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<document id="FA59C4614646631E501DF64867472083" ID-CLB-Dataset="20891" ID-DOI="10.11646/zootaxa.4991.2.8" ID-GBIF-Dataset="5d6789a3-94ef-445a-be9f-41a15128625a" ID-ISSN="1175-5326" ID-Zenodo-Dep="5030538" ID-ZooBank="5F7027D6-07E4-4447-93F3-C3FBEC45D630" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624558585460" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2021" docId="0579878DFB299279FF23FC6DE610D1AF" docLanguage="en" docName="zootaxa.4991.2.8.pdf" docOrigin="Zootaxa 4991 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichodromeus kaliyuga Shavrin 2021, sp.n." docType="treatment" docVersion="7" lastPageNumber="360" masterDocId="F940FFF5FB2F927EFFB4FFBFE535D05A" masterDocTitle="Four new species of the genus Trichodromeus Luze, 1903 from Nepal and Pakistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="362" masterPageNumber="353" pageNumber="359" updateTime="1699020750815" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8BB492361C55C0EA5229C9C18EE6F1CA">Four new species of the genus Trichodromeus Luze, 1903 from Nepal and Pakistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="1BC0B24F664596FA6EC5D5F6146F2C2D">Shavrin, Alexey V.</mods:namePart>
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<mods:date id="71241E2A5E8495BCC72A55C1A2C743AD">2021</mods:date>
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<mods:number id="E84E6BE17BF770976B72BD2D23F8C43C">4991</mods:number>
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<treatment id="0579878DFB299279FF23FC6DE610D1AF" ID-DOI="http://doi.org/10.5281/zenodo.5916681" ID-GBIF-Taxon="192745371" ID-Zenodo-Dep="5916681" LSID="urn:lsid:plazi:treatment:0579878DFB299279FF23FC6DE610D1AF" httpUri="http://treatment.plazi.org/id/0579878DFB299279FF23FC6DE610D1AF" lastPageId="7" lastPageNumber="360" pageId="6" pageNumber="359">
<subSubSection id="C5CA6510FB299278FF23FC6DE741D3B7" box="[151,628,978,1005]" pageId="6" pageNumber="359" type="nomenclature">
<paragraph id="8D6F369BFB299278FF23FC6DE741D3B7" blockId="6.[151,628,978,1041]" box="[151,628,978,1005]" pageId="6" pageNumber="359">
<heading id="D62781F7FB299278FF23FC6DE741D3B7" bold="true" box="[151,628,978,1005]" fontSize="11" level="1" pageId="6" pageNumber="359" reason="1">
<emphasis id="BFA4EA89FB299278FF23FC6DE741D3B7" bold="true" box="[151,628,978,1005]" pageId="6" pageNumber="359">
<taxonomicName id="4AD04D18FB299278FF23FC6DE719D3B6" ID-CoL="8YBYN" authority="Shavrin" authorityName="Shavrin" authorityYear="2021" box="[151,556,978,1005]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="species" species="kaliyuga" status="sp. nov.">
<emphasis id="BFA4EA89FB299278FF23FC6DE48AD3B7" bold="true" box="[151,447,978,1005]" italics="true" pageId="6" pageNumber="359">Trichodromeus kaliyuga</emphasis>
Shavrin
</taxonomicName>
,
<taxonomicNameLabel id="A49757F2FB299278FD8DFC6CE741D3B7" box="[569,628,979,1005]" pageId="6" pageNumber="359" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C5CA6510FB299278FF23FC48E40ED44B" box="[151,315,1015,1041]" pageId="6" pageNumber="359" type="description">
<paragraph id="8D6F369BFB299278FF23FC48E40ED44B" blockId="6.[151,628,978,1041]" box="[151,315,1015,1041]" pageId="6" pageNumber="359">
(
<figureCitation id="15EB2A1EFB299278FF2BFC48E5DBD44B" box="[159,238,1015,1041]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1946,1970]" captionTargetBox="[168,1419,192,1909]" captionTargetId="figure-28@2.[152,1436,179,1920]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus of Trichodromeus: 1—T. acutipenis (holotype), 2—T. jaegeri (paratype), 3—T. kaliyuga (holotype), 4—T. trilobatus (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5030540" httpUri="https://zenodo.org/record/5030540/files/figure.png" pageId="6" pageNumber="359">Figs. 3</figureCitation>
,
<figureCitation id="15EB2A1EFB299278FF4FFC48E406D44B" box="[251,307,1015,1041]" captionStart-0="FIGURES 5–10" captionStart-1="FIGURE 11" captionStartId-0="3.[151,264,1943,1967]" captionStartId-1="5.[151,250,1679,1703]" captionTargetBox-0="[173,1420,193,1912]" captionTargetBox-1="[151,1436,704,1656]" captionTargetId-0="figure-16@3.[155,1432,181,1919]" captionTargetId-1="figure-216@5.[151,1436,704,1656]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="FIGURES 5–10. Aedeagus of Trichodromeus (5–6: T. acutipenis (holotype), 7–8: T. jaegeri (holotype), 9–10: T. kaliyuga (holotype): 5, 7, 9–parameral view, 6, 8, 10 lateral view. Scale bar: 1.0 mm." captionText-1="FIGURE 11. Distribution of Trichodromeus in Nepal: T. acutipenis (circles), T. jaegeri (square), T. kaliyuga (triangle)." figureDoi-0="http://doi.org/10.5281/zenodo.5030542" figureDoi-1="http://doi.org/10.5281/zenodo.5030544" httpUri-0="https://zenodo.org/record/5030542/files/figure.png" httpUri-1="https://zenodo.org/record/5030544/files/figure.png" pageId="6" pageNumber="359">9–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C5CA6510FB299278FF23FB80E1E5D4FB" pageId="6" pageNumber="359" type="materials_examined">
<paragraph id="8D6F369BFB299278FF23FB80E1E5D4FB" blockId="6.[151,1437,1086,2013]" pageId="6" pageNumber="359">
<emphasis id="BFA4EA89FB299278FF23FB80E4F2D402" bold="true" box="[151,455,1086,1113]" pageId="6" pageNumber="359">Type material examined:</emphasis>
<typeStatus id="526B8839FB299278FE65FB80E709D403" box="[465,572,1087,1113]" pageId="6" pageNumber="359" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9BD6FD12FB299278FDFEFB80E76AD403" box="[586,607,1087,1113]" pageId="6" pageNumber="359" type="male">♂</specimenCount>
[dissected; specimen without left antennomeres 3–11]: ‘
<collectingCountry id="F5C7760BFB299278FB5CFB80E07DD403" box="[1256,1352,1087,1113]" name="Nepal" pageId="6" pageNumber="359">NEPAL</collectingCountry>
, Dhau- lagiri-Himal | Kali Gand. vall., Yak Kharka | (upp. Marpha),
<quantity id="4A289B7EFB299278FCFEFBDCE6F0D427" box="[842,965,1123,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.3205" metricValueMax="4.6" metricValueMin="0.040999999999999995" pageId="6" pageNumber="359" unit="m" value="2320.5" valueMax="4600.0" valueMin="41.0">41–4600m</quantity>
, | 12./
<date id="F96E105BFB299278FBBDFBDCE1A0D427" box="[1033,1173,1123,1149]" pageId="6" pageNumber="359" value="1998-07-13">13.VII.1998</date>
, leg. Jãger ’ &lt;printed&gt;, ‘
<typeStatus id="526B8839FB299278FF2FFB38E401D4FB" box="[155,308,1159,1185]" pageId="6" pageNumber="359" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="4AD04D18FB299278FEFCFB38E4C4D4FB" authorityName="Luze" authorityYear="1903" box="[328,497,1159,1185]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="genus">
<emphasis id="BFA4EA89FB299278FEFCFB38E4C4D4FB" box="[328,497,1159,1185]" italics="true" pageId="6" pageNumber="359">Trichodromeus</emphasis>
</taxonomicName>
|
<taxonomicName id="4AD04D18FB299278FDB0FB38E751D4FB" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[516,612,1159,1185]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="359" phylum="Arthropoda" rank="species" species="kaliyuga" status="sp. nov.">
<emphasis id="BFA4EA89FB299278FDB0FB38E751D4FB" box="[516,612,1159,1185]" italics="true" pageId="6" pageNumber="359">kaliyuga</emphasis>
</taxonomicName>
<emphasis id="BFA4EA89FB299278FDDFFB38E796D4FB" bold="true" box="[619,675,1159,1185]" pageId="6" pageNumber="359">
<taxonomicNameLabel id="A49757F2FB299278FDDFFB38E796D4FB" box="[619,675,1159,1185]" pageId="6" pageNumber="359" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="EBC1AE5EFB299278FC99FB38E676D4FB" box="[813,835,1159,1185]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="359" type="Museum">V</collectionCode>
. des. 2021’ &lt;red, printed&gt; (
<collectionCode id="EBC1AE5EFB299278FBC3FB38E1F1D4FB" box="[1143,1220,1159,1185]" pageId="6" pageNumber="359">SNSD</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C5CA6510FB299279FF73FB15E7BED1D3" lastPageId="7" lastPageNumber="360" pageId="6" pageNumber="359" type="description">
<paragraph id="8D6F369BFB299278FF73FB15E4FAD557" blockId="6.[151,1437,1086,2013]" pageId="6" pageNumber="359">
<emphasis id="BFA4EA89FB299278FF73FB15E46CD49E" bold="true" box="[199,345,1194,1220]" pageId="6" pageNumber="359">Description.</emphasis>
Measurements of the
<typeStatus id="526B8839FB299278FDE9FB14E7F6D49F" box="[605,707,1195,1221]" pageId="6" pageNumber="359" type="holotype">holotype</typeStatus>
: HW: 0.97; HL: 0.75; OL: 0.22; LT: 0.15; AL: 3.12; PL: 0.88; PWmax: 0.97; PWmin: 0.72; ESL: 1.46; EW: 1.72; MTbL: 1.27; MTrL: 0.52 (MTrL 1–4: 0.27; MTrL 5: 0.25); AW: 1.60; AedL: 1.12; BL: 5.00.
</paragraph>
<paragraph id="8D6F369BFB299278FF73FAA8E703D523" blockId="6.[151,1437,1086,2013]" pageId="6" pageNumber="359">
Body, mouthparts and antennae reddish-brown (elytra slightly paler); legs yellow-brown. Head with very fine transverse microsculpture, coarser in middle; pronotum with fine isodiametric microreticulation, mediobasal third without meshes. Habitus as in
<figureCitation id="15EB2A1EFB299278FE59FAE0E704D523" box="[493,561,1375,1401]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1946,1970]" captionTargetBox="[168,1419,192,1909]" captionTargetId="figure-28@2.[152,1436,179,1920]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus of Trichodromeus: 1—T. acutipenis (holotype), 2—T. jaegeri (paratype), 3—T. kaliyuga (holotype), 4—T. trilobatus (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5030540" httpUri="https://zenodo.org/record/5030540/files/figure.png" pageId="6" pageNumber="359">Fig. 3</figureCitation>
.
</paragraph>
<paragraph id="8D6F369BFB299278FF73FA3CE01DD62F" blockId="6.[151,1437,1086,2013]" pageId="6" pageNumber="359">Head 1.2 times as broad as long, with strongly elevated portions between ocelli and eyes; anterio-median depression moderately deep and wide; temples slightly shorter than longitudinal length of eyes; interocellar depression very deep, moderately narrow, subrectangular, separated from infraorbital ridges by deep anteocellar foveae. Ocelli indistinct, distance between ocelli slightly shorter than distance between ocellus and posterior margin of eye. Punctation dense, large and deep, finer in middle. Antennae reaching apical third of elytra when reclined; basal antennomere wide, distinctly more than twice as long as broad, antennomere 2 siginificantly shorter and narrower than basal antennomere, 3 longer than 2, 4–8 slightly shorter and broader than 3, 9–10 slightly shorter than 8.</paragraph>
<paragraph id="8D6F369BFB299278FF73F9C0E628D773" blockId="6.[151,1437,1086,2013]" pageId="6" pageNumber="359">Pronotum slightly broader than long, as broad as head, from widest anterior portion gradually narrowed posteriad, anterior margin rounded, not protruded anteriad; posterior margin slightly concaved; middle portion with long and moderately deep longitudinal depression, connected with indistinct transverse mediobasal depression. Punctation about as that on head, but denser and finer, sparser in mediobasal third, interspaces between punctures in middle as broad as an average diameter of two nearest punctures.</paragraph>
<paragraph id="8D6F369BFB299278FF73F88CE037D72B" blockId="6.[151,1437,1086,2013]" pageId="6" pageNumber="359">Elytra slightly longer than broad, slightly widened posteriad; hind margins somewhat straight. Punctation regular, distinctly finer and sparser than that on pronotum, coarser in basal portion and finer along suture.</paragraph>
<paragraph id="8D6F369BFB299279FF73F8C4E072D08F" blockId="6.[151,1437,1086,2013]" lastBlockId="7.[151,1436,151,501]" lastPageId="7" lastPageNumber="360" pageId="6" pageNumber="359">
Male. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII concaved. Aedeagus with wide basal portion, slightly narrowed toward middle of median lobe and slightly widened and rounded in apical portion, with moderately small apex and narrow, moderately deep median excision; parameres very long, significantly exceeding apex of median lobe, slightly widened apically, with four long apical setae and a short additional seta on inner margins of each paramere in middle (
<figureCitation id="15EB2A1EFB289279FC98FF04E644D08F" box="[812,881,187,213]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[173,1420,193,1912]" captionTargetId="figure-16@3.[155,1432,181,1919]" captionTargetPageId="3" captionText="FIGURES 5–10. Aedeagus of Trichodromeus (5–6: T. acutipenis (holotype), 7–8: T. jaegeri (holotype), 9–10: T. kaliyuga (holotype): 5, 7, 9–parameral view, 6, 8, 10 lateral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5030542" httpUri="https://zenodo.org/record/5030542/files/figure.png" pageId="7" pageNumber="360">Fig. 9</figureCitation>
). Lateral aspect of aedeagus as in
<figureCitation id="15EB2A1EFB289279FB44FF04E076D08F" box="[1264,1347,187,213]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[173,1420,193,1912]" captionTargetId="figure-16@3.[155,1432,181,1919]" captionTargetPageId="3" captionText="FIGURES 5–10. Aedeagus of Trichodromeus (5–6: T. acutipenis (holotype), 7–8: T. jaegeri (holotype), 9–10: T. kaliyuga (holotype): 5, 7, 9–parameral view, 6, 8, 10 lateral view. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5030542" httpUri="https://zenodo.org/record/5030542/files/figure.png" pageId="7" pageNumber="360">Fig. 10</figureCitation>
.
</paragraph>
<paragraph id="8D6F369BFB289279FF73FF60E4A4D0A3" blockId="7.[151,1436,151,501]" box="[199,401,223,249]" pageId="7" pageNumber="360">Female unknown.</paragraph>
<paragraph id="8D6F369BFB289279FF73FEBDE7BED1D3" blockId="7.[151,1436,151,501]" pageId="7" pageNumber="360">
<emphasis id="BFA4EA89FB289279FF73FEBDE486D147" bold="true" box="[199,435,258,285]" pageId="7" pageNumber="360">Comparative notes.</emphasis>
Based on the general shape of the body,
<taxonomicName id="4AD04D18FB289279FC24FEBCE125D147" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[912,1040,259,285]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="360" phylum="Arthropoda" rank="species" species="kaliyuga" status="sp. nov.">
<emphasis id="BFA4EA89FB289279FC24FEBCE695D146" box="[912,928,259,284]" italics="true" pageId="7" pageNumber="360">T</emphasis>
.
<emphasis id="BFA4EA89FB289279FC04FEBCE125D147" box="[944,1040,259,285]" italics="true" pageId="7" pageNumber="360">kaliyuga</emphasis>
</taxonomicName>
<emphasis id="BFA4EA89FB289279FBADFEBCE164D147" bold="true" box="[1049,1105,259,285]" pageId="7" pageNumber="360">
<taxonomicNameLabel id="A49757F2FB289279FBADFEBCE164D147" box="[1049,1105,259,285]" pageId="7" pageNumber="360" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4AD04D18FB289279FB58FEBCE06ED146" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[1260,1371,259,285]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="360" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
<emphasis id="BFA4EA89FB289279FB58FEBCE1C9D146" box="[1260,1276,259,284]" italics="true" pageId="7" pageNumber="360">T</emphasis>
.
<emphasis id="BFA4EA89FB289279FAB8FEBCE06ED146" box="[1292,1371,259,284]" italics="true" pageId="7" pageNumber="360">jaegeri</emphasis>
</taxonomicName>
<emphasis id="BFA4EA89FB289279FAD0FEBCE0A9D147" bold="true" box="[1380,1436,259,285]" pageId="7" pageNumber="360">
<taxonomicNameLabel id="A49757F2FB289279FAD0FEBCE0A9D147" box="[1380,1436,259,285]" pageId="7" pageNumber="360" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4AD04D18FB289279FF7DFE98E463D11B" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[201,342,295,321]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="360" phylum="Arthropoda" rank="species" species="trilobatus" status="sp. nov.">
<emphasis id="BFA4EA89FB289279FF7DFE98E5ECD11A" box="[201,217,295,320]" italics="true" pageId="7" pageNumber="360">T</emphasis>
.
<emphasis id="BFA4EA89FB289279FF5DFE98E463D11B" box="[233,342,295,321]" italics="true" pageId="7" pageNumber="360">trilobatus</emphasis>
</taxonomicName>
<emphasis id="BFA4EA89FB289279FEEBFE98E4A3D11B" bold="true" box="[351,406,295,321]" pageId="7" pageNumber="360">
<taxonomicNameLabel id="A49757F2FB289279FEEBFE98E4A3D11B" box="[351,406,295,321]" pageId="7" pageNumber="360" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, from which it can be distinguished by the smaller body and different morphology of the aedeagus.
<taxonomicName id="4AD04D18FB289279FEB8FEF4E729D13F" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[268,540,331,357]" class="Insecta" family="Staphylinidae" genus="Trichodromeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="360" phylum="Arthropoda" rank="species" species="kaliyuga" status="sp. nov.">
<emphasis id="BFA4EA89FB289279FEB8FEF4E729D13F" box="[268,540,331,357]" italics="true" pageId="7" pageNumber="360">Trichodromeus kaliyuga</emphasis>
</taxonomicName>
<emphasis id="BFA4EA89FB289279FD97FEF4E76ED13F" bold="true" box="[547,603,331,357]" pageId="7" pageNumber="360">
<taxonomicNameLabel id="A49757F2FB289279FD97FEF4E76ED13F" box="[547,603,331,357]" pageId="7" pageNumber="360" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
differs from all known species of the genus by the shape of apical portion of the median lobe and very long parameres.
</paragraph>
</subSubSection>
<subSubSection id="C5CA6510FB289279FF73FE2DE1EED1F7" box="[199,1243,402,429]" pageId="7" pageNumber="360" type="distribution">
<paragraph id="8D6F369BFB289279FF73FE2DE1EED1F7" blockId="7.[151,1436,151,501]" box="[199,1243,402,429]" pageId="7" pageNumber="360">
<emphasis id="BFA4EA89FB289279FF73FE2DE454D1F6" bold="true" box="[199,353,402,428]" pageId="7" pageNumber="360">Distribution.</emphasis>
The species is known only from the
<typeStatus id="526B8839FB289279FD4AFE2CE61BD1F7" box="[766,814,403,429]" pageId="7" pageNumber="360">type</typeStatus>
locality in the central
<collectingCountry id="F5C7760BFB289279FB9DFE2CE159D1F7" box="[1065,1132,403,429]" name="Nepal" pageId="7" pageNumber="360">Nepal</collectingCountry>
(
<figureCitation id="15EB2A1EFB289279FBCFFE2CE1F8D1F7" box="[1147,1229,403,429]" captionStart="FIGURE 11" captionStartId="5.[151,250,1679,1703]" captionTargetBox="[151,1436,704,1656]" captionTargetId="figure-216@5.[151,1436,704,1656]" captionTargetPageId="5" captionText="FIGURE 11. Distribution of Trichodromeus in Nepal: T. acutipenis (circles), T. jaegeri (square), T. kaliyuga (triangle)." figureDoi="http://doi.org/10.5281/zenodo.5030544" httpUri="https://zenodo.org/record/5030544/files/figure.png" pageId="7" pageNumber="360">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C5CA6510FB289279FF73FE08E610D1AF" pageId="7" pageNumber="360" type="etymology">
<paragraph id="8D6F369BFB289279FF73FE08E610D1AF" blockId="7.[151,1436,151,501]" pageId="7" pageNumber="360">
<emphasis id="BFA4EA89FB289279FF73FE08E47BD18B" bold="true" box="[199,334,439,465]" pageId="7" pageNumber="360">Etymology.</emphasis>
The name of a new species is devoted to the Kali Yuga. This is the fourth and worst of the four yugas in Hinduism, the dark epoch full of conflict and sin.
</paragraph>
</subSubSection>
</treatment>
</document>
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