1394 lines
115 KiB
XML
1394 lines
115 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/vz.73.e91111" ID-Pensoft-Pub="2625-8498-73-127" ID-Pensoft-UUID="083B431D3FFD57A89E7572A415F0622C" ID-ZooBank="1B781DFC3B2E4833BBF289AF72A195FC" ModsDocID="2625-8498-73-127" checkinTime="1674741901026" checkinUser="pensoft" docAuthor="Oliver, Paul M., Iskandar, Djoko T. & Richards, Stephen J." docDate="2023" docId="60AD9422E29A5F319112726FADC2753D" docLanguage="en" docName="VertZool 73: 127-139" docOrigin="Vertebrate Zoology 73" docPubDate="2023-01-26" docSource="http://dx.doi.org/10.3897/vz.73.e91111" docTitle="Litoria hastula Oliver & Iskandar & Richards 2023, sp. nov." docType="treatment" docVersion="2" id="083B431D3FFD57A89E7572A415F0622C" lastPageNumber="127" masterDocId="083B431D3FFD57A89E7572A415F0622C" masterDocTitle="A new species of torrent-breeding treefrog (Pelodryadidae: Litoria) from the mountains of Papua, Indonesia, with new records and observations of Litoria dorsivena (Tyler, 1968)" masterLastPageNumber="139" masterPageNumber="127" pageNumber="127" updateTime="1674742202005" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new species of torrent-breeding treefrog (Pelodryadidae: Litoria) from the mountains of Papua, Indonesia, with new records and observations of Litoria dorsivena (Tyler, 1968)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Oliver, Paul M.</mods:namePart>
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<mods:affiliation>Centre for Planetary Health and Food Security, Griffith University, 170 Kessels Rd, Brisbane, Queensland 4121, and Biodiversity and Geosciences Program, Queensland Museum, South Brisbane, Queensland, 4101, Australia</mods:affiliation>
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<mods:nameIdentifier type="email">p.oliver@griffith.edu.au</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Iskandar, Djoko T.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8923-6081</mods:nameIdentifier>
|
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<mods:affiliation>School of Life Sciences and Technology, Institut Teknologi Bandung, 10 Jalan Ganesha, Bandung 40132, Indonesia & Basic Sciences Commission, Indonesian Academy of Sciences. 11, Jl. Medan Merdeka Selatan Jakarta 10110, Indonesia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Richards, Stephen J.</mods:namePart>
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<mods:affiliation>Herpetology Department, South Australian Museum, North Terrace, Adelaide, South Australia 5000, Australia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Vertebrate Zoology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-01-26</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>73</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>127</mods:start>
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<mods:end>139</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/vz.73.e91111</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/vz.73.e91111</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2625-8498-73-127</mods:identifier>
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<mods:identifier type="ZooBank">1B781DFC3B2E4833BBF289AF72A195FC</mods:identifier>
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<mods:identifier type="Pensoft-UUID">083B431D3FFD57A89E7572A415F0622C</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:ACt/5492FB0B-D70C-4620-86BB-8CF7BB1092D9" httpUri="http://treatment.plazi.org/id/60AD9422E29A5F319112726FADC2753D" lastPageNumber="127" pageId="0" pageNumber="127">
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<subSubSection pageId="0" pageNumber="127" type="nomenclature">
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<paragraph pageId="0" pageNumber="127">
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<taxonomicName LSID="https://zoobank.org/act/5492FB0B-D70C-4620-86BB-8CF7BB1092D9" authority="Oliver & Iskandar & Richards, 2023" authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula" status="sp. nov.">Litoria hastula</taxonomicName>
|
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<taxonomicNameLabel pageId="0" pageNumber="127">sp. nov.</taxonomicNameLabel>
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</paragraph>
|
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</subSubSection>
|
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<subSubSection pageId="0" pageNumber="127" type="description">
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Type series of Litoria hastula sp. nov. showing distinctly acuminate snout tip and marked sexual size dimorphism: A dorsal view and B ventral view. Smaller males from top left to bottom right are: MZB Amph. 32873 (holotype); SAMA R 72334, SAMA R 72335, MZB Amph. 32874. Much larger female on far right is MZB Amph. 32872. Tags on the specimens are field numbers. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e91111.figure1" httpUri="https://binary.pensoft.net/fig/800713" pageId="0" pageNumber="127">Figs 1</figureCitation>
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Details of holotype of Litoria hastula sp. nov. MZB Amph. 32873: A lateral view of head; B dorsal view of head; C heels with conical tubercles, D ventral view of hand, and E) ventral view of foot." figureDoi="10.3897/vz.73.e91111.figure2" httpUri="https://binary.pensoft.net/fig/800714" pageId="0" pageNumber="127">, 2</figureCitation>
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Colouration of Litoria hastula sp. nov. in life: A lateral view of adult male paratype SAMA R 72334; B hidden coloration in thighs of male paratype SAMA R 72335; C lateral view of adult female paratype MZB Amph. 32872; D ventral view of adult female paratype MZB Amph. 32872. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure3" httpUri="https://binary.pensoft.net/fig/800715" pageId="0" pageNumber="127">, 3</figureCitation>
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Comparison of head profile of a male paratype of Litoria dorsivena SAMA R 7907 (left) and holotype of Litoria hastula MZB Amph. 32873 (right). Note much more sharply pointed tip of snout on Litoria hastula sp. nov." figureDoi="10.3897/vz.73.e91111.figure7" httpUri="https://binary.pensoft.net/fig/800719" pageId="0" pageNumber="127">, 7</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="reference_group">
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<taxonomicName authority="Oliver & Iskandar & Richards, 2023" authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="genus" status="sp. nov.">Litoria</taxonomicName>
|
||
sp. 10 in
|
||
<bibRefCitation author="Richards, SJ" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="127" refId="B18" refString="Richards, SJ, Iskandar, DT, Allison, A, 2000. Amphibian and reptiles recorded on the RAP survey in the Wapoga area, Irian Jaya, Indonesia. In: Mack AL, Alonso L (Eds) A Biological Assessment of the Wapoga River area of northwestern Irian Jaya, Indonesia. Conservation International, Arlington, pp. 113-115." title="Amphibian and reptiles recorded on the RAP survey in the Wapoga area, Irian Jaya, Indonesia. In: Mack AL, Alonso L (Eds) A Biological Assessment of the Wapoga River area of northwestern Irian Jaya, Indonesia. Conservation International, Arlington, pp. 113 - 115." year="2000">Richards et al. (2000)</bibRefCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="holotype">
|
||
<paragraph pageId="0" pageNumber="127">Holotype.</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
MZB Amph.32873 (SJR[JCUNQ]4141), adult male, ~2090 m altitude, unnamed mountain range, Derewo River Basin, Papua Province, Indonesia, (
|
||
<geoCoordinate degrees="3.4416" direction="south" orientation="latitude" precision="5" value="-3.4416">3.4416°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="136.4738" direction="east" orientation="longitude" precision="5" value="136.4738">136.4738°E</geoCoordinate>
|
||
), collected by Stephen Richards and Djoko Iskandar, 4th April 1998.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="paratypes (n = 4)">
|
||
<paragraph pageId="0" pageNumber="127">Paratypes (n = 4).</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">SAMA R72334-5 (SJR[JCUNQ]4142, 4144), MZB Amph.32874 (SJR[JCUNQ]4152) all adult males, MZB Amph.32872 (SJR[JCUNQ4140]) adult female, same locality and collectors as holotype, between 4-5 April 1998.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="127">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
A species of
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria</emphasis>
|
||
</taxonomicName>
|
||
that can be distinguished from all congeners by the following unique combination of characters: moderate size and strong sexual size dimorphism (4 adult males 30.4-31.8 mm SVL, one adult female 48.9 mm SVL); snout moderately long (EN/IN 0.69-0.80), with sharply acuminate tip and with concave dorsal surface; canthus rostralis nearly straight, sharply defined; limbs moderately long (TL/SVL 0.55-0.60); finger webbing moderate, not extending beyond third phalanx between Finger 3 and Finger 4; toe webbing extensive, extending to penultimate phalanx between all digits except Toe 1 and Toe 2; dorsal skin relatively smooth with scattered small tubercles; heel with 2-3 distinct conical tubercles; vomeropalatines prominent; vocal slits present in males; dorsal colouration predominately light to mid-brown with scattered darker-brown spots and or blotches; venter largely buff with little to no pattern; and advertisement calls produced in series of 5-7 calls, each comprising a single short note that is unpulsed or slightly pulsatile but may become longer, with discrete pulses, in terminal calls of the series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="description">
|
||
<paragraph pageId="0" pageNumber="127">Description of the holotype.</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
Adult male with vocal slits and pale-brown nuptial pads. Body moderately slender, limbs long (TL/SVL 0.60), head moderately wide (HW/SVL 0.34), slightly longer than wide (HL/SVL 0.34, HL/HW = 1.04). Vomerine teeth in two small but prominent clumps, each approximately 0.5 mm in diameter. Tongue large, broadly oval; lateral margins with distinct indentations, posterior margin with deep notch; vocal slits laterally in floor of mouth, extending from near angle of jaws to about one third distance to front of jaw. Snout protruding distinctly beyond lower jaw, tip sharply acuminate in both dorsal and lateral views, only slightly less so in lateral view, dorsal surface of snout distinctly concave (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Type series of Litoria hastula sp. nov. showing distinctly acuminate snout tip and marked sexual size dimorphism: A dorsal view and B ventral view. Smaller males from top left to bottom right are: MZB Amph. 32873 (holotype); SAMA R 72334, SAMA R 72335, MZB Amph. 32874. Much larger female on far right is MZB Amph. 32872. Tags on the specimens are field numbers. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e91111.figure1" httpUri="https://binary.pensoft.net/fig/800713" pageId="0" pageNumber="127">1</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Details of holotype of Litoria hastula sp. nov. MZB Amph. 32873: A lateral view of head; B dorsal view of head; C heels with conical tubercles, D ventral view of hand, and E) ventral view of foot." figureDoi="10.3897/vz.73.e91111.figure2" httpUri="https://binary.pensoft.net/fig/800714" pageId="0" pageNumber="127">2A</figureCitation>
|
||
). Canthus rostralis nearly straight, sharply defined (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Details of holotype of Litoria hastula sp. nov. MZB Amph. 32873: A lateral view of head; B dorsal view of head; C heels with conical tubercles, D ventral view of hand, and E) ventral view of foot." figureDoi="10.3897/vz.73.e91111.figure2" httpUri="https://binary.pensoft.net/fig/800714" pageId="0" pageNumber="127">2B</figureCitation>
|
||
); loreal region steeply sloping, slightly concave in dorsal view; nostrils slightly closer to tip of snout than to eyes; internarial distance greater than distance from external naris to eye (EN/IN 0.73, IN/SVL 0.12, EN/SVL 0.086). Eyes large (EYE/SVL 0.13), prominent, protruding in dorsal and ventral views; pupil horizontal, pigmentation on nictitating membrane restricted to narrow band along dorsal edge. Tympanum small (TYM/SVL 0.04), less than half diameter of eye (TYM/EYE = 0.35), annulus poorly defined, particularly on right side, dorsal ~1/3 obscured by moderately thick, slightly curved supratympanic ridge.
|
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</paragraph>
|
||
<caption doi="10.3897/vz.73.e91111.figure1" httpUri="https://binary.pensoft.net/fig/800713" pageId="0" pageNumber="127" start="Figure 1" startId="F1">
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">Figure 1.</emphasis>
|
||
Type series of
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
showing distinctly acuminate snout tip and marked sexual size dimorphism:
|
||
<emphasis bold="true" pageId="0" pageNumber="127">A</emphasis>
|
||
dorsal view and
|
||
<emphasis bold="true" pageId="0" pageNumber="127">B</emphasis>
|
||
ventral view. Smaller males from top left to bottom right are: MZB Amph.32873 (holotype); SAMA R72334, SAMA R72335, MZB Amph.32874. Much larger female on far right is MZB Amph.32872. Tags on the specimens are field numbers. Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/vz.73.e91111.figure2" httpUri="https://binary.pensoft.net/fig/800714" pageId="0" pageNumber="127" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">Figure 2.</emphasis>
|
||
Details of holotype of
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
MZB Amph.32873:
|
||
<emphasis bold="true" pageId="0" pageNumber="127">A</emphasis>
|
||
lateral view of head;
|
||
<emphasis bold="true" pageId="0" pageNumber="127">B</emphasis>
|
||
dorsal view of head;
|
||
<emphasis bold="true" pageId="0" pageNumber="127">C</emphasis>
|
||
heels with conical tubercles,
|
||
<emphasis bold="true" pageId="0" pageNumber="127">D</emphasis>
|
||
ventral view of hand, and E) ventral view of foot.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
Skin of dorsum and dorsal surfaces of limbs smooth except for scattered low, indistinct tubercles (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Type series of Litoria hastula sp. nov. showing distinctly acuminate snout tip and marked sexual size dimorphism: A dorsal view and B ventral view. Smaller males from top left to bottom right are: MZB Amph. 32873 (holotype); SAMA R 72334, SAMA R 72335, MZB Amph. 32874. Much larger female on far right is MZB Amph. 32872. Tags on the specimens are field numbers. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e91111.figure1" httpUri="https://binary.pensoft.net/fig/800713" pageId="0" pageNumber="127">1</figureCitation>
|
||
); skin on throat and chest loose, folded and with patches of low granules; abdomen coarsely granular; patch of low, pale buff tubercles extends posteriorly from behind and below eye, under lower edge of tympanum to axilla. Ventral surfaces of forelimbs smooth with scattered tubercles; ventral surfaces of thighs coarsely granular proximally, becoming less so distally, with patch of large pale buff tubercles below vent; tarsus and tibia smooth ventrally. Low, pale buff tubercles form indistinct ridge along outer edge of forearm; heels both with one enlarged, conical and two smaller but distinct tubercles (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Details of holotype of Litoria hastula sp. nov. MZB Amph. 32873: A lateral view of head; B dorsal view of head; C heels with conical tubercles, D ventral view of hand, and E) ventral view of foot." figureDoi="10.3897/vz.73.e91111.figure2" httpUri="https://binary.pensoft.net/fig/800714" pageId="0" pageNumber="127">2C</figureCitation>
|
||
); eyelids with two prominent and several smaller conical tubercles dorsally.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
Fingers moderately long with prominently expanded terminal discs (3FD/3FP 1.85; 3FD/SVL 0.08) with distinct circum-marginal grooves (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Details of holotype of Litoria hastula sp. nov. MZB Amph. 32873: A lateral view of head; B dorsal view of head; C heels with conical tubercles, D ventral view of hand, and E) ventral view of foot." figureDoi="10.3897/vz.73.e91111.figure2" httpUri="https://binary.pensoft.net/fig/800714" pageId="0" pageNumber="127">2D</figureCitation>
|
||
); relative lengths of fingers 3>4>2>1, subarticular tubercles unilobed. Webbing moderate, reaching halfway to subarticular tubercle at base of penultimate phalanx on inside of Finger 4; to tubercle at base of ultipenultimate phalanx before continuing as narrow flange to subarticular tubercle at base of penultimate phalanx on outside of Finger 3; in a narrow basal flange between fingers 2 and 3; and absent between fingers 1 and 2. Hand with indistinct round (0.6 mm diameter), unpigmented outer, and oval (1.5 mm long) inner metacarpal tubercles. Narrow pale-brown nuptial pad on first finger low, granular, extending 2.5 mm along outer edge of Finger 1, wider proximally than distally, distal ~25% angled at nearly right angles across base of thumb. Toes with expanded terminal discs with terminal grooves (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Details of holotype of Litoria hastula sp. nov. MZB Amph. 32873: A lateral view of head; B dorsal view of head; C heels with conical tubercles, D ventral view of hand, and E) ventral view of foot." figureDoi="10.3897/vz.73.e91111.figure2" httpUri="https://binary.pensoft.net/fig/800714" pageId="0" pageNumber="127">2E</figureCitation>
|
||
), disc on Toe 4 slightly narrower than on Finger 3 (3FD/4TD 1.14). Webbing between toes reaches to proximal edge of subarticular tubercle at base of penultimate phalanges on both sides of Toe 4, nearly to base of disc on all other toes except Toe 1 where it reaches only to subarticular tubercle at base of penultimate phalanx; relative lengths of toes 4>5>3>2>1. Inner metatarsal tubercle oval, small (1.3 mm long) but prominent; outer metatarsal tubercle absent (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Details of holotype of Litoria hastula sp. nov. MZB Amph. 32873: A lateral view of head; B dorsal view of head; C heels with conical tubercles, D ventral view of hand, and E) ventral view of foot." figureDoi="10.3897/vz.73.e91111.figure2" httpUri="https://binary.pensoft.net/fig/800714" pageId="0" pageNumber="127">2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="colouration in preservative">
|
||
<paragraph pageId="0" pageNumber="127">Colouration in preservative.</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
Dorsal base colouration mottled varying shades of light brown, patterned with scattered darker-brown maculations across snout, back and limbs, with larger indistinct darker-brown blotches on hindblimbs and posterior portion of torso (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Type series of Litoria hastula sp. nov. showing distinctly acuminate snout tip and marked sexual size dimorphism: A dorsal view and B ventral view. Smaller males from top left to bottom right are: MZB Amph. 32873 (holotype); SAMA R 72334, SAMA R 72335, MZB Amph. 32874. Much larger female on far right is MZB Amph. 32872. Tags on the specimens are field numbers. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e91111.figure1" httpUri="https://binary.pensoft.net/fig/800713" pageId="0" pageNumber="127">1A</figureCitation>
|
||
). Dark colouration of eyeball visible through dorsal skin of orbital. Ventral surfaces buff, largely unpatterned except for dense light-brown maculations on outer edges of lower forelimbs and lower hindlimbs (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Type series of Litoria hastula sp. nov. showing distinctly acuminate snout tip and marked sexual size dimorphism: A dorsal view and B ventral view. Smaller males from top left to bottom right are: MZB Amph. 32873 (holotype); SAMA R 72334, SAMA R 72335, MZB Amph. 32874. Much larger female on far right is MZB Amph. 32872. Tags on the specimens are field numbers. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e91111.figure1" httpUri="https://binary.pensoft.net/fig/800713" pageId="0" pageNumber="127">1B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="variation">
|
||
<paragraph pageId="0" pageNumber="127">Variation in the type series.</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
Measurements and proportions of males in the type series show relatively limited variation (Table
|
||
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Summary of measurement data for the type series of Litoria hastula sp. nov." httpUri="http://table.plazi.org/id/B1520D4F3E781CD1D3B4DA42704EAAA3" pageId="0" pageNumber="127" tableUuid="B1520D4F3E781CD1D3B4DA42704EAAA3">1</tableCitation>
|
||
), and all have similar extents of webbing on the hands and feet and conspicuous conical tubercles above the eyes and on the heel. Dorsal base colouration varies from light to medium brown, and the amount and extent of darker-brown maculations and blotching varies from a few blotches on the hindlimbs, to darker patches forming broad bands across the torso and forelimbs (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Type series of Litoria hastula sp. nov. showing distinctly acuminate snout tip and marked sexual size dimorphism: A dorsal view and B ventral view. Smaller males from top left to bottom right are: MZB Amph. 32873 (holotype); SAMA R 72334, SAMA R 72335, MZB Amph. 32874. Much larger female on far right is MZB Amph. 32872. Tags on the specimens are field numbers. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e91111.figure1" httpUri="https://binary.pensoft.net/fig/800713" pageId="0" pageNumber="127">1A</figureCitation>
|
||
). Base colouration of venter is always buff, generally without pigmentation (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Type series of Litoria hastula sp. nov. showing distinctly acuminate snout tip and marked sexual size dimorphism: A dorsal view and B ventral view. Smaller males from top left to bottom right are: MZB Amph. 32873 (holotype); SAMA R 72334, SAMA R 72335, MZB Amph. 32874. Much larger female on far right is MZB Amph. 32872. Tags on the specimens are field numbers. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e91111.figure1" httpUri="https://binary.pensoft.net/fig/800713" pageId="0" pageNumber="127">1B</figureCitation>
|
||
). When compared against the holotype, paratype SAMA R72335 has tubercles below vent larger and more elongate, lateral margins of tongue less emarginate, and vomerine teeth that are somewhat less conspicuous. Paratype SAMA R72334 has lateral margins of tongue that are smooth and a narrower and straighter supratympanic fold. Paratype MZB Amph.32874 is noticeably darker brown on the dorsum, and the pale tubercles along outer edge of forearm are more conspicuous.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="B1520D4F3E781CD1D3B4DA42704EAAA3" httpUri="http://table.plazi.org/id/B1520D4F3E781CD1D3B4DA42704EAAA3" pageId="0" pageNumber="127" start="Table 1" startId="T1">
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">Table 1.</emphasis>
|
||
Summary of measurement data for the type series of
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<table pageId="0" pageNumber="127">
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="127" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">MZB</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">MZB</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">SAMA</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">SAMA</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">MZB</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="127" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">Amph.32873</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">Amph.32874</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">R72334</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">R72335</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">Amph.32872</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="127" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">holotype</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">paratype</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">paratype</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">paratype</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">paratype</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">Sex</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">m</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">m</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">m</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">m</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">f</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">31.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">30.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">31.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">31.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">48.9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">TL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">18.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">16.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">17.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">19</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">29.2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">HW</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">15</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">HL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">11.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">11.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">14.8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">EYE</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">TYM</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">EN</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">IN</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">5.1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">4TD</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">4TP</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3FD</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.9</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3FP</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">TL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.6</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">HW/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.31</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">HL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.3</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">HL/HW</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.09</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.03</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.99</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">EN/IN</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.73</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.71</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.71</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.69</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">EYE/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">TYM/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.04</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">TYM/EYE</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">4TD/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.07</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3FD/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">4TD/4TP</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.45</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.55</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.32</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3FD/3FP</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.85</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.67</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.61</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3FD/4TD</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.18</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.16</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
The single female paratype, MZB Amph.32872, is considerably longer than the males, more robust in body form and has a less acute snout tip (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Type series of Litoria hastula sp. nov. showing distinctly acuminate snout tip and marked sexual size dimorphism: A dorsal view and B ventral view. Smaller males from top left to bottom right are: MZB Amph. 32873 (holotype); SAMA R 72334, SAMA R 72335, MZB Amph. 32874. Much larger female on far right is MZB Amph. 32872. Tags on the specimens are field numbers. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e91111.figure1" httpUri="https://binary.pensoft.net/fig/800713" pageId="0" pageNumber="127">1A</figureCitation>
|
||
) but does not show obvious differences in ratios or proportions from the males (Table
|
||
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Summary of measurement data for the type series of Litoria hastula sp. nov." httpUri="http://table.plazi.org/id/B1520D4F3E781CD1D3B4DA42704EAAA3" pageId="0" pageNumber="127" tableUuid="B1520D4F3E781CD1D3B4DA42704EAAA3">1</tableCitation>
|
||
). Compared to males, the ventral surfaces of the female paratype show more extensive dark-brown blotches and maculations across the limbs, torso and throat (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Type series of Litoria hastula sp. nov. showing distinctly acuminate snout tip and marked sexual size dimorphism: A dorsal view and B ventral view. Smaller males from top left to bottom right are: MZB Amph. 32873 (holotype); SAMA R 72334, SAMA R 72335, MZB Amph. 32874. Much larger female on far right is MZB Amph. 32872. Tags on the specimens are field numbers. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e91111.figure1" httpUri="https://binary.pensoft.net/fig/800713" pageId="0" pageNumber="127">1B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="colouration in life">
|
||
<paragraph pageId="0" pageNumber="127">Colouration in life.</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
The following description of colour in life is based on images of three different paratypes (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Colouration of Litoria hastula sp. nov. in life: A lateral view of adult male paratype SAMA R 72334; B hidden coloration in thighs of male paratype SAMA R 72335; C lateral view of adult female paratype MZB Amph. 32872; D ventral view of adult female paratype MZB Amph. 32872. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure3" httpUri="https://binary.pensoft.net/fig/800715" pageId="0" pageNumber="127">3</figureCitation>
|
||
). Dorsal and lateral surfaces of male paratype SAMA R72334 light brown, with variable amounts of medium-brown mottling and small dark-brown spots (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Colouration of Litoria hastula sp. nov. in life: A lateral view of adult male paratype SAMA R 72334; B hidden coloration in thighs of male paratype SAMA R 72335; C lateral view of adult female paratype MZB Amph. 32872; D ventral view of adult female paratype MZB Amph. 32872. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure3" httpUri="https://binary.pensoft.net/fig/800715" pageId="0" pageNumber="127">3A</figureCitation>
|
||
). Female paratype with dorsal pattern comprising extensive areas of dark-green pigmentation across head, torso and limbs (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Colouration of Litoria hastula sp. nov. in life: A lateral view of adult male paratype SAMA R 72334; B hidden coloration in thighs of male paratype SAMA R 72335; C lateral view of adult female paratype MZB Amph. 32872; D ventral view of adult female paratype MZB Amph. 32872. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure3" httpUri="https://binary.pensoft.net/fig/800715" pageId="0" pageNumber="127">3C</figureCitation>
|
||
) and ventral colouration buff with numerous dark-brown maculations that in some areas coalesce to form indistinct spots (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Colouration of Litoria hastula sp. nov. in life: A lateral view of adult male paratype SAMA R 72334; B hidden coloration in thighs of male paratype SAMA R 72335; C lateral view of adult female paratype MZB Amph. 32872; D ventral view of adult female paratype MZB Amph. 32872. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure3" httpUri="https://binary.pensoft.net/fig/800715" pageId="0" pageNumber="127">3D</figureCitation>
|
||
). Hidden surfaces of thighs of SAMA R72335 dark orange with extensive dark-brown spots and flecks (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Colouration of Litoria hastula sp. nov. in life: A lateral view of adult male paratype SAMA R 72334; B hidden coloration in thighs of male paratype SAMA R 72335; C lateral view of adult female paratype MZB Amph. 32872; D ventral view of adult female paratype MZB Amph. 32872. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure3" httpUri="https://binary.pensoft.net/fig/800715" pageId="0" pageNumber="127">3B</figureCitation>
|
||
). Iris predominately light brown, with extensive thin darker-brown reticulations.
|
||
</paragraph>
|
||
<caption doi="10.3897/vz.73.e91111.figure3" httpUri="https://binary.pensoft.net/fig/800715" pageId="0" pageNumber="127" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">Figure 3.</emphasis>
|
||
Colouration of
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
in life:
|
||
<emphasis bold="true" pageId="0" pageNumber="127">A</emphasis>
|
||
lateral view of adult male paratype SAMA R72334;
|
||
<emphasis bold="true" pageId="0" pageNumber="127">B</emphasis>
|
||
hidden coloration in thighs of male paratype SAMA R72335;
|
||
<emphasis bold="true" pageId="0" pageNumber="127">C</emphasis>
|
||
lateral view of adult female paratype MZB Amph.32872;
|
||
<emphasis bold="true" pageId="0" pageNumber="127">D</emphasis>
|
||
ventral view of adult female paratype MZB Amph.32872. All photographs by S.J. Richards.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="vocalisation">
|
||
<paragraph pageId="0" pageNumber="127">Vocalisation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
We analysed two call series produced by SAMA R72335 at an air temperature of 16.4° C. Calls were recorded against a background of loud rushing water. The first series contained seven calls and lasted six seconds, and the second series, separated from the first by 23 seconds, lasted 3.4 seconds and contained five calls (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Two complete advertisement call series produced by paratype of Litoria hastula sp. nov. SAMA R 72334, top = waveform, bottom = spectrogram. Time between the two call series has been shortened to allow comparisons." figureDoi="10.3897/vz.73.e91111.figure4" httpUri="https://binary.pensoft.net/fig/800716" pageId="0" pageNumber="127">4</figureCitation>
|
||
). Inter-call intervals for both series were relatively uniform (0.58-1.03 s, mean 0.78, SD 0.16, n = 10). Dominant frequency was 2530-2730 Hz (mean 2650, SD 67.4, n = 12). Length of individual calls in Series 1 increased substantially from 0.06 s in the first call to 0.35 s in the last call, and calls 5-7 in this series were characterized by increasingly pulsatile structure and subdivision into groups of variably well-defined pulses (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Two complete advertisement call series produced by paratype of Litoria hastula sp. nov. SAMA R 72334, top = waveform, bottom = spectrogram. Time between the two call series has been shortened to allow comparisons." figureDoi="10.3897/vz.73.e91111.figure4" httpUri="https://binary.pensoft.net/fig/800716" pageId="0" pageNumber="127">4</figureCitation>
|
||
). In contrast, call length in Series 2 increased only slightly during the series, from 0.071 s in Call 1 to 0.091 s in Call 5. Excluding calls 5-7 in Series 1, call length for all calls combined was 0.06-0.09 s (mean 0.07, SD 0.01, n = 9).
|
||
</paragraph>
|
||
<caption doi="10.3897/vz.73.e91111.figure4" httpUri="https://binary.pensoft.net/fig/800716" pageId="0" pageNumber="127" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">Figure 4.</emphasis>
|
||
Two complete advertisement call series produced by paratype of
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
SAMA R72334, top = waveform, bottom = spectrogram. Time between the two call series has been shortened to allow comparisons.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="etymology">
|
||
<paragraph pageId="0" pageNumber="127">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
Latin, "little
|
||
<normalizedToken originalValue="spear’">spear'</normalizedToken>
|
||
, combining spear "
|
||
<emphasis italics="true" pageId="0" pageNumber="127">hasta</emphasis>
|
||
" with the diminutive suffix "-
|
||
<emphasis italics="true" pageId="0" pageNumber="127">ula</emphasis>
|
||
" in reference to the elongate and sharply pointed snout of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="distribution">
|
||
<paragraph pageId="0" pageNumber="127">Distribution and ecological notes.</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
is known only from the type locality in the mountains of Papua Province, Indonesia (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Known distribution of Litoria hastula sp. nov. in Indonesia (red star) and L. dorsivena in western Papua New Guinea (blue circles)." figureDoi="10.3897/vz.73.e91111.figure5" httpUri="https://binary.pensoft.net/fig/800717" pageId="0" pageNumber="127">5</figureCitation>
|
||
). The habitat at this locality consisted of very mossy wet mid-montane rainforest at about 2000 m a.s.l (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Details of habitat at the type locality for Litoria hastula sp. nov.: A dense and very mossy mid-montane forest, and B small fast-flowing rocky stream. The type series was collected from low vegetation along this stream at night. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure6" httpUri="https://binary.pensoft.net/fig/800718" pageId="0" pageNumber="127">6A</figureCitation>
|
||
). The type series was collected on vegetation along a fast-flowing stream in very rugged and steep terrain (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Details of habitat at the type locality for Litoria hastula sp. nov.: A dense and very mossy mid-montane forest, and B small fast-flowing rocky stream. The type series was collected from low vegetation along this stream at night. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure6" httpUri="https://binary.pensoft.net/fig/800718" pageId="0" pageNumber="127">6B</figureCitation>
|
||
) where males called from leaves at night. The single female paratype is gravid and contains numerous large (2.4-2.6 mm) yellow eggs. Nine species of frog were collected at the type locality, including three other species of torrent-breeding pelodryadid treefrogs (
|
||
<taxonomicName baseAuthorityName="Boulenger" baseAuthorityYear="1915" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria angiana" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="angiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria angiana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. fuscula" pageId="0" pageNumber="127" rank="species" species="fuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. fuscula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="L. pratti" pageId="0" pageNumber="127" rank="species" species="pratti">L. cf. pratti</taxonomicName>
|
||
), four species of microhylid frogs in the genus
|
||
<taxonomicName authorityName="Boettger" authorityYear="1895" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" lsidName="Oreophryne" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Oreophryne</emphasis>
|
||
</taxonomicName>
|
||
and the limnodynastid frog
|
||
<taxonomicName authorityName="Parker" authorityYear="1940" class="Amphibia" family="Limnodynastidae" genus="Lechriodus" kingdom="Animalia" lsidName="Lechriodus platyceps" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="platyceps">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Lechriodus platyceps</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Richards, SJ" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="127" refId="B18" refString="Richards, SJ, Iskandar, DT, Allison, A, 2000. Amphibian and reptiles recorded on the RAP survey in the Wapoga area, Irian Jaya, Indonesia. In: Mack AL, Alonso L (Eds) A Biological Assessment of the Wapoga River area of northwestern Irian Jaya, Indonesia. Conservation International, Arlington, pp. 113-115." title="Amphibian and reptiles recorded on the RAP survey in the Wapoga area, Irian Jaya, Indonesia. In: Mack AL, Alonso L (Eds) A Biological Assessment of the Wapoga River area of northwestern Irian Jaya, Indonesia. Conservation International, Arlington, pp. 113 - 115." year="2000">Richards et. al. 2000</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/vz.73.e91111.figure5" httpUri="https://binary.pensoft.net/fig/800717" pageId="0" pageNumber="127" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">Figure 5.</emphasis>
|
||
Known distribution of
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
in Indonesia (red star) and
|
||
<taxonomicName lsidName="L. dorsivena" pageId="0" pageNumber="127" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. dorsivena</emphasis>
|
||
</taxonomicName>
|
||
in western Papua New Guinea (blue circles).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/vz.73.e91111.figure6" httpUri="https://binary.pensoft.net/fig/800718" pageId="0" pageNumber="127" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">Figure 6.</emphasis>
|
||
Details of habitat at the type locality for
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
:
|
||
<emphasis bold="true" pageId="0" pageNumber="127">A</emphasis>
|
||
dense and very mossy mid-montane forest, and
|
||
<emphasis bold="true" pageId="0" pageNumber="127">B</emphasis>
|
||
small fast-flowing rocky stream. The type series was collected from low vegetation along this stream at night. All photographs by S.J. Richards.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="suggested iucn status">
|
||
<paragraph pageId="0" pageNumber="127">Suggested IUCN status.</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
is currently only known from a single locality. However, large areas of suitable habitat at similar elevations remain in nearby areas. This species also occurs at elevations around which
|
||
<taxonomicName class="Rhizophydiomycetes" genus="Batrachochytrium" kingdom="Fungi" lsidName="Batrachochytrium dendrobatidis" order="Rhizophydiales" pageId="0" pageNumber="127" phylum="Chytridiomycota" rank="species" species="dendrobatidis">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Batrachochytrium dendrobatidis</emphasis>
|
||
</taxonomicName>
|
||
has devastated communities of ecologically similar
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria</emphasis>
|
||
</taxonomicName>
|
||
in Australia, and is predicted to be very sensitive to this frog pathogen. Given the
|
||
<normalizedToken originalValue="species’">species'</normalizedToken>
|
||
poorly known distribution and that threats are poorly understood, we recommend that this species be considered as Data Deficient at this stage.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="comparisons">
|
||
<paragraph pageId="0" pageNumber="127">Comparison with other species.</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
The combination of predominantly brown dorsal colouration, canthus rostralis sharply defined and straight, snout tip pointed in dorsal view, moderate webbing on fingers, and moderate size (adult male SVL 30-35 mm) readily distinguishes
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
from all other non-torrent-breeding
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria</emphasis>
|
||
</taxonomicName>
|
||
in New Guinea. It differs from all species we currently ascribe to the genus
|
||
<taxonomicName authorityName="Stejneger" authorityYear="1916" class="Amphibia" family="Pelodryadidae" genus="Nyctimystes" kingdom="Animalia" lsidName="Nyctimystes" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Nyctimystes</emphasis>
|
||
</taxonomicName>
|
||
(most of which are also torrent-breeders) in these same characters, in having a horizontal (versus vertical) pupil, and in lacking palpebral venation.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
In comparison to other torrent-breeding
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria</emphasis>
|
||
</taxonomicName>
|
||
from New Guinea,
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
is smaller than
|
||
<taxonomicName lsidName="L. angiana" pageId="0" pageNumber="127" rank="species" species="angiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. angiana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. arfakiana" pageId="0" pageNumber="127" rank="species" species="arfakiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. arfakiana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. becki" pageId="0" pageNumber="127" rank="species" species="becki">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. becki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. macki" pageId="0" pageNumber="127" rank="species" species="macki">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. macki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. oenicolen" pageId="0" pageNumber="127" rank="species" species="oenicolen">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. oenicolen</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. spartacus" pageId="0" pageNumber="127" rank="species" species="spartacus">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. spartacus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. spinifera" pageId="0" pageNumber="127" rank="species" species="spinifera">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. spinifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="L. wollastoni" pageId="0" pageNumber="127" rank="species" species="wollastoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. wollastoni</emphasis>
|
||
</taxonomicName>
|
||
(max male SVL <35mm versus>35mm); and further differs from
|
||
<taxonomicName lsidName="L. arfakiana" pageId="0" pageNumber="127" rank="species" species="arfakiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. arfakiana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. becki" pageId="0" pageNumber="127" rank="species" species="becki">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. becki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. macki" pageId="0" pageNumber="127" rank="species" species="macki">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. macki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. oenicolen" pageId="0" pageNumber="127" rank="species" species="oenicolen">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. oenicolen</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. spinifera" pageId="0" pageNumber="127" rank="species" species="spinifera">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. spinifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="L. wollastoni" pageId="0" pageNumber="127" rank="species" species="wollastoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. wollastoni</emphasis>
|
||
</taxonomicName>
|
||
in having moderately extensive webbing between fingers 2 to 4 (versus absent, or at most a thin basal strip between fingers 3 and 4); from
|
||
<taxonomicName lsidName="L. angiana" pageId="0" pageNumber="127" rank="species" species="angiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. angiana</emphasis>
|
||
</taxonomicName>
|
||
in having a snout that is sharply pointed in dorsal and lateral profile (versus slightly pointed in dorsal profile only); from
|
||
<taxonomicName lsidName="L. macki" pageId="0" pageNumber="127" rank="species" species="macki">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. macki</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. spartacus" pageId="0" pageNumber="127" rank="species" species="spartacus">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. spartacus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="L. spinifera" pageId="0" pageNumber="127" rank="species" species="spinifera">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. spinifera</emphasis>
|
||
</taxonomicName>
|
||
in having a predominately brown dorsum (versus mottled green and brown) and in lacking prominent tubercles along the legs and much of the body (versus spiniform and/or conical tubercles prominent and widespread on body and/or limbs).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
is larger than
|
||
<taxonomicName lsidName="L. amnicola" pageId="0" pageNumber="127" rank="species" species="amnicola">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. amnicola</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. brongersmai" pageId="0" pageNumber="127" rank="species" species="brongersmai">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. brongersmai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. megalops" pageId="0" pageNumber="127" rank="species" species="megalops">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. megalops</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. napaea" pageId="0" pageNumber="127" rank="species" species="napaea">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. napaea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. rara" pageId="0" pageNumber="127" rank="species" species="rara">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. rara</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="L. rivicola" pageId="0" pageNumber="127" rank="species" species="rivicola">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. rivicola</emphasis>
|
||
</taxonomicName>
|
||
(male SVL>30 mm versus <25 mm); and further differs from
|
||
<taxonomicName lsidName="L. amnicola" pageId="0" pageNumber="127" rank="species" species="amnicola">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. amnicola</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. brongersmai" pageId="0" pageNumber="127" rank="species" species="brongersmai">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. brongersmai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="L. napaea" pageId="0" pageNumber="127" rank="species" species="napaea">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. napaea</emphasis>
|
||
</taxonomicName>
|
||
in having prominent vomerine teeth (versus indistinct and detectable only as slight bumps) and prominent heel tubercles (versus absent); from
|
||
<taxonomicName lsidName="L. megalops" pageId="0" pageNumber="127" rank="species" species="megalops">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. megalops</emphasis>
|
||
</taxonomicName>
|
||
in having webbing on the hand (versus absent), canthus rostralis relatively straight in dorsal profile (versus curved), and in its smaller eye (EYE/SVL 0.13 versus 0.14-0.17); from
|
||
<taxonomicName lsidName="L. rivicola" pageId="0" pageNumber="127" rank="species" species="rivicola">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. rivicola</emphasis>
|
||
</taxonomicName>
|
||
by it smooth dorsal skin (versus strongly tuberculate); and from
|
||
<taxonomicName lsidName="L. rara" pageId="0" pageNumber="127" rank="species" species="rara">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. rara</emphasis>
|
||
</taxonomicName>
|
||
by its sharply pointed snout in dorsal and lateral views (versus rounded).
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
is slightly larger than
|
||
<taxonomicName lsidName="L. scabra" pageId="0" pageNumber="127" rank="species" species="scabra">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. scabra</emphasis>
|
||
</taxonomicName>
|
||
(male SVL 30.4-31.8 versus 23.6-27.2.1, female SVL 48.9 versus 27.2-30.6 mm), and further differs in having a relatively smooth dorsum (versus covered in distinct tubercles), relatively straight canthus rostralis (versus curved) and pointed snout tip in dorsal and lateral views (versus rounded and truncate).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
differs from five similar-sized (max male SVL between 30-40 mm) torrent-breeding taxa (
|
||
<taxonomicName lsidName="L. bulmeri" pageId="0" pageNumber="127" rank="species" species="bulmeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. bulmeri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. fuscula" pageId="0" pageNumber="127" rank="species" species="fuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. fuscula</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. micromembrana" pageId="0" pageNumber="127" rank="species" species="micromembrana">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. micromembrana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="L. modica" pageId="0" pageNumber="127" rank="species" species="modica">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. modica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="L. pratti" pageId="0" pageNumber="127" rank="species" species="pratti">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. pratti</emphasis>
|
||
</taxonomicName>
|
||
) in having moderately extensive finger webbing that extends to the third phalanx on fingers 2, 3 and 4 (versus at most basal webbing between fingers 3 and 4) and in having a sharply pointed snout and relatively straight canthus rostralis (versus typically rounded snout and curved canthus rostralis). It further differs from the sympatric
|
||
<taxonomicName lsidName="L. fuscula" pageId="0" pageNumber="127" rank="species" species="fuscula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. fuscula</emphasis>
|
||
</taxonomicName>
|
||
in having prominent vomerine teeth (versus indistinct and detectable only as slight bumps), dorsum light brown with dark-brown spots and blotches (versus dark brown with no obvious pattern), and in having small conical tubercles on the heel (versus no tubercles).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
is most similar to
|
||
<taxonomicName baseAuthorityName="Tyler" baseAuthorityYear="1968" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria dorsivena" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria dorsivena</emphasis>
|
||
</taxonomicName>
|
||
, a species known from about 700 km to the east in the Telefomin and upper Strickland River areas of Papua New Guinea (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Known distribution of Litoria hastula sp. nov. in Indonesia (red star) and L. dorsivena in western Papua New Guinea (blue circles)." figureDoi="10.3897/vz.73.e91111.figure5" httpUri="https://binary.pensoft.net/fig/800717" pageId="0" pageNumber="127">5</figureCitation>
|
||
) but differs from that taxon in its slightly larger size (male SVL 30.4-31.8 versus 27.2-29.1, female SVL 48.9 versus 41.0-48 mm); proportionately longer snout (EN/IN ratio 0.71-0.80 versus 0.57-0.65); more angular canthus rostralis and more sharply pointed snout (Fig.
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Comparison of head profile of a male paratype of Litoria dorsivena SAMA R 7907 (left) and holotype of Litoria hastula MZB Amph. 32873 (right). Note much more sharply pointed tip of snout on Litoria hastula sp. nov." figureDoi="10.3897/vz.73.e91111.figure7" httpUri="https://binary.pensoft.net/fig/800719" pageId="0" pageNumber="127">7</figureCitation>
|
||
); dorsal surface of snout concave (versus not obviously concave); dorsum with only sparse tubercles (versus tubercles scattered across the dorsum); and in having at least two moderately prominent conical tubercules on the heel (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Details of holotype of Litoria hastula sp. nov. MZB Amph. 32873: A lateral view of head; B dorsal view of head; C heels with conical tubercles, D ventral view of hand, and E) ventral view of foot." figureDoi="10.3897/vz.73.e91111.figure2" httpUri="https://binary.pensoft.net/fig/800714" pageId="0" pageNumber="127">2C</figureCitation>
|
||
) (versus one).
|
||
</paragraph>
|
||
<caption doi="10.3897/vz.73.e91111.figure7" httpUri="https://binary.pensoft.net/fig/800719" pageId="0" pageNumber="127" start="Figure 7" startId="F7">
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">Figure 7.</emphasis>
|
||
Comparison of head profile of a male paratype of
|
||
<taxonomicName baseAuthorityName="Tyler" baseAuthorityYear="1968" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria dorsivena" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria dorsivena</emphasis>
|
||
</taxonomicName>
|
||
SAMA R7907 (left) and holotype of
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
MZB Amph.32873 (right). Note much more sharply pointed tip of snout on
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria hastula" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="127" type="distribution">
|
||
<paragraph pageId="0" pageNumber="127">
|
||
Notes on the distribution, ecology and colour in life of
|
||
<taxonomicName baseAuthorityName="Tyler" baseAuthorityYear="1968" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria dorsivena" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria dorsivena</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
Material of
|
||
<taxonomicName baseAuthorityName="Tyler" baseAuthorityYear="1968" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria dorsivena" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria dorsivena</emphasis>
|
||
</taxonomicName>
|
||
collected from localities in the Muller Range (
|
||
<geoCoordinate degrees="5.657" direction="south" orientation="latitude" precision="55" value="-5.657">5.657°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="142.305" direction="east" orientation="longitude" precision="55" value="142.305">142.305°E</geoCoordinate>
|
||
,~1600 m a.s.l.) and Upper Strickland River region (
|
||
<geoCoordinate degrees="5.288" direction="south" orientation="latitude" precision="55" value="-5.288">5.288°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="142.494" direction="east" orientation="longitude" precision="55" value="142.494">142.494°E</geoCoordinate>
|
||
, ~1100 m a.s.l.) (
|
||
<bibRefCitation author="Richards, SJ" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="127" refId="B16" refString="Richards, SJ, Dahl, C, 2011. Herpetofauna of the Strickland Basin and Muller Range, Papua New Guinea. In: Richards SJ, Gamui BG (Eds), Rapid Biological Assessments of the Nakanai Mountains and the upper Strickland Basin: Surveying the biodiversity of Papua New Guinea's sublime karst environments. Conservation International, Arlington, pp. 190-202." title="Herpetofauna of the Strickland Basin and Muller Range, Papua New Guinea. In: Richards SJ, Gamui BG (Eds), Rapid Biological Assessments of the Nakanai Mountains and the upper Strickland Basin: Surveying the biodiversity of Papua New Guinea's sublime karst environments. Conservation International, Arlington, pp. 190 - 202." year="2011">Richards and Dahl 2011</bibRefCitation>
|
||
) (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Known distribution of Litoria hastula sp. nov. in Indonesia (red star) and L. dorsivena in western Papua New Guinea (blue circles)." figureDoi="10.3897/vz.73.e91111.figure5" httpUri="https://binary.pensoft.net/fig/800717" pageId="0" pageNumber="127">5</figureCitation>
|
||
) extends the range of this species approximately 80 km east of the type locality at Telefomin. The specimen from the Muller Range (SAMA R65133) is a subadult female. In contrast the collections from the Upper Strickland region comprise four adult males, allowing us to present new information on the colouration and variation in adult males of this species. In life (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Colour variation and habitats details for male Litoria dorsivena from Tualapa in the upper Strickland River basin, Hela Province, Papua New Guinea: A SAMA R 72320; B SAMA R 72321; C SAMA R 72321 ventral surfaces; D preserved samples (SAMA R 72321 - 4) illustrating variation in dorsal pattern; and E swift-flowing stream habitat in the upper Strickland River region. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure8" httpUri="https://binary.pensoft.net/fig/800720" pageId="0" pageNumber="127">8A-C</figureCitation>
|
||
) and preservative (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Colour variation and habitats details for male Litoria dorsivena from Tualapa in the upper Strickland River basin, Hela Province, Papua New Guinea: A SAMA R 72320; B SAMA R 72321; C SAMA R 72321 ventral surfaces; D preserved samples (SAMA R 72321 - 4) illustrating variation in dorsal pattern; and E swift-flowing stream habitat in the upper Strickland River region. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure8" httpUri="https://binary.pensoft.net/fig/800720" pageId="0" pageNumber="127">8D</figureCitation>
|
||
) these frogs are predominately light brown on the dorsum, with dark-brown spotting and blotching of greatly varying extent. The canthus rostralis is highlighted by either a sharp boundary between dark-brown lateral colouration on the snout and paler-brown dorsal colouration (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Colour variation and habitats details for male Litoria dorsivena from Tualapa in the upper Strickland River basin, Hela Province, Papua New Guinea: A SAMA R 72320; B SAMA R 72321; C SAMA R 72321 ventral surfaces; D preserved samples (SAMA R 72321 - 4) illustrating variation in dorsal pattern; and E swift-flowing stream habitat in the upper Strickland River region. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure8" httpUri="https://binary.pensoft.net/fig/800720" pageId="0" pageNumber="127">8A</figureCitation>
|
||
) or by a thin yellowish-brown canthal stripe (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Colour variation and habitats details for male Litoria dorsivena from Tualapa in the upper Strickland River basin, Hela Province, Papua New Guinea: A SAMA R 72320; B SAMA R 72321; C SAMA R 72321 ventral surfaces; D preserved samples (SAMA R 72321 - 4) illustrating variation in dorsal pattern; and E swift-flowing stream habitat in the upper Strickland River region. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure8" httpUri="https://binary.pensoft.net/fig/800720" pageId="0" pageNumber="127">8B</figureCitation>
|
||
). The venter and hidden surfaces of limbs have a buff base colouration, overlain with light orange on the posterior portion of torso and most of the hindlimbs and groin, and sparse to dense patches of light-brown maculations (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Colour variation and habitats details for male Litoria dorsivena from Tualapa in the upper Strickland River basin, Hela Province, Papua New Guinea: A SAMA R 72320; B SAMA R 72321; C SAMA R 72321 ventral surfaces; D preserved samples (SAMA R 72321 - 4) illustrating variation in dorsal pattern; and E swift-flowing stream habitat in the upper Strickland River region. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure8" httpUri="https://binary.pensoft.net/fig/800720" pageId="0" pageNumber="127">8C</figureCitation>
|
||
). Iris is predominately silvery grey, with light-brown vermiculations and dark-brown blotching across the ventral half.
|
||
</paragraph>
|
||
<caption doi="10.3897/vz.73.e91111.figure8" httpUri="https://binary.pensoft.net/fig/800720" pageId="0" pageNumber="127" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">Figure 8.</emphasis>
|
||
Colour variation and habitats details for male
|
||
<taxonomicName baseAuthorityName="Tyler" baseAuthorityYear="1968" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria dorsivena" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria dorsivena</emphasis>
|
||
</taxonomicName>
|
||
from Tualapa in the upper Strickland River basin, Hela Province, Papua New Guinea:
|
||
<emphasis bold="true" pageId="0" pageNumber="127">A</emphasis>
|
||
SAMA R72320;
|
||
<emphasis bold="true" pageId="0" pageNumber="127">B</emphasis>
|
||
SAMA R72321;
|
||
<emphasis bold="true" pageId="0" pageNumber="127">C</emphasis>
|
||
SAMA R72321 ventral surfaces;
|
||
<emphasis bold="true" pageId="0" pageNumber="127">D</emphasis>
|
||
preserved samples (SAMA R72321-4) illustrating variation in dorsal pattern; and
|
||
<emphasis bold="true" pageId="0" pageNumber="127">E</emphasis>
|
||
swift-flowing stream habitat in the upper Strickland River region. All photographs by S.J. Richards.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
Summary measurements of the new adult male
|
||
<taxonomicName baseAuthorityName="Tyler" baseAuthorityYear="1968" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria dorsivena" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria dorsivena</emphasis>
|
||
</taxonomicName>
|
||
specimens are presented in Table
|
||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Summary of measurement data for adult Litoria dorsivena, including male specimens from the type series (SAMA R 7901, R 7907 - 11), recently collected males from the upper Strickland River region (SAMA R 72319 - 22) and the female holotype (SAMA R 7901)." httpUri="http://table.plazi.org/id/B5A44361F93CB5D830BE6C181E08F275" pageId="0" pageNumber="127" tableUuid="B5A44361F93CB5D830BE6C181E08F275">2</tableCitation>
|
||
along with newly obtained measurements of the female holotype (SAMA R7901) and five male paratypes (SAMA R7907-7911). Measurements of the newly documented specimens agree closely with those taken from the type series and confirm that males of this species are consistently smaller than males of
|
||
<taxonomicName lsidName="L. hastula" pageId="0" pageNumber="127" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
(see above and Table
|
||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Summary of measurement data for adult Litoria dorsivena, including male specimens from the type series (SAMA R 7901, R 7907 - 11), recently collected males from the upper Strickland River region (SAMA R 72319 - 22) and the female holotype (SAMA R 7901)." httpUri="http://table.plazi.org/id/B5A44361F93CB5D830BE6C181E08F275" pageId="0" pageNumber="127" tableUuid="B5A44361F93CB5D830BE6C181E08F275">2</tableCitation>
|
||
). The specimen from the Muller Range (SAMA R65133) is a subadult female with an SVL of 32.9 mm).
|
||
</paragraph>
|
||
<caption ID-Table-UUID="B5A44361F93CB5D830BE6C181E08F275" httpUri="http://table.plazi.org/id/B5A44361F93CB5D830BE6C181E08F275" pageId="0" pageNumber="127" start="Table 2" startId="T2">
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">Table 2.</emphasis>
|
||
Summary of measurement data for adult
|
||
<taxonomicName baseAuthorityName="Tyler" baseAuthorityYear="1968" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria dorsivena" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria dorsivena</emphasis>
|
||
</taxonomicName>
|
||
, including male specimens from the type series (SAMA R7901, R7907-11), recently collected males from the upper Strickland River region (SAMA R72319-22) and the female holotype (SAMA R7901).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<table pageId="0" pageNumber="127">
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="127" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">R7901</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">R7907</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">R7908</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">R7909</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">R7910</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">R7911</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">R72319</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">R72320</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">R72321</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">R72322</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="127" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">holotype</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">paratype</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">paratype</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">paratype</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">paratype</emphasis>
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="127">paratype</emphasis>
|
||
</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="127" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="127" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="127" rowspan="1">-</td>
|
||
<td colspan="1" isEmpty="true" pageId="0" pageNumber="127" rowspan="1">-</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">Sex</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">F</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">M</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">M</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">M</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">M</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">M</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">M</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">M</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">M</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">M</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">44.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">27</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">27.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">27.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">28</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">28</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">27.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">27.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">29.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">28</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">TL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">27.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">16.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">17.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">16.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">17</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">17.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">16.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">16.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">16.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">17.6</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">HW</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">14.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">9.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">9.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">9.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">9.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">9.8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">HL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">14.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">9.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">9.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">9.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">11</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">10.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">EYE</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">4.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">TYM</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">EN</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.3</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">IN</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3.6</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">4TD</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">4TP</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3FD</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3FP</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.8</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">TL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.63</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.59</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.62</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.58</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.63</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">HW/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.32</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.39</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.37</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.35</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">HL/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.33</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.34</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.36</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.37</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.36</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">HL/HW</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.03</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.97</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.02</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.03</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.01</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.03</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.06</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">EN/IN</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.57</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.61</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.65</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.63</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.64</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.65</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.64</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">EYE/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.11</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.12</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.13</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.13</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">TYM/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.04</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">TYM/EYE</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.38</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.39</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.42</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.35</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.43</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.31</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.4</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">4TD/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3FD/SVL</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.07</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.05</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">0.06</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">4TD/4TP</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.41</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.3</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.4</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.44</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.36</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3FD/3FP</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">2</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.9</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.89</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.72</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.6</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.5</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.56</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.88</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.7</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.7</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="127">
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">3FD/4TD</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.17</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.27</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.21</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.19</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.14</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.07</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.08</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.15</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.06</td>
|
||
<td colspan="1" pageId="0" pageNumber="127" rowspan="1">1.13</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
The new specimens from Tualapa in the upper Strickland River basin were collected from trees along a large torrential stream with a rocky substrate flowing through relatively undisturbed lower montane forest at an altitude of 1,100 m a.s.l. (Fig.
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Colour variation and habitats details for male Litoria dorsivena from Tualapa in the upper Strickland River basin, Hela Province, Papua New Guinea: A SAMA R 72320; B SAMA R 72321; C SAMA R 72321 ventral surfaces; D preserved samples (SAMA R 72321 - 4) illustrating variation in dorsal pattern; and E swift-flowing stream habitat in the upper Strickland River region. All photographs by S. J. Richards." figureDoi="10.3897/vz.73.e91111.figure8" httpUri="https://binary.pensoft.net/fig/800720" pageId="0" pageNumber="127">8E</figureCitation>
|
||
). Although we did not observe reproduction in the field, examination of two adult females in the type series of
|
||
<taxonomicName lsidName="L. dorsivena" pageId="0" pageNumber="127" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. dorsivena</emphasis>
|
||
</taxonomicName>
|
||
revealed that they contain large yellow eggs typical of torrent-breeding
|
||
<taxonomicName authorityName="Oliver & Iskandar & Richards" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Litoria" order="Anura" pageId="0" pageNumber="127" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">Litoria</emphasis>
|
||
</taxonomicName>
|
||
in New Guinea. SAMA R7904 (SVL 45mm) had eggs approximately 2 mm in diameter and SAMA R7903 (SVL 46 mm) had eggs between 1.4-1.6 mm in diameter.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="127">
|
||
<bibRefCitation author="Menzies, J" journalOrPublisher="Annali dei Museo Civico di Storia Naturale Genova" pageId="0" pageNumber="127" pagination="17 - 21" refId="B9" refString="Menzies, J, Zweifel, RG, 1976. Specific identity of specimens in the type series of the Papuan hylid frog, Litoria arfakiana. Annali dei Museo Civico di Storia Naturale Genova 81: 17 - 21" title="Specific identity of specimens in the type series of the Papuan hylid frog, Litoria arfakiana." volume="81" year="1976">Menzies and Zweifel (1976)</bibRefCitation>
|
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reported specimens of
|
||
<taxonomicName lsidName="L. dorsivena" pageId="0" pageNumber="127" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. dorsivena</emphasis>
|
||
</taxonomicName>
|
||
in the paralectotype series of
|
||
<taxonomicName lsidName="L. arfakiana" pageId="0" pageNumber="127" rank="species" species="arfakiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. arfakiana</emphasis>
|
||
</taxonomicName>
|
||
, which was collected from the Arfak Mountains and stored in the Museo Civico di Storia Naturale (MSNG). One of us (SJR) has examined this material. None of these specimens from far western New Guinea identified as
|
||
<taxonomicName lsidName="L. dorsivena" pageId="0" pageNumber="127" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. dorsivena</emphasis>
|
||
</taxonomicName>
|
||
overlap the size range of either the type series of
|
||
<taxonomicName lsidName="L. dorsivena" pageId="0" pageNumber="127" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. dorsivena</emphasis>
|
||
</taxonomicName>
|
||
, additional material confirmed by us to be
|
||
<taxonomicName lsidName="L. dorsivena" pageId="0" pageNumber="127" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. dorsivena</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName lsidName="L. hastula" pageId="0" pageNumber="127" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. hastula</emphasis>
|
||
</taxonomicName>
|
||
; and the snouts of the MSNG material identified by Menzies and Zweifel are much more bluntly rounded than those of both
|
||
<taxonomicName lsidName="L. dorsivena" pageId="0" pageNumber="127" rank="species" species="dorsivena">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. dorsivena</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="L. hastula" pageId="0" pageNumber="127" rank="species" species="hastula">
|
||
<emphasis italics="true" pageId="0" pageNumber="127">L. hastula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="0" pageNumber="127">sp. nov.</emphasis>
|
||
On this basis we consider that neither species is represented in this series, and the specific status of this far western material is unresolved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |