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<document ID-CLB-Dataset="276125" ID-DOI="http://dx.doi.org/10.3897/phytokeys.235.112133" ID-GBIF-Dataset="8fc4dde9-3bc0-45b7-a930-9e88dc8684dd" ID-Pensoft-Pub="1314-2003-235-1" ID-Pensoft-UUID="71E754B19C905AFBAAA33F59CA927F7F" ModsDocID="1314-2003-235-1" checkinTime="1699037580884" checkinUser="pensoft" docAuthor="Dai, Jin-Hong, Van Do, Truong &amp; Liu, Ying" docDate="2023" docId="48251918F33D5760BF8DA2895F2356A8" docLanguage="en" docName="PhytoKeys 235: 1-19" docOrigin="PhytoKeys 235" docPubDate="2023-11-03" docSource="http://dx.doi.org/10.3897/phytokeys.235.112133" docTitle="Perilimnastes multisepala J. H. Dai, T. V. Do &amp; Ying Liu 2023, sp. nov." docType="treatment" docVersion="2" id="71E754B19C905AFBAAA33F59CA927F7F" lastPageNumber="1" masterDocId="71E754B19C905AFBAAA33F59CA927F7F" masterDocTitle="Four new species of Perilimnastes (Sonerileae, Melastomataceae) from Vietnam" masterLastPageNumber="19" masterPageNumber="1" pageNumber="1" updateTime="1699056822319" updateUser="ExternalLinkService">
<mods:mods id="9D09E88017081E6CD0A0F29ED18034CE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5DE99FC8C2026E23C3A111A06292D325">
<mods:title id="C5D399927028688EEEE9EC5C0EF4907C">Four new species of Perilimnastes (Sonerileae, Melastomataceae) from Vietnam</mods:title>
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<mods:name id="5F06944964467FA7E41B2819C438BBC9" type="personal">
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<mods:namePart id="7B64945D580B3F76584B83B1548E7818">Dai, Jin-Hong</mods:namePart>
<mods:nameIdentifier id="AE1EAE125233F493C34C853DDA9AAAAC" type="ORCID">https://orcid.org/0000-0001-5069-6016</mods:nameIdentifier>
<mods:affiliation id="7A30C53E7A7DDC3F83BA78F387CFF0C7">School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
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<mods:namePart id="0CFF167258AF3EB6CA68A21E741A8253">Van Do, Truong</mods:namePart>
<mods:nameIdentifier id="99A7266607CED953F2CDF2EE1E9279E3" type="ORCID">https://orcid.org/0000-0002-0585-5513</mods:nameIdentifier>
<mods:affiliation id="2282CBA31988734435043DFD69F33B1E">State Key Laboratory of Biocontrol and Guangdong Key Laboratory of Plant Resources, Sun Yat-sen University, No. 135, Xin-Gang-Xi Road, Guangzhou 510275, China &amp; Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 th Hoang Quoc Viet Road, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:name id="2A1740E4E5E4C1B5BEDC941D31EBFA9E" type="personal">
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<mods:namePart id="8969C62A94BE5E5D360849A5F4E6F1D8">Liu, Ying</mods:namePart>
<mods:nameIdentifier id="B8E4C72EB9D1F2645CDB0B357D092E5D" type="ORCID">https://orcid.org/0000-0003-0613-837X</mods:nameIdentifier>
<mods:affiliation id="C6C569CEFFE3A6763409B8206C92BDB5">School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
<mods:nameIdentifier id="0A9D13EBCD43949666B115B6BA171095" type="email">liliumrosa@163.com</mods:nameIdentifier>
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<mods:typeOfResource id="01B71611922D56FED3DEA5EB4A668A5D">text</mods:typeOfResource>
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<mods:title id="6461943BED8F80E7DE185E4B2A8C0544">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="0B9421E25A0BA3BEA641D07628090DC7">
<mods:date id="77F001527C563EFBAAE4A6862FB38F8B">2023</mods:date>
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<mods:number id="00A754E9253A531C397AC2C0118AEBC9">2023-11-03</mods:number>
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<mods:number id="3EE8CAE7404074FAF66FA6E20096B612">235</mods:number>
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<mods:url id="FF49DE9831CBCB18251CDF10E3495ACB">http://dx.doi.org/10.3897/phytokeys.235.112133</mods:url>
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<mods:classification id="9326E43160409931934F7F27937BDB26">journal article</mods:classification>
<mods:identifier id="82133FD5A6328F10C35430A10CA3A98C" type="DOI">http://dx.doi.org/10.3897/phytokeys.235.112133</mods:identifier>
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<treatment id="48251918F33D5760BF8DA2895F2356A8" ID-GBIF-Taxon="217222401" LSID="urn:lsid:plazi:treatment:48251918F33D5760BF8DA2895F2356A8" httpUri="http://treatment.plazi.org/id/48251918F33D5760BF8DA2895F2356A8" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="4DF8971FF149429C33F106AE23440B46" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="BC6F9A69AD16FAA49EE2E0EDEE43BD3F" pageId="0" pageNumber="1">
<taxonomicName id="6A91F65870B8C19A11E079B357FCD599" LSID="48251918-F33D-5760-BF8D-A2895F2356A8" authority="J. H. Dai, T. V. Do &amp; Ying Liu" authorityName="J. H. Dai, T. V. Do &amp; Ying Liu" authorityYear="2023" class="Magnoliopsida" family="Melastomataceae" genus="Perilimnastes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Perilimnastes multisepala" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="multisepala" status="sp. nov.">Perilimnastes multisepala J.H.Dai, T.V.Do &amp; Ying Liu</taxonomicName>
<taxonomicNameLabel id="4F3E424F4B095017C385D1954F7EBEBF" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="4643841CBF1E7EFF10F18DBBCCCB2C87" pageId="0" pageNumber="1" type="description">
<paragraph id="A353EB41A2F083230DD9C52EC8D63363" pageId="0" pageNumber="1">
<figureCitation id="E6218F6E07C13ADFABC62C6CA27B9B23" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Holotype of Perilimnastes multisepala, Jin-hong Dai and Ying Liu 821 (PE). The inset shows druses (as white spots) on adaxial leaf surface under stereoscope. Scale bars: 5 cm, 1 mm (inset)." figureDoi="10.3897/phytokeys.235.112133.figure2" httpUri="https://binary.pensoft.net/fig/929967" pageId="0" pageNumber="1">Figs 2</figureCitation>
<figureCitation id="0F1D697CE9C1BB01C5B4143A91AB70F6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Perilimnastes multisepala A habit B adaxial (top) and abaxial (bottom) leaf surfaces C close-up of a branchlet showing a 2 - flowered cyme. All from Jin-hong Dai and Ying Liu 821 (A, PE, SYS)." figureDoi="10.3897/phytokeys.235.112133.figure3" httpUri="https://binary.pensoft.net/fig/929968" pageId="0" pageNumber="1">, 3</figureCitation>
<figureCitation id="B48F24B0AE0E432E9AF73FA66E9536F8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Perilimnastes multisepala A top view of a flower B longitudinal section of a flower showing the isomorphic stamens C top view of a young capsule D lateral view of a young capsule E top view of an old capsule F longitudinal section of an old capsule showing enlarged ovary crown and morphology of the placental column and placentas. Scale bars: 5 mm (B); 3 mm (F). All from Jin-hong Dai and Ying Liu 821 (A, PE, SYS)." figureDoi="10.3897/phytokeys.235.112133.figure4" httpUri="https://binary.pensoft.net/fig/929969" pageId="0" pageNumber="1">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5C0A900E41C2BDED97180BD10CA6D683" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="7D9DA8CA2C5977A13406E36AF9B17C9E" pageId="0" pageNumber="1">
<typeStatus id="5A7409F223C18248E9F6E31E1136BD4F">Type</typeStatus>
.
</paragraph>
<paragraph id="23376E784C0B907F71CBB3F7C561CF58" pageId="0" pageNumber="1">
<materialsCitation id="9D834AF75D982613FA7323D3DCC27412" collectingDate="2019-11-23" collectorName="Khu Du Lich Sinh Thai Khe Lim" country="Vietnam" county="Khu Du Lich Sinh Thai" elevation="400" location="Qu" municipality="Nam Province" specimenCount="1" typeStatus="holotype">
<collectingCountry id="8B6690A340FC0B1267B509E256C543AF" name="Vietnam">Vietnam</collectingCountry>
.
<location id="5BFBE408354BD0D854105C88313AB937" LSID="urn:lsid:plazi:treatment:48251918F33D5760BF8DA2895F2356A8:5BFBE408354BD0D854105C88313AB937" country="Vietnam" county="Khu Du Lich Sinh Thai" municipality="Nam Province" name="Qu">Qu</location>
ảng
<collectingMunicipality id="21A703940B6C13D07D29E608088B5B36">Nam Province</collectingMunicipality>
:
<normalizedToken id="F061B3011B28FA3C3AD19328BDDACF8D" originalValue="Đ">Đ</normalizedToken>
ại Lộc, about
<quantity id="FDB18417F925F361101FADD0462367D8" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">
<elevation id="A273428D6D2BBC1AC4E12CA34745827E" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</elevation>
</quantity>
south of
<collectorName id="FF8A0DC47431DF4D7FB71CDCA1C28276">
<collectingCounty id="B5728CE7C5841D7753677F1D643903F2">Khu Du Lich Sinh Thai</collectingCounty>
Khe Lim
</collectorName>
, along newly opened road,
<quantity id="63FB32B4D29D0BE452DA7090ABF5BDF4" metricMagnitude="2" metricUnit="m" metricValue="5.74" unit="m" value="574.0">
<elevation id="017FBA50495D8038957178BBF8070841" metricMagnitude="2" metricUnit="m" metricValue="5.74" unit="m" value="574.0">574 m</elevation>
</quantity>
elevation, on rocks along a stream,
<collectingDate id="A46F998748DC8AEDE59E953BEF5F6619" value="2019-11-23">23 Nov 2019</collectingDate>
, Jin-hong Dai and Ying Liu 821 (
<typeStatus id="14FDADC8B37DEA45A5FC6A37FC9E3F6E">holotype</typeStatus>
: PE; isotypes: A, SYS, VNMN)
</materialsCitation>
.
</paragraph>
<caption id="5FD7A185A0283944DB3B581CC51F01D9" doi="10.3897/phytokeys.235.112133.figure2" httpUri="https://binary.pensoft.net/fig/929967" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
<paragraph id="07AB8F9B947AB2B58B5B04C2621F4D39" pageId="0" pageNumber="1">
<emphasis id="6AF13BC4C1613B58B90C38BD4B4C361C" bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
<typeStatus id="03A79EA2CFB0A495847A67C5B9648303">Holotype</typeStatus>
of
<taxonomicName id="58EAF14FFA43EEB18E299DEB5C388EA0" authorityName="J. H. Dai, T. V. Do &amp; Ying Liu" authorityYear="2023" class="Magnoliopsida" family="Melastomataceae" genus="Perilimnastes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Perilimnastes multisepala" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="multisepala">
<emphasis id="E0FD5CA06394DF297734F65C913265A9" italics="true" pageId="0" pageNumber="1">Perilimnastes multisepala</emphasis>
</taxonomicName>
, Jin-hong Dai and Ying Liu 821 (PE). The inset shows druses (as white spots) on adaxial leaf surface under stereoscope. Scale bars:
<quantity id="FFCD6281CD998FD5C7D303E037D55BD7" metricMagnitude="-2" metricUnit="m" metricValue="5.0" unit="cm" value="5.0">5 cm</quantity>
,
<quantity id="396CE5D6D6FE7A428057773BBAC72302" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
(inset).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="E47A76F2A6C96616F9FD3DB573841D98" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="5C1BFF5113D5DC4D0DACBAAB5E58D46E" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="279DEAC94F116F984BC36DF67C87438A" pageId="0" pageNumber="1">
Resembles
<taxonomicName id="115342FB65532B34E3B9DAC8B92C1A1E" lsidName="P. fruticosa" pageId="0" pageNumber="1" rank="species" species="fruticosa">
<emphasis id="C60682042E1F9CC956E11896CE5249A2" italics="true" pageId="0" pageNumber="1">P. fruticosa</emphasis>
</taxonomicName>
and
<taxonomicName id="AA27898E33420B8FCF071F3429C0C265" lsidName="P. stenophylla" pageId="0" pageNumber="1" rank="species" species="stenophylla">
<emphasis id="29FFAE5D0266EE1EEBD15B485FAA164F" italics="true" pageId="0" pageNumber="1">P. stenophylla</emphasis>
</taxonomicName>
in the habitat preference, habit, leaf and inflorescence morphology but differs from these species in the petals 28
<normalizedToken id="E04758EBF20C9D65C7361A712E7D9449" originalValue="×">x</normalizedToken>
9 mm, apex long acuminate (vs. 8.5-16
<normalizedToken id="5F3FC8125D497CB91ACCF417A0A3D4F1" originalValue="×">x</normalizedToken>
3.5-5 mm, acuminate, and 12
<normalizedToken id="A680953E332F72070BBC456AE2714E15" originalValue="×">x</normalizedToken>
6 mm, short acuminate), calyx lobes 4-8 (vs. 4) and pedicels only 1-2 mm at fruiting stage (vs. 10-15 mm).
</paragraph>
<caption id="C0F33C7DD54138A0DC0FA4CA44EC89B6" doi="10.3897/phytokeys.235.112133.figure3" httpUri="https://binary.pensoft.net/fig/929968" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
<paragraph id="B387226E1F24ECB3C94ED3DB1462BC9B" pageId="0" pageNumber="1">
<emphasis id="8FB3A746F91C102E5A3E5CED9BCFB63B" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
<taxonomicName id="779AE1BCA5C5B808B67AC668B65D0EBB" authorityName="J. H. Dai, T. V. Do &amp; Ying Liu" authorityYear="2023" class="Magnoliopsida" family="Melastomataceae" genus="Perilimnastes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Perilimnastes multisepala" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="multisepala">
<emphasis id="A0B40DA008D17093DAB690E840D3FE80" italics="true" pageId="0" pageNumber="1">Perilimnastes multisepala</emphasis>
</taxonomicName>
<emphasis id="B7A9DB569FD91BE466DA5781B3BB3E87" bold="true" pageId="0" pageNumber="1">A</emphasis>
habit
<emphasis id="F66676DD150FDE95D58D00A2D0C8B4A0" bold="true" pageId="0" pageNumber="1">B</emphasis>
adaxial (top) and abaxial (bottom) leaf surfaces
<emphasis id="CE852117AB594F94D1BA62CA790659B0" bold="true" pageId="0" pageNumber="1">C</emphasis>
close-up of a branchlet showing a 2-flowered cyme. All from Jin-hong Dai and Ying Liu 821 (A, PE, SYS).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="54E39C7E49268A3D6D7E22F2BD434F59" pageId="0" pageNumber="1" type="description">
<paragraph id="DCDF5ACAE8D8272C79EC31A32A0EF5B0" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="546BD2EE0C18CACE0D17F6EC702E1FA5" pageId="0" pageNumber="1">
Shrubs, much-branched, up to 0.8 m tall, with druses in many parts. Stems obtusely 4-sided, slightly compressed when young; branchlets glabrous, sulcate, nodes only pubescent with uniseriate hairs when young. Leaves opposite, equal to distinctly unequal in a pair, pubescent with brownish-yellow stellate hairs only when young, glabrous when mature; petiole 0-10 mm; leaf blade obovate-lanceolate, oblong-lanceolate to oblanceolate, 2.4-8
<normalizedToken id="167CD3655C40CEC007B02E061518AB1B" originalValue="×">x</normalizedToken>
0.7-2.4 cm, subcoriaceous, 3-veined with the lateral two veins often diverged from the midvein above the base, dark green adaxially, pale green abaxially, base cuneate, margin entire, apex obtuse, acute, rarely shortly acuminate. Inflorescences terminal, cymose contracted to umbellate, solitary or 2-flowered, rarely 3-flowered; peduncle ca. 1 mm long, sometimes sessile, subtended by a pair of bracts to 5 mm long. Flowers 4-merous; pedicel 1-2 mm long, glabrous; hypanthium funnel-shaped, 7-8 mm long, sparsely pubescent with stellate hairs; calyx lobes linear, laterally compressed, alternipetalous 4, 8-10 mm long, antepetalous 0-4, 3-8 mm long, sparsely pubescent with stellate hairs; petals pinkish purple, 28
<normalizedToken id="CE44C62210E09D3B107323D57A62DDC6" originalValue="×">x</normalizedToken>
9 mm, ovate, slightly oblique, apex long acuminate, abaxially very sparsely pubescent with stellate hairs; stamens 8, isomorphic, filaments 7-9 mm long, glabrous, anthers lanceolate, yellow, 9 mm long, connective decurrent, tuberculate ventrally, forming a spur dorsally; ovary half as long as hypanthium (crown excluded), ovary crown wedge-like, 4-lobed; style 22 mm long. Capsule cup-shaped, 7-8
<normalizedToken id="3F1C094023BD9D95A3BAA0FDB39F16BE" originalValue="×">x</normalizedToken>
7 mm, 4-sided; hypanthium 8-ribbed; crown enlarged enclosing an obpyramidal space; placental column unbeaked, 4-horned; placenta thready.
</paragraph>
<caption id="1395F1733B5BB49BEBB50D8169FCC206" doi="10.3897/phytokeys.235.112133.figure4" httpUri="https://binary.pensoft.net/fig/929969" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph id="C7899CF4B797A015CCFA9FF83F91E2BA" pageId="0" pageNumber="1">
<emphasis id="BD0C679AA015CAAF0FEEB3E0FB1247AF" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
<taxonomicName id="92C42803A795801E173329897D8F4664" authorityName="J. H. Dai, T. V. Do &amp; Ying Liu" authorityYear="2023" class="Magnoliopsida" family="Melastomataceae" genus="Perilimnastes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Perilimnastes multisepala" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="multisepala">
<emphasis id="CC9F720E6E337D2F3A44767453889B3B" italics="true" pageId="0" pageNumber="1">Perilimnastes multisepala</emphasis>
</taxonomicName>
<emphasis id="DB8D44D567E41347AC1D6C4D0E54FA8C" bold="true" pageId="0" pageNumber="1">A</emphasis>
top view of a flower
<emphasis id="0FC345FEACF7858FBAE56F8CB5DEC244" bold="true" pageId="0" pageNumber="1">B</emphasis>
longitudinal section of a flower showing the isomorphic stamens
<emphasis id="EDCEF758D6CBBB3122BC7CF3F287E1AE" bold="true" pageId="0" pageNumber="1">C</emphasis>
top view of a young capsule
<emphasis id="835AD834B76839B2FCCBC3956516D76E" bold="true" pageId="0" pageNumber="1">D</emphasis>
lateral view of a young capsule
<emphasis id="A37ADED5D4D49BBF67C941899BA0632C" bold="true" pageId="0" pageNumber="1">E</emphasis>
top view of an old capsule
<emphasis id="DCBBCEFD482BFB7160292F5E785AD086" bold="true" pageId="0" pageNumber="1">F</emphasis>
longitudinal section of an old capsule showing enlarged ovary crown and morphology of the placental column and placentas. Scale bars: 5 mm (
<emphasis id="532532483714F300272BF641B37DA9BB" bold="true" pageId="0" pageNumber="1">B</emphasis>
); 3 mm (
<emphasis id="DBBE5C59A3246F578EB5805B38E4FC44" bold="true" pageId="0" pageNumber="1">F</emphasis>
). All from Jin-hong Dai and Ying Liu 821 (A, PE, SYS).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="09836A0E86C1B768B5D14C53E222B4C0" pageId="0" pageNumber="1" type="phenology">
<paragraph id="35EE182CBB96DA3F802E2E0296341ADC" pageId="0" pageNumber="1">Phenology.</paragraph>
<paragraph id="F16BA34EF2B22D1759190FB3280D2126" pageId="0" pageNumber="1">Flowers, young fruits and old fruits in November.</paragraph>
</subSubSection>
<subSubSection id="B266A0061ACEA211A0B2588D65A40FEE" pageId="0" pageNumber="1" type="etymology">
<paragraph id="0BC5DF69ED6F8743785FD60BA9450DA6" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="2E0A0188FFCCAD2B81C1277EE73D2A65" pageId="0" pageNumber="1">The specific epithet is based on the 4-8 calyx lobes of this species.</paragraph>
</subSubSection>
<subSubSection id="F8584C88A0B80AE389ABB41AA235DF7E" pageId="0" pageNumber="1" type="distribution">
<paragraph id="692ABC5500F3B58006DC250468CC674B" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="D07B04359F9C6788CA85365A7F40ECA3" pageId="0" pageNumber="1">
<taxonomicName id="54B0ED12800B3B6D15EC1A0250275068" authorityName="J. H. Dai, T. V. Do &amp; Ying Liu" authorityYear="2023" class="Magnoliopsida" family="Melastomataceae" genus="Perilimnastes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Perilimnastes multisepala" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="multisepala">
<emphasis id="BF8B689AB8BCE46E3ECD733220405497" italics="true" pageId="0" pageNumber="1">Perilimnastes multisepala</emphasis>
</taxonomicName>
is currently known from
<normalizedToken id="E1CDBF4C1C42A1CC2BE39B4ABA807144" originalValue="Đ">Đ</normalizedToken>
ại Lộc, Quảng Nam Province, Vietnam (Fig.
<figureCitation id="371F802DABE7148F72D9DD343A4D652B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Perilimnastes multisepala (triangle), P. setipetiola (square), and P. uniflora and P. banaensis (solid circle)." figureDoi="10.3897/phytokeys.235.112133.figure1" httpUri="https://binary.pensoft.net/fig/929966" pageId="0" pageNumber="1">1</figureCitation>
). It grows on rocks along streams in the forest, at 574 m elevation.
</paragraph>
</subSubSection>
</treatment>
</document>