691 lines
77 KiB
XML
691 lines
77 KiB
XML
<document ID-DOI="10.7717/peerj.11514" ID-GBIF-Dataset="e56413f9-7dc9-4ebc-9085-008cb38b4dc1" ID-ISSN="2167-8359" ID-PMC="PMC8254503" ID-PubMed="34249486" ID-Zenodo-Dep="5102026" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1626269878614" checkinUser="diego" docAuthor="Prebus, Matthew M." docDate="2021" docId="054FDB70FFBEFFA5B49C1CA17A5F22E7" docLanguage="en" docName="PeerJ.9.e11514.pdf.imf" docOrigin="PeerJ (e 11514) 9" docSource="http://dx.doi.org/10.7717/peerj.11514" docStyle="DocumentStyle:B6346867C9D49AB2DD3A1D607021D2E9:PeerJ.2021-.journal_article" docStyleId="B6346867C9D49AB2DD3A1D607021D2E9" docStyleName="PeerJ.2021-.journal_article" docTitle="Temnothorax augusti" docType="treatment" docVersion="4" lastPageNumber="128" masterDocId="F976A308FFC7FFDAB56E1A1D79322757" masterDocTitle="Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax" masterLastPageNumber="462" masterPageNumber="1" pageNumber="122" updateTime="1668138851842" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Temnothorax salvini clade (Hymenoptera: Formicidae), with a key to the clades of New World Temnothorax</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prebus, Matthew M.</mods:namePart>
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<mods:affiliation>School of Life Sciences, Arizona State University, Tempe, Arizona, United States & Department of Entomology & Nematology, University of California, Davis, Davis, California, United States</mods:affiliation>
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<mods:nameIdentifier type="email">mprebus@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>PeerJ</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="series">
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<mods:title>e 11514</mods:title>
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</mods:detail>
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<mods:detail type="pubDate">
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<mods:number>2021-06-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>9</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>462</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.7717/peerj.11514</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.7717/peerj.11514</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e56413f9-7dc9-4ebc-9085-008cb38b4dc1</mods:identifier>
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<mods:identifier type="ISSN">2167-8359</mods:identifier>
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<mods:identifier type="PMC">PMC8254503</mods:identifier>
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<mods:identifier type="PubMed">34249486</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5102026</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5102503" ID-GBIF-Taxon="183542583" ID-Zenodo-Dep="5102503" LSID="urn:lsid:plazi:treatment:054FDB70FFBEFFA5B49C1CA17A5F22E7" httpUri="http://treatment.plazi.org/id/054FDB70FFBEFFA5B49C1CA17A5F22E7" lastPageId="127" lastPageNumber="128" pageId="121" pageNumber="122">
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<subSubSection box="[498,1140,1724,1754]" pageId="121" pageNumber="122" type="nomenclature">
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<paragraph blockId="121.[498,1140,1724,1796]" box="[498,1140,1724,1754]" pageId="121" pageNumber="122">
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<heading box="[498,1140,1724,1754]" level="2" pageId="121" pageNumber="122">
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<taxonomicName authority="(Baroni Urbani, 1978)" baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[498,1140,1724,1754]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="augusti">
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<emphasis bold="true" box="[498,1140,1724,1754]" pageId="121" pageNumber="122">
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<emphasis bold="true" box="[498,817,1724,1754]" italics="true" pageId="121" pageNumber="122">Temnothorax augusti</emphasis>
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(
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<bibRefCitation author="Baroni Urbani C." box="[836,1130,1724,1754]" journalOrPublisher="Entomologica Basiliensia" pageId="121" pageNumber="122" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">
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<emphasis bold="true" box="[836,1130,1724,1754]" italics="true" pageId="121" pageNumber="122">Baroni Urbani, 1978</emphasis>
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</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[498,1075,1770,1796]" pageId="121" pageNumber="122" type="distribution">
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<paragraph blockId="121.[498,1140,1724,1796]" box="[498,1075,1770,1796]" pageId="121" pageNumber="122">
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Distribution:
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<figureCitation box="[657,770,1770,1796]" captionStart="Figure 101" captionStartId="121.[524,589,1194,1216]" captionTargetBox="[498,1541,234,1181]" captionTargetId="figure-149@121.[498,1534,234,1166]" captionTargetPageId="121" captionText="Figure 101 Geographical and elevational distribution of the augusti group. (A) Temnothorax augusti (B) T. aureus sp. nov. (C) T. leucacanthoides sp. nov. (D) T. leucacanthus. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-101" figureDoi="http://doi.org/10.5281/zenodo.5102293" httpUri="https://zenodo.org/record/5102293/files/figure.png" pageId="121" pageNumber="122">Fig. 101A</figureCitation>
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; worker & gyne:
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<figureCitation box="[975,1069,1770,1796]" captionStart="Figure 102" captionStartId="122.[524,589,1266,1288]" captionTargetBox="[498,1541,234,1253]" captionTargetId="figure-113@122.[498,1534,234,1238]" captionTargetPageId="122" captionText="Figure 102 Temnothorax augusti. (A–C) Lectotype worker (MCZENT00021005). (A) Full-face view. (B) Profile view. (C) Dorsal view. (D-F) Gyne (MCZENT00577105). (D) Full-face view. (E) Profile view. (F) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-102" figureDoi="http://doi.org/10.5281/zenodo.5102295" httpUri="https://zenodo.org/record/5102295/files/figure.png" pageId="121" pageNumber="122">Fig. 102</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection box="[498,1436,1832,1859]" pageId="121" pageNumber="122" type="reference_group">
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<paragraph blockId="121.[498,1436,1832,1899]" box="[498,1436,1832,1859]" pageId="121" pageNumber="122">
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<treatmentCitationGroup box="[498,1436,1832,1859]" pageId="121" pageNumber="122">
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<treatmentCitation author="Forel A." box="[498,969,1832,1859]" page="129" pageId="121" pageNumber="122" year="1901">
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<taxonomicName authority="Forel, 1901 a: 129" authorityName="Forel" authorityPageNumber="129" authorityYear="1901" box="[498,969,1832,1859]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="petiolatus">
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<emphasis box="[498,910,1832,1858]" italics="true" pageId="121" pageNumber="122">
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Leptothorax petiolatus
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<bibRefCitation author="Forel A." box="[767,969,1832,1859]" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="121" pageNumber="122" pagination="123 - 141" part="45" refId="ref271024" refString="Forel A. 1901 a. I. Fourmis mexicaines recoltees par M. le professeur W. - M. Wheeler. II. A propos de la classification des fourmis. Annales de la Societe Entomologique de Belgique 45: 123 - 141." title="I. Fourmis mexicaines recoltees par M. le professeur W. - M. Wheeler. II. A propos de la classification des fourmis" type="journal article" year="1901">
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<author box="[767,828,1832,1858]" fns="*" inm="true" npo="*" pageId="121" pageNumber="122">Forel</author>
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,
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<year box="[840,895,1832,1858]" pageId="121" pageNumber="122">1901</year>
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a: 129
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</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus box="[981,1073,1832,1858]" pageId="121" pageNumber="122">Syntype</typeStatus>
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workers.
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<location LSID="urn:lsid:plazi:treatment:054FDB70FFBEFFA5B49C1CA17A5F22E7:88393CBDFFBEFFA3B1C91D357C002015" box="[1191,1330,1832,1858]" country="Mexico" name="Cuernavaca" pageId="121" pageNumber="122">Cuernavaca</location>
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,
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<collectingCountry box="[1342,1431,1832,1858]" name="Mexico" pageId="121" pageNumber="122">Mexico</collectingCountry>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection box="[498,954,1872,1899]" pageId="121" pageNumber="122" type="materials_examined">
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<paragraph blockId="121.[498,1436,1832,1899]" box="[498,954,1872,1899]" pageId="121" pageNumber="122">
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One worker here designated
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<emphasis bold="true" box="[839,948,1872,1898]" pageId="121" pageNumber="122">
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<typeStatus box="[839,948,1872,1898]" pageId="121" pageNumber="122">lectotype</typeStatus>
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</emphasis>
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.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.5102295" ID-Zenodo-Dep="5102295" httpUri="https://zenodo.org/record/5102295/files/figure.png" pageId="122" pageNumber="123" startId="122.[524,589,1266,1288]" targetBox="[498,1541,234,1253]" targetPageId="122">
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<paragraph blockId="122.[524,1516,1266,1347]" pageId="122" pageNumber="123">
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<emphasis bold="true" box="[524,863,1266,1288]" pageId="122" pageNumber="123">
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Figure 102
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<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[644,857,1266,1288]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="augusti">
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<emphasis bold="true" box="[644,857,1266,1288]" italics="true" pageId="122" pageNumber="123">Temnothorax augusti</emphasis>
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</taxonomicName>
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||
.
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</emphasis>
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||
(A–C) Lectotype worker (MCZENT00021005). (A) Full-face view. (B) Profile view. (C) Dorsal view. (D-F) Gyne (MCZENT00577105). (D) Full-face view. (E) Profile view. (F) Dorsal view. Scale bars 0.2 mm. Full-size DOI: 10.7717/peerj.11514/fig-102
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||
</paragraph>
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||
</caption>
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||
<subSubSection pageId="122" pageNumber="123" type="reference_group">
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||
<paragraph blockId="122.[498,1542,1402,1707]" pageId="122" pageNumber="123">
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||
<treatmentCitationGroup pageId="122" pageNumber="123">
|
||
<taxonomicName authority="(Forel)" baseAuthorityName="Forel" baseAuthorityYear="1901" box="[498,853,1402,1429]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="petiolata">
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<emphasis box="[498,762,1402,1428]" italics="true" pageId="122" pageNumber="123">Macromischa petiolata</emphasis>
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(Forel)
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</taxonomicName>
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:
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<treatmentCitation author="Emery C." box="[865,1068,1402,1428]" page="247" pageId="122" pageNumber="123" year="1924">
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||
<bibRefCitation author="Emery C." box="[865,1068,1402,1428]" journalOrPublisher="Subfam. Myrmicinae. Genera Insectorum" pageId="122" pageNumber="123" pagination="207 - 397" part="174" refId="ref270715" refString="Emery C. 1924. Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. Genera Insectorum 174 C: 207 - 397." title="Hymenoptera. Fam. Formicidae" type="journal article" year="1924">
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||
<emphasis box="[865,1009,1402,1428]" italics="true" pageId="122" pageNumber="123">
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||
<author box="[865,941,1403,1428]" fns="*" inm="true" npo="*" pageId="122" pageNumber="123">Emery</author>
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,
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<year box="[954,1009,1402,1428]" pageId="122" pageNumber="123">1924</year>
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||
</emphasis>
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||
: 247
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||
</bibRefCitation>
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||
</treatmentCitation>
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||
. First combination in
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<taxonomicName authorityName="Roger" authorityYear="1863" box="[1328,1484,1402,1428]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="genus">
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<emphasis box="[1328,1484,1402,1428]" italics="true" pageId="122" pageNumber="123">Macromischa</emphasis>
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</taxonomicName>
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(nec
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<treatmentCitation author="Mayr G." box="[498,948,1442,1469]" page="15" pageId="122" pageNumber="123" year="1868">
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<taxonomicName authority="Mayr, 1868: 15" authorityName="Mayr" authorityPageNumber="15" authorityYear="1868" box="[498,948,1442,1469]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="petiolata">
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<emphasis box="[498,906,1442,1468]" italics="true" pageId="122" pageNumber="123">
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Macromischa petiolata
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<bibRefCitation author="Mayr G." box="[772,948,1442,1469]" journalOrPublisher="Beitrage zur Naturkunde Preussens" pageId="122" pageNumber="123" pagination="1 - 102" part="1" refId="ref271911" refString="Mayr G. 1868. Die Ameisen des baltischen Bernsteins. Beitrage zur Naturkunde Preussens 1: 1 - 102." title="Die Ameisen des baltischen Bernsteins" type="journal article" year="1868">
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<author box="[772,838,1443,1468]" fns="*" inm="true" npo="*" pageId="122" pageNumber="123">Mayr</author>
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,
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<year box="[851,906,1442,1468]" pageId="122" pageNumber="123">1868</year>
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: 15
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</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</treatmentCitation>
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(fossil species)).
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</treatmentCitationGroup>
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||
</paragraph>
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<paragraph blockId="122.[498,1542,1402,1707]" pageId="122" pageNumber="123">
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<treatmentCitationGroup pageId="122" pageNumber="123">
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||
<treatmentCitation author="Aguayo CG" box="[498,947,1481,1508]" page="182" pageId="122" pageNumber="123" year="1931">
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<taxonomicName authority="Aguayo, 1931: 182" authorityName="Aguayo" authorityPageNumber="182" authorityYear="1931" box="[498,947,1481,1508]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="foreli">
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||
<emphasis box="[498,887,1481,1508]" italics="true" pageId="122" pageNumber="123">
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||
Macromischa foreli
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<bibRefCitation author="Aguayo CG" box="[730,947,1482,1508]" journalOrPublisher="Psyche: A Journal of Entomology" pageId="122" pageNumber="123" pagination="175 - 183" part="38" refId="ref269639" refString="Aguayo CG. 1931. New ants of the genus Macromischa. Psyche: A Journal of Entomology 38 (4): 175 - 183 DOI 10.1155 / 1931 / 72094." title="New ants of the genus Macromischa" type="journal article" year="1931">
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||
<author box="[730,820,1482,1508]" fns="*" inm="true" npo="*" pageId="122" pageNumber="123">Aguayo</author>
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||
,
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||
<year box="[832,887,1482,1508]" pageId="122" pageNumber="123">1931</year>
|
||
: 182
|
||
</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
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||
</treatmentCitation>
|
||
. Nomen novum for
|
||
<taxonomicName authority="(Forel)" baseAuthorityName="Forel" baseAuthorityYear="1901" box="[1189,1542,1481,1508]" class="Insecta" family="Formicidae" genus="Macromischa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="petiolata">
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||
<emphasis box="[1189,1453,1481,1507]" italics="true" pageId="122" pageNumber="123">Macromischa petiolata</emphasis>
|
||
(Forel)
|
||
</taxonomicName>
|
||
(nec Mayr).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="122.[498,1542,1402,1707]" pageId="122" pageNumber="123">
|
||
<treatmentCitationGroup pageId="122" pageNumber="123">
|
||
<treatmentCitation author="Baroni Urbani C." box="[498,1032,1561,1588]" page="423" pageId="122" pageNumber="123" year="1978">
|
||
<taxonomicName authority="Baroni Urbani, 1978: 423" authorityName="Baroni Urbani" authorityPageNumber="423" authorityYear="1978" box="[498,1032,1561,1588]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[498,974,1561,1588]" italics="true" pageId="122" pageNumber="123">
|
||
Leptothorax augusti
|
||
<bibRefCitation author="Baroni Urbani C." box="[736,1032,1561,1588]" journalOrPublisher="Entomologica Basiliensia" pageId="122" pageNumber="123" pagination="395 - 618" part="3" refId="ref269776" refString="Baroni Urbani C. 1978. Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae). Entomologica Basiliensia 3: 395 - 618." title="Materiali per una revisione dei Leptothorax neotropicali appartenenti al sottogenere Macromischa Roger, n. comb. (Hymenoptera: Formicidae)" type="journal article" year="1978">
|
||
<author box="[736,907,1561,1587]" case="TC" fns="*" inm="true" npo="*" pageId="122" pageNumber="123">Baroni Urbani</author>
|
||
,
|
||
<year box="[919,974,1562,1588]" pageId="122" pageNumber="123">1978</year>
|
||
: 423
|
||
</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
. Re-transfer to
|
||
<taxonomicName authority="Mayr." authorityName="Mayr." box="[1211,1351,1561,1587]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1211,1351,1561,1587]" italics="true" pageId="122" pageNumber="123">Leptothorax</emphasis>
|
||
</taxonomicName>
|
||
. Nomen novum for
|
||
<taxonomicName authority="(Aguayo)" baseAuthorityName="Aguayo" box="[541,866,1601,1627]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="foreli">
|
||
<emphasis box="[541,748,1601,1627]" italics="true" pageId="122" pageNumber="123">Leptothorax foreli</emphasis>
|
||
(Aguayo)
|
||
</taxonomicName>
|
||
(nec
|
||
<treatmentCitation author="Santschi F." box="[934,1376,1601,1628]" page="330" pageId="122" pageNumber="123" year="1907">
|
||
<taxonomicName authority="Santschi, 1907: 330" authorityName="Santschi" authorityPageNumber="330" authorityYear="1907" box="[934,1376,1601,1628]" class="Insecta" family="Formicidae" genus="Leptothorax" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="foreli">
|
||
<emphasis box="[934,1315,1601,1627]" italics="true" pageId="122" pageNumber="123">
|
||
Leptothorax foreli
|
||
<bibRefCitation author="Santschi F." box="[1150,1376,1601,1628]" journalOrPublisher="Revue Suisse de Zoologie" pageId="122" pageNumber="123" pagination="305 - 334" part="15" refId="ref272576" refString="Santschi F. 1907. Fourmis de Tunisie capturees en 1906. Revue Suisse de Zoologie 15: 305 - 334 DOI 10.5962 / bhl. part. 75189." title="Fourmis de Tunisie capturees en 1906" type="journal article" year="1907">
|
||
<author box="[1150,1248,1601,1627]" fns="*" inm="true" npo="*" pageId="122" pageNumber="123">Santschi</author>
|
||
,
|
||
<year box="[1260,1315,1601,1627]" pageId="122" pageNumber="123">1907</year>
|
||
: 330
|
||
</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph blockId="122.[498,1542,1402,1707]" pageId="122" pageNumber="123">
|
||
<treatmentCitationGroup pageId="122" pageNumber="123">
|
||
<taxonomicName authority="(Baroni Urbani)" baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[498,949,1641,1668]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[498,748,1641,1667]" italics="true" pageId="122" pageNumber="123">Temnothorax augusti</emphasis>
|
||
(Baroni Urbani)
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation author="Bolton B." box="[962,1168,1641,1668]" page="271" pageId="122" pageNumber="123" year="2003">
|
||
<bibRefCitation author="Bolton B." box="[962,1168,1641,1668]" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="122" pageNumber="123" pagination="370" part="71" refId="ref270076" refString="Bolton B. 2003. Synopsis and Classification of Formicidae. Memoirs of the American Entomological Institute 71: 370." title="Synopsis and Classification of Formicidae" type="journal article" year="2003">
|
||
<emphasis box="[962,1108,1641,1667]" italics="true" pageId="122" pageNumber="123">
|
||
<collectingRegion box="[962,1040,1641,1667]" country="United Kingdom" name="Bolton" pageId="122" pageNumber="123">
|
||
<author box="[962,1040,1641,1667]" fns="*" inm="true" npo="*" pageId="122" pageNumber="123">Bolton</author>
|
||
</collectingRegion>
|
||
,
|
||
<year box="[1053,1108,1641,1667]" pageId="122" pageNumber="123">2003</year>
|
||
</emphasis>
|
||
: 271
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
. First combination in
|
||
<taxonomicName authorityName="Mayr" authorityYear="1861" box="[498,655,1681,1707]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[498,655,1681,1707]" italics="true" pageId="122" pageNumber="123">Temnothorax</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="123" lastPageNumber="124" pageId="122" pageNumber="123" type="materials_examined">
|
||
<paragraph blockId="122.[498,1542,1743,1890]" pageId="122" pageNumber="123">
|
||
<emphasis bold="true" box="[530,834,1743,1770]" pageId="122" pageNumber="123">Type material examined:</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="3321861660" collectionCode="MCZC" collectorName="Menozzi C" country="Mexico" location="Morelos" pageId="122" pageNumber="123" specimenCode="MCZENT00021005" specimenCount="1" stateProvince="Morelos" typeStatus="lectotype">
|
||
<emphasis box="[843,1045,1743,1769]" italics="true" pageId="122" pageNumber="123">
|
||
<typeStatus box="[843,951,1744,1769]" pageId="122" pageNumber="123">Lectotype</typeStatus>
|
||
<specimenCount box="[959,1039,1743,1769]" pageId="122" pageNumber="123" type="worker">worker</specimenCount>
|
||
:
|
||
</emphasis>
|
||
<collectingCountry box="[1054,1169,1744,1770]" name="Mexico" pageId="122" pageNumber="123">MEXICO</collectingCountry>
|
||
:
|
||
<collectingRegion box="[1181,1280,1743,1769]" country="Mexico" name="Morelos" pageId="122" pageNumber="123">Morelos</collectingRegion>
|
||
:
|
||
<collectingRegion box="[1292,1431,1744,1770]" pageId="122" pageNumber="123">Cuernavaca</collectingRegion>
|
||
, [collection date unknown], [collector unspecified, but
|
||
<bibRefCitation author="Wheeler WM" box="[1132,1308,1783,1809]" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="122" pageNumber="123" pagination="1 - 34" part="72" refId="ref273428" refString="Wheeler WM. 1931. New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex. Bulletin of the Museum of Comparative Zoology 72: 1 - 34." title="New and little-known ants of the genera Macromischa, Creosomyrmex and Antillaemyrmex" type="journal article" year="1931">
|
||
<emphasis box="[1132,1308,1783,1809]" italics="true" pageId="122" pageNumber="123">Wheeler (1931)</emphasis>
|
||
</bibRefCitation>
|
||
states that he made the collection], in
|
||
<taxonomicName box="[711,827,1823,1849]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="122" pageNumber="123" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[711,827,1823,1849]" italics="true" pageId="122" pageNumber="123">Tillandsia</emphasis>
|
||
</taxonomicName>
|
||
, M.C.Z. cotype 21005 (
|
||
<specimenCode box="[1100,1335,1823,1850]" pageId="122" pageNumber="123">MCZENT00021005</specimenCode>
|
||
, top specimen on pin) [
|
||
<collectionCode box="[563,654,1862,1889]" pageId="122" pageNumber="123">MCZC</collectionCode>
|
||
].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="123.[498,1542,234,1935]" pageId="123" pageNumber="124">
|
||
<materialsCitation ID-GBIF-Occurrence="3321861713" collectionCode="MCZC" pageId="123" pageNumber="124" specimenCount="2" typeStatus="paralectotype">
|
||
<emphasis box="[530,773,234,260]" italics="true" pageId="123" pageNumber="124">
|
||
<typeStatus box="[530,684,234,260]" pageId="123" pageNumber="124">Paralectotype</typeStatus>
|
||
worker
|
||
</emphasis>
|
||
: same pin as
|
||
<typeStatus box="[933,1038,234,260]" pageId="123" pageNumber="124">lectotype</typeStatus>
|
||
(MCZENT00021005, bottom specimen on pin) [
|
||
<collectionCode box="[563,654,273,300]" pageId="123" pageNumber="124">MCZC</collectionCode>
|
||
]
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="123.[498,1542,234,1935]" pageId="123" pageNumber="124">
|
||
<emphasis bold="true" box="[530,888,314,340]" pageId="123" pageNumber="124">Non-type material examined:</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="3321861553" pageId="123" pageNumber="124">
|
||
same data as previous, 1 dealate gyne (MCZENT00577105) [
|
||
<collectionCode box="[764,855,353,380]" pageId="123" pageNumber="124">MCZC</collectionCode>
|
||
].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[530,1510,393,421]" pageId="123" pageNumber="124" type="distribution">
|
||
<paragraph blockId="123.[498,1542,234,1935]" box="[530,1510,393,421]" pageId="123" pageNumber="124">
|
||
<emphasis bold="true" box="[530,755,393,420]" pageId="123" pageNumber="124">Geographic range:</emphasis>
|
||
Mid elevations in South-Central
|
||
<collectingCountry box="[1150,1236,394,420]" name="Mexico" pageId="123" pageNumber="124">Mexico</collectingCountry>
|
||
(
|
||
<collectingRegion box="[1253,1353,394,421]" country="Mexico" name="Morelos" pageId="123" pageNumber="124">Morelos</collectingRegion>
|
||
) (
|
||
<figureCitation box="[1380,1494,394,420]" captionStart="Figure 101" captionStartId="121.[524,589,1194,1216]" captionTargetBox="[498,1541,234,1181]" captionTargetId="figure-149@121.[498,1534,234,1166]" captionTargetPageId="121" captionText="Figure 101 Geographical and elevational distribution of the augusti group. (A) Temnothorax augusti (B) T. aureus sp. nov. (C) T. leucacanthoides sp. nov. (D) T. leucacanthus. Colored scale to the left of each map represents elevation in meters. Points in black represent the species named in each subfigure, while points in red represent other members of the species group. Type localities are represented by triangles, non-type localities are represented by circles. Bounding box in inset map shows location of main map. Full-size DOI: 10.7717/peerj.11514/fig-101" figureDoi="http://doi.org/10.5281/zenodo.5102293" httpUri="https://zenodo.org/record/5102293/files/figure.png" pageId="123" pageNumber="124">Fig. 101A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="123" pageNumber="124" type="diagnosis">
|
||
<paragraph blockId="123.[498,1542,234,1935]" pageId="123" pageNumber="124">
|
||
<emphasis bold="true" box="[530,754,433,460]" pageId="123" pageNumber="124">Worker diagnosis:</emphasis>
|
||
The following character combination separates
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis italics="true" pageId="123" pageNumber="124">Temnothorax augusti</emphasis>
|
||
</taxonomicName>
|
||
from all other species in the
|
||
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[922,997,473,499]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis box="[922,997,473,499]" italics="true" pageId="123" pageNumber="124">salvini</emphasis>
|
||
</taxonomicName>
|
||
clade: moderately long antennal scapes, which surpass the posterior margin of the head by about the maximum width of the scape (SI 103–117); mesosoma about one and a half times as long as the width of the head (WLI 143–150); propodeal spines shorter than the length of the propodeal declivity (PSI 29); petiolar node subquadrate, and overhangs the caudal cylinder of the petiole; postpetiole narrow, less than two times the width of the petiole in dorsal view (PWI 187–190); first gastral tergite with very faint areolate sculpture; erect setae on the dorsal surface of the head; integument medium brown.
|
||
</paragraph>
|
||
<paragraph blockId="123.[498,1542,234,1935]" pageId="123" pageNumber="124">
|
||
<emphasis bold="true" box="[530,719,792,818]" pageId="123" pageNumber="124">Similar species:</emphasis>
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[727,1046,792,818]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
|
||
<emphasis box="[727,1046,792,818]" italics="true" pageId="123" pageNumber="124">Temnothorax acutispinosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1055,1148,793,819]" pageId="123" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<emphasis italics="true" pageId="123" pageNumber="124">
|
||
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1161,1313,792,818]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="subditivus">T. subditivus</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1326,1512,792,818]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="tenuisculptus">T. tenuisculptus</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[498,640,832,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">T. tuxtlanus</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel box="[649,742,832,858]" pageId="123" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, species of the
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[921,1017,833,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="annexus">
|
||
<emphasis box="[921,1017,833,858]" italics="true" pageId="123" pageNumber="124">annexus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[1032,1116,833,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[1032,1116,833,858]" italics="true" pageId="123" pageNumber="124">augusti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[1131,1223,832,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="fuscatus">
|
||
<emphasis box="[1131,1223,832,858]" italics="true" pageId="123" pageNumber="124">fuscatus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1292,1367,832,858]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis box="[1292,1367,832,858]" italics="true" pageId="123" pageNumber="124">salvini</emphasis>
|
||
</taxonomicName>
|
||
groups.
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[498,747,872,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[498,747,872,898]" italics="true" pageId="123" pageNumber="124">Temnothorax augusti</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from the first three species by the shape of the dorsal margin of the mesosoma in profile, which is predominantly flat: in
|
||
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[1346,1492,912,938]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="subditivus">
|
||
<emphasis box="[1346,1492,912,938]" italics="true" pageId="123" pageNumber="124">T. subditivus</emphasis>
|
||
</taxonomicName>
|
||
, the mesosoma is strongly arched, while
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[925,1109,951,977]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="tenuisculptus">
|
||
<emphasis box="[925,1109,951,977]" italics="true" pageId="123" pageNumber="124">T. tenuisculptus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1170,1311,951,977]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">
|
||
<emphasis box="[1170,1311,951,977]" italics="true" pageId="123" pageNumber="124">T. tuxtlanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1320,1411,952,978]" pageId="123" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
have a sinuate profile. Petiolar node shape is also a useful character for separating
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[1363,1478,992,1017]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[1363,1478,992,1017]" italics="true" pageId="123" pageNumber="124">T. augusti</emphasis>
|
||
</taxonomicName>
|
||
from the above taxa:
|
||
<taxonomicName baseAuthorityName="Wheeler" baseAuthorityYear="1903" box="[675,821,1031,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="subditivus">
|
||
<emphasis box="[675,821,1031,1057]" italics="true" pageId="123" pageNumber="124">T. subditivus</emphasis>
|
||
</taxonomicName>
|
||
and the potentially co-occurring members of the
|
||
<taxonomicName baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1391,1466,1031,1057]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="salvini">
|
||
<emphasis box="[1391,1466,1031,1057]" italics="true" pageId="123" pageNumber="124">salvini</emphasis>
|
||
</taxonomicName>
|
||
group have a squamiform petiolar node, which is much broader than the peduncle in dorsal view (only slightly wider in
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[824,940,1112,1137]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[824,940,1112,1137]" italics="true" pageId="123" pageNumber="124">T. augusti</emphasis>
|
||
</taxonomicName>
|
||
), and the nodes of the
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1931" box="[1211,1307,1112,1137]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="annexus">
|
||
<emphasis box="[1211,1307,1112,1137]" italics="true" pageId="123" pageNumber="124">annexus</emphasis>
|
||
</taxonomicName>
|
||
group are erect and subquadrate, as opposed to leaning posteriorly in
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[1087,1204,1152,1177]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[1087,1204,1152,1177]" italics="true" pageId="123" pageNumber="124">T. augusti</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1978" box="[1219,1403,1151,1177]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="tenuisculptus">
|
||
<emphasis box="[1219,1403,1151,1177]" italics="true" pageId="123" pageNumber="124">T. tenuisculptus</emphasis>
|
||
</taxonomicName>
|
||
, like
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[498,613,1191,1216]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[498,613,1191,1216]" italics="true" pageId="123" pageNumber="124">T. augusti</emphasis>
|
||
</taxonomicName>
|
||
, has a subquadrate petiolar node, but it does not lean posteriorly over the caudal cylinder;
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[605,745,1230,1256]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">
|
||
<emphasis box="[605,745,1230,1256]" italics="true" pageId="123" pageNumber="124">T. tuxtlanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[751,840,1231,1257]" pageId="123" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[897,1080,1231,1256]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="acutispinosus" status="sp. nov.">
|
||
<emphasis box="[897,1080,1231,1256]" italics="true" pageId="123" pageNumber="124">T. acutispinosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1087,1176,1231,1257]" pageId="123" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
have rounded petiolar nodes in profile view.
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[651,900,1270,1296]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[651,900,1270,1296]" italics="true" pageId="123" pageNumber="124">Temnothorax augusti</emphasis>
|
||
</taxonomicName>
|
||
can also be distinguished from
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1278,1420,1270,1296]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="tuxtlanus" status="sp. nov.">
|
||
<emphasis box="[1278,1420,1270,1296]" italics="true" pageId="123" pageNumber="124">T. tuxtlanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1429,1520,1271,1297]" pageId="123" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authorityName="Prebus" authorityYear="2021" box="[551,737,1311,1336]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="acutispinosus">
|
||
<emphasis box="[551,737,1311,1336]" italics="true" pageId="123" pageNumber="124">T. acutispinosus</emphasis>
|
||
</taxonomicName>
|
||
sp nov. by the presence of erect setae on the dorsum of the propodeum. The potentially co-occurring members of the
|
||
<taxonomicName baseAuthorityName="Mann" baseAuthorityYear="1920" box="[1188,1280,1350,1376]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="fuscatus">
|
||
<emphasis box="[1188,1280,1350,1376]" italics="true" pageId="123" pageNumber="124">fuscatus</emphasis>
|
||
</taxonomicName>
|
||
group have a low, elongate petiolar node in profile view. Finally,
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[1047,1164,1391,1416]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[1047,1164,1391,1416]" italics="true" pageId="123" pageNumber="124">T. augusti</emphasis>
|
||
</taxonomicName>
|
||
can be separated from fellow members of the
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[692,776,1431,1456]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[692,776,1431,1456]" italics="true" pageId="123" pageNumber="124">augusti</emphasis>
|
||
</taxonomicName>
|
||
group by the combination of short propodeal spines, which are shorter than the length of the propodeal declivity (as long as or longer than the declivity in other
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[569,653,1510,1535]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[569,653,1510,1535]" italics="true" pageId="123" pageNumber="124">augusti</emphasis>
|
||
</taxonomicName>
|
||
group members, except
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[945,1091,1510,1535]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="casanovai" status="sp. nov.">
|
||
<emphasis box="[945,1091,1510,1535]" italics="true" pageId="123" pageNumber="124">T. casanovai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1100,1194,1510,1536]" pageId="123" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), and the shining, faintly areolate first gastral tergite (which is strongly areolate and dull in
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[1278,1424,1550,1575]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="casanovai" status="sp. nov.">
|
||
<emphasis box="[1278,1424,1550,1575]" italics="true" pageId="123" pageNumber="124">T. casanovai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1433,1528,1550,1576]" pageId="123" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="127" lastPageNumber="128" pageId="123" pageNumber="124" type="description">
|
||
<paragraph blockId="123.[498,1542,234,1935]" lastBlockId="124.[498,1542,234,1934]" lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124">
|
||
<emphasis bold="true" box="[530,1018,1589,1615]" pageId="123" pageNumber="124">
|
||
Worker measurements & indices (
|
||
<emphasis bold="true" box="[941,957,1589,1615]" italics="true" pageId="123" pageNumber="124">n</emphasis>
|
||
= 2):
|
||
</emphasis>
|
||
SL =
|
||
<geoCoordinate box="[1084,1146,1589,1616]" degrees="0.605" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.605">0.605</geoCoordinate>
|
||
<geoCoordinate box="[1146,1223,1589,1615]" degrees="0.662" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.662">–0.662</geoCoordinate>
|
||
(0.634); FRS =
|
||
<geoCoordinate box="[1402,1464,1589,1615]" degrees="0.183" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.183">0.183</geoCoordinate>
|
||
<geoCoordinate box="[1464,1541,1589,1615]" degrees="0.190" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.19">–0.190</geoCoordinate>
|
||
(0.187); CW =
|
||
<geoCoordinate box="[678,740,1629,1655]" degrees="0.624" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.624">0.624</geoCoordinate>
|
||
<geoCoordinate box="[740,817,1629,1655]" degrees="0.630" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.63">–0.630</geoCoordinate>
|
||
(0.627); CWb =
|
||
<geoCoordinate box="[1020,1082,1629,1656]" degrees="0.566" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.566">0.566</geoCoordinate>
|
||
<geoCoordinate box="[1082,1159,1629,1656]" degrees="0.588" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.588">–0.588</geoCoordinate>
|
||
(0.577); PoOC =
|
||
<geoCoordinate box="[1372,1434,1629,1655]" degrees="0.278" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.278">0.278</geoCoordinate>
|
||
<geoCoordinate box="[1434,1511,1629,1655]" degrees="0.297" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.297">–0.297</geoCoordinate>
|
||
(0.288); CL =
|
||
<geoCoordinate box="[665,727,1669,1695]" degrees="0.693" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.693">0.693</geoCoordinate>
|
||
<geoCoordinate box="[727,804,1669,1695]" degrees="0.737" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.737">–0.737</geoCoordinate>
|
||
(0.715); EL =
|
||
<geoCoordinate box="[977,1039,1669,1696]" degrees="0.156" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.156">0.156</geoCoordinate>
|
||
<geoCoordinate box="[1039,1116,1669,1695]" degrees="0.160" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.16">–0.160</geoCoordinate>
|
||
(0.158); EW =
|
||
<geoCoordinate box="[1302,1364,1669,1695]" degrees="0.118" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.118">0.118</geoCoordinate>
|
||
<geoCoordinate box="[1364,1441,1669,1696]" degrees="0.153" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.153">–0.153</geoCoordinate>
|
||
(0.136); MD =
|
||
<geoCoordinate box="[580,642,1709,1735]" degrees="0.162" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.162">0.162</geoCoordinate>
|
||
<geoCoordinate box="[642,719,1709,1735]" degrees="0.173" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.173">–0.173</geoCoordinate>
|
||
(0.168); WL =
|
||
<geoCoordinate box="[904,966,1709,1735]" degrees="0.812" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.812">0.812</geoCoordinate>
|
||
<geoCoordinate box="[966,1043,1709,1735]" degrees="0.884" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.884">–0.884</geoCoordinate>
|
||
(0.848); SPST =
|
||
<geoCoordinate box="[1245,1307,1709,1735]" degrees="0.239" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.239">0.239</geoCoordinate>
|
||
<geoCoordinate box="[1308,1385,1708,1735]" degrees="0.252" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.252">–0.252</geoCoordinate>
|
||
(0.246); MPST =
|
||
<geoCoordinate box="[607,669,1749,1775]" degrees="0.247" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.247">0.247</geoCoordinate>
|
||
<geoCoordinate box="[669,746,1748,1775]" degrees="0.263" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.263">–0.263</geoCoordinate>
|
||
(0.255); PEL =
|
||
<geoCoordinate box="[935,997,1748,1774]" degrees="0.306" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.306">0.306</geoCoordinate>
|
||
<geoCoordinate box="[997,1074,1749,1775]" degrees="0.340" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.34">–0.340</geoCoordinate>
|
||
(0.323); NOL =
|
||
<geoCoordinate box="[1274,1336,1749,1775]" degrees="0.177" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.177">0.177</geoCoordinate>
|
||
<geoCoordinate box="[1336,1413,1749,1775]" degrees="0.212" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.212">–0.212</geoCoordinate>
|
||
(0.195); NOH =
|
||
<geoCoordinate box="[598,660,1788,1814]" degrees="0.116" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.116">0.116</geoCoordinate>
|
||
<geoCoordinate box="[660,737,1788,1814]" degrees="0.124" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.124">–0.124</geoCoordinate>
|
||
(0.120); PEH =
|
||
<geoCoordinate box="[933,995,1788,1814]" degrees="0.208" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.208">0.208</geoCoordinate>
|
||
<geoCoordinate box="[995,1072,1788,1815]" degrees="0.225" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.225">–0.225</geoCoordinate>
|
||
(0.217); PPL =
|
||
<geoCoordinate box="[1262,1324,1788,1814]" degrees="0.137" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.137">0.137</geoCoordinate>
|
||
<geoCoordinate box="[1324,1401,1788,1815]" degrees="0.153" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.153">–0.153</geoCoordinate>
|
||
(0.145); PPH =
|
||
<geoCoordinate box="[588,650,1828,1854]" degrees="0.186" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.186">0.186</geoCoordinate>
|
||
<geoCoordinate box="[650,727,1828,1854]" degrees="0.193" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.193">–0.193</geoCoordinate>
|
||
(0.190); PW =
|
||
<geoCoordinate box="[912,974,1828,1854]" degrees="0.383" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.383">0.383</geoCoordinate>
|
||
<geoCoordinate box="[975,1052,1828,1854]" degrees="0.418" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.418">–0.418</geoCoordinate>
|
||
(0.401); SBPA =
|
||
<geoCoordinate box="[1260,1322,1828,1854]" degrees="0.144" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.144">0.144</geoCoordinate>
|
||
<geoCoordinate box="[1322,1399,1828,1855]" degrees="0.152" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.152">–0.152</geoCoordinate>
|
||
(0.148); SPTI =
|
||
<geoCoordinate box="[590,652,1868,1894]" degrees="0.161" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.161">0.161</geoCoordinate>
|
||
<geoCoordinate box="[653,730,1868,1894]" degrees="0.180" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.18">–0.180</geoCoordinate>
|
||
(0.171); PEW =
|
||
<geoCoordinate box="[932,994,1868,1894]" degrees="0.128" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.128">0.128</geoCoordinate>
|
||
<geoCoordinate box="[994,1071,1868,1894]" degrees="0.129" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.129">–0.129</geoCoordinate>
|
||
(0.129); PNW =
|
||
<geoCoordinate box="[1278,1340,1868,1894]" degrees="0.172" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.172">0.172</geoCoordinate>
|
||
<geoCoordinate box="[1340,1417,1868,1894]" degrees="0.184" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.184">–0.184</geoCoordinate>
|
||
(0.178); PPW =
|
||
<geoCoordinate box="[590,652,1908,1934]" degrees="0.239" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.239">0.239</geoCoordinate>
|
||
<geoCoordinate box="[653,730,1908,1935]" degrees="0.245" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.245">–0.245</geoCoordinate>
|
||
(0.242); HFL =
|
||
<geoCoordinate box="[917,979,1908,1934]" degrees="0.681" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.681">0.681</geoCoordinate>
|
||
<geoCoordinate box="[979,1056,1908,1934]" degrees="0.684" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.684">–0.684</geoCoordinate>
|
||
(0.683); HFWmax =
|
||
<geoCoordinate box="[1306,1368,1908,1935]" degrees="0.155" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="55" value="0.155">0.155</geoCoordinate>
|
||
<geoCoordinate box="[1368,1445,1908,1934]" degrees="0.161" direction="west" orientation="longitude" pageId="123" pageNumber="124" precision="55" value="-0.161">–0.161</geoCoordinate>
|
||
(0.158); HFWmin =
|
||
<geoCoordinate box="[645,707,234,260]" degrees="0.046" direction="north" orientation="latitude" pageId="124" pageNumber="125" precision="55" value="0.046">0.046</geoCoordinate>
|
||
<geoCoordinate box="[708,785,234,260]" degrees="0.049" direction="west" orientation="longitude" pageId="124" pageNumber="125" precision="55" value="-0.049">–0.049</geoCoordinate>
|
||
(0.048); CS =
|
||
<geoCoordinate box="[958,1020,234,260]" degrees="0.913" direction="north" orientation="latitude" pageId="124" pageNumber="125" precision="55" value="0.913">0.913</geoCoordinate>
|
||
<geoCoordinate box="[1021,1098,234,261]" degrees="0.957" direction="west" orientation="longitude" pageId="124" pageNumber="125" precision="55" value="-0.957">–0.957</geoCoordinate>
|
||
(0.935); ES =
|
||
<geoCoordinate box="[1269,1331,234,261]" degrees="0.215" direction="north" orientation="latitude" pageId="124" pageNumber="125" precision="55" value="0.215">0.215</geoCoordinate>
|
||
<geoCoordinate box="[1331,1408,234,260]" degrees="0.237" direction="west" orientation="longitude" pageId="124" pageNumber="125" precision="55" value="-0.237">–0.237</geoCoordinate>
|
||
(0.226); SI = 103–117 (110); OI = 24–25 (24); CI = 80–82 (81); WLI = 143–150 (147); SBI = 25–26 (26); PSI = 29; PWI = 187–190 (188); PLI = 222–223 (223); NI = 153–171 (162); PNWI = 134–143 (139); NLI = 58–62 (60); FI = 329–337 (333).
|
||
</paragraph>
|
||
<paragraph blockId="124.[498,1542,234,1934]" pageId="124" pageNumber="125">
|
||
<emphasis bold="true" box="[530,775,393,420]" pageId="124" pageNumber="125">Worker description:</emphasis>
|
||
In full-face view, head subquadrate, longer than broad (CI 80–82). Mandibles densely, finely striate, weakly shining and armed with five teeth: the apical-most well developed and acute, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin weakly emarginated medially. Antennal scapes moderately long: when fully retracted, surpassing the posterior margin of the head capsule by about the maximum width of the scape (SI 103–117). Antennae 12-segmented; antennal club of composed of three segments, with the apical-most segment longer than the preceding two in combination. Frontal carinae moderately long, extending past the antennal toruli by about two and a half times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head convex, forming a continuous arc from the mandibular insertions to the posterior margin of the head. Posterior head margin convex but rounding evenly into the lateral margins.
|
||
</paragraph>
|
||
<paragraph blockId="124.[498,1542,234,1934]" pageId="124" pageNumber="125">
|
||
In profile view, compound eyes ovular and moderately large (OI 24–25), with 11 ommatidia in longest row. Pronotal declivity indistinct, but neck and anterior face of pronotum forming a ~120
|
||
<emphasis box="[809,819,987,1005]" italics="true" pageId="124" pageNumber="125">
|
||
<superScript attach="left" box="[809,819,987,1005]" fontSize="8" pageId="124" pageNumber="125">°</superScript>
|
||
</emphasis>
|
||
angle. Mesosoma convex from where it joins the pronotal neck to the pronotal dorsum, then flat to the base of the propodeal spines. Promesonotal suture extending from the posterior margin of the procoxal insertion only to the mesothoracic spiracle, which is moderately well developed. Metanotal groove visible as a disruption of the sculpture laterally from where it arises between the mid- and hind coxae to where it ends in the poorly developed metathoracic spiracle, which is nearly indistinguishable against the ground sculpture, continuing dorsally as a disruption in the sculpture. Propodeal spiracle poorly developed, directed posterolaterally, and separated from the propodeal declivity by about four and a half spiracle diameters. Propodeal spines moderately well developed, but short (PSI 29), slightly shorter than the propodeal declivity, tapering evenly from the base, straight, and acute. Propodeal declivity straight, forming a rounded ~120
|
||
<emphasis box="[665,675,1425,1443]" italics="true" pageId="124" pageNumber="125">
|
||
<superScript attach="left" box="[665,675,1425,1443]" fontSize="8" pageId="124" pageNumber="125">°</superScript>
|
||
</emphasis>
|
||
angle with the base of the propodeal spines. Propodeal lobes rounded and weakly developed. Metapleural gland bulla small, extending from the metacoxal insertion a third of the way to the propodeal spiracle. Petiole long (PLI 222–223), without tubercles anterodorsally. Subpetiolar process in the form of a very small, acute tooth; ventral margin of petiole weakly bulging medially. Petiolar peduncle long: comprising about two thirds of the petiole. Petiolar node robust and erect, nearly squamiform: transition between peduncle and node marked by a rounded angle of ~120
|
||
<emphasis box="[1364,1374,1664,1682]" italics="true" pageId="124" pageNumber="125">
|
||
<superScript attach="left" box="[1364,1374,1664,1682]" fontSize="8" pageId="124" pageNumber="125">°</superScript>
|
||
</emphasis>
|
||
, resulting in a concave anterior node face; anterior face forming a ~100
|
||
<emphasis box="[1169,1179,1704,1722]" italics="true" pageId="124" pageNumber="125">
|
||
<superScript attach="left" box="[1169,1179,1704,1722]" fontSize="8" pageId="124" pageNumber="125">°</superScript>
|
||
</emphasis>
|
||
angle with the dorsal face, which is weakly convex; dorsal face rounding evenly into the short posterior face, which forms a ~85
|
||
<emphasis box="[641,651,1784,1802]" italics="true" pageId="124" pageNumber="125">
|
||
<superScript attach="left" box="[641,651,1784,1802]" fontSize="8" pageId="124" pageNumber="125">°</superScript>
|
||
</emphasis>
|
||
angle with the caudal cylinder, overhanging it. Postpetiole evenly rounded anteriorly, flattened dorsally, and weakly lobed ventrally.
|
||
</paragraph>
|
||
<paragraph blockId="124.[498,1542,234,1934]" lastBlockId="125.[498,1542,234,1934]" lastPageId="125" lastPageNumber="126" pageId="124" pageNumber="125">In dorsal view, humeri developed: evenly rounded and wider than the rest of the mesosoma; mesothoracic spiracles not protruding past the lateral margins of the mesosoma. Metanotal groove absent: mesonotum and propodeum completely fused and lateral margins converging evenly to the bases of the propodeal spines. Propodeal spines narrowly approximated basally, joined by a transverse welt, and diverging apically, their apices separated from each other by about their length, the negative space between them “U” shaped. Petiolar peduncle with spiracles protruding past the lateral margins, broadened where they arise. Petiolar node campaniform: flattened posteriorly. Petiolar node broader than the peduncle, and about one and a quarter times as wide as the caudal cylinder. Postpetiole narrow (PWI 187–190) and campaniform, articulating with most of the anterior margin of the gaster, leaving small, angulate margins on each side exposed. Anterior margin of the postpetiole convex and evenly rounds into the lateral margins, which diverge to the angulate posterior corners; posterior margin flat. Metafemur strongly incrassate (FI 329–337).</paragraph>
|
||
<paragraph blockId="125.[498,1542,234,1934]" pageId="125" pageNumber="126">Sculpture: median clypeal carina present, extending posteriorly nearly to the frontal triangle, and flanked on either side by two slightly weaker carinae. Lateral clypeal lobes with additional, weaker carinae; ground sculpture weakly areolate. Antennal scapes areolate. Cephalic dorsum areolate; costulae flanking the frontal carinae, and weak concentric costulae surrounding the antennal insertions. Lateral surfaces of head areolate, and weak rugulose sculpture between the compound eye and the mandibular insertion. Ventral surface of head weakly areolate. Mesosoma areolate-rugulose, with weak rugulae overlying it dorsally. Femora weakly areolate. Petiole and postpetiole predominantly smooth and shining, with traces of weak areolate sculpture. Gaster smooth and shining, with traces of areolate sculpture and spectral iridescence on the basal half of the first gastral tergite.</paragraph>
|
||
<paragraph blockId="125.[498,1542,234,1934]" pageId="125" pageNumber="126">Setae: antennal scapes and funiculi with short, decumbent pilosity. Dorsum of the head, pronotum, waist segments, and gaster with moderately abundant, erect, blunt-tipped setae, the longest of which are about the width of the compound eye. The head bears ~40, mesosoma ~26, petiole 6, postpetiole ~10, and first gastral tergite ~40 setae. Short, sparse pubescence present over the entire body, but difficult to detect against the ground sculpture.</paragraph>
|
||
<paragraph blockId="125.[498,1542,234,1934]" box="[530,1225,1390,1416]" pageId="125" pageNumber="126">Color: predominantly medium brown, with yellowish tarsi.</paragraph>
|
||
<paragraph blockId="125.[498,1542,234,1934]" pageId="125" pageNumber="126">
|
||
<emphasis bold="true" box="[530,1001,1429,1456]" pageId="125" pageNumber="126">
|
||
Gyne measurements & indices (
|
||
<emphasis bold="true" box="[919,935,1430,1456]" italics="true" pageId="125" pageNumber="126">n</emphasis>
|
||
= 1):
|
||
</emphasis>
|
||
SL = 0.746; FRS = 0.288; CW = 0.874; CWb = 0.819; PoOC = 0.361; CL = 0.931; EL = 0.255; EW = 0.197; MD = 0.204; WL = 1.437; SPST = 0.323; MPST = 0.416; PEL = 0.483; NOL = 0.224; NOH = 0.216; PEH = 0.373; PPL = 0.279; PPH = 0.311; PW = 0.859; SBPA = 0.371; SPTI = 0.378; PEW = 0.213; PNW = 0.300; PPW = 0.432; HFL = 0.900; HFWmax = 0.191; HFWmin = 0.070; CS = 1.285; ES = 0.354; SI = 91; OI = 28; CI = 88; WLI = 175; SBI = 45; PSI = 22; PWI = 203; PLI = 173; NI = 104; PNWI = 141; NLI = 46; FI = 273.
|
||
</paragraph>
|
||
<paragraph blockId="125.[498,1542,234,1934]" lastBlockId="126.[498,1542,234,1934]" lastPageId="126" lastPageNumber="127" pageId="125" pageNumber="126">
|
||
<emphasis bold="true" box="[530,747,1708,1734]" pageId="125" pageNumber="126">Gyne description:</emphasis>
|
||
In full-face view, head subquadrate, longer than broad (CI 88). Mandibles densely, finely striate but weakly shining and armed with five teeth: the apical-most well developed, followed by a less developed preapical tooth and three equally developed smaller teeth. Anterior clypeal margin weakly emarginated medially. Antennal scapes moderately long: when fully retracted, just reaching the posterior margin of the head capsule (SI 91). Antennae 12-segmented; antennal club composed of three segments, with the apical-most segment longer than the preceding two in combination. Frontal carinae long, extending past the antennal toruli by about three times the maximum width of the antennal scape. Compound eyes moderately protruding past the lateral margins of the head capsule. Lateral margin of head evenly convex, converging from below the compound eyes to the mandibular insertions. Posterior head margin convex, rounding evenly into the lateral margins.
|
||
</paragraph>
|
||
<paragraph blockId="126.[498,1542,234,1934]" pageId="126" pageNumber="127">
|
||
In profile view, compound eyes ovular and large (OI 28), with 22 ommatidia in longest row. Mesoscutum rounded evenly anteriorly, covering the dorsal surface of the pronotum, and flat dorsally. Mesoscutellum at the same level as the mesoscutum. Posterior margin of metanotum extending slightly past the posterior margin of the mesoscutum. Propodeal spiracle poorly developed, directed posterolaterally, and separated from the propodeal declivity by about five spiracle diameters. Propodeal spines stout and well developed, but short (PSI 22), about half as long as the propodeal declivity, tapering evenly from the base, directed posteriorly, straight, and blunt. Propodeal declivity straight and flat, forming a rounded ~100
|
||
<emphasis box="[663,673,787,805]" italics="true" pageId="126" pageNumber="127">
|
||
<superScript attach="left" box="[663,673,787,805]" fontSize="8" pageId="126" pageNumber="127">°</superScript>
|
||
</emphasis>
|
||
angle with the base of the propodeal spines. Propodeal lobes rounded and very weakly developed. Metapleural gland bulla small, extending from the metacoxal insertion a third of the way to the propodeal spiracle. Petiole moderately long (PLI 173), without tubercles anterodorsally. Subpetiolar process in the form of a very small, acute tooth; ventral margin of the petiole bulging slightly medially. Petiolar peduncle long: comprising about two thirds of the petiole. Petiolar node robust and erect, nearly squamiform: transition between peduncle and node marked by a rounded angle of ~120
|
||
<emphasis box="[1524,1534,1027,1045]" italics="true" pageId="126" pageNumber="127">
|
||
<superScript attach="none" box="[1524,1534,1027,1045]" fontSize="8" pageId="126" pageNumber="127">°</superScript>
|
||
</emphasis>
|
||
, resulting in a concave anterior node face; anterior face forming a ~100
|
||
<emphasis box="[1333,1343,1067,1085]" italics="true" pageId="126" pageNumber="127">
|
||
<superScript attach="left" box="[1333,1343,1067,1085]" fontSize="8" pageId="126" pageNumber="127">°</superScript>
|
||
</emphasis>
|
||
angle with the dorsal face, which is weakly convex; dorsal face rounding evenly into the short posterior face, which forms a ~85
|
||
<emphasis box="[781,791,1146,1164]" italics="true" pageId="126" pageNumber="127">
|
||
<superScript attach="left" box="[781,791,1146,1164]" fontSize="8" pageId="126" pageNumber="127">°</superScript>
|
||
</emphasis>
|
||
angle with the caudal cylinder, overhanging it. Postpetiole subquadrate: flat anteriorly, transitioning into the flattened dorsal face through a rounded angle; ventral surface weakly lobed.
|
||
</paragraph>
|
||
<paragraph blockId="126.[498,1542,234,1934]" pageId="126" pageNumber="127">In dorsal view, mesoscutum covering pronotum anteriorly, but humeri visible laterally as slightly angulate sclerites. Propodeal spines broadly approximated basally and diverging apically, their apices separated from each other by about twice their length. Petiolar peduncle with spiracles protruding past the lateral margins, the peduncle constricted anterior to them. Petiolar node subovate and transversely broad. Petiolar node broader than the peduncle, and about one and a half times as wide as the caudal cylinder. Postpetiole moderately broad (PWI 203), anteroposteriorly compressed, and subquadrate, articulating with most of the anterior margin of the gaster, leaving small, angulate margins on each side exposed. Anterior margin of postpetiole flat, with corners marked by rounded angles as it transitions to the lateral margins, which weakly diverge to the angulate posterior corners; posterior margin flat. Metafemur moderately incrassate (FI 273).</paragraph>
|
||
<paragraph blockId="126.[498,1542,234,1934]" lastBlockId="127.[498,1542,234,1456]" lastPageId="127" lastPageNumber="128" pageId="126" pageNumber="127">Sculpture: median clypeal carina present but flanked by numerous equally strong carinae. Lateral clypeal lobes with additional weaker carinae; ground sculpture weakly areolate. Antennal scapes areolate. Cephalic dorsum costulate, with areolate-rugulose ground sculpture; weak concentric costulae surrounding the antennal insertions. Lateral surfaces of head sculptured similarly to the dorsum. Ventral surface of head sculptured similarly to the rest of the head, but weaker. Pronotal neck areolate. Pronotum, metapleuron and lateral face of the propodeum sculptured similarly to the head dorsum. Anepisternum and katepisternum weakly areolate on their anterior quarters, transitioning into stronger areolate sculpture posteriorly. Propodeal declivity weakly areolate. Mesoscutum and mesoscutellum with longitudinal striae over areolate ground sculpture. Femora areolate. Petiole and postpetiole weakly areolate. First gastral tergite weakly areolate, with weak spectral iridescence. Surface of the first gastral sternite with weak spectral iridescence.</paragraph>
|
||
<paragraph blockId="127.[498,1542,234,1456]" pageId="127" pageNumber="128">Setae: antennal scapes and funiculi with short, decumbent pilosity. Dorsum of the head, pronotum, waist segments, and gaster with moderately abundant, erect, blunt-tipped setae, the longest of which are about a third of the width of the compound eye. Short, sparse pubescence present over the entire body, but difficult to detect against the ground sculpture.</paragraph>
|
||
<paragraph blockId="127.[498,1542,234,1456]" box="[530,1225,712,738]" pageId="127" pageNumber="128">Color: predominantly medium brown, with yellowish tarsi.</paragraph>
|
||
<paragraph blockId="127.[498,1542,234,1456]" box="[530,730,752,778]" pageId="127" pageNumber="128">
|
||
<emphasis bold="true" box="[530,597,752,778]" pageId="127" pageNumber="128">Male:</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="127" pageNumber="128" type="etymology">
|
||
<paragraph blockId="127.[498,1542,234,1456]" pageId="127" pageNumber="128">
|
||
<emphasis bold="true" box="[530,668,792,818]" pageId="127" pageNumber="128">Etymology:</emphasis>
|
||
Patronym, for the Swiss myrmecologist Auguste Forel, who originally described this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="127" pageNumber="128" type="discussion">
|
||
<paragraph blockId="127.[498,1542,234,1456]" pageId="127" pageNumber="128">
|
||
<emphasis bold="true" box="[530,670,872,898]" pageId="127" pageNumber="128">Comments:</emphasis>
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[679,929,872,898]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[679,929,872,898]" italics="true" pageId="127" pageNumber="128">Temnothorax augusti</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus box="[1225,1275,872,898]" pageId="127" pageNumber="128">type</typeStatus>
|
||
series, which was collected from
|
||
<taxonomicName box="[670,786,912,938]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="127" pageNumber="128" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[670,786,912,938]" italics="true" pageId="127" pageNumber="128">Tillandsia</emphasis>
|
||
</taxonomicName>
|
||
epiphytes near Cuernavaca,
|
||
<collectingRegion box="[1116,1214,912,938]" country="Mexico" name="Morelos" pageId="127" pageNumber="128">Morelos</collectingRegion>
|
||
,
|
||
<collectingRegion box="[1224,1344,912,938]" country="Mexico" name="Distrito Federal" pageId="127" pageNumber="128">Mexico by</collectingRegion>
|
||
William Morton Wheeler. The epiphyte in which the nest series was collected was shared with two other ant species:
|
||
<taxonomicName authority="Mayr" authorityName="Mayr" authorityYear="1862" box="[640,973,992,1018]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="crinosa">
|
||
<emphasis box="[640,899,992,1018]" italics="true" pageId="127" pageNumber="128">Crematogaster crinosa</emphasis>
|
||
Mayr
|
||
</taxonomicName>
|
||
, a common inhabitant of the canopy and disturbed areas in the Neotropics, and
|
||
<taxonomicName authority="(Forel)" baseAuthorityName="Forel" baseAuthorityYear="1901" box="[958,1276,1031,1058]" class="Insecta" family="Formicidae" genus="Cephalotes" kingdom="Animalia" order="Hymenoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="wheeleri">
|
||
<emphasis box="[958,1186,1031,1057]" italics="true" pageId="127" pageNumber="128">Cephalotes wheeleri</emphasis>
|
||
(Forel)
|
||
</taxonomicName>
|
||
, a much less common arboreal species known only from Central
|
||
<collectingCountry box="[1002,1088,1071,1097]" name="Mexico" pageId="127" pageNumber="128">Mexico</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Wheeler WM" box="[1107,1272,1071,1097]" journalOrPublisher="New York: Columbia University Press" pageId="127" pageNumber="128" refId="ref273372" refString="Wheeler WM. 1910. Ants: their structure, development and behavior. New York: Columbia University Press." title="Ants: their structure, development and behavior" type="book" year="1910">
|
||
<emphasis box="[1107,1272,1071,1097]" italics="true" pageId="127" pageNumber="128">Wheeler, 1910</emphasis>
|
||
</bibRefCitation>
|
||
). The morphologically closest species to
|
||
<taxonomicName baseAuthorityName="Baroni Urbani" baseAuthorityYear="1901" box="[705,822,1112,1137]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="augusti">
|
||
<emphasis box="[705,822,1112,1137]" italics="true" pageId="127" pageNumber="128">T. augusti</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<taxonomicName authority="Prebus, 2021" authorityName="Prebus" authorityYear="2021" box="[857,1003,1112,1137]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="casanovai" status="sp. nov.">
|
||
<emphasis box="[857,1003,1112,1137]" italics="true" pageId="127" pageNumber="128">T. casanovai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1012,1106,1111,1137]" pageId="127" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, which was collected in neighboring
|
||
<collectingRegion box="[498,641,1151,1177]" country="Mexico" name="Puebla" pageId="127" pageNumber="128">Puebla state</collectingRegion>
|
||
,
|
||
<collectingCountry box="[654,740,1151,1177]" name="Mexico" pageId="127" pageNumber="128">Mexico</collectingCountry>
|
||
from the limb of a pitahaya cactus,
|
||
<taxonomicName authorityName="Britton & Rose" authorityYear="1918" baseAuthorityName="Haw." box="[1170,1398,1151,1177]" class="Magnoliopsida" family="Cactaceae" genus="Hylocereus" kingdom="Plantae" order="Caryophyllales" pageId="127" pageNumber="128" phylum="Tracheophyta" rank="species" species="undatus">
|
||
<emphasis box="[1170,1398,1151,1177]" italics="true" pageId="127" pageNumber="128">Hylocereus undatus</emphasis>
|
||
</taxonomicName>
|
||
, a hemiepiphytic species commonly cultivated for its fruit. Based on multiple morphological disparities between the two, including gross size, distance between the base of the propodeal spines, degree of sculpture on the first gastral tergite, and color, I choose to keep these species separate in the current revision. This issue should be revisited as more collections become available. This species is reported from
|
||
<collectingRegion box="[1195,1362,1350,1377]" country="Mexico" name="Quintana Roo" pageId="127" pageNumber="128">Quintana Roo</collectingRegion>
|
||
by
|
||
<bibRefCitation author="Lachaud JP & Lachaud P." editor="Vasquez- Bolanos M & Castano-Meneses G & Cisneros-Caballero A & Quiroz-Rocha GA & Navarrete- Heredia JL" journalOrPublisher="Formicidae de Mexico. Guadalajara: Cuerpo Academico de Zoologia" pageId="127" pageNumber="128" pagination="23 - 32" refId="ref271640" refString="Lachaud JP, Lachaud P. 2013. Revision preliminar de las hormigas de Campeche y Quintana Roo, Mexico, con base en la coleccion de Arthropoda del Colegio de la Frontera Sur. In: Vasquez- Bolanos M, Castano-Meneses G, Cisneros-Caballero A, Quiroz-Rocha GA, Navarrete- Heredia JL, eds. Formicidae de Mexico. Guadalajara: Cuerpo Academico de Zoologia, 23 - 32." title="Revision preliminar de las hormigas de Campeche y Quintana Roo, Mexico, con base en la coleccion de Arthropoda del Colegio de la Frontera Sur" type="book chapter" year="2013">
|
||
<emphasis italics="true" pageId="127" pageNumber="128">Lachaud & Lachaud (2013)</emphasis>
|
||
</bibRefCitation>
|
||
; I have not personally examined these specimens, so I have excluded it from the range for the moment.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |