treatments-xml/data/C6/6D/87/C66D8798782B591CF6A0FCFDFC8EFF4E.xml
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<document id="1F0C8C8665847D1CFCC6FAF559DBF3A2" ID="10.11646/zootaxa.4184.2.2" ID-DOI="10.11646/zootaxa.4184.2.2" ID-GBIF-Dataset="6dce6de7-19d7-462d-a123-e94ad5135e89" ID-ISSN="1175-5326" ID-Zenodo-Dep="164749" ID-ZooBank="5B8F792E-EEDB-494D-9EB3-48CA6174CC6E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1478243740653" checkinUser="plazi" docAuthor="Leon, Stephanie &amp; Weirauch, Christiane" docDate="2016" docId="C66D8798782B591CF6A0FCFDFC8EFF4E" docLanguage="en" docName="zootaxa.4184.2.2.pdf" docOrigin="Zootaxa 4184 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Schizoptera (Odontorhagus) quasicompleta Leon &amp; Weirauch, 2016, n. sp." docType="treatment" docVersion="9" lastPageNumber="275" masterDocId="3A54FFE078385908F637FFBFFFD8FF9B" masterDocTitle="Scratching the surface? Taxonomic revision of the subgenus Schizoptera (Odontorhagus) reveals vast undocumented biodiversity in the largest litter bug genus Schizoptera Fieber (Hemiptera: Dipsocoromorpha)" masterLastPageNumber="284" masterPageNumber="255" pageNumber="274" updateTime="1698391976884" updateUser="plazi">
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<mods:title id="D3421030F4426CB3D142E1AAE4973671">Scratching the surface? Taxonomic revision of the subgenus Schizoptera (Odontorhagus) reveals vast undocumented biodiversity in the largest litter bug genus Schizoptera Fieber (Hemiptera: Dipsocoromorpha)</mods:title>
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<mods:namePart id="B2852FE88EFC620107A54089A82656B1">Leon, Stephanie</mods:namePart>
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<mods:namePart id="2BB86F6E9CAC9910CC9607C732F2B096">Weirauch, Christiane</mods:namePart>
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<treatment id="C66D8798782B591CF6A0FCFDFC8EFF4E" ID-DOI="http://doi.org/10.5281/zenodo.6066344" ID-GBIF-Taxon="124562898" ID-Zenodo-Dep="6066344" LSID="urn:lsid:plazi:treatment:C66D8798782B591CF6A0FCFDFC8EFF4E" httpUri="http://treatment.plazi.org/id/C66D8798782B591CF6A0FCFDFC8EFF4E" lastPageId="20" lastPageNumber="275" pageId="19" pageNumber="274">
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<taxonomicName id="89C44D0D782B591BF6A0FCFDFD7BFCC7" ID-CoL="036bf7da-24e0-425f-829e-b0f0583e979f" box="[151,675,834,860]" class="Insecta" family="Schizopteridae" genus="Schizoptera" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="274" phylum="Arthropoda" rank="species" species="quasicompleta" status="sp. nov." subGenus="Odontorhagus">
<emphasis id="7CB0EA9C782B591BF6A0FCFDFEFDFCC7" bold="true" box="[151,293,834,860]" italics="true" pageId="19" pageNumber="274">Schizoptera</emphasis>
(
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)
<emphasis id="7CB0EA9C782B591BF7C2FCFDFD7BFCC7" bold="true" box="[501,675,834,860]" italics="true" pageId="19" pageNumber="274">quasicompleta</emphasis>
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,
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<paragraph id="4E7B368E782B591BF6A0FCDAFE97FCE5" blockId="19.[151,756,834,894]" box="[151,335,869,894]" pageId="19" pageNumber="274">
(
<figureCitation id="D6FF2A0B782B591BF6A8FCDAFF3FFCE5" box="[159,231,869,894]" captionStart="FIGURE 3" captionStartId="7.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1664]" captionTargetId="figure@7.[151,1436,193,1668]" captionTargetPageId="7" captionText="FIGURE 3. Habitus images of S. (Odontorhagus) species." httpUri="https://zenodo.org/record/164752/files/figure.png" pageId="19" pageNumber="274">Figs 3</figureCitation>
,
<figureCitation id="D6FF2A0B782B591BF6C4FCDAFED9FCE5" box="[243,257,869,894]" captionStart="FIGURE 6 M W" captionStartId="21.[151,250,1861,1883]" captionTargetBox="[170,1427,199,1825]" captionTargetId="figure@21.[151,1436,193,1840]" captionTargetPageId="21" captionText="FIGURE 6 M W. Line drawings of dorsal (above) and ventral (below) views of male genitalia, different colors correspond to different structures labeled on S. (Odonotorhagus) acuta, n. sp. (Fig. 5 A)." httpUri="https://zenodo.org/record/256164/files/figure.png" pageId="19" pageNumber="274">6</figureCitation>
and
<figureCitation id="D6FF2A0B782B591BF70FFCD9FE9FFCE5" box="[312,327,870,894]" captionStart="FIGURE 7" captionStartId="22.[151,250,1896,1918]" captionText="FIGURE 7. Distribution of S. (Odontorhagus) species." pageId="19" pageNumber="274">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="06DE6505782B591BF6A0FC15FD33FA44" pageId="19" pageNumber="274" type="diagnosis">
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<emphasis id="7CB0EA9C782B591BF6A0FC15FEC9FC58" bold="true" box="[151,273,938,963]" pageId="19" pageNumber="274">Diagnosis.</emphasis>
Distinguished among species of
<taxonomicName id="89C44D0D782B591BF499FC14FC2DFC58" authority="Odontorhagus" authorityName="Odontorhagus" box="[686,1013,939,963]" class="Insecta" family="Schizopteridae" genus="Schizoptera" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="7CB0EA9C782B591BF499FC14FCEAFC58" box="[686,818,939,963]" italics="true" pageId="19" pageNumber="274">Schizoptera</emphasis>
(
<emphasis id="7CB0EA9C782B591BF57CFC14FC34FC58" box="[843,1004,939,963]" italics="true" pageId="19" pageNumber="274">Odontorhagus</emphasis>
)
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by the rounded head, the weakly asymmetrical subgenital plate, the tab-like process of stVI, the small, blunt process of the subgenital plate, the needlelike right conjunctival appendage, and the uniformly thin stem of the right paramere.
</paragraph>
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:
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10.43333
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84.01667
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. La Selva Biol. Sta., 10.43056
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N 84.00667
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, B. Brown, L. Gonzales, and K. Walker, 1♂ (00096271) (UCR);
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, J. Mottern, 2♀ (00081614, 00082318) (UCR).
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trail,
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10.43028
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(00122238, 00122239) (
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</collectingMunicipality>
,
<location id="4B1B6055782B591BF747FB50FE61FA9C" LSID="urn:lsid:plazi:treatment:C66D8798782B591CF6A0FCFDFC8EFF4E:4B1B6055782B591BF747FB50FE61FA9C" box="[368,441,1263,1287]" country="Costa Rica" latitude="10.43111" longLatPrecision="1" longitude="84.02389" municipality="La Selva Biological Station" name="Jaguar" pageId="19" pageNumber="274" stateProvince="Heredia">Jaguar</location>
trail entrance,
<geoCoordinate id="2BF05049782B591BF455FB50FD30FA9D" box="[610,744,1262,1287]" direction="north" orientation="latitude" pageId="19" pageNumber="274" precision="1" value="10.43111">
10.43111
<emphasis id="7CB0EA9C782B591BF4FEFB51FD0CFA9C" bold="true" box="[713,724,1262,1287]" pageId="19" pageNumber="274">°</emphasis>
N
</geoCoordinate>
<geoCoordinate id="2BF05049782B591BF4C7FB50FCAFFA9D" box="[752,887,1262,1287]" direction="east" orientation="longitude" pageId="19" pageNumber="274" precision="1" value="84.02389">
84.02389
<emphasis id="7CB0EA9C782B591BF56EFB51FCBCFA9C" bold="true" box="[857,868,1262,1287]" pageId="19" pageNumber="274">°</emphasis>
E
</geoCoordinate>
,
<quantity id="893C9B6B782B591BF5B3FB51FC67FA9D" box="[900,959,1262,1287]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="274" unit="m" value="60.0">
<elevation id="C5E9D1BD782B591BF5B3FB51FC67FA9D" box="[900,959,1262,1287]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="274" unit="m" value="60.0">60 m</elevation>
</quantity>
,
<date id="3A7A104E782B591BF5FCFB50FBBBFA9C" box="[971,1123,1263,1287]" pageId="19" pageNumber="274" value="2011-05-21">
<collectingDate id="2A3EE9A6782B591BF5FCFB50FBBBFA9C" box="[971,1123,1263,1287]" pageId="19" pageNumber="274" value="2011-05-21">21 May 2011</collectingDate>
</date>
,
<collectorName id="E3315358782B591BF258FB50FB0CFA9C" box="[1135,1236,1263,1287]" pageId="19" pageNumber="274">J. Heraty</collectorName>
&amp;
<collectorName id="E3315358782B591BF2CEFB50FAB5FA9C" box="[1273,1389,1263,1287]" pageId="19" pageNumber="274">E. Murray</collectorName>
,
<specimenCount id="58C2FD07782B591BF34EFB50FA44FA9C" box="[1401,1436,1263,1287]" pageId="19" pageNumber="274" type="female">1♀</specimenCount>
(00084362) (
<collectionCode id="28D5AE4B782B591BF71FFAACFEB0FAB0" box="[296,360,1299,1323]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15736" name="University of California" pageId="19" pageNumber="274">UCR</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="FEAC3CD3782B591BF74DFAACFD33FAD4" ID-GBIF-Occurrence="1324639527" collectionCode="INBIO" collectorName="La Selva Biological &amp; No Date" country="Costa Rica" elevation="52" latitude="10.43086" location="La Selva Biological Station" longLatPrecision="1" longitude="-84.00647" pageId="19" pageNumber="274" specimenCount="2" specimenCount-male="2" stateProvince="Heredia" typeStatus="paratype">
<location id="4B1B6055782B591BF74DFAACFD68FAB0" LSID="urn:lsid:plazi:treatment:C66D8798782B591CF6A0FCFDFC8EFF4E:4B1B6055782B591BF74DFAACFD68FAB0" box="[378,688,1298,1323]" country="Costa Rica" latitude="10.43086" longLatPrecision="1" longitude="-84.00647" name="La Selva Biological Station" pageId="19" pageNumber="274" stateProvince="Heredia">
<collectorName id="E3315358782B591BF74DFAACFD80FAB0" box="[378,600,1298,1323]" pageId="19" pageNumber="274">La Selva Biological</collectorName>
Station
</location>
, nr Puerto Viejo,
<geoCoordinate id="2BF05049782B591BF544FAADFC23FAB1" box="[883,1019,1298,1323]" direction="north" orientation="latitude" pageId="19" pageNumber="274" precision="1" value="10.43086">
10.43086
<emphasis id="7CB0EA9C782B591BF5EBFAADFC3FFAB0" bold="true" box="[988,999,1298,1323]" pageId="19" pageNumber="274">°</emphasis>
N
</geoCoordinate>
<geoCoordinate id="2BF05049782B591BF235FAADFB49FAB1" box="[1026,1169,1298,1323]" direction="west" orientation="longitude" pageId="19" pageNumber="274" precision="1" value="-84.00647">
84.00647
<emphasis id="7CB0EA9C782B591BF25CFAADFBAEFAB0" bold="true" box="[1131,1142,1298,1323]" pageId="19" pageNumber="274">°</emphasis>
W
</geoCoordinate>
,
<quantity id="893C9B6B782B591BF2ACFAADFB0EFAB1" box="[1179,1238,1298,1323]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="19" pageNumber="274" unit="m" value="52.0">
<elevation id="C5E9D1BD782B591BF2ACFAADFB0EFAB1" box="[1179,1238,1298,1323]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="19" pageNumber="274" unit="m" value="52.0">52 m</elevation>
</quantity>
,
<collectorName id="E3315358782B591BF2D6FAACFA9AFAB1" box="[1249,1346,1299,1322]" pageId="19" pageNumber="274">No Date</collectorName>
, INBio- OET-ALAS,
<specimenCount id="58C2FD07782B591BF701FA89FE80FAD4" box="[310,344,1334,1359]" pageId="19" pageNumber="274" type="male">2♂</specimenCount>
(00109180, 00109181) (
<collectionCode id="28D5AE4B782B591BF4B8FA89FD3BFAD4" box="[655,739,1334,1359]" pageId="19" pageNumber="274">INBIO</collectionCode>
)
</materialsCitation>
;
<date id="3A7A104E782B591BF536FA89FC7DFAD4" box="[769,933,1334,1359]" pageId="19" pageNumber="274" value="1998-02-16">16 Feb 1998</date>
, INBio-OET-ALAS, 1♂ (00109178), 1♀ (00109179) (INBIO);
<date id="3A7A104E782B591BF7A3FAE5FDF0FAE8" box="[404,552,1370,1395]" pageId="19" pageNumber="274" value="1998-04-16">16 Apr 1998</date>
, INBio-OET-ALAS, 1♂ (00109172), 1♀ (00109173) (INBIO);
<date id="3A7A104E782B591BF334FAE5FA40FAE8" box="[1283,1432,1370,1395]" pageId="19" pageNumber="274" value="1998-04-16">16 Apr 1998</date>
, INBio-OET-ALAS, 4♂ (00109146- 00109149) (INBIO);
<date id="3A7A104E782B591BF571FAC1FC36FA0C" box="[838,1006,1406,1431]" pageId="19" pageNumber="274" value="1998-05-19">19 May 1998</date>
, INBio-OET-ALAS,
<collectingMethod id="97854E99782B591BF2DAFAC1FABEFA0D" box="[1261,1382,1406,1431]" pageId="19" pageNumber="274">light Trap</collectingMethod>
, 4♂ (0010915800109161), 4♀ (0010916200109165), 1♂ (00109154) (INBIO);
<date id="3A7A104E782B591BF220FA1DFB69FA20" box="[1047,1201,1442,1467]" pageId="19" pageNumber="274" value="1999-12-30">30 Dec 1999</date>
, INBio-OET-ALAS, 2♀ (00109121, 00109122), 1Ƌ (00109123) (INBIO).
</paragraph>
</subSubSection>
<subSubSection id="06DE6505782B591BF6F0FA55FCCBF828" pageId="19" pageNumber="274" type="description">
<paragraph id="4E7B368E782B591BF6F0FA55FEECF814" blockId="19.[151,1436,938,2007]" pageId="19" pageNumber="274">
<emphasis id="7CB0EA9C782B591BF6F0FA55FE81F998" bold="true" box="[199,345,1514,1539]" pageId="19" pageNumber="274">Description.</emphasis>
<emphasis id="7CB0EA9C782B591BF753FA54FE46F998" box="[356,414,1515,1539]" italics="true" pageId="19" pageNumber="274">Male</emphasis>
: length:
<quantity id="893C9B6B782B591BF43DFA54FD6DF999" box="[522,693,1515,1539]" metricMagnitude="-3" metricUnit="m" metricValue="1.4049999999999998" metricValueMax="1.42" metricValueMin="1.39" pageId="19" pageNumber="274" unit="mm" value="1.4049999999999998" valueMax="1.42" valueMin="1.39">1.391.42 mm</quantity>
; body broadly ovate;
<emphasis id="7CB0EA9C782B591BF58DFA55FB5BF998" bold="true" box="[954,1155,1514,1539]" pageId="19" pageNumber="274">COLORATION:</emphasis>
general coloration dark brown; fuscous patch covering more than 1/2 of wing membrane; Sc+C concolorous with remainder of hemelytra (
<figureCitation id="D6FF2A0B782B591BF6A8F98DFF3CF9D0" box="[159,228,1586,1611]" captionStart="FIGURE 3" captionStartId="7.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1664]" captionTargetId="figure@7.[151,1436,193,1668]" captionTargetPageId="7" captionText="FIGURE 3. Habitus images of S. (Odontorhagus) species." httpUri="https://zenodo.org/record/164752/files/figure.png" pageId="19" pageNumber="274">Fig. 3</figureCitation>
).
<emphasis id="7CB0EA9C782B591BF6CCF98DFDAEF9D0" bold="true" box="[251,630,1586,1611]" pageId="19" pageNumber="274">SURFACE AND VESTITURE:</emphasis>
as in subgeneric description.
<emphasis id="7CB0EA9C782B591BF5FFF98DFBA6F9D0" bold="true" box="[968,1150,1586,1611]" pageId="19" pageNumber="274">STRUCTURE:</emphasis>
HEAD: slightly rounded in ventral view (
<figureCitation id="D6FF2A0B782B591BF779F9E9FE4BF9F4" box="[334,403,1622,1647]" captionStart="FIGURE 3" captionStartId="7.[151,250,1689,1711]" captionTargetBox="[151,1436,193,1664]" captionTargetId="figure@7.[151,1436,193,1668]" captionTargetPageId="7" captionText="FIGURE 3. Habitus images of S. (Odontorhagus) species." httpUri="https://zenodo.org/record/164752/files/figure.png" pageId="19" pageNumber="274">Fig. 3</figureCitation>
); eyes about 1/3 as wide as synthlipsis. THORAX: as in subgeneric description; hemelytron as in subgeneric description; M2 vein distinctly curving; anterior membranal cell wedge-shaped. ABDOMEN: as in subgeneric description; stVI with a tab-like process on posterior margin; subgenital plate weakly asymmetrical; process of subgenital plate blunt, not fully differentiated from subgenital plate (
<figureCitation id="D6FF2A0B782B591BF21AF97DFB5DF940" box="[1069,1157,1730,1755]" captionStart="FIGURE 6 M W" captionStartId="21.[151,250,1861,1883]" captionTargetBox="[170,1427,199,1825]" captionTargetId="figure@21.[151,1436,193,1840]" captionTargetPageId="21" captionText="FIGURE 6 M W. Line drawings of dorsal (above) and ventral (below) views of male genitalia, different colors correspond to different structures labeled on S. (Odonotorhagus) acuta, n. sp. (Fig. 5 A)." httpUri="https://zenodo.org/record/256164/files/figure.png" pageId="19" pageNumber="274">Fig. 6P</figureCitation>
); tVII rectangular; tVIII weakly rectangular and sclerotized; length of tVII about 1/3 length of tVII. GENITALIA: py as in subgeneric description; rp as in subgeneric description with a uniformly thin stem; lp as in subgeneric description, with a blunt triangular dorsal process; rca needlelike; lca with a short, about 1/3 length of rca, tapering tip (
<figureCitation id="D6FF2A0B782B591BF2D1F891FAE7F8DC" box="[1254,1343,1838,1863]" captionStart="FIGURE 6 M W" captionStartId="21.[151,250,1861,1883]" captionTargetBox="[170,1427,199,1825]" captionTargetId="figure@21.[151,1436,193,1840]" captionTargetPageId="21" captionText="FIGURE 6 M W. Line drawings of dorsal (above) and ventral (below) views of male genitalia, different colors correspond to different structures labeled on S. (Odonotorhagus) acuta, n. sp. (Fig. 5 A)." httpUri="https://zenodo.org/record/256164/files/figure.png" pageId="19" pageNumber="274">Fig. 6P</figureCitation>
); vesica uniformly thin with a needlelike tip, forming two coils; anophore as in subgeneric description with a tab-like dorsal flap (
<figureCitation id="D6FF2A0B782B591BF6E5F8C9FEFFF814" box="[210,295,1910,1935]" captionStart="FIGURE 6 M W" captionStartId="21.[151,250,1861,1883]" captionTargetBox="[170,1427,199,1825]" captionTargetId="figure@21.[151,1436,193,1840]" captionTargetPageId="21" captionText="FIGURE 6 M W. Line drawings of dorsal (above) and ventral (below) views of male genitalia, different colors correspond to different structures labeled on S. (Odonotorhagus) acuta, n. sp. (Fig. 5 A)." httpUri="https://zenodo.org/record/256164/files/figure.png" pageId="19" pageNumber="274">Fig. 6P</figureCitation>
).
</paragraph>
<paragraph id="4E7B368E782B591BF6F0F824FCCBF828" blockId="19.[151,1436,938,2007]" box="[199,787,1946,1971]" pageId="19" pageNumber="274">
<emphasis id="7CB0EA9C782B591BF6F0F824FEC3F828" box="[199,283,1947,1971]" italics="true" pageId="19" pageNumber="274">Female</emphasis>
: length
<quantity id="893C9B6B782B591BF741F824FE05F829" box="[374,477,1947,1971]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="19" pageNumber="274" unit="mm" value="1.42">1.42 mm</quantity>
; general habitus as in male.
</paragraph>
</subSubSection>
<subSubSection id="06DE6505782B591BF6F0F801FA4DF84C" box="[199,1429,1982,2007]" pageId="19" pageNumber="274" type="distribution">
<paragraph id="4E7B368E782B591BF6F0F801FA4DF84C" blockId="19.[151,1436,938,2007]" box="[199,1429,1982,2007]" pageId="19" pageNumber="274">
<emphasis id="7CB0EA9C782B591BF6F0F801FEB8F84C" bold="true" box="[199,352,1982,2007]" pageId="19" pageNumber="274">Distribution.</emphasis>
Known from multiple collection events at the La Selva Biological station in
<collectingCountry id="36D3761E782B591BF28DF800FAECF84C" box="[1210,1332,1982,2007]" name="Costa Rica" pageId="19" pageNumber="274">Costa Rica</collectingCountry>
(
<figureCitation id="D6FF2A0B782B591BF374F801FA5FF84C" box="[1347,1415,1982,2007]" captionStart="FIGURE 7" captionStartId="22.[151,250,1896,1918]" captionText="FIGURE 7. Distribution of S. (Odontorhagus) species." pageId="19" pageNumber="274">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="06DE6505782C591CF6F0FF28FC8EFF4E" pageId="20" pageNumber="275" type="etymology">
<paragraph id="4E7B368E782C591CF6F0FF28FC8EFF4E" blockId="20.[151,1436,151,213]" pageId="20" pageNumber="275">
<emphasis id="7CB0EA9C782C591CF6F0FF28FE96FF2B" bold="true" box="[199,334,151,176]" pageId="20" pageNumber="275">Etymology.</emphasis>
Derived from the Latin “quasi” meaning as if and “completus” meaning completed because of the seemingly “entire” look of the subgenital plate in ventral view.
</paragraph>
</subSubSection>
</treatment>
</document>