) Carapace, lateral margins of DS, spines of scutal area III and legs dark brown. Reddish chelicera.
<taxonomicName genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">Pedipalpus</taxonomicName>
yellowish with black spots. Posterior margin of DS and free tergites I-III light yellow. Scutal areas light yellow, with series of dark spots.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
: (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3A</figureCitation>
) Gamma-type DSS. Anterior margin of the DS with a median elevation, with few granules sparsely distributed. Ocularium with medial depression; with a pair of spines and few granules near the eyes. Areas I-IV with few granules concentrated in the lateral regions; area III with a pair of spines directed backwards, located at the highest point in elevations of the integument, totally granular; area IV with a pair of small lateral tubercles. Posterior margins of DS smooth, with a pair of medium tubercles. Free tergites I-II with a pair of median tubercles; and a lateral pair of tiny tubercles. Free tergite III with two elongated median tubercles, larger than those of the free tergites I-II.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
: (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3A</figureCitation>
) Swollen in large males (swollen in holotype); slightly larger than that of the females and small males. Segment I granulate; II with few granules; III with two teeth.
<taxonomicName genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
</taxonomicName>
: Trochanter with two ventroapical setiferous tubercles, one large and one smaller. Femur with a row of six ventral setiferous tubercles in the basal ⅔. Patella with a small retroapical tubercle. Tibia: prolateral iiIii, retrolateral IIi. Tarsus: prolateral iiIiIi, retrolateral iIiIi.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
: Coxa I with a row of 4-5 tubercles. Coxae II-IV with granules throughout their length. Genital area, free sternites and anal operculum with few granules.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
: (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3A</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7O, P</figureCitation>
) Coxae I-II each one with a retrolateral and prolateral apophyses. Coxa III with a prolateral apophysis. Coxa IV with setiferous granules distributed over the entire length. Trochanters I-III unarmed and with few granules. Trochanter IV with few granules and an apical retrolateral blunt tubercle. Femora I-III unarmed and with few granules. Femur IV with sparse granules; a retrolateral row of 15-17 acuminate tubercles over the entire length of the segment, except the base; a prolateral row of 10-12 acuminate tubercles along the distal ⅔; with four ventrodistal tubercles. Patellae I-III unarmed, with few granules. Patella IV with seven dorsal acuminate tubercles, the two largest apical ones; several granules on the ventral surface. Tibiae I-IV unarmed, with few granules. Tarsal formula: (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) 7-8 (8), 12-13 (1 3), 10-15 (10), 11-13 (13).
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
: (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D Ayacucho inermis (Roewer, 1957) comb. nov.; E - F Ayacucho pasolinii sp. nov.; G - H Ayacucho spielbergi sp. nov.; I - J Ayacucho titschacki Roewer, 1949; K - L Ayacucho triarmatus nom. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure12" httpUri="https://binary.pensoft.net/fig/729886" pageId="0" pageNumber="309">12G, H</figureCitation>
) VP subrectangular, with distal margin straight and lateral projections; straight in lateral view. MS C1-C3 subapical long and straight; MS A1 median long and straight; MS B1 sub basal long and straight. Lateral sacs long, apically acuminate, with long T3-like microsetae. Stylus with a non-swollen apex and without apical projections. Dorsal process conical, with acuminated apex. Promontory straight. -
<emphasis bold="true" pageId="0" pageNumber="309">
FEMALE:
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=2) DSW: 4.2-4.5; DSL: 4.5-4.9; CL: 1.5-1.7. FIVL: 7.2-8.1. ChL: 1.7-1.8. Chelicerae not swollen. Femur IV with retrolateral row of 13-14 tubercles, smaller than those of the retrolateral row of the males; prolateral row and ventrodistal tubercles absent or in the form of granules. Patella IV with dorsodistal acuminate tubercles, but smaller than in males. Tarsal formula: (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=2) 7, 11-12, 9-10, 10-12.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="diagnosis">
<paragraph pageId="0" pageNumber="309">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="309">
It differs from other species of the genus by the combination of the following characteristics: Gamma-type DSS (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3A</figureCitation>
); area III of DS with a pair of large spines (also present in
<taxonomicName genus="A." lsidName="A. pasolinii" pageId="0" pageNumber="309" rank="species" species="pasolinii">
<emphasis italics="true" pageId="0" pageNumber="309">A. pasolinii</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
); male femur IV armed (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7O, P</figureCitation>
); longer legs (ratio between FIVL and DSL greater than 1; 1.65 in
<taxonomicName genus="A." lsidName="A. spielbergi" pageId="0" pageNumber="309" rank="species" species="spielbergi">
<emphasis italics="true" pageId="0" pageNumber="309">A. spielbergi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
); and scutal areas light yellow, with series of dark spots (Fig.
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Live specimens of Ayacucho. A: A. glauberrochai sp. nov., male. B: female. C: A. pomacocha sp. nov., male. D, female. E: A. silvae sp. nov., male. F: female. G: A. pasolinii sp. nov., male. H: A. spielbergi sp. nov., male." figureDoi="10.3897/asp.80.e73829.figure22" httpUri="https://binary.pensoft.net/fig/729896" pageId="0" pageNumber="309">22H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="etymology">
<paragraph pageId="0" pageNumber="309">Derivatio nominis.</paragraph>
<paragraph pageId="0" pageNumber="309">The specific epithet of masculine gender, in the genitive form, dedicated to the American filmmaker, producer and screenwriter Steven Allen Spielberg.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="distribution">
<paragraph pageId="0" pageNumber="309">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="309">
(Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">28</figureCitation>
) PERU.
<taxonomicName authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
. Parque Nacional Cutervo.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="materials_examined">
<paragraph pageId="0" pageNumber="309">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">
<emphasis italics="true" pageId="0" pageNumber="309">Type material</emphasis>
:
<materialsCitation collectingDate="2010-05-22" collectorName="R. Pinto-da-Rocha, D. Silva" country="PERU" latitude="-6.2027774" location="Parque Nacional Cutervo" longLatPrecision="21" longitude="-78.73944" municipality="Puente Suro" specimenCount="♂" typeStatus="Holotype">
<typeStatus>Holotype</typeStatus>
<specimenCount>♂</specimenCount>
, '
<collectingCountry name="Peru">PERU</collectingCountry>
,
<taxonomicName authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
,
<location LSID="urn:lsid:plazi:treatment:40D7F7A9EF4F57D094D22FA682E04E4C:CD1E62CCFD9A42E26A48F3FF0E65DA81" country="PERU" latitude="-6.2027774" longLatPrecision="21" longitude="-78.73944" municipality="Puente Suro" name="Parque Nacional Cutervo">Parque Nacional Cutervo</location>
,
<collectingMunicipality>Puente Suro</collectingMunicipality>
,
<geoCoordinate degrees="6" direction="south" minutes="12" orientation="latitude" precision="15" seconds="10" value="-6.2027774">6°12′10″S</geoCoordinate>
<geoCoordinate degrees="78" direction="west" minutes="44" orientation="longitude" precision="15" seconds="22" value="-78.73944">78°44′22″W</geoCoordinate>
,
<collectingDate value="2010-05-22">22/V/2010</collectingDate>
,
<collectorName>R. Pinto-da-Rocha</collectorName>
&amp;
<collectorName>D. Silva</collectorName>
leg. (MUSM)
</materialsCitation>
</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Paratypes</typeStatus>
</emphasis>
<specimenCount type="male">2 ♂</specimenCount>
,
<specimenCount type="female">1 ♀</specimenCount>
,
<normalizedToken originalValue="‘ditto’">'ditto'</normalizedToken>
(MZSP 36995);
<materialsCitation collectingDate="2010-05-22" collectorName="R. Pinto-da-Rocha, D. Silva" country="PERU" location="Parque Nacional Cutervo" municipality="Cueva San Andreas" specimenCount="1" specimenCount-female="1" typeStatus="Paratype">
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Paratype</typeStatus>
</emphasis>
<specimenCount type="female">1 ♀</specimenCount>
, '
<collectingCountry name="Peru">PERU</collectingCountry>
,
<taxonomicName authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
,
<location LSID="urn:lsid:plazi:treatment:40D7F7A9EF4F57D094D22FA682E04E4C:3D8AD7553A13C2823FC7CA6398482815" country="PERU" municipality="Cueva San Andreas" name="Parque Nacional Cutervo">Parque Nacional Cutervo</location>
, near to
<collectingMunicipality>Cueva San Andreas</collectingMunicipality>
,
<collectingDate value="2010-05-22">22/V/2010</collectingDate>
,
<collectorName>R. Pinto-da-Rocha</collectorName>
&amp;
<collectorName>D. Silva</collectorName>
leg. (MZSP 36996)
</materialsCitation>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Additional material</emphasis>
:
<materialsCitation collectingDate="1996-06-13" collectorName="S. Cordova" country="PERU" location="Parque Nacional Cutervo" municipality="San Andreas de Cutervo" specimenCount="1" specimenCount-male="1">
<specimenCount type="male">1 ♂</specimenCount>
, '
<collectingCountry name="Peru">PERU</collectingCountry>
,
<taxonomicName authorityName="Roewer" authorityYear="1952" class="Arachnida" family="Metasarcidae" genus="Cajamarca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cajamarca" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Cajamarca</taxonomicName>
,
<location LSID="urn:lsid:plazi:treatment:40D7F7A9EF4F57D094D22FA682E04E4C:2116D8AFD75667397F2DBFCD12CD1CD6" country="PERU" municipality="San Andreas de Cutervo" name="Parque Nacional Cutervo">Parque Nacional Cutervo</location>
,
<collectingMunicipality>San Andreas de Cutervo</collectingMunicipality>
,
<collectingDate value="1996-06-13">13/VI/1996</collectingDate>
,
<collectorName>
S.
<normalizedToken originalValue="Córdova">Cordova</normalizedToken>
</collectorName>
leg. (MUSM 0501248)
</materialsCitation>
;
<specimenCount type="female">1 ♀</specimenCount>
,
<normalizedToken originalValue="‘ditto’">'ditto'</normalizedToken>
(MUSM 0501247).
</paragraph>
</subSubSection>
</treatment>
</document>
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