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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.974.54794" ID-GBIF-Dataset="5f0b0760-7a15-49c0-9e80-fc6747f8046e" ID-PMC="PMC7557337" ID-Pensoft-Pub="1313-2970-974-31" ID-Pensoft-UUID="20CAFA6BA01359369DA3F27F90E22660" ID-PubMed="33110378" ID-ZooBank="9B8888660F074DECAE7B88DFB0A4621C" ModsDocID="1313-2970-974-31" checkinTime="1602130124706" checkinUser="pensoft" docAuthor="Wood, Thomas James, Michez, Denis, Cejas, Diego, Lhomme, Patrick &amp; Rasmont, Pierre" docDate="2020" docId="05CF51C20CB95B71AF9A0DFFE6251846" docLanguage="en" docName="ZooKeys 974: 31-92" docOrigin="ZooKeys 974" docSource="http://dx.doi.org/10.3897/zookeys.974.54794" docTitle="Andrena tenebricorpus Wood 2020, sp. nov." docType="treatment" docUuid="6C1EA635-A7DD-418E-BC59-CF97AB02F508" docUuidSource="ZooBank" docVersion="4" id="20CAFA6BA01359369DA3F27F90E22660" lastPageNumber="31" masterDocId="20CAFA6BA01359369DA3F27F90E22660" masterDocTitle="An update and revision of the Andrena fauna of Morocco (Hymenoptera, Apoidea, Andrenidae) with the description of eleven new North African species" masterLastPageNumber="92" masterPageNumber="31" pageNumber="31" updateTime="1668169591898" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>An update and revision of the Andrena fauna of Morocco (Hymenoptera, Apoidea, Andrenidae) with the description of eleven new North African species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wood, Thomas James</mods:namePart>
<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5653-224X</mods:nameIdentifier>
<mods:nameIdentifier type="email">thomasjames.wood@umons.ac.be</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Michez, Denis</mods:namePart>
<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-8880-1838</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cejas, Diego</mods:namePart>
<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lhomme, Patrick</mods:namePart>
<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium &amp; International Center of Agricultural Research in the Dry Areas, Rabat, Morocco</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6735-9104</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rasmont, Pierre</mods:namePart>
<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>974</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>31</mods:start>
<mods:end>92</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.974.54794</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.974.54794</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-974-31</mods:identifier>
<mods:identifier type="ZooBank">9B8888660F074DECAE7B88DFB0A4621C</mods:identifier>
<mods:identifier type="Pensoft-UUID">20CAFA6BA01359369DA3F27F90E22660</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="168243733" LSID="urn:lsid:zoobank.org:act:6C1EA635-A7DD-418E-BC59-CF97AB02F508" httpUri="http://treatment.plazi.org/id/05CF51C20CB95B71AF9A0DFFE6251846" lastPageNumber="31" pageId="0" pageNumber="31">
<subSubSection pageId="0" pageNumber="31" type="nomenclature">
<paragraph pageId="0" pageNumber="31">
<taxonomicName LSID="05CF51C2-0CB9-5B71-AF9A-0DFFE6251846" authority="(incertae sedis) tenebricorpus Wood" authorityName="tenebricorpus Wood" authorityYear="2020" baseAuthorityName="incertae sedis" class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus" status="sp. nov.">Andrena (incertae sedis) tenebricorpus Wood</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="31">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 6366" captionStartId="F10" captionText="Figures 63 - 66. Andrena (Nobandrena) tenebricorpus sp. nov. 63 female profile 64 female face 65 female propodeum 66 female tergites." figureDoi="10.3897/zookeys.974.54794.figures63-66" httpUri="https://binary.pensoft.net/fig/460651" pageId="0" pageNumber="31">Figures 63-66</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="materials_examined">
<paragraph pageId="0" pageNumber="31">Material.</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Holotype</emphasis>
: Morocco: Guelmim-Oued Noun, 10 km E Guelmim, 15.iv.1995, 1♀, leg. Ma. Halada. Deposited in the
<normalizedToken originalValue="OÖLM">OOELM</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="diagnosis">
<paragraph pageId="0" pageNumber="31">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="31">
<taxonomicName class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena tenebricorpus" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="tenebricorpus">
<emphasis italics="true" pageId="0" pageNumber="31">Andrena tenebricorpus</emphasis>
</taxonomicName>
is very similar to
<taxonomicName lsidName="A. gafsensis" pageId="0" pageNumber="31" rank="species" species="gafsensis">
<emphasis italics="true" pageId="0" pageNumber="31">A. gafsensis</emphasis>
</taxonomicName>
and faces the same problems of subgeneric classification, and is therefore also not
<taxonomicName lsidName="currently" pageId="0" pageNumber="31" rank="species" species="currently">currently</taxonomicName>
placed in one until molecular data are available. Both species have a flattened and shiny clypeus (Fig.
<figureCitation captionStart="Figures 6366" captionStartId="F10" captionText="Figures 63 - 66. Andrena (Nobandrena) tenebricorpus sp. nov. 63 female profile 64 female face 65 female propodeum 66 female tergites." figureDoi="10.3897/zookeys.974.54794.figures63-66" httpUri="https://binary.pensoft.net/fig/460651" pageId="0" pageNumber="31">64</figureCitation>
) and a short propodeum (Fig.
<figureCitation captionStart="Figures 6366" captionStartId="F10" captionText="Figures 63 - 66. Andrena (Nobandrena) tenebricorpus sp. nov. 63 female profile 64 female face 65 female propodeum 66 female tergites." figureDoi="10.3897/zookeys.974.54794.figures63-66" httpUri="https://binary.pensoft.net/fig/460651" pageId="0" pageNumber="31">65</figureCitation>
), but differ most obviously in colour, with
<taxonomicName lsidName="A. tenebricorpus" pageId="0" pageNumber="31" rank="species" species="tenebricorpus">
<emphasis italics="true" pageId="0" pageNumber="31">A. tenebricorpus</emphasis>
</taxonomicName>
having dark brown terga (Fig.
<figureCitation captionStart="Figures 6366" captionStartId="F10" captionText="Figures 63 - 66. Andrena (Nobandrena) tenebricorpus sp. nov. 63 female profile 64 female face 65 female propodeum 66 female tergites." figureDoi="10.3897/zookeys.974.54794.figures63-66" httpUri="https://binary.pensoft.net/fig/460651" pageId="0" pageNumber="31">66</figureCitation>
) in strong contrast to the red terga of
<taxonomicName lsidName="A. gafsensis" pageId="0" pageNumber="31" rank="species" species="gafsensis">
<emphasis italics="true" pageId="0" pageNumber="31">A. gafsensis</emphasis>
</taxonomicName>
. In terms of structural difference, in
<taxonomicName lsidName="A. tenebricorpus" pageId="0" pageNumber="31" rank="species" species="tenebricorpus">
<emphasis italics="true" pageId="0" pageNumber="31">A. tenebricorpus</emphasis>
</taxonomicName>
the foveae are slightly wider, the hairs on the scutum and scutellum are relatively much shorter (compare Figs
<figureCitation captionStart="Figures 5562" captionStartId="F9" captionText="Figures 55 - 62. Andrena (Nobandrena) gafsensis sp. nov. 55 female profile 56 female face 57 female propodeum 58 female tergites 59 male profile 60 male face 61 male tergites 62 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures55-62" httpUri="https://binary.pensoft.net/fig/460650" pageId="0" pageNumber="31">57</figureCitation>
,
<figureCitation captionStart="Figures 6366" captionStartId="F10" captionText="Figures 63 - 66. Andrena (Nobandrena) tenebricorpus sp. nov. 63 female profile 64 female face 65 female propodeum 66 female tergites." figureDoi="10.3897/zookeys.974.54794.figures63-66" httpUri="https://binary.pensoft.net/fig/460651" pageId="0" pageNumber="31">65</figureCitation>
) at the same time that the hair bands on the apical margins of the tergites are relatively much thicker (compare Figs
<figureCitation captionStart="Figures 5562" captionStartId="F9" captionText="Figures 55 - 62. Andrena (Nobandrena) gafsensis sp. nov. 55 female profile 56 female face 57 female propodeum 58 female tergites 59 male profile 60 male face 61 male tergites 62 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures55-62" httpUri="https://binary.pensoft.net/fig/460650" pageId="0" pageNumber="31">58</figureCitation>
,
<figureCitation captionStart="Figures 6366" captionStartId="F10" captionText="Figures 63 - 66. Andrena (Nobandrena) tenebricorpus sp. nov. 63 female profile 64 female face 65 female propodeum 66 female tergites." figureDoi="10.3897/zookeys.974.54794.figures63-66" httpUri="https://binary.pensoft.net/fig/460651" pageId="0" pageNumber="31">66</figureCitation>
), suggesting that this is not as a result of abrasion, and the scopa are a dirty brown (Fig.
<figureCitation captionStart="Figures 6366" captionStartId="F10" captionText="Figures 63 - 66. Andrena (Nobandrena) tenebricorpus sp. nov. 63 female profile 64 female face 65 female propodeum 66 female tergites." figureDoi="10.3897/zookeys.974.54794.figures63-66" httpUri="https://binary.pensoft.net/fig/460651" pageId="0" pageNumber="31">63</figureCitation>
) rather than white (Fig.
<figureCitation captionStart="Figures 5562" captionStartId="F9" captionText="Figures 55 - 62. Andrena (Nobandrena) gafsensis sp. nov. 55 female profile 56 female face 57 female propodeum 58 female tergites 59 male profile 60 male face 61 male tergites 62 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures55-62" httpUri="https://binary.pensoft.net/fig/460650" pageId="0" pageNumber="31">55</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="31">
In terms of overall colouration and appearance
<taxonomicName lsidName="A. tenebricorpus" pageId="0" pageNumber="31" rank="species" species="tenebricorpus">
<emphasis italics="true" pageId="0" pageNumber="31">A. tenebricorpus</emphasis>
</taxonomicName>
is extremely similar to
<taxonomicName lsidName="A. guichardi" pageId="0" pageNumber="31" rank="species" species="guichardi">
<emphasis italics="true" pageId="0" pageNumber="31">A. guichardi</emphasis>
</taxonomicName>
Warncke, 1980 (also found only in south-western Morocco) with the same dark to dark brownish tergal colouration with contrasting white hair bands (see Fig.
<figureCitation captionStart="Figures 99104" captionStartId="F17" captionText="Figures 99 - 104. Andrena (Aciandrena) fulica Warncke, 1974 99 male face 100 male genitalia. Andrena (Aciandrena) fulica astrella Warncke, 1975 (Morocco) 101 male face 102 male genitalia (Spain) 103 male face 104 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures99-104" httpUri="https://binary.pensoft.net/fig/460658" pageId="0" pageNumber="31">104</figureCitation>
). However,
<taxonomicName lsidName="A. tenebricorpus" pageId="0" pageNumber="31" rank="species" species="tenebricorpus">
<emphasis italics="true" pageId="0" pageNumber="31">A. tenebricorpus</emphasis>
</taxonomicName>
can be clearly separated by the flattened clypeus (domed in
<taxonomicName lsidName="A. guichardi" pageId="0" pageNumber="31" rank="species" species="guichardi">
<emphasis italics="true" pageId="0" pageNumber="31">A. guichardi</emphasis>
</taxonomicName>
), more densely punctate scutum (punctures separated by 1-2 puncture diameters, by 2-4 puncture diameters in
<taxonomicName lsidName="A. guichardi" pageId="0" pageNumber="31" rank="species" species="guichardi">
<emphasis italics="true" pageId="0" pageNumber="31">A. guichardi</emphasis>
</taxonomicName>
, see Fig.
<figureCitation captionStart="Figures 99104" captionStartId="F17" captionText="Figures 99 - 104. Andrena (Aciandrena) fulica Warncke, 1974 99 male face 100 male genitalia. Andrena (Aciandrena) fulica astrella Warncke, 1975 (Morocco) 101 male face 102 male genitalia (Spain) 103 male face 104 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures99-104" httpUri="https://binary.pensoft.net/fig/460658" pageId="0" pageNumber="31">103</figureCitation>
), and by the propodeal triangle which is weakly differentiated only by sculpturing (clearly marked by a small raised carina in
<taxonomicName lsidName="A. guichardi" pageId="0" pageNumber="31" rank="species" species="guichardi">
<emphasis italics="true" pageId="0" pageNumber="31">A. guichardi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="description">
<paragraph pageId="0" pageNumber="31">Description.</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Female</emphasis>
: Body length 10 mm (Fig.
<figureCitation captionStart="Figures 6366" captionStartId="F10" captionText="Figures 63 - 66. Andrena (Nobandrena) tenebricorpus sp. nov. 63 female profile 64 female face 65 female propodeum 66 female tergites." figureDoi="10.3897/zookeys.974.54794.figures63-66" httpUri="https://binary.pensoft.net/fig/460651" pageId="0" pageNumber="31">63</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Head</emphasis>
: Dark, as wide as long. Clypeus slightly arched laterally, broadly flattened on the disc, underlying surface laterally with shagreenation, the majority of disc shiny (Fig.
<figureCitation captionStart="Figures 6366" captionStartId="F10" captionText="Figures 63 - 66. Andrena (Nobandrena) tenebricorpus sp. nov. 63 female profile 64 female face 65 female propodeum 66 female tergites." figureDoi="10.3897/zookeys.974.54794.figures63-66" httpUri="https://binary.pensoft.net/fig/460651" pageId="0" pageNumber="31">64</figureCitation>
). Moderately punctured, punctures separated by 1-2 puncture diameters, with generally impunctate line in centre of clypeus. Process of labrum trapezoidal with faint lateral striations, fore margin weakly emarginate, slightly bulging. Gena equal to width of compound eye. Gena, face, and scape with white hairs, longest not exceeding length of the scape. Vertex with mixture of whitish and yellowish hairs of same length as on rest of face. Foveae of average width, occupying slightly more than half of distance between top of compound eye and lateral ocellus. Antennae dark, A4 apically and A5-12 lightened orange below, A3 equalling A4+5. Ocelloccipital distance equalling width of lateral ocellus.
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Mesosoma</emphasis>
: Scutum and scutellum densely punctured, punctures separated by 1-1.5 puncture diameters, underlying surface of the scutum shagreened anteriorly, elsewhere and on the scutellum shiny. Episternum microreticulate, dull. Lateral faces of the propodeum shagreened, weakly shining. Hind face of propodeum microreticulate and weakly rugose, rugosity outlining the propodeal triangle which is itself rugose and therefore defined by the change in texture between rugosities (Fig.
<figureCitation captionStart="Figures 6366" captionStartId="F10" captionText="Figures 63 - 66. Andrena (Nobandrena) tenebricorpus sp. nov. 63 female profile 64 female face 65 female propodeum 66 female tergites." figureDoi="10.3897/zookeys.974.54794.figures63-66" httpUri="https://binary.pensoft.net/fig/460651" pageId="0" pageNumber="31">65</figureCitation>
). Scutum and scutellum with very short and fine hairs, scarcely longer than width of lateral ocellus. Episternum and propodeum with longer white hair, never exceeding length of the scape. Legs dark, tarsi brown, pubescence brown. Femoral scopa white, tibial scopa dirty brown. Wings hyaline, venation brown, stigma dark brown. Nervulus slightly antefurcal.
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Metasoma</emphasis>
: Tergites dark brown throughout with exception of two lateral black spots on T2 (Fig.
<figureCitation captionStart="Figures 6366" captionStartId="F10" captionText="Figures 63 - 66. Andrena (Nobandrena) tenebricorpus sp. nov. 63 female profile 64 female face 65 female propodeum 66 female tergites." figureDoi="10.3897/zookeys.974.54794.figures63-66" httpUri="https://binary.pensoft.net/fig/460651" pageId="0" pageNumber="31">66</figureCitation>
). Tergites densely, finely, and evenly punctured, punctures separated by one puncture diameter. T2-4 with dense white hair bands that obscure underlying surface, on T2 interrupted medially, on T3-4 complete. T5-6 with thick brown hairs flanking pygidial plate, with a few white hairs laterally. Pygidial plate with slightly raised margin, centrally flat.
</paragraph>
<caption doi="10.3897/zookeys.974.54794.figures63-66" httpUri="https://binary.pensoft.net/fig/460651" pageId="0" pageNumber="31" start="Figures 6366" startId="F10">
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Figures 63-66.</emphasis>
<taxonomicName class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Nobandrena) tenebricorpus" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="tenebricorpus" subGenus="Nobandrena">Andrena (Nobandrena) tenebricorpus</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="31">63</emphasis>
female profile
<emphasis bold="true" pageId="0" pageNumber="31">64</emphasis>
female face
<emphasis bold="true" pageId="0" pageNumber="31">65</emphasis>
female propodeum
<emphasis bold="true" pageId="0" pageNumber="31">66</emphasis>
female tergites.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="distribution">
<paragraph pageId="0" pageNumber="31">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="31">
South-western Morocco (Fig.
<figureCitation captionStart="Figure 145" captionStartId="F24" captionText="Figure 145. Distribution of sampling locations of newly described species A Andrena hebescens (circles), Andrena semiadesus (squares) B Andrena tenebricorpus (diamond), Andrena niveofacies (circle), Andrena triangulivalvis (triangle) C Andrena sparsipunctata (circles), Andrena breviceps (diamonds) D Andrena acutidentis (squares), Andrena farinosoides (triangles), Andrena nigriclypeus (circle). Relief is indicated by shading, measured in meters above sea level." figureDoi="10.3897/zookeys.974.54794.figure145" httpUri="https://binary.pensoft.net/fig/460665" pageId="0" pageNumber="31">145b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="floral preferences">
<paragraph pageId="0" pageNumber="31">Floral preferences.</paragraph>
<paragraph pageId="0" pageNumber="31">None recorded.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="remarks">
<paragraph pageId="0" pageNumber="31">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="31">
Whilst
<taxonomicName lsidName="A. tenebricorpus" pageId="0" pageNumber="31" rank="species" species="tenebricorpus">
<emphasis italics="true" pageId="0" pageNumber="31">A. tenebricorpus</emphasis>
</taxonomicName>
is structurally very similar to
<taxonomicName lsidName="A. gafsensis" pageId="0" pageNumber="31" rank="species" species="gafsensis">
<emphasis italics="true" pageId="0" pageNumber="31">A. gafsensis</emphasis>
</taxonomicName>
, the subtle morphological differences suggest that
<taxonomicName lsidName="A. tenebricorpus" pageId="0" pageNumber="31" rank="species" species="tenebricorpus">
<emphasis italics="true" pageId="0" pageNumber="31">A. tenebricorpus</emphasis>
</taxonomicName>
is distinct. Moreover, the Souss valley to the Guelmim region is bio-climatically quite distinct to the deserts of southern Tunisia and hosts many unique bee species within Morocco itself, further suggesting a distinct specific identity.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="etymology">
<paragraph pageId="0" pageNumber="31">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="31">
The name
<taxonomicName lsidName="tenebricorpus" pageId="0" pageNumber="31" rank="species" species="tenebricorpus">
<emphasis italics="true" pageId="0" pageNumber="31">tenebricorpus</emphasis>
</taxonomicName>
from
<emphasis italics="true" pageId="0" pageNumber="31">tenebris</emphasis>
(dark) +
<emphasis italics="true" pageId="0" pageNumber="31">corpus</emphasis>
(body) was chosen because of the structural similarities between
<taxonomicName lsidName="A. tenebricorpus" pageId="0" pageNumber="31" rank="species" species="tenebricorpus">
<emphasis italics="true" pageId="0" pageNumber="31">A. tenebricorpus</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="A. gafsensis" pageId="0" pageNumber="31" rank="species" species="gafsensis">
<emphasis italics="true" pageId="0" pageNumber="31">A. gafsensis</emphasis>
</taxonomicName>
, but without the same reddish colouration.
</paragraph>
</subSubSection>
</treatment>
</document>